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<document id="680323EF6DD2C4B32DBD3ADD69469D6C" ID-DOI="10.11646/zootaxa.3872.2.1" ID-GBIF-Dataset="e9b8d555-d661-4f1a-86e8-f2a0acc38798" ID-ISSN="1175-5326" ID-Zenodo-Dep="253224" ID-ZooBank="F9F81CC8-E033-46B7-B73B-9FB777DF4116" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460926510372" checkinUser="plazi" docAuthor="Ponder, Winston F., Fukuda, Hiroshi &amp; Hallan, Anders" docDate="2014" docId="03CDAD65575F3C0CFF05FA24BF10AB09" docLanguage="en" docName="zt03872p153.pdf" docOrigin="Zootaxa 3872 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coliracemata" docType="treatment" docVersion="9" lastPageNumber="140" masterDocId="FFF4D51D57783C24FF92FFB2BB3EA911" masterDocTitle="A review of the family Clenchiellidae (Mollusca: Caenogastropoda: Truncatelloidea)" masterLastPageNumber="153" masterPageNumber="101" pageNumber="140" updateTime="1698362634141" updateUser="plazi">
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<mods:title id="68530C5D6AC977A2E6321D8702B458A2">A review of the family Clenchiellidae (Mollusca: Caenogastropoda: Truncatelloidea)</mods:title>
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<mods:name id="1E928C1D6D8737993EE7A8917E599257" type="personal">
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<mods:namePart id="2C0DE611CA0277D5D2C61B07BB0D7C16">Ponder, Winston F.</mods:namePart>
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<mods:namePart id="468D479ED7EBD20D502EBB4773B8D10A">Fukuda, Hiroshi</mods:namePart>
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<mods:namePart id="DADFBBEE61518774429E00B13AAA0D4E">Hallan, Anders</mods:namePart>
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<mods:title id="E8A6ECA92B2D3DE98AA43135A8C57410">Zootaxa</mods:title>
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<mods:part id="2B198BFE1FCD8F8ADC042FDB88C7D29B">
<mods:date id="471B947FD8A8FFCB94070B41577FFC7B">2014</mods:date>
<mods:detail id="717F2783386E531F94E40B0A6B888344" type="volume">
<mods:number id="575885EAD16D98094A01031EA3456BF6">3872</mods:number>
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<mods:classification id="D7EAE637BB83ED098B71CCF04F314D5F">journal article</mods:classification>
<mods:identifier id="9EC0B94B8E3ADF7C0B347E92A31ADC43" type="DOI">10.11646/zootaxa.3872.2.1</mods:identifier>
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<treatment id="03CDAD65575F3C0CFF05FA24BF10AB09" ID-DOI="http://doi.org/10.5281/zenodo.5631029" ID-GBIF-Taxon="119589570" ID-Zenodo-Dep="5631029" LSID="urn:lsid:plazi:treatment:03CDAD65575F3C0CFF05FA24BF10AB09" httpUri="http://treatment.plazi.org/id/03CDAD65575F3C0CFF05FA24BF10AB09" lastPageId="40" lastPageNumber="140" pageId="39" pageNumber="140">
<subSubSection id="C37E4FF8575F3C03FF05FA24BA3DACC3" pageId="39" pageNumber="140" type="nomenclature">
<paragraph id="8BDB1C73575F3C03FF05FA24B9BEACA0" blockId="39.[151,640,1430,1490]" box="[151,640,1430,1457]" pageId="39" pageNumber="140">
<heading id="D093AB1F575F3C03FF05FA24B9BEACA0" bold="true" box="[151,640,1430,1457]" fontSize="11" level="1" pageId="39" pageNumber="140" reason="1">
<emphasis id="B910C061575F3C03FF05FA24B9BEACA0" bold="true" box="[151,640,1430,1457]" pageId="39" pageNumber="140">
<taxonomicName id="4C6467F0575F3C03FF05FA24BA07ACA1" ID-CoL="7P24W" box="[151,313,1430,1456]" class="Gastropoda" family="Clenchiellidae" genus="Coliracemata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="39" pageNumber="140" phylum="Mollusca" rank="genus">
<emphasis id="B910C061575F3C03FF05FA24BA07ACA1" bold="true" box="[151,313,1430,1456]" italics="true" pageId="39" pageNumber="140">Coliracemata</emphasis>
</taxonomicName>
?
<taxonomicName id="4C6467F0575F3C03FEDDFA24B9BEACA0" authority="Preston, 1915" authorityName="Preston" authorityYear="1915" box="[335,640,1430,1457]" class="Gastropoda" family="Clenchiellidae" genus="Coliracemata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="39" pageNumber="140" phylum="Mollusca" rank="species" species="innocens">
<emphasis id="B910C061575F3C03FEDDFA24BA82ACA1" bold="true" box="[335,444,1430,1456]" italics="true" pageId="39" pageNumber="140">innocens</emphasis>
(
<bibRefCitation id="EFF56182575F3C03FE59FA25B946ACA0" author="Preston" box="[459,632,1431,1457]" pageId="39" pageNumber="140" refString="Preston, H. B. (1915) A further report on Mollusca from Lake Chilka on the east coast of India. Records of the Indian Museum, 11, 289 - 310." type="journal article" year="1915">Preston, 1915</bibRefCitation>
)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDB1C73575F3C03FF05FA0BBA3DACC3" blockId="39.[151,640,1430,1490]" box="[151,259,1465,1490]" pageId="39" pageNumber="140">
<figureCitation id="135F00F6575F3C03FF05FA0BBA3DACC3" box="[151,259,1465,1490]" captionStart="FIGURE 27" captionStartId="39.[151,250,1303,1325]" captionTargetBox="[498,1102,515,1284]" captionTargetId="figure@39.[494,1103,509,1288]" captionTargetPageId="39" captionText="FIGURE 27. Coliracemata? microscopica (A, B) modified from the original illustrations by Annandale and Kemp (1916, fig. 3) and Coliracemata? innocens (C – E) modified from the original illustrations by Preston (1915, fig. 9 a – c). Scale bars: 500 µm." httpUri="https://zenodo.org/record/253251/files/figure.png" pageId="39" pageNumber="140">Figure 27</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37E4FF8575F3C03FF05F9B3BF7DAF90" pageId="39" pageNumber="140" type="reference_group">
<paragraph id="8BDB1C73575F3C03FF05F9B3B814AF06" blockId="39.[151,810,1537,1559]" box="[151,810,1537,1559]" pageId="39" pageNumber="140">
<treatmentCitationGroup id="AB743B5D575F3C03FF05F9B3B814AF06" box="[151,810,1537,1559]" pageId="39" pageNumber="140">
<taxonomicName id="4C6467F0575F3C03FF05F9B3B99CAF06" authority="Preston, 1915: 296" authorityName="Preston" authorityPageNumber="296" authorityYear="1915" box="[151,674,1537,1559]" class="Gastropoda" family="Liotiidae" genus="Cyclostrema" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="39" pageNumber="140" phylum="Mollusca" rank="species" species="innocens" subGenus="Tubiola">
<emphasis id="B910C061575F3C03FF05F9B3BA28AF06" box="[151,278,1537,1559]" italics="true" pageId="39" pageNumber="140">Cyclostrema</emphasis>
(
<emphasis id="B910C061575F3C03FEB6F9B3BA4FAF06" box="[292,369,1537,1559]" italics="true" pageId="39" pageNumber="140">Tubiola</emphasis>
)
<emphasis id="B910C061575F3C03FE13F9B0BAE5AF06" box="[385,475,1538,1559]" italics="true" pageId="39" pageNumber="140">innocens</emphasis>
<treatmentCitation id="0AC53A62575F3C03FE73F9B0B99CAF06" author="Preston" box="[481,674,1537,1559]" page="296" pageId="39" pageNumber="140" year="1915">
<bibRefCitation id="EFF56182575F3C03FE73F9B0B94EAF06" author="Preston" box="[481,624,1537,1559]" pageId="39" pageNumber="140" refString="Preston, H. B. (1915) A further report on Mollusca from Lake Chilka on the east coast of India. Records of the Indian Museum, 11, 289 - 310." type="journal article" year="1915">Preston, 1915</bibRefCitation>
: 296
</treatmentCitation>
</taxonomicName>
, figs 9, 9a–b.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDB1C73575F3C03FF05F9F6BF7DAF90" blockId="39.[151,1437,1604,2025]" pageId="39" pageNumber="140">
<emphasis id="B910C061575F3C03FF05F9F6BA8CAF4C" bold="true" box="[151,434,1604,1629]" pageId="39" pageNumber="140">
<typeStatus id="54DFA2D1575F3C03FF05F9F6BBE2AF4C" box="[151,220,1604,1629]" pageId="39" pageNumber="140">Types</typeStatus>
and
<typeStatus id="54DFA2D1575F3C03FE88F9F6BA70AF4C" box="[282,334,1604,1629]" pageId="39" pageNumber="140">type</typeStatus>
locality.
</emphasis>
<typeStatus id="54DFA2D1575F3C03FE29F9F6B916AF4C" box="[443,552,1604,1629]" pageId="39" pageNumber="140" type="holotype">Holotype</typeStatus>
: Serua Nadi, Lake Chilka (=Chilika Lake) (eastern
<collectingCountry id="F3735CE3575F3C03FBE0F9F6BF8EAF4C" box="[1138,1200,1604,1629]" name="India" pageId="39" pageNumber="140">India</collectingCountry>
),
<quantity id="4C9CB196575F3C03FB57F9F6BE35AF4C" box="[1221,1291,1604,1629]" metricMagnitude="0" metricUnit="m" metricValue="2.1336" metricValueMax="2.7432" metricValueMin="1.524" pageId="39" pageNumber="140" unit="ft" value="7.0" valueMax="9.0" valueMin="5.0">5–9 ft</quantity>
. (
<quantity id="4C9CB196575F3C03FA8DF9F6BEAAAF4C" box="[1311,1428,1604,1629]" metricMagnitude="0" metricUnit="m" metricValue="2.1" metricValueMax="2.7" metricValueMin="1.5" pageId="39" pageNumber="140" unit="m" value="2.1" valueMax="2.7" valueMin="1.5">1.5–2.7 m</quantity>
) depth (Indian Museum, Calcutta,
<bibRefCitation id="EFF56182575F3C03FD9DF9DBB993AF90" author="Preston" box="[527,685,1640,1665]" pageId="39" pageNumber="140" refString="Preston, H. B. (1915) A further report on Mollusca from Lake Chilka on the east coast of India. Records of the Indian Museum, 11, 289 - 310." type="journal article" year="1915">Preston, 1915</bibRefCitation>
: 289 (
<typeStatus id="54DFA2D1575F3C03FD63F9DBB81DAF90" box="[753,803,1641,1665]" pageId="39" pageNumber="140">type</typeStatus>
material apparently lost).
</paragraph>
</subSubSection>
<subSubSection id="C37E4FF8575F3C03FF55F93EBF13AFB4" box="[199,1069,1676,1701]" pageId="39" pageNumber="140" type="distribution">
<paragraph id="8BDB1C73575F3C03FF55F93EBF13AFB4" blockId="39.[151,1437,1604,2025]" box="[199,1069,1676,1701]" pageId="39" pageNumber="140">
<emphasis id="B910C061575F3C03FF55F93EBA5EAFB4" bold="true" box="[199,352,1676,1701]" pageId="39" pageNumber="140">Distribution.</emphasis>
Known only from the
<typeStatus id="54DFA2D1575F3C03FDF2F93FB9AEAFB4" box="[608,656,1677,1701]" pageId="39" pageNumber="140">type</typeStatus>
locality, a large brackish-water lake.
</paragraph>
</subSubSection>
<subSubSection id="C37E4FF8575F3C03FF55F902BAC2AFD9" box="[199,508,1712,1737]" pageId="39" pageNumber="140" type="materials_examined">
<paragraph id="8BDB1C73575F3C03FF55F902BAC2AFD9" blockId="39.[151,1437,1604,2025]" box="[199,508,1712,1737]" pageId="39" pageNumber="140">
<emphasis id="B910C061575F3C03FF55F902BA8EAFD8" bold="true" box="[199,432,1712,1737]" pageId="39" pageNumber="140">Material examined.</emphasis>
None.
</paragraph>
</subSubSection>
<subSubSection id="C37E4FF8575F3C03FF55F966B9F2AED4" pageId="39" pageNumber="140" type="description">
<paragraph id="8BDB1C73575F3C03FF55F966B822AFFC" blockId="39.[151,1437,1604,2025]" box="[199,796,1748,1773]" pageId="39" pageNumber="140">
<emphasis id="B910C061575F3C03FF55F966BA6FAFFC" bold="true" box="[199,337,1748,1773]" pageId="39" pageNumber="140">Description</emphasis>
(based on Preston 1914: 296, figs 9a–c).
</paragraph>
<paragraph id="8BDB1C73575F3C03FF55F94ABA52AEB0" blockId="39.[151,1437,1604,2025]" pageId="39" pageNumber="140">
<emphasis id="B910C061575F3C03FF55F94ABBC1AE00" box="[199,255,1784,1809]" italics="true" pageId="39" pageNumber="140">Shell</emphasis>
. Minute (2.0 mm in diameter;
<tableCitation id="C6E629C8575F3C03FDF6F94AB984AE00" box="[612,698,1784,1809]" captionStart="TABLE 3" captionStartId="13.[151,239,656,679]" captionText="TABLE 3. Shell dimensions of type specimens of species of Clenchiellidae." pageId="39" pageNumber="140">Table 3</tableCitation>
), spire low (
<figureCitation id="135F00F6575F3C03FCDFF94AB89EAE00" box="[845,928,1784,1809]" captionStart="FIGURE 27" captionStartId="39.[151,250,1303,1325]" captionTargetBox="[498,1102,515,1284]" captionTargetId="figure@39.[494,1103,509,1288]" captionTargetPageId="39" captionText="FIGURE 27. Coliracemata? microscopica (A, B) modified from the original illustrations by Annandale and Kemp (1916, fig. 3) and Coliracemata? innocens (C – E) modified from the original illustrations by Preston (1915, fig. 9 a – c). Scale bars: 500 µm." httpUri="https://zenodo.org/record/253251/files/figure.png" pageId="39" pageNumber="140">Fig. 27</figureCitation>
C–E). Protoconch not elevated above line of spire, details unknown. Teleoconch of about 3 convex whorls, surface smooth except for growth lines. Periphery evenly convex. Base evenly convex, umbilicus moderately wide (slightly more than half width of base). Sutures impressed. Aperture near circular, prosocline, with simple, slightly thickened peristome, no obvious external varix. Colour milk white.
</paragraph>
<paragraph id="8BDB1C73575F3C03FF55F81EB9F2AED4" blockId="39.[151,1437,1604,2025]" box="[199,716,1964,1989]" pageId="39" pageNumber="140">Operculum, head-foot and anatomy unknown.</paragraph>
</subSubSection>
<subSubSection id="C37E4FF8575F3C0CFF55F862BF10AB09" lastPageId="40" lastPageNumber="141" pageId="39" pageNumber="140" type="discussion">
<paragraph id="8BDB1C73575F3C0CFF55F862BF10AB09" blockId="39.[151,1437,1604,2025]" lastBlockId="40.[151,1437,151,536]" lastPageId="40" lastPageNumber="141" pageId="39" pageNumber="140">
<emphasis id="B910C061575F3C03FF55F862BA05AEF8" bold="true" box="[199,315,2000,2025]" pageId="39" pageNumber="140">Remarks.</emphasis>
Preston’s
<taxonomicName id="4C6467F0575F3C03FE23F862B826AEF9" box="[433,792,2000,2025]" class="Gastropoda" family="Liotiidae" genus="Cyclostrema" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="39" pageNumber="140" phylum="Mollusca" rank="species" species="innocens" subGenus="Tubiola">
<emphasis id="B910C061575F3C03FE23F862B900AEF8" box="[433,574,2000,2025]" italics="true" pageId="39" pageNumber="140">Cyclostrema</emphasis>
(
<emphasis id="B910C061575F3C03FDDCF862B99DAEF8" box="[590,675,2000,2025]" italics="true" pageId="39" pageNumber="140">Tubiola</emphasis>
)
<emphasis id="B910C061575F3C03FD26F863B826AEF9" box="[692,792,2001,2024]" italics="true" pageId="39" pageNumber="140">innocens</emphasis>
</taxonomicName>
was compared with
<taxonomicName id="4C6467F0575F3C03FB90F863BFF6AEF9" box="[1026,1224,2001,2024]" class="Gastropoda" family="Vanikoridae" genus="Tubiola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="39" pageNumber="140" phylum="Mollusca" rank="species" species="microscopica">
<emphasis id="B910C061575F3C03FB90F863BF1DAEF9" box="[1026,1059,2001,2024]" italics="true" pageId="39" pageNumber="140">Co</emphasis>
.
<emphasis id="B910C061575F3C03FBA0F863BFF6AEF9" box="[1074,1224,2001,2024]" italics="true" pageId="39" pageNumber="140">microscopica</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFF56182575F3C0CFB61F862BA57A9A1" author="Annandale" lastPageId="40" lastPageNumber="141" pageId="39" pageNumber="140" refString="Annandale, N. &amp; Kemp, S. (1916) Fauna of the Chilka Lake: Mollusca Gastropoda and Lamellibranchia. Memoirs of the Indian Museum, 5, 329 - 366." type="journal article" year="1916">Annandale and Kemp (1916: 347)</bibRefCitation>
with the comment that it “clearly belongs to the same genus” and “appears to differ from that species in its smaller size, reddish colour and in the sculpture on its surface; but the
<typeStatus id="54DFA2D157503C0CFBF4FF0FBFA8A9C4" box="[1126,1174,189,213]" pageId="40" pageNumber="141">type</typeStatus>
of Preston’s species is bleached and perhaps somewhat eroded.” Preston’s description stated that his specimen (from Serua Nadi, Lake Chilka, a brackish water locality) is smooth. He gives a size of 2.0 mm maximum diameter, whereas
<taxonomicName id="4C6467F057503C0CFA94FEB7BEA2A80D" box="[1286,1436,261,284]" class="Gastropoda" family="Vanikoridae" genus="Tubiola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="40" pageNumber="141" phylum="Mollusca" rank="species" species="microscopica">
<emphasis id="B910C06157503C0CFA94FEB7BEA2A80D" box="[1286,1436,261,284]" italics="true" pageId="40" pageNumber="141">microscopica</emphasis>
</taxonomicName>
is, according to Nevill’s original description,
<quantity id="4C9CB19657503C0CFD36FE95B9C2A82E" box="[676,764,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="40" pageNumber="141" unit="mm" value="1.5">1.5 mm</quantity>
in diameter. Annandale later (1924: 858) stated a slightly different opinion - “Dr Kemp and I were inclined to think [that
<taxonomicName id="4C6467F057503C0CFC14FEFFB8D4A875" box="[902,1002,333,356]" class="Gastropoda" family="Liotiidae" genus="Cyclostrema" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="40" pageNumber="141" phylum="Mollusca" rank="species" species="innocens">
<emphasis id="B910C06157503C0CFC14FEFFB8D4A875" box="[902,1002,333,356]" italics="true" pageId="40" pageNumber="141">innocens</emphasis>
</taxonomicName>
] is a small denuded shell of Nevill’s species”. These authors are the only ones who had access to the
<typeStatus id="54DFA2D157503C0CFCE4FEC2B898A899" box="[886,934,368,392]" pageId="40" pageNumber="141">type</typeStatus>
material of both and that have published an opinion on these taxa.
<bibRefCitation id="EFF5618257503C0CFE04FE26B908A8BD" author="Abbott" box="[406,566,404,429]" pageId="40" pageNumber="141" refString="Abbott, R. T. (1949) An Indian species of Clenchiella. The Nautilus, 63, 62." type="journal article" year="1949">Abbott (1949)</bibRefCitation>
was uncertain regarding the synonymy of these two species. Because, on the available evidence,
<emphasis id="B910C06157503C0CFEE1FE0ABAA5A8DE" box="[371,411,440,463]" italics="true" pageId="40" pageNumber="141">Co.</emphasis>
?
<taxonomicName id="4C6467F057503C0CFE3FFE0BB92FA8C1" box="[429,529,441,464]" class="Gastropoda" family="Liotiidae" genus="Cyclostrema" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="40" pageNumber="141" phylum="Mollusca" rank="species" species="innocens">
<emphasis id="B910C06157503C0CFE3FFE0BB92FA8C1" box="[429,529,441,464]" italics="true" pageId="40" pageNumber="141">innocens</emphasis>
</taxonomicName>
differs from
<emphasis id="B910C06157503C0CFD31FE0AB9F5A8DE" box="[675,715,440,463]" italics="true" pageId="40" pageNumber="141">Co.</emphasis>
?
<taxonomicName id="4C6467F057503C0CFD4CFE0BB84AA8C1" box="[734,884,441,464]" class="Gastropoda" family="Vanikoridae" genus="Tubiola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="40" pageNumber="141" phylum="Mollusca" rank="species" species="microscopica">
<emphasis id="B910C06157503C0CFD4CFE0BB84AA8C1" box="[734,884,441,464]" italics="true" pageId="40" pageNumber="141">microscopica</emphasis>
</taxonomicName>
in its larger size, apparent lack of spiral sculpture and slightly wider umbilicus, we tentatively treat both as separate species pending the discovery of new topotypic material.
<emphasis id="B910C06157503C0CFE92FDB2BA1FAB06" box="[256,289,512,535]" italics="true" pageId="40" pageNumber="141">Co</emphasis>
.?
<taxonomicName id="4C6467F057503C0CFEAEFDB3BA9EAB09" box="[316,416,513,536]" class="Gastropoda" family="Liotiidae" genus="Cyclostrema" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="40" pageNumber="141" phylum="Mollusca" rank="species" species="innocens">
<emphasis id="B910C06157503C0CFEAEFDB3BA9EAB09" box="[316,416,513,536]" italics="true" pageId="40" pageNumber="141">innocens</emphasis>
</taxonomicName>
could also conceivably be a species of
<taxonomicName id="4C6467F057503C0CFCCBFE4DB8E7AB09" box="[857,985,511,536]" class="Gastropoda" family="Clenchiellidae" genus="Coleglabra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="40" pageNumber="141" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B910C06157503C0CFCCBFE4DB8E7AB09" box="[857,985,511,536]" italics="true" pageId="40" pageNumber="141">Coleglabra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2237D1A57503C0CFC4DFE4DBF10AB09" box="[991,1070,511,536]" pageId="40" pageNumber="141" rank="genus">
<emphasis id="B910C06157503C0CFC4DFE4DBF10AB09" bold="true" box="[991,1070,511,536]" pageId="40" pageNumber="141">n. gen.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>
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