treatments-xml/data/03/C0/87/03C087934476FF99FCF7ED50A480FB0E.xml

<document id="5382E9AD0B987677CB831D4B51330D96" ID-DOI="10.12651/JSR.2017.6.2.181" ID-ISSN="2713-8615" ID-Zenodo-Dep="13161080" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722544677625" checkinUser="felipe" docAuthor="Hartini, Sri &amp; Dwibadra, Dhian" docDate="2017" docId="03C087934476FF99FCF7ED50A480FB0E" docLanguage="en" docName="JSpeciesRes.6.2.181-189.pdf" docOrigin="Journal of Species Research 6 (2)" docStyle="DocumentStyle:4D398BFF45FF8CA9E08F546BB06AE372.2:JSpeciesRes.2012-.journal_article" docStyleId="4D398BFF45FF8CA9E08F546BB06AE372" docStyleName="JSpeciesRes.2012-.journal_article" docStyleVersion="2" docTitle="Macrocheles gedeensis Hartini &amp; Dwibadra, 2017, sp. nov." docType="treatment" docUuid="04CDD3EB-1F37-492E-B28A-363AC0F793DA" docUuidSource="ZooBank" docVersion="2" lastPageNumber="183" masterDocId="FFF9FFEB4477FF9BFFDEEC55A103FFF9" masterDocTitle="Macrochelid mites (Acari: Mesostigmata) associated with dung beetles in Mount Gede-Pangrango National Park, West Java, Indonesia" masterLastPageNumber="189" masterPageNumber="181" pageNumber="182" updateTime="1722567086849" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="58F51905172ADBF1AC93E4EB49424776">Macrochelid mites (Acari: Mesostigmata) associated with dung beetles in Mount Gede-Pangrango National Park, West Java, Indonesia</mods:title>
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<mods:namePart id="BE382B123AA5A2197AF4ECCC76CA7E43">Hartini, Sri</mods:namePart>
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<mods:namePart id="F7061213C09959B8F86FE59AC3D22D51">Dwibadra, Dhian</mods:namePart>
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<mods:date id="BF53933273B07B8BBD5341C9B1243AC1">2017</mods:date>
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<treatment id="03C087934476FF99FCF7ED50A480FB0E" LSID="urn:lsid:zoobank.org:act:04CDD3EB-1F37-492E-B28A-363AC0F793DA" httpUri="http://treatment.plazi.org/id/03C087934476FF99FCF7ED50A480FB0E" lastPageId="2" lastPageNumber="183" pageId="1" pageNumber="182">
<subSubSection id="C373650E4476FF9AFCF7ED50A5FBFEE4" box="[809,1272,261,285]" pageId="1" pageNumber="182" type="nomenclature">
<paragraph id="8BD636854476FF9AFCF7ED50A5FBFEE4" blockId="1.[809,1272,261,285]" box="[809,1272,261,285]" pageId="1" pageNumber="182">
<heading id="D09E81E94476FF9AFCF7ED50A5FBFEE4" bold="true" box="[809,1272,261,285]" fontSize="10" level="2" pageId="1" pageNumber="182" reason="4">
<emphasis id="B91DEA974476FF9AFCF7ED50A5FBFEE4" bold="true" box="[809,1272,261,285]" pageId="1" pageNumber="182">
<taxonomicName id="4C694D064476FF9AFCF7ED50A526FEE4" authorityName="Hartini &amp; Dwibadra" authorityYear="2017" box="[809,1061,261,285]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="182" phylum="Arthropoda" rank="species" species="gedeensis" status="sp. nov.">
<emphasis id="B91DEA974476FF9AFCF7ED50A526FEE4" bold="true" box="[809,1061,261,285]" italics="true" pageId="1" pageNumber="182">Macrocheles gedeensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57EC4476FF9AFBF2ED50A581FEE4" box="[1068,1154,261,285]" pageId="1" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="13522A004476FF9AFB53ED50A5ECFEE4" box="[1165,1263,261,285]" captionStart="Figs" captionStartId="1.[144,186,1952,1974]" captionTargetBox="[318,1234,800,1918]" captionTargetId="figure-243@1.[317,1240,794,1920]" captionTargetPageId="1" captionText="Figs. 1-6. Macrocheles gedeensis sp. nov., holotype female(MZB.Acar.7091.2). 1, dorsum; 2, venter; 3, ventral view of gnathosoma; 4, epi- stome; 5, chelicerae; 6, sacculus foemineus." figureDoi="http://doi.org/10.5281/zenodo.13161082" httpUri="https://zenodo.org/record/13161082/files/figure.png" pageId="1" pageNumber="182">Figs. 1-6</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C373650E4476FF99FCF7ED13A318F865" lastPageId="2" lastPageNumber="183" pageId="1" pageNumber="182" type="materials_examined">
<paragraph id="8BD636854476FF9AFCF7ED13A5AFFE06" blockId="1.[809,1414,325,737]" pageId="1" pageNumber="182">
<emphasis id="B91DEA974476FF9AFCF7ED13A2A5FEA4" bold="true" box="[809,934,325,350]" pageId="1" pageNumber="182">Type series</emphasis>
: 
<typeStatus id="54D288274476FF9AFC6AED10A518FEA4" box="[948,1051,325,349]" pageId="1" pageNumber="182" type="holotype">Holotype</typeStatus>
: female (
<collectionCode id="ED78AE404476FF9AFBA4ED13A5BBFEA7" box="[1146,1208,326,350]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="182" type="Museum">MZB</collectionCode>
. Acar.7091.2) 
<quantity id="4C919B604476FF9AFA8FED13A23DFE84" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="1" pageNumber="182" unit="m" value="1130.0">1130 m</quantity>
above sea level (a.s.l.), Selabintana, Mount Gede-Pangrango, Sukabumi, 
<collectingRegion id="49ADF8674476FF9AFBF5EDD2A59FFE67" box="[1067,1180,391,414]" country="Indonesia" name="Jawa Barat" pageId="1" pageNumber="182">West Java</collectingRegion>
, 
<geoCoordinate id="EE5D50424476FF9AFB74EDD3A42CFE67" box="[1194,1327,389,414]" degrees="06" direction="south" minutes="52" orientation="latitude" pageId="1" pageNumber="182" precision="15" seconds="14" value="-6.870556">S 06°52′14″</geoCoordinate>
<geoCoordinate id="EE5D50424476FF9AFAE7EDD2A271FE47" degrees="106" direction="east" minutes="57" orientation="longitude" pageId="1" pageNumber="182" precision="15" seconds="09" value="106.9525">E 106° 57′09″</geoCoordinate>
, 
<date id="FFD710454476FF9AFC5AEDF3A523FE47" box="[900,1056,422,447]" pageId="1" pageNumber="182" value="2010-05-15">15 May 2010</date>
, Hartini and Dwibadra leg., 
<emphasis id="B91DEA974476FF9AFAB3EDF2A2E9FE27" italics="true" pageId="1" pageNumber="182">
ex 
<taxonomicName id="4C694D064476FF9AFCF7ED92A2E9FE27" authorityName="Lansberge" authorityYear="1874" box="[809,1002,455,478]" class="Insecta" family="Scarabaeidae" genus="Synapsis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="182" phylum="Arthropoda" rank="species" species="ritsemae">Synapsis ritsemae</taxonomicName>
</emphasis>
(
<taxonomicName id="4C694D064476FF9AFC29ED92A571FE26" authorityName="Linnaeus" authorityYear="1758" box="[1015,1138,455,479]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="182" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
: 
<taxonomicName id="4C694D064476FF9AFBA3ED92A40CFE26" authorityName="Latreille" authorityYear="1802" box="[1149,1295,455,479]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="182" phylum="Arthropoda" rank="family">Scarabaeidae</taxonomicName>
). 
<typeStatus id="54D288274476FF9AFAFFED92A480FE26" box="[1313,1411,455,479]" pageId="1" pageNumber="182" type="paratype">Paratype</typeStatus>
: 14$$, other data same as 
<typeStatus id="54D288274476FF9AFB99EDB2A5ABFE06" box="[1095,1192,487,511]" pageId="1" pageNumber="182" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BD636854476FF9AFCF7EE52A2C9FD99" blockId="1.[809,1414,325,737]" pageId="1" pageNumber="182">
<emphasis id="B91DEA974476FF9AFCF7EE52A27FFDE6" bold="true" box="[809,892,519,543]" pageId="1" pageNumber="182">Female</emphasis>
: Length of dorsal shield 1021 (880-1110), width at level of coxae II 613 (500-690) (n 
<emphasis id="B91DEA974476FF9AFB1AEE72A5DBFDC6" bold="true" box="[1220,1240,551,575]" pageId="1" pageNumber="182">=</emphasis>
15). Specimens reddish brown.
</paragraph>
<paragraph id="8BD636854476FF9AFCF7EE3DA486FD18" blockId="1.[809,1414,325,737]" pageId="1" pageNumber="182">
<emphasis id="B91DEA974476FF9AFCF7EE3DA286FD79" bold="true" box="[809,901,616,640]" pageId="1" pageNumber="182">Dorsum</emphasis>
(
<figureCitation id="13522A004476FF9AFC4CEE3DA2D1FD79" box="[914,978,616,640]" captionStart="Figs" captionStartId="1.[144,186,1952,1974]" captionTargetBox="[318,1234,800,1918]" captionTargetId="figure-243@1.[317,1240,794,1920]" captionTargetPageId="1" captionText="Figs. 1-6. Macrocheles gedeensis sp. nov., holotype female(MZB.Acar.7091.2). 1, dorsum; 2, venter; 3, ventral view of gnathosoma; 4, epi- stome; 5, chelicerae; 6, sacculus foemineus." figureDoi="http://doi.org/10.5281/zenodo.13161082" httpUri="https://zenodo.org/record/13161082/files/figure.png" pageId="1" pageNumber="182">Fig. 1</figureCitation>
): Dorsal shield oval, attenuated posteri- orly, surface ornamented with distinct reticulate pattern and punctation; lateral margin smooth; shield with 28 pairs of dorsal setae and 22 pairs of pores; setae j1-4
</paragraph>
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<paragraph id="8BD636854476FF9AFF4EEBF5A31CF828" blockId="1.[144,1414,1952,2001]" pageId="1" pageNumber="182">
<emphasis id="B91DEA974476FF9AFF4EEBF5A1E4F84F" bold="true" box="[144,231,1952,1974]" pageId="1" pageNumber="182">Figs. 1-6.</emphasis>
<taxonomicName id="4C694D064476FF9AFF33EBF5A0BCF84C" authorityName="Hartini &amp; Dwibadra" authorityYear="2017" box="[237,447,1952,1973]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="182" phylum="Arthropoda" rank="species" species="gedeensis" status="sp. nov.">
<emphasis id="B91DEA974476FF9AFF33EBF5A0BCF84C" box="[237,447,1952,1973]" italics="true" pageId="1" pageNumber="182">Macrocheles gedeensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57EC4476FF9AFE1AEBF4A30EF84F" box="[452,525,1953,1974]" pageId="1" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
, holotype female (MZB.Acar.7091.2). 1, dorsum; 2, venter; 3, ventral view of gnathosoma; 4, epistome; 5, chelicerae; 6, sacculus foemineus.
</paragraph>
</caption>
<paragraph id="8BD636854475FF99FEF7ED50A558FEDA" blockId="2.[297,720,261,291]" box="[297,1115,261,291]" lastBlockId="2.[1091,1115,261,291]" pageId="2" pageNumber="183">
<emphasis id="B91DEA974475FF99FEF7ED50A558FEDA" bold="true" box="[297,1115,261,291]" pageId="2" pageNumber="183">A B C</emphasis>
</paragraph>
<caption id="DF16660D4475FF99FF70EE76A07CFDAA" ID-DOI="http://doi.org/10.5281/zenodo.13161084" ID-Zenodo-Dep="13161084" httpUri="https://zenodo.org/record/13161084/files/figure.png" pageId="2" pageNumber="183" startId="2.[174,207,547,569]" targetBox="[245,1376,280,509]" targetPageId="2" targetType="figure">
<paragraph id="8BD636854475FF99FF70EE76A07CFDAA" blockId="2.[174,1444,547,595]" pageId="2" pageNumber="183">
<emphasis id="B91DEA974475FF99FF70EE76A1E8FDC1" bold="true" box="[174,235,547,569]" pageId="2" pageNumber="183">Fig. 7.</emphasis>
Epigynial setae of A. 
<taxonomicName id="4C694D064475FF99FE66EE76A388FDC1" authorityName="Hartini &amp; Dwibadra" authorityYear="2017" box="[440,651,547,568]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="183" phylum="Arthropoda" rank="species" species="gedeensis">
<emphasis id="B91DEA974475FF99FE66EE76A388FDC1" box="[440,651,547,568]" italics="true" pageId="2" pageNumber="183">Macrocheles gedeensis</emphasis>
</taxonomicName>
(pilose distally), B. 
<taxonomicName id="4C694D064475FF99FC98EE76A2C3FDC1" authorityName="Takaku and Hartini" authorityYear="2001" box="[838,960,547,568]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="183" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="B91DEA974475FF99FC98EE76A2C3FDC1" box="[838,960,547,568]" italics="true" pageId="2" pageNumber="183">M. monticola</emphasis>
</taxonomicName>
(simple), and C. 
<taxonomicName id="4C694D064475FF99FB80EE76A5D5FDC1" authorityName="Hartini, Dwibadra and Takaku" authorityYear="2007" box="[1118,1238,547,568]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="183" phylum="Arthropoda" rank="species" species="persimilis">
<emphasis id="B91DEA974475FF99FB80EE76A5D5FDC1" box="[1118,1238,547,568]" italics="true" pageId="2" pageNumber="183">M. persimilis</emphasis>
</taxonomicName>
(simple). Scale bar: a, c 
<emphasis id="B91DEA974475FF99FF65EE68A1CFFDAB" bold="true" box="[187,204,573,594]" pageId="2" pageNumber="183">=</emphasis>
100 μm, b 
<emphasis id="B91DEA974475FF99FEF2EE68A03EFDAB" bold="true" box="[300,317,573,594]" pageId="2" pageNumber="183">=</emphasis>
50 μm.
</paragraph>
</caption>
<paragraph id="8BD636854475FF99FF70EECFA0FFFB8C" blockId="2.[174,779,665,1981]" pageId="2" pageNumber="183">
pilose distally; j5, j6, z1, z5, z6, and J2 simple, but in some cases z1 and z6 slightly pilose; J5 bipectinate entirely; other dorsal setae pilose in distal half to distal 2/3. 
<emphasis id="B91DEA974475FF99FF70EEAEA1FEFCEA" bold="true" box="[174,253,763,787]" pageId="2" pageNumber="183">Venter</emphasis>
(
<figureCitation id="13522A004475FF99FED2EEAFA056FCEA" box="[268,341,762,787]" captionStart="Figs" captionStartId="1.[144,186,1952,1974]" captionTargetBox="[318,1234,800,1918]" captionTargetId="figure-243@1.[317,1240,794,1920]" captionTargetPageId="1" captionText="Figs. 1-6. Macrocheles gedeensis sp. nov., holotype female(MZB.Acar.7091.2). 1, dorsum; 2, venter; 3, ventral view of gnathosoma; 4, epi- stome; 5, chelicerae; 6, sacculus foemineus." figureDoi="http://doi.org/10.5281/zenodo.13161082" httpUri="https://zenodo.org/record/13161082/files/figure.png" pageId="2" pageNumber="183">Fig. 2</figureCitation>
): Length of sternal shield, length 192 (170-207.5), width at level of coxae II 167 (150-175) (n 
<emphasis id="B91DEA974475FF99FF16EF6EA1DFFCAA" bold="true" box="[200,220,827,851]" pageId="2" pageNumber="183">=</emphasis>
15); shield ornamented with lines and punctations; linea angulata (l.ang.), two linea arcuata (l.arc.), linea oblique anterior (l.o.a.), linea media transversa (l.m.t.) present and with punctations; l.m.t. not straight; anterior l.arc. faint; linea oblique posteriore (l.o.p.) distinct, with punctation, bifurcated, and connect to l.m.t. Epigynial shield ornamented with arched lines and punctations; shield with a pair of pilose distally setae on posteri- or corner. Length of epigynial shield 180 (162.5-195), width 216 (185-237.5) (n 
<emphasis id="B91DEA974475FF99FE64E808A0CDFB8C" bold="true" box="[442,462,1117,1141]" pageId="2" pageNumber="183">=</emphasis>
15).
</paragraph>
<paragraph id="8BD636854475FF99FF1BE82BA0BAFA61" blockId="2.[174,779,665,1981]" pageId="2" pageNumber="183">
Ventrianal shield pentagonal, shield ornamented with semiconcentric line punctations; length 329 (267.5- 370), width 311 (247.5-365) (n 
<emphasis id="B91DEA974475FF99FE24E8EBA30DFB2F" bold="true" box="[506,526,1214,1238]" pageId="2" pageNumber="183">=</emphasis>
15); shield with 3 pairs of preanal simple setae, pair of paranal simple setae, and 1 postanal pilose seta. Cribrum located posterior to postanal seta. Ophisthogastric setae simple and pilose. A pair of metapodal shield oblong. Postcoxal pore free from podal shield. Anterior extremities of peritreme located at level of setae z1.
</paragraph>
<paragraph id="8BD636854475FF99FF70E9F5A052F980" blockId="2.[174,779,665,1981]" pageId="2" pageNumber="183">
<emphasis id="B91DEA974475FF99FF70E9F5A043FA41" bold="true" box="[174,320,1440,1464]" pageId="2" pageNumber="183">Gnathosoma</emphasis>
(
<figureCitation id="13522A004475FF99FE90E9F5A091FA41" box="[334,402,1440,1464]" captionStart="Figs" captionStartId="1.[144,186,1952,1974]" captionTargetBox="[318,1234,800,1918]" captionTargetId="figure-243@1.[317,1240,794,1920]" captionTargetPageId="1" captionText="Figs. 1-6. Macrocheles gedeensis sp. nov., holotype female(MZB.Acar.7091.2). 1, dorsum; 2, venter; 3, ventral view of gnathosoma; 4, epi- stome; 5, chelicerae; 6, sacculus foemineus." figureDoi="http://doi.org/10.5281/zenodo.13161082" httpUri="https://zenodo.org/record/13161082/files/figure.png" pageId="2" pageNumber="183">Fig. 3</figureCitation>
): Developed and sclerotized. Epistome (
<figureCitation id="13522A004475FF99FF24E995A039FA21" box="[250,314,1472,1496]" captionStart="Figs" captionStartId="1.[144,186,1952,1974]" captionTargetBox="[318,1234,800,1918]" captionTargetId="figure-243@1.[317,1240,794,1920]" captionTargetPageId="1" captionText="Figs. 1-6. Macrocheles gedeensis sp. nov., holotype female(MZB.Acar.7091.2). 1, dorsum; 2, venter; 3, ventral view of gnathosoma; 4, epi- stome; 5, chelicerae; 6, sacculus foemineus." figureDoi="http://doi.org/10.5281/zenodo.13161082" httpUri="https://zenodo.org/record/13161082/files/figure.png" pageId="2" pageNumber="183">Fig. 4</figureCitation>
) and chelicera (
<figureCitation id="13522A004475FF99FE3CE995A321FA21" box="[482,546,1472,1496]" captionStart="Figs" captionStartId="1.[144,186,1952,1974]" captionTargetBox="[318,1234,800,1918]" captionTargetId="figure-243@1.[317,1240,794,1920]" captionTargetPageId="1" captionText="Figs. 1-6. Macrocheles gedeensis sp. nov., holotype female(MZB.Acar.7091.2). 1, dorsum; 2, venter; 3, ventral view of gnathosoma; 4, epi- stome; 5, chelicerae; 6, sacculus foemineus." figureDoi="http://doi.org/10.5281/zenodo.13161082" httpUri="https://zenodo.org/record/13161082/files/figure.png" pageId="2" pageNumber="183">Fig. 5</figureCitation>
) fixed digit with simple dorsal seta, robust median tooth, a distal tooth, 
<emphasis id="B91DEA974475FF99FD09E9B5A1FEF9E0" italics="true" pageId="2" pageNumber="183">pilus dentilis</emphasis>
and terminal hook; movable digit with bidentate tooth, small distal tooth and terminal hook. Length of fixed digit 259 (225-272.5), and movable digit 105 (100- 112.5) (n 
<emphasis id="B91DEA974475FF99FECEEA34A027F980" bold="true" box="[272,292,1633,1657]" pageId="2" pageNumber="183">=</emphasis>
15).
</paragraph>
<paragraph id="8BD636854475FF99FF70EAD7A09AF8C2" blockId="2.[174,779,665,1981]" pageId="2" pageNumber="183">
<emphasis id="B91DEA974475FF99FF70EAD7A1E4F963" bold="true" box="[174,231,1666,1690]" pageId="2" pageNumber="183">Legs</emphasis>
: Most leg segments with simple, pilose, and plumose setae, except for coxa, trochanter, and tarsus I with simple setae; and coxa, genu IV with pilose setae. Leg length (except ambulacrum, n 
<emphasis id="B91DEA974475FF99FDC0EAB7A331F903" bold="true" box="[542,562,1762,1786]" pageId="2" pageNumber="183">=</emphasis>
15); leg I, 763 (640- 930); leg II, 720 (580-850); leg III, 738 (680-800); leg IV, 1159 (1040-1260).
</paragraph>
<paragraph id="8BD636854475FF99FF70EB16A318F865" blockId="2.[174,779,665,1981]" pageId="2" pageNumber="183">
<emphasis id="B91DEA974475FF99FF70EB16A08BF8A2" bold="true" box="[174,392,1859,1883]" pageId="2" pageNumber="183">Sacculus foemineus</emphasis>
: (
<figureCitation id="13522A004475FF99FE43EB16A0DFF8A2" box="[413,476,1859,1883]" captionStart="Figs" captionStartId="1.[144,186,1952,1974]" captionTargetBox="[318,1234,800,1918]" captionTargetId="figure-243@1.[317,1240,794,1920]" captionTargetPageId="1" captionText="Figs. 1-6. Macrocheles gedeensis sp. nov., holotype female(MZB.Acar.7091.2). 1, dorsum; 2, venter; 3, ventral view of gnathosoma; 4, epi- stome; 5, chelicerae; 6, sacculus foemineus." figureDoi="http://doi.org/10.5281/zenodo.13161082" httpUri="https://zenodo.org/record/13161082/files/figure.png" pageId="2" pageNumber="183">Fig. 6</figureCitation>
). Pair of sacculi fused; small cornu rounded distally and sclerotized; spermateca oval. 
<emphasis id="B91DEA974475FF99FF70EBD1A09EF865" bold="true" box="[174,413,1924,1948]" pageId="2" pageNumber="183">Male and other stage</emphasis>
: Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C373650E4475FF99FF70EBF0A28BFD4B" box="[174,904,666,1981]" pageId="2" pageNumber="183" type="etymology">
<paragraph id="8BD636854475FF99FF70EBF0A28BFD4B" blockId="2.[174,779,665,1981]" box="[174,904,666,1981]" lastBlockId="2.[839,1444,666,1271]" pageId="2" pageNumber="183">
<emphasis id="B91DEA974475FF99FF70EBF0A02AF844" bold="true" box="[174,297,1957,1981]" pageId="2" pageNumber="183">Etymology</emphasis>
: The species name is derived from 
<typeStatus id="54D288274475FF99FD68EBF0A3E7F844" box="[694,740,1957,1981]" pageId="2" pageNumber="183">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C373650E4475FF99FC99EEEFA480FB0E" pageId="2" pageNumber="183" type="discussion">
<paragraph id="8BD636854475FF99FC99EEEFA5F7FBEC" blockId="2.[839,1444,666,1271]" pageId="2" pageNumber="183">
<emphasis id="B91DEA974475FF99FC99EEEFA2B3FD2B" bold="true" box="[839,944,698,722]" pageId="2" pageNumber="183">Remarks</emphasis>
: This species is very similar to 
<taxonomicName id="4C694D064475FF99FAC7EEEFA467FD0B" authority="Hartini, Dwibadra and Takaku, 2007" authorityName="Hartini, Dwibadra and Takaku" authorityYear="2007" class="Arachnida" family="Macrochelidae" genus="Macrocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="183" phylum="Arthropoda" rank="species" species="persimilis">
<emphasis id="B91DEA974475FF99FAC7EEEFA2B5FD0B" italics="true" pageId="2" pageNumber="183">Macrocheles persimilis</emphasis>
<bibRefCitation id="EFF84B744475FF99FC1EEE8FA467FD0B" author="Hartini, S. &amp; D. Dwibadra &amp; G. Takaku" box="[960,1380,730,754]" pageId="2" pageNumber="183" pagination="73 - 96" refId="ref6646" refString="Hartini, S., D. Dwibadra and G. Takaku. 2007. Macrochelid mites (Acari: Gamasida: Macrochelidae) associated with dung beetles in Sulawesi, Indonesia. The Acarological Society of Japan 16 (2): 73 - 96." type="journal article" year="2007">Hartini, Dwibadra and Takaku, 2007</bibRefCitation>
</taxonomicName>
from 
<collectingRegion id="49ADF8674475FF99FC99EEAFA2A9FCEB" box="[839,938,762,786]" country="Indonesia" name="Sulawesi Utara" pageId="2" pageNumber="183">Sulawesi</collectingRegion>
, 
<collectingCountry id="F37E76154475FF99FC68EEAFA523FCEB" box="[950,1056,762,786]" name="Indonesia" pageId="2" pageNumber="183">Indonesia</collectingCountry>
, in its dorsal setae and sternal ornamentation. However, 
<taxonomicName id="4C694D064475FF99FBE6EF4EA43BFCCA" authorityName="Hartini &amp; Dwibadra" authorityYear="2017" box="[1080,1336,795,819]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="183" phylum="Arthropoda" rank="species" species="gedeensis">
<emphasis id="B91DEA974475FF99FBE6EF4EA43BFCCA" box="[1080,1336,795,819]" italics="true" pageId="2" pageNumber="183">Macrocheles gedeensis</emphasis>
</taxonomicName>
is distinguishable from 
<taxonomicName id="4C694D064475FF99FC30EF69A578FCAA" authorityName="Hartini, Dwibadra and Takaku" authorityYear="2007" box="[1006,1147,827,851]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="183" phylum="Arthropoda" rank="species" species="persimilis">
<emphasis id="B91DEA974475FF99FC30EF69A578FCAA" box="[1006,1147,827,851]" italics="true" pageId="2" pageNumber="183">M. persimilis</emphasis>
</taxonomicName>
by the following characters as shown in 
<figureCitation id="13522A004475FF99FC0EEF0EA51AFC8A" box="[976,1049,859,883]" captionStart="Fig" captionStartId="2.[174,207,547,569]" captionTargetBox="[245,1376,280,509]" captionTargetId="figure-719@2.[216,1401,249,514]" captionTargetPageId="2" captionText="Fig. 7. Epigynial setae of A. Macrocheles gedeensis (pilose distally), B. M. monticola (simple), and C. M. persimilis (simple). Scale bar: a, c=100μm, b=50μm." figureDoi="http://doi.org/10.5281/zenodo.13161084" httpUri="https://zenodo.org/record/13161084/files/figure.png" pageId="2" pageNumber="183">Figs. 7</figureCitation>
and 
<figureCitation id="13522A004475FF99FB8EEF0EA563FC8A" box="[1104,1120,859,883]" captionStart="Fig" captionStartId="3.[144,177,766,788]" captionTargetBox="[214,1343,257,733]" captionTargetId="figure-337@3.[214,1343,257,733]" captionTargetPageId="3" captionText="Fig. 8. Sternal shield of A. Macrocheles gedeensis with l.o.p. connected to l.m.t. and B. of M. persimilis which is disjunct to l.m.t." figureDoi="http://doi.org/10.5281/zenodo.13161088" httpUri="https://zenodo.org/record/13161088/files/figure.png" pageId="2" pageNumber="183">8</figureCitation>
: (corresponding conditions of 
<taxonomicName id="4C694D064475FF99FC99EF29A2DBFC6A" authorityName="Hartini, Dwibadra and Takaku" authorityYear="2007" box="[839,984,891,915]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="183" phylum="Arthropoda" rank="species" species="persimilis">
<emphasis id="B91DEA974475FF99FC99EF29A2DBFC6A" box="[839,984,891,915]" italics="true" pageId="2" pageNumber="183">M. persimilis</emphasis>
</taxonomicName>
in parentheses): 1) epigynial setae pilose (simple); 2) linea oblique posterior (l.o.p.) bifurcated and connected to linea media transversa (l.m.t.) (not bifurcated and disjunct to l.m.t; 3) length of dorsal shield more than 1000 μm (less than 1000 μm).
</paragraph>
<paragraph id="8BD636854475FF99FC80E848A480FB0E" blockId="2.[839,1444,666,1271]" pageId="2" pageNumber="183">
The species also resembles 
<taxonomicName id="4C694D064475FF99FB42E848A559FBAC" authority="Takaku and Hartini, 2001" authorityName="Takaku and Hartini" authorityYear="2001" class="Arachnida" family="Macrochelidae" genus="Macrocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="183" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="B91DEA974475FF99FB42E848A4A7FBCC" box="[1180,1444,1053,1077]" italics="true" pageId="2" pageNumber="183">Macrocheles monticola</emphasis>
<bibRefCitation id="EFF84B744475FF99FC99E868A559FBAC" author="Takaku, G. &amp; S. Hartini" box="[839,1114,1085,1109]" pageId="2" pageNumber="183" pagination="323 - 345" refId="ref7343" refString="Takaku, G., and S. Hartini. 2001. Macrochelid mites (Arachnida: Acari: Macrochelidae: Glyptholaspis, Macrocheles, Neopodocinum) associated with dung beetles in Bali, Indonesia. Species Diversity 6 (4): 323 - 345." type="journal article" year="2001">Takaku and Hartini, 2001</bibRefCitation>
</taxonomicName>
, recorded from 
<collectingRegion id="49ADF8674475FF99FADDE868A431FBAC" box="[1283,1330,1085,1109]" country="Indonesia" name="Bali" pageId="2" pageNumber="183">Bali</collectingRegion>
, 
<collectingCountry id="F37E76154475FF99FAE2E868A4A7FBAC" box="[1340,1444,1085,1109]" name="Indonesia" pageId="2" pageNumber="183">Indonesia</collectingCountry>
in the sternal ornamentation but it differs from the latter in the following characteristics of the female (corresponding conditions of 
<taxonomicName id="4C694D064475FF99FBDCE8CAA58CFB4F" authorityName="Takaku and Hartini" authorityYear="2001" box="[1026,1167,1182,1206]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="183" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="B91DEA974475FF99FBDCE8CAA58CFB4F" box="[1026,1167,1182,1206]" italics="true" pageId="2" pageNumber="183">M. monticola</emphasis>
</taxonomicName>
in parentheses), 1) epigynial setae pilose (simple), as are shown in 
<figureCitation id="13522A004475FF99FAD9E8EBA44BFB2F" box="[1287,1352,1214,1238]" captionStart="Fig" captionStartId="2.[174,207,547,569]" captionTargetBox="[245,1376,280,509]" captionTargetId="figure-719@2.[216,1401,249,514]" captionTargetPageId="2" captionText="Fig. 7. Epigynial setae of A. Macrocheles gedeensis (pilose distally), B. M. monticola (simple), and C. M. persimilis (simple). Scale bar: a, c=100μm, b=50μm." figureDoi="http://doi.org/10.5281/zenodo.13161084" httpUri="https://zenodo.org/record/13161084/files/figure.png" pageId="2" pageNumber="183">Fig. 7</figureCitation>
; 
<figureCitation id="13522A004475FF99FA8DE8EBA462FB2F" box="[1363,1377,1214,1238]" captionStart="Figs" captionStartId="1.[144,186,1952,1974]" captionTargetBox="[318,1234,800,1918]" captionTargetId="figure-243@1.[317,1240,794,1920]" captionTargetPageId="1" captionText="Figs. 1-6. Macrocheles gedeensis sp. nov., holotype female(MZB.Acar.7091.2). 1, dorsum; 2, venter; 3, ventral view of gnathosoma; 4, epi- stome; 5, chelicerae; 6, sacculus foemineus." figureDoi="http://doi.org/10.5281/zenodo.13161082" httpUri="https://zenodo.org/record/13161082/files/figure.png" pageId="2" pageNumber="183">2</figureCitation>
) most of dorsal setae pilose (simple), as are shown in 
<figureCitation id="13522A004475FF99FAE0E88BA47CFB0E" box="[1342,1407,1246,1271]" captionStart="Fig" captionStartId="3.[144,177,1285,1307]" captionTargetBox="[199,1359,856,1252]" captionTargetId="figure-364@3.[197,1361,852,1253]" captionTargetPageId="3" captionText="Fig. 9. Dorsal shield of A. Macrocheles gedeensis with pilose setae and B. of M. monticola with simple setae. Scale bar: 100μm." figureDoi="http://doi.org/10.5281/zenodo.13161090" httpUri="https://zenodo.org/record/13161090/files/figure.png" pageId="2" pageNumber="183">Fig. 9</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>