<documentid="010C669F719C9A8B7FE71DC6CAEDD647"ID-DOI="10.11646/phytotaxa.651.1.1"ID-ISSN="1179-3163"ID-Zenodo-Dep="13216497"IM.bibliography_approvedBy="felipe"IM.illustrations_approvedBy="felipe"IM.materialsCitations_approvedBy="felipe"IM.tables_requiresApprovalFor="GgImagineBatch"IM.taxonomicNames_approvedBy="felipe"IM.treatmentCitations_approvedBy="felipe"IM.treatments_approvedBy="felipe"checkinTime="1722602978082"checkinUser="felipe"docAuthor="Valdemarin, Karinne Sampaio, Mazine, Fiorella F. & Souza, Vinicius Castro"docDate="2024"docId="03BD8E6EFFA7FF94248CFEFEFD87FF00"docLanguage="en"docName="phytotaxa.651.1.1.pdf"docOrigin="Phytotaxa 651 (1)"docSource="http://dx.doi.org/10.11646/phytotaxa.651.1.1"docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.4:Phytotaxa.2014-.monograph"docStyleId="F08184CE06D8A97EA3E6DE35D99648B0"docStyleName="Phytotaxa.2014-.monograph"docStyleVersion="4"docTitle="Eugenia pachnantha O. Berg 1859"docType="treatment"docVersion="2"lastPageNumber="56"masterDocId="FF84F616FF90FFAC2404FFACFFA7FFDC"masterDocTitle="Eugenia (Myrtaceae) from Reserva Natural Vale, Espírito Santo, a center of plant endemism in the Brazilian Atlantic Rainforest"masterLastPageNumber="79"masterPageNumber="1"pageNumber="56"updateTime="1722809275918"updateUser="ExternalLinkService">
<mods:titleid="E87EAA2D307CF70A12109FCC0D46322B">Eugenia (Myrtaceae) from Reserva Natural Vale, Espírito Santo, a center of plant endemism in the Brazilian Atlantic Rainforest</mods:title>
<mods:affiliationid="5F32544E02E2EDB011DBEE49F589A77A">Departamento de Ciências Biológicas, Escola Superior de Agricultura “ Luiz de Queiroz ”, Universidade de São Paulo, Caixa Postal 9, 13418 - 900, Piracicaba, SP, Brazil. & kvaldemarin @ gmail. com; https: // orcid. org / 0000 - 0002 - 9564 - 1163</mods:affiliation>
<mods:affiliationid="1E658252466C1790F47DDC7DDA5DCD37">Departamento de Ciências Ambientais, Universidade Federal de São Carlos - Campus Sorocaba, Rod. João Leme dos Santos, km 110, 18052 - 780, Sorocaba, SP, Brazil. & fiorella @ ufscar. br; https: // orcid. org / 0000 - 0002 - 2604 - 6088</mods:affiliation>
<mods:affiliationid="10B5AB647EA0745CDDC5888E580829D4">Departamento de Ciências Biológicas, Escola Superior de Agricultura “ Luiz de Queiroz ”, Universidade de São Paulo, Caixa Postal 9, 13418 - 900, Piracicaba, SP, Brazil. & Departamento de Ciências Biológicas, Escola Superior de Agricultura “ Luiz de Queiroz ”, Universidade de São Paulo, Caixa Postal 9, 13418 - 900, Piracicaba, SP, Brazil. & vcsouza @ usp. br; https: // orcid. org / 0000 - 0002 - 3733 - 7892 * Corresponding author</mods:affiliation>
<figureCitationid="132F23FDFFA7FF9B269AFEFEFCA1FEB0"box="[670,774,338,364]"captionStart="FIGURE 22"captionStartId="53.[136,229,1759,1781]"captionTargetBox="[151,1433,194,1733]"captionTargetId="figure-18@53.[150,1435,193,1735]"captionTargetPageId="53"captionText="FIGURE 22. Fruiting branch of Eugenia neosilvestris (K.S. Valdemarin 344) B. Flowering branch of Eugenia ochracea (K.S. Valdemarin 467), C. Fruiting branch of Eugenia pachnantha (K.S. Valdemarin 776), D. Flowering branch of Eugenia pisiformis (G. Siqueira, Vale S.A.)."figureDoi="http://doi.org/10.5281/zenodo.13216533"httpUri="https://zenodo.org/record/13216533/files/figure.png"pageId="55"pageNumber="56">Fig. 22C</figureCitation>
, elliptic or rare narrow-lanceolate, discolorous when dry, lighter abaxially, not glaucous and glabrous on both surfaces; base acute, obtuse or attenuate; apex acute; midvein sulcate, plane or rare slightly raised adaxially and raised abaxially, glabrous on both surfaces; secondary veins 8–12 at each side, raised on both surfaces, first pair confluent with the innermost marginal vein; marginal veins two, the innermost
, ovate or oblong, apex obtuse, puberulent or glabrous and ciliate; petals 4, obovate, oil glands evident; staminal ring puberulent; stamens with filaments
<collectingDateid="EFEEE050FFA7FF9B248CFC52FEE2FBC4"box="[136,325,1022,1049]"pageId="55"pageNumber="56"value="2017-01-26">26 January 2017</collectingDate>
<materialsCitationid="3B7C3525FFA7FF9B26C2FC52FD8AFBE0"collectingDate="2016-12-10"collectorName="K. S. Valdemarin"country="Brazil"county="Linhares"location="Estrada Roxinho"municipality="Aceiro"pageId="55"pageNumber="56"specimenCount="1"stateProvince="Espirito Santo">
<collectingDateid="EFEEE050FFA7FF9B20ACFC52FA26FBC4"box="[1192,1409,1022,1048]"pageId="55"pageNumber="56"value="2016-12-10">10 December 2016</collectingDate>
<materialsCitationid="3B7C3525FFA7FF9B263DFB8EFA01FBE0"box="[569,1446,1058,1085]"collectingDate="2004-04-12"collectorName="Folli"country="Brazil"county="Linhares"location="Aceiro do Viveiro"municipality="Aceiro"pageId="55"pageNumber="56"specimenCount="1"stateProvince="Espirito Santo">
<locationid="8ECB69A3FFA7FF9B267CFB8EFCE2FBE0"LSID="urn:lsid:plazi:treatment:03BD8E6EFFA7FF94248CFEFEFD87FF00:8ECB69A3FFA7FF9B267CFB8EFCE2FBE0"box="[632,837,1058,1084]"country="Brazil"name="Aceiro do Viveiro"pageId="55"pageNumber="56"stateProvince="Espirito Santo">Aceiro do Viveiro</location>
<collectingDateid="EFEEE050FFA7FF9B2754FB8EFC48FBE0"box="[848,1007,1058,1084]"pageId="55"pageNumber="56"value="2004-04-12">12 April 2004</collectingDate>
<collectingDateid="EFEEE050FFA7FF9B2566FBEAFD81FBBC"box="[354,550,1094,1120]"pageId="55"pageNumber="56"value="1983-10-31">31 October 1983</collectingDate>
<materialsCitationid="3B7C3525FFA7FF9B2078FBEAFCABFB58"collectingDate="2003-09-25"collectorName="D. A. Folli"country="Brazil"county="Linhares"location="Estrada Carneiro"municipality="Aceiro"pageId="55"pageNumber="56"specimenCount="1"stateProvince="Espirito Santo">
<materialsCitationid="3B7C3525FFA7FF9B271EFBC6FDC1FB74"collectingDate="2008-11-21"collectorName="G. S. Siqueira"country="Brazil"county="Linhares"location="Estrada Cinco Folhas"municipality="Aceiro"pageId="55"pageNumber="56"specimenCount="1"stateProvince="Espirito Santo">
<locationid="8ECB69A3FFA7FF9B275AFBC6FBF2FB58"LSID="urn:lsid:plazi:treatment:03BD8E6EFFA7FF94248CFEFEFD87FF00:8ECB69A3FFA7FF9B275AFBC6FBF2FB58"box="[862,1109,1130,1156]"country="Brazil"name="Estrada Cinco Folhas"pageId="55"pageNumber="56"stateProvince="Espirito Santo">Estrada Cinco Folhas</location>
<collectingDateid="EFEEE050FFA7FF9B2060FBC6FAE3FB58"box="[1124,1348,1130,1156]"pageId="55"pageNumber="56"value="2008-11-21">21 November 2008</collectingDate>
<materialsCitationid="3B7C3525FFA7FF9B2672FB22FAA3FB74"box="[630,1284,1166,1193]"collectingDate="2017-01-27"collectorName="Valdemarin"country="Brazil"county="Linhares"location="Estrada Cinco Folhas"municipality="Aceiro"pageId="55"pageNumber="56"specimenCount="1"stateProvince="Espirito Santo">
<collectingDateid="EFEEE050FFA7FF9B26BEFB22FCDAFB74"box="[698,893,1166,1193]"pageId="55"pageNumber="56"value="2017-01-27">27 January 2017</collectingDate>
<collectingDateid="EFEEE050FFA7FF9B2502FB1EFE02FB10"box="[262,421,1202,1228]"pageId="55"pageNumber="56"value="2006-04-07">07 April 2006</collectingDate>
<collectingDateid="EFEEE050FFA7FF9B27E8FB1EFB64FB10"box="[1004,1219,1202,1228]"pageId="55"pageNumber="56"value="2016-12-11">11 December 2016</collectingDate>
<collectingDateid="EFEEE050FFA7FF9B2560FB7AFDAEFB2C"box="[356,521,1238,1265]"pageId="55"pageNumber="56"value="2011-04-15">15 April 2011</collectingDate>
<collectingDateid="EFEEE050FFA7FF9B251FFB56FE5EFAC8"box="[283,505,1274,1300]"pageId="55"pageNumber="56"value="2006-11-08">08 November 2006</collectingDate>
<collectingDateid="EFEEE050FFA7FF9B270CFAB2FC61FAE4"box="[776,966,1310,1337]"pageId="55"pageNumber="56"value="2017-01-24">24 January 2017</collectingDate>
<materialsCitationid="3B7C3525FFA7FF9B26AEFAEEFF4FFA5C"collectingDate="2002-08-07"collectorName="D. A. Folli"country="Brazil"county="Linhares"location="Estrada Gavea"municipality="Aceiro"pageId="55"pageNumber="56"specimenCount="1"stateProvince="Espirito Santo">
<collectingDateid="EFEEE050FFA7FF9B279BFAEEFBF1FA80"box="[927,1110,1346,1372]"pageId="55"pageNumber="56"value="2002-08-07">07 August 2002</collectingDate>
<collectingDateid="EFEEE050FFA7FF9B253CFACAFDB4FA5C"box="[312,531,1382,1408]"pageId="55"pageNumber="56"value="2016-12-12">12 December 2016</collectingDate>
<materialsCitationid="3B7C3525FFA7FF9B2798FACAFDB5FA78"collectingDate="2016-12-12"collectorName="de Janeiro & K. S. Valdemarin"country="Brazil"county="Linhares"location="Estrada Jequitiba Rosa"municipality="Aceiro"pageId="55"pageNumber="56"specimenCount="1"stateProvince="Espirito Santo">
<collectingDateid="EFEEE050FFA7FF9B20AAFA26FAEBFA78"box="[1198,1356,1418,1444]"pageId="55"pageNumber="56"value="2011-04-20">20 April 2011</collectingDate>
<materialsCitationid="3B7C3525FFA7FF9B269BFA02FF4FFA30"collectingDate="2012-06-20"collectorName="G. S. Siqueira"country="Brazil"county="Linhares"location="Estrada Louro"municipality="Aceiro"pageId="55"pageNumber="56"specimenCount="1"stateProvince="Espirito Santo">
<collectingDateid="EFEEE050FFA7FF9B2792FA02FB92FA14"box="[918,1077,1454,1481]"pageId="55"pageNumber="56"value="2012-06-20">20 June 2012</collectingDate>
<locationid="8ECB69A3FFA7FF9B2530FA7EFD03FA30"LSID="urn:lsid:plazi:treatment:03BD8E6EFFA7FF94248CFEFEFD87FF00:8ECB69A3FFA7FF9B2530FA7EFD03FA30"box="[308,676,1490,1516]"country="Brazil"name="Estrada Macanaiba Pele de Sapo"pageId="55"pageNumber="56"stateProvince="Espirito Santo">Estrada Macanaíba Pele de Sapo</location>
<collectingDateid="EFEEE050FFA7FF9B26ABFA7EFCE5FA30"box="[687,834,1490,1517]"pageId="55"pageNumber="56"value="2003-07-15">15 July 2003</collectingDate>
<materialsCitationid="3B7C3525FFA7FF9B248CFA5AFBBAF9CC"box="[136,1053,1526,1553]"collectingDate="2016-12-11"collectorName="K. S. Valdemarin"country="Brazil"county="Linhares"location="Estrada Mantegueira"municipality="Aceiro"pageId="55"pageNumber="56"specimenCount="1"stateProvince="Espirito Santo">
<collectingDateid="EFEEE050FFA7FF9B25C1FA5AFD07F9CC"box="[453,672,1526,1552]"pageId="55"pageNumber="56"value="2016-12-11">11 December 2016</collectingDate>
<collectingDateid="EFEEE050FFA7FF9B2069FA5AFAEFF9CC"box="[1133,1352,1526,1552]"pageId="55"pageNumber="56"value="2016-12-11">11 December 2016</collectingDate>
<collectingDateid="EFEEE050FFA7FF9B25DAF9B6FD13F9E8"box="[478,692,1562,1588]"pageId="55"pageNumber="56"value="2016-12-11">11 December 2016</collectingDate>
<collectingDateid="EFEEE050FFA7FF9B206BF9B6FAEFF9E8"box="[1135,1352,1562,1588]"pageId="55"pageNumber="56"value="2006-11-14">14 November 2006</collectingDate>
<materialsCitationid="3B7C3525FFA7FF9B2558F9CEFB4BF9A0"box="[348,1260,1634,1661]"collectingDate="2017-01-23"collectorName="Valdemarin"country="Brazil"county="Linhares"location="Trilha da Peroba Amarela"municipality="Aceiro"pageId="55"pageNumber="56"specimenCount="1"stateProvince="Espirito Santo">
<locationid="8ECB69A3FFA7FF9B259FF9CEFD1CF9A0"LSID="urn:lsid:plazi:treatment:03BD8E6EFFA7FF94248CFEFEFD87FF00:8ECB69A3FFA7FF9B259FF9CEFD1CF9A0"box="[411,699,1634,1660]"country="Brazil"name="Trilha da Peroba Amarela"pageId="55"pageNumber="56"stateProvince="Espirito Santo">Trilha da Peróba Amarela</location>
<collectingDateid="EFEEE050FFA7FF9B26C1F9CEFC27F9A0"box="[709,896,1634,1661]"pageId="55"pageNumber="56"value="2017-01-23">23 January 2017</collectingDate>
<materialsCitationid="3B7C3525FFA7FF9B20F6F9CEFD96F97C"collectingDate="2017-01-23"collectorName="Valdemarin"country="Brazil"county="Linhares"location="Trilha da Peroba Amarela"municipality="Aceiro"pageId="55"pageNumber="56"specimenCount="1"stateProvince="Espirito Santo">
<materialsCitationid="3B7C3525FFA7FF9B2638F92AFB03F97C"box="[572,1188,1670,1697]"collectingDate="2017-01-23"collectorName="Valdemarin"country="Brazil"county="Linhares"location="Trilha da Peroba Amarela"municipality="Aceiro"pageId="55"pageNumber="56"specimenCount="1"stateProvince="Espirito Santo">
<collectingDateid="EFEEE050FFA7FF9B267FF92AFC92F97C"box="[635,821,1670,1697]"pageId="55"pageNumber="56"value="2017-01-23">23 January 2017</collectingDate>
<materialsCitationid="3B7C3525FFA7FF9B20AAF92AFE4AF918"collectingDate="2017-01-23"collectorName="Valdemarin"country="Brazil"county="Linhares"location="Trilha da Peroba Amarela"municipality="Aceiro"pageId="55"pageNumber="56"specimenCount="1"stateProvince="Espirito Santo">
<collectingDateid="EFEEE050FFA7FF9B20E9F92AFA00F97C"box="[1261,1447,1670,1697]"pageId="55"pageNumber="56"value="2017-01-23">23 January 2017</collectingDate>
<materialsCitationid="3B7C3525FFA7FF9B25FCF906FBC5F918"box="[504,1122,1706,1733]"collectingDate="2017-01-23"collectorName="Valdemarin"country="Brazil"county="Linhares"location="Trilha da Peroba Amarela"municipality="Aceiro"pageId="55"pageNumber="56"specimenCount="1"stateProvince="Espirito Santo">
<collectingDateid="EFEEE050FFA7FF9B263CF906FD55F918"box="[568,754,1706,1733]"pageId="55"pageNumber="56"value="2017-01-23">23 January 2017</collectingDate>
<materialsCitationid="3B7C3525FFA7FF9B2068F906FE6AF934"collectingDate="2016-12-13"collectorName="Valdemarin"country="Brazil"county="Linhares"location="Trilha da Peroba Amarela"municipality="Aceiro"pageId="55"pageNumber="56"specimenCount="1"stateProvince="Espirito Santo">
<collectingDateid="EFEEE050FFA7FF9B20A8F906FA25F918"box="[1196,1410,1706,1732]"pageId="55"pageNumber="56"value="2016-12-13">13 December 2016</collectingDate>
<materialsCitationid="3B7C3525FFA7FF9B25DDF962FA00F934"box="[473,1447,1742,1769]"collectingDate="2016-12-13"collectorName="Valdemarin"country="Brazil"county="Linhares"location="Trilha do Pequi Vinagreiro"municipality="Aceiro"pageId="55"pageNumber="56"specimenCount="1"stateProvince="Espirito Santo">
<locationid="8ECB69A3FFA7FF9B261EF962FCECF934"LSID="urn:lsid:plazi:treatment:03BD8E6EFFA7FF94248CFEFEFD87FF00:8ECB69A3FFA7FF9B261EF962FCECF934"box="[538,843,1742,1768]"country="Brazil"name="Trilha do Pequi Vinagreiro"pageId="55"pageNumber="56"stateProvince="Espirito Santo">Trilha do Pequi Vinagreiro</location>
<collectingDateid="EFEEE050FFA7FF9B2752F962FB88F934"box="[854,1071,1742,1768]"pageId="55"pageNumber="56"value="2016-12-13">13 December 2016</collectingDate>
<materialsCitationid="3B7C3525FFA7FF9B248CF95EFF57F8EC"collectingDate="1986-03-23"collectingDateMax="2017-01-27"collectingDateMin="1986-03-23"collectorName="K. S. Valdemarin & Sobral"country="Brazil"county="Linhares"location="Trilha do Pequi Vinagreiro"municipality="Aceiro"pageId="55"pageNumber="56"specimenCount="1"stateProvince="Espirito Santo">
<collectingDateid="EFEEE050FFA7FF9B24CFF95EFE2CF8D0"box="[203,395,1778,1805]"pageId="55"pageNumber="56"value="2017-01-27">27 January 2017</collectingDate>
<collectingDateid="EFEEE050FFA7FF9B279BF95EFBF5F8D0"box="[927,1106,1778,1804]"pageId="55"pageNumber="56"value="1986-03-23">23 March 1986</collectingDate>
<figureCitationid="132F23FDFFA7FF9B20CFF80AFAB7F81D"box="[1227,1296,1958,1985]"captionStart="FIGURE 5"captionStartId="9.[136,229,1993,2015]"captionTargetBox="[150,1434,193,1969]"captionTargetId="figure-18@9.[150,1435,193,1970]"captionTargetPageId="9"captionText="FIGURE 5. Phenological diagram of Eugenia species from Reserva Natural Vale, Espírito Santo, Brazil. Flowering periods are represented by dark red border triangles, fruiting in grey filled triangles, and “n” is the number of observations."figureDoi="http://doi.org/10.5281/zenodo.13216507"httpUri="https://zenodo.org/record/13216507/files/figure.png"pageId="55"pageNumber="56">Fig. 5</figureCitation>
due to its inflorescences in glomerules or fascicles, and bracteoles persistent in the fruit. The species is easily distinguished from all other species in RNV by its leaf blades with acute, obtuse or attenuate base, two marginal veins, inflorescences in glomerules or fascicles with pedicels up to