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<document id="33A320B8536AA76C7D803C409CBFBCEA" ID-DOI="10.5281/zenodo.5279282" ID-ISSN="0253-116X" ID-Zenodo-Dep="5279282" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630025850813" checkinUser="carolina" docAuthor="Jaeger, B. &amp; H, Ban" docDate="2009" docId="039F194AC346EC26FF061AC1FC36FEE2" docLanguage="en" docName="LinzerbiolBeitr.2009.41.2.1635-1657.pdf" docOrigin="Linzer biologische Beiträge 41 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Psychristus discretus ANDREWES. Habitus 1930" docType="treatment" docVersion="3" lastPageNumber="1644" masterDocId="FFA66132C341EC2FFFB41804FFDEFFA6" masterDocTitle="Two new species of Psychristus subgenus Psychristus ANDREWES 1930 from Southeast Asia, India and Pakistan, with additions to the Psychristus discretus group (Col., Carabidae)" masterLastPageNumber="1657" masterPageNumber="1635" pageNumber="1642" updateTime="1699137677022" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2216FCE49E76516346163ABB5AE8F2F0">Two new species of Psychristus subgenus Psychristus ANDREWES 1930 from Southeast Asia, India and Pakistan, with additions to the Psychristus discretus group (Col., Carabidae)</mods:title>
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<mods:namePart id="F8DCAE4F5E708475540564506B4E666E">Jaeger, B.</mods:namePart>
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<mods:namePart id="B85F2794972BC85D5B3F0DF415EABF8D">H, Ban</mods:namePart>
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<mods:title id="51A42F8E0B6DE177246FCAC3E0105101">Linzer biologische Beiträge</mods:title>
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<mods:date id="2C8461DC400662DA82A42C2DE9EF4D20">2009</mods:date>
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<mods:number id="8A7994F4EF15F3D670560BA97DD30410">2009-12-18</mods:number>
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<mods:number id="46BE99697A4A7F354D22F5BE3C5FD71A">41</mods:number>
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<mods:classification id="42911A3D16BF3027E447E743312884A6">journal article</mods:classification>
<mods:identifier id="6E6C89BF05AF9B8822C41DEF96977D00" type="DOI">10.5281/zenodo.5279282</mods:identifier>
<mods:identifier id="510D769D1588FF79BE9C75890F63A24C" type="ISSN">0253-116X</mods:identifier>
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<treatment id="039F194AC346EC26FF061AC1FC36FEE2" LSID="urn:lsid:plazi:treatment:039F194AC346EC26FF061AC1FC36FEE2" httpUri="http://treatment.plazi.org/id/039F194AC346EC26FF061AC1FC36FEE2" lastPageId="9" lastPageNumber="1644" pageId="7" pageNumber="1642">
<subSubSection id="C32CFBD7C346EC28FF061AC1FC6FFD7D" box="[178,945,709,731]" pageId="7" pageNumber="1642" type="nomenclature">
<paragraph id="8B89A85CC346EC28FF061AC1FC6FFD7D" blockId="7.[178,945,709,731]" box="[178,945,709,731]" pageId="7" pageNumber="1642">
<heading id="D0C11F30C346EC28FF061AC1FC6FFD7D" box="[178,945,709,731]" fontSize="8" level="2" pageId="7" pageNumber="1642" reason="2">
<taxonomicName id="4C36D3DFC346EC28FF061AC1FD52FD7D" ID-CoL="4PR2W" authority="ANDREWES 1930" authorityName="ANDREWES. Habitus" authorityYear="1930" box="[178,652,709,731]" class="Insecta" family="Carabidae" genus="Psychristus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1642" phylum="Arthropoda" rank="species" species="discretus">
<emphasis id="B942744EC346EC28FF061AC1FD52FD7D" bold="true" box="[178,652,709,731]" pageId="7" pageNumber="1642">
<emphasis id="B942744EC346EC28FF061AC1FEF7FD7D" bold="true" box="[178,297,709,731]" italics="true" pageId="7" pageNumber="1642">Psychristus</emphasis>
(s.str.)
<emphasis id="B942744EC346EC28FECF1AC1FE09FD7D" bold="true" box="[379,471,709,731]" italics="true" pageId="7" pageNumber="1642">discretus</emphasis>
<bibRefCitation id="EFA7D5ADC346EC28FE691AC1FD8BFD7C" author="ANDREWES H" box="[477,597,709,731]" pageId="7" pageNumber="1642" pagination="1 - 44" refId="ref8886" refString="ANDREWES H. E. (1930): The Carabidae of the third Mount Everest Expedition, 1924. - The Transactions of the Royal Entomological Society of London 78: 1 - 44 + 1 map." type="book chapter" year="1930">ANDREWES</bibRefCitation>
1930
</emphasis>
</taxonomicName>
(
<figureCitation id="130DB4D9C346EC28FD2E1AC1FD2FFD7D" box="[666,753,709,731]" captionStart="Figs 7-9" captionStartId="16.[178,218,1480,1500]" captionTargetBox="[183,1077,232,1449]" captionTargetId="figure-0@16.[174,1081,219,1617]" captionTargetPageId="16" captionText="Figs 7-9. Psychristus discretus ANDREWES. Habitus, head and pronotum. (7) Nepal, Basantapur; (8) Vietnam, Sa Pa; (9) India, Bomdila." figureDoi="http://doi.org/10.5281/zenodo.5279290" httpUri="https://zenodo.org/record/5279290/files/figure.png" pageId="7" pageNumber="1642">Figs 7-9</figureCitation>
,
<figureCitation id="130DB4D9C346EC28FD4F1AC1FCE7FD7D" box="[763,825,709,731]" captionStart="Figs 18-21" captionStartId="19.[202,242,1272,1292]" captionTargetBox="[256,1458,411,1190]" captionTargetId="figure-32@19.[229,1539,397,1270]" captionTargetPageId="19" captionText="Figs 18-21. Median lobe of aedoagus, lateral and dorsal aspect. (18-19) Psychristus discretus ANDREWES (India, Bomdila); (20-21) P. hibatai ITO (Paratype, Taiwan, Wushe)." figureDoi="http://doi.org/10.5281/zenodo.5279300" httpUri="https://zenodo.org/record/5279300/files/figure.png" pageId="7" pageNumber="1642">18-19</figureCitation>
,
<figureCitation id="130DB4D9C346EC28FCF71AC1FC81FD7D" box="[835,863,709,731]" captionStart="Figs" captionStartId="20.[178,220,1499,1519]" captionTargetBox="[198,1064,239,1458]" captionTargetId="figure-0@20.[178,1086,219,1505]" captionTargetPageId="20" captionText="Figs. 22-25. Female genitalia. Hemisternite and gonocoxa. (22) Psychristus dentatus nov.sp., (Holotype); (23) P. brunneus nov.sp. (Paratype, Laos, Ban Khoun Ngeun); (24) P. discretus ANDREWES (India, Bomdila); (25) P. shibatai ITO (Taiwan, Mt. Kwena)." figureDoi="http://doi.org/10.5281/zenodo.5279302" httpUri="https://zenodo.org/record/5279302/files/figure.png" pageId="7" pageNumber="1642">24</figureCitation>
,
<figureCitation id="130DB4D9C346EC28FCDD1AC1FC5AFD7D" box="[873,900,709,731]" captionStart="Figs 26-27" captionStartId="21.[178,218,1483,1503]" captionTargetBox="[178,1058,219,1459]" captionTargetId="figure-33@21.[178,1085,219,1460]" captionTargetPageId="21" captionText="Figs 26-27. Distribution. (26) P. dentatus nov.sp. (open circles), P. brunneus nov.sp. (filled circles) and P. shibatai ITO (filled squares); (27) P. discretus ANDREWES." figureDoi="http://doi.org/10.5281/zenodo.5279304" httpUri="https://zenodo.org/record/5279304/files/figure.png" pageId="7" pageNumber="1642">27</figureCitation>
,
<figureCitation id="130DB4D9C346EC28FC3B1AC1FC77FD7D" box="[911,937,709,731]" captionStart="Figs 28-32" captionStartId="22.[178,218,966,986]" captionTargetBox="[178,1085,219,947]" captionTargetId="figure-45@22.[178,1085,219,950]" captionTargetPageId="22" captionText="Figs 28-32. Mentum and definition of measurements. (28) Psychristus dentatus nov.sp. (Holotype); (29) P. brunneus nov.sp. (Paratype, India, Similipal); (30) P. discretus ANDREWES (Nepal, Lamjung Himal); (31) P. shibatai ITO (Taiwan, Mt. Kwena)." figureDoi="http://doi.org/10.5281/zenodo.5279306" httpUri="https://zenodo.org/record/5279306/files/figure.png" pageId="7" pageNumber="1642">30</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32CFBD7C346EC28FF061AE8FE39FC95" pageId="7" pageNumber="1642" type="reference_group">
<paragraph id="8B89A85CC346EC28FF061AE8FBE3FCBD" blockId="7.[178,1085,748,819]" pageId="7" pageNumber="1642">
<treatmentCitationGroup id="AB268F72C346EC28FF061AE8FBE3FCBD" pageId="7" pageNumber="1642">
<treatmentCitation id="0A978E4DC346EC28FF061AE8FDE6FCA6" author="ANDREWES H" box="[178,568,748,768]" page="22" pageId="7" pageNumber="1642" year="1930">
<taxonomicName id="4C36D3DFC346EC28FF061AE8FDE6FCA6" ID-CoL="4PR2W" authority="ANDREWES 1930: 22" authorityName="ANDREWES. Habitus" authorityPageNumber="22" authorityYear="1930" box="[178,568,748,768]" class="Insecta" family="Carabidae" genus="Psychristus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1642" phylum="Arthropoda" rank="species" species="discretus">
<emphasis id="B942744EC346EC28FF061AE8FEACFCA6" box="[178,370,748,768]" italics="true" pageId="7" pageNumber="1642">Psychristus discretus</emphasis>
<bibRefCitation id="EFA7D5ADC346EC28FEC31AE8FDE6FCA6" author="ANDREWES H" box="[375,568,748,768]" pageId="7" pageNumber="1642" pagination="1 - 44" refId="ref8886" refString="ANDREWES H. E. (1930): The Carabidae of the third Mount Everest Expedition, 1924. - The Transactions of the Royal Entomological Society of London 78: 1 - 44 + 1 map." type="book chapter" year="1930">ANDREWES 1930: 22</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="548D16FEC346EC28FDF71AE8FDB5FCA6" box="[579,619,748,768]" pageId="7" pageNumber="1642">type</typeStatus>
locality:
<collectingCountry id="F321E8CCC346EC28FD761AE8FD2AFCA6" box="[706,756,748,768]" name="India" pageId="7" pageNumber="1642">India</collectingCountry>
:
<collectingRegion id="49F266BEC346EC28FD481AE8FC9FFCA6" box="[764,833,748,768]" country="India" name="Sikkim" pageId="7" pageNumber="1642">Sikkim</collectingRegion>
: Phadam Chen).
<taxonomicName id="4C36D3DFC346EC28FF061B03FD7CFCBD" ID-CoL="N56M" authority="LINDROTH 1939: 117 - 119" authorityName="LINDROTH" authorityPageNumber="117 - 119" authorityYear="1939" box="[178,674,775,795]" class="Insecta" family="Carabidae" genus="Bradycellus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1642" phylum="Arthropoda" rank="species" species="ponderosus">
<emphasis id="B942744EC346EC28FF061B03FE4BFCBD" box="[178,405,775,795]" italics="true" pageId="7" pageNumber="1642">Bradycellus ponderosus</emphasis>
LINDROTH 1939: 117-119
</taxonomicName>
(
<typeStatus id="548D16FEC346EC28FD021B03FD00FCBD" box="[694,734,775,795]" pageId="7" pageNumber="1642">type</typeStatus>
locality:
<collectingCountry id="F321E8CCC346EC28FCF31B03FC59FCBD" box="[839,903,775,795]" name="Russia" pageId="7" pageNumber="1642">Russia</collectingCountry>
:
<collectingRegion id="49F266BEC346EC28FC2C1B03FC00FCBD" box="[920,990,775,795]" country="Russia" name="Kareliya" pageId="7" pageNumber="1642">Karelia</collectingRegion>
: &quot;Fennia,
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B89A85CC346EC28FF521B1BFE39FC95" blockId="7.[178,1085,748,819]" box="[230,487,799,819]" pageId="7" pageNumber="1642">Kuusamo, See Paanajärvi&quot;).</paragraph>
</subSubSection>
<subSubSection id="C32CFBD7C346EC26FF061B47FC36FEE2" lastPageId="9" lastPageNumber="1644" pageId="7" pageNumber="1642" type="description">
<paragraph id="8B89A85CC346EC28FF061B47FC98FC68" blockId="7.[178,1084,835,974]" pageId="7" pageNumber="1642">
The original description given by
<bibRefCitation id="EFA7D5ADC346EC28FDB81B47FD2FFCFF" author="ANDREWES H" box="[524,753,835,857]" pageId="7" pageNumber="1642" pagination="1 - 44" refId="ref8886" refString="ANDREWES H. E. (1930): The Carabidae of the third Mount Everest Expedition, 1924. - The Transactions of the Royal Entomological Society of London 78: 1 - 44 + 1 map." type="book chapter" year="1930">ANDREWES (1930: 22)</bibRefCitation>
and the re-description and illustrations given by
<bibRefCitation id="EFA7D5ADC346EC28FED11B65FDE1FCD1" author="JAEGER B" box="[357,575,865,887]" pageId="7" pageNumber="1642" pagination="63 - 93" refId="ref8994" refString="JAEGER B. (1997): Revision der Himalaya-Arten der Gattung Psychristus ANDREWES 1930 (Col., Carabidae) - Linzer biologische Beitrage 29 (1): 63 - 93." type="journal article" year="1997">JAEGER (1997: 70-71</bibRefCitation>
, 87, and 89) include most main characters important for recognition of the species. Therefore, only a diagnosis, including characters that separate the species from others of the
<taxonomicName id="4C36D3DFC346EC28FD891B98FD6BFC17" authorityName="ANDREWES. Habitus" authorityYear="1930" box="[573,693,923,945]" class="Insecta" family="Carabidae" genus="Psychristus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1642" phylum="Arthropoda" rank="species" species="discretus">
<emphasis id="B942744EC346EC28FD891B98FD6BFC17" box="[573,693,923,945]" italics="true" pageId="7" pageNumber="1642">P. discretus</emphasis>
</taxonomicName>
group, additional images and ratios of various body dimensions (see table 1 and 2) are presented below.
</paragraph>
<paragraph id="8B89A85CC346EC27FF061BDAFC5EFDB3" blockId="7.[178,1086,990,1597]" lastBlockId="8.[178,1085,219,533]" lastPageId="8" lastPageNumber="1643" pageId="7" pageNumber="1642">
D i a g n o s i s:Thelargestspeciesinthe
<taxonomicName id="4C36D3DFC346EC28FDCA1BDBFD29FC52" authorityName="ANDREWES. Habitus" authorityYear="1930" box="[638,759,990,1012]" class="Insecta" family="Carabidae" genus="Psychristus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1642" phylum="Arthropoda" rank="species" species="discretus">
<emphasis id="B942744EC346EC28FDCA1BDBFD29FC52" box="[638,759,990,1012]" italics="true" pageId="7" pageNumber="1642">P. discretus</emphasis>
</taxonomicName>
group with general appearance as figured (
<figureCitation id="130DB4D9C346EC28FE9E1BFFFEB2FBB7" box="[298,364,1019,1041]" captionStart="Figs 7-9" captionStartId="16.[178,218,1480,1500]" captionTargetBox="[183,1077,232,1449]" captionTargetId="figure-0@16.[174,1081,219,1617]" captionTargetPageId="16" captionText="Figs 7-9. Psychristus discretus ANDREWES. Habitus, head and pronotum. (7) Nepal, Basantapur; (8) Vietnam, Sa Pa; (9) India, Bomdila." figureDoi="http://doi.org/10.5281/zenodo.5279290" httpUri="https://zenodo.org/record/5279290/files/figure.png" pageId="7" pageNumber="1642">Fig. 7</figureCitation>
). Body length
<quantity id="4CCE05B9C346EC28FDBF1BFFFD5DFBB7" box="[523,643,1019,1041]" metricMagnitude="-3" metricUnit="m" metricValue="4.199999999999999" metricValueMax="4.6" metricValueMin="3.8" pageId="7" pageNumber="1642" unit="mm" value="4.199999999999999" valueMax="4.6" valueMin="3.8">3.8-4.6 mm</quantity>
and width
<quantity id="4CCE05B9C346EC28FD4A1BFFFCA7FBB7" box="[766,889,1019,1041]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="7" pageNumber="1642" unit="mm" value="1.75" valueMax="1.9" valueMin="1.6">1.6-1.9 mm</quantity>
. Ground colour of dorsal surface usually blackish brown, sometimes dark reddish brown, with elytral interval 1 (often indistinct or partly), pronotal base and apex medially, and lateral margins of elytra and pronotum paler reddish. Clypeus, labrum and mandibles reddish (with inner and partly outer margins, and apex of mandibles blackish), palpi yellowish. Ventral surface dark reddish- to blackish brown. Legs and antennae yellowish to pale reddish brown. Head including eyes (
<figureCitation id="130DB4D9C346EC28FD981CAFFD5FFB67" box="[556,641,1195,1217]" captionStart="Figs 7-9" captionStartId="16.[178,218,1480,1500]" captionTargetBox="[183,1077,232,1449]" captionTargetId="figure-0@16.[174,1081,219,1617]" captionTargetPageId="16" captionText="Figs 7-9. Psychristus discretus ANDREWES. Habitus, head and pronotum. (7) Nepal, Basantapur; (8) Vietnam, Sa Pa; (9) India, Bomdila." figureDoi="http://doi.org/10.5281/zenodo.5279290" httpUri="https://zenodo.org/record/5279290/files/figure.png" pageId="7" pageNumber="1642">Figs 7-9</figureCitation>
) of medium width, 0.75-0.81 times as wide as pronotum, with eyes distinctly prominent (head 1.54-1.63 times as wide as head between eyes). Apical margin of labrum usually weakly emarginated, sometimes almost rectilinear.
<collectingRegion id="49F266BEC346EC28FE9D1D07FEA0FABE" box="[297,382,1283,1304]" country="Myanmar" name="Chin" pageId="7" pageNumber="1642">Mentum</collectingRegion>
with epilobes (
<figureCitation id="130DB4D9C346EC28FD921D06FDAAFABE" box="[550,628,1282,1304]" captionStart="Figs 28-32" captionStartId="22.[178,218,966,986]" captionTargetBox="[178,1085,219,947]" captionTargetId="figure-45@22.[178,1085,219,950]" captionTargetPageId="22" captionText="Figs 28-32. Mentum and definition of measurements. (28) Psychristus dentatus nov.sp. (Holotype); (29) P. brunneus nov.sp. (Paratype, India, Similipal); (30) P. discretus ANDREWES (Nepal, Lamjung Himal); (31) P. shibatai ITO (Taiwan, Mt. Kwena)." figureDoi="http://doi.org/10.5281/zenodo.5279306" httpUri="https://zenodo.org/record/5279306/files/figure.png" pageId="7" pageNumber="1642">Fig. 30</figureCitation>
) strongly projecting forward and distinctly acuminate at apices. Mental tooth sharp, usually about half as long as epilobes. Ligular sclerite with apex, narrowly or widely rounded, sometimes somewhat acuminate. Mandibles of average size, not markedly prolonged and curved in apical half. Antennae 0.56-0.64 times as long as elytra and 1.70-2.04 times as long as pronotum, with antennomere 3 with scattered pubescence in apical half, at least with few additional setae. Pronotum with shape and punctation as figured (
<figureCitation id="130DB4D9C346EC28FD6B1DB6FCE9FA6E" box="[735,823,1458,1480]" captionStart="Figs 7-9" captionStartId="16.[178,218,1480,1500]" captionTargetBox="[183,1077,232,1449]" captionTargetId="figure-0@16.[174,1081,219,1617]" captionTargetPageId="16" captionText="Figs 7-9. Psychristus discretus ANDREWES. Habitus, head and pronotum. (7) Nepal, Basantapur; (8) Vietnam, Sa Pa; (9) India, Bomdila." figureDoi="http://doi.org/10.5281/zenodo.5279290" httpUri="https://zenodo.org/record/5279290/files/figure.png" pageId="7" pageNumber="1642">Figs 7-9</figureCitation>
), 1.23-1.35 times as wide as long and 1.24-1.33 times as wide as head. Base about 1.1-1.2 times as wide as apical margin between anterios angles and 0.83-0.91 times as wide as maximum pronotal width. Median line distinctly punctate in basal third, impunctate, or rarely very faintly punctate in medial third, and moderately punctate in apical third. Macropterous. Elytra (
<figureCitation id="130DB4D9C346EC28FC411E23FBEAF99B" box="[1013,1076,1575,1597]" captionStart="Figs 7-9" captionStartId="16.[178,218,1480,1500]" captionTargetBox="[183,1077,232,1449]" captionTargetId="figure-0@16.[174,1081,219,1617]" captionTargetPageId="16" captionText="Figs 7-9. Psychristus discretus ANDREWES. Habitus, head and pronotum. (7) Nepal, Basantapur; (8) Vietnam, Sa Pa; (9) India, Bomdila." figureDoi="http://doi.org/10.5281/zenodo.5279290" httpUri="https://zenodo.org/record/5279290/files/figure.png" pageId="7" pageNumber="1642">Fig. 7</figureCitation>
) 1.48-1.60 times as long as wide, 3.00-3.25 times as long and 1.51-1.63 times as wide as pronotum, with surface strongly convex. Humeri less angulate than in other species, with humeral tooth very small, usually no more than suggested. Setiferous pore in posterior half of interval 3 present. Elytral intervals 2-3 comparatively wide and flat at apex, 3 often somewhat dilated. Microsculpture almost absent, except for isodiametric mesh patterns on scutellum and anterior to basal margin. Pro- and mesotarsomeres in both sexes ventrally without a dentiform process, protarsomeres 1-4 of males only very weakly dilated, with typically dilated adhesive hairs beneath. Number and size of spines in pro- and mesotibiae unmodified. Median lobe of aedoeagus (
<figureCitation id="130DB4D9C349EC27FCE519C1FC1DFE7D" box="[849,963,453,475]" captionStart="Figs 18-21" captionStartId="19.[202,242,1272,1292]" captionTargetBox="[256,1458,411,1190]" captionTargetId="figure-32@19.[229,1539,397,1270]" captionTargetPageId="19" captionText="Figs 18-21. Median lobe of aedoagus, lateral and dorsal aspect. (18-19) Psychristus discretus ANDREWES (India, Bomdila); (20-21) P. hibatai ITO (Paratype, Taiwan, Wushe)." figureDoi="http://doi.org/10.5281/zenodo.5279300" httpUri="https://zenodo.org/record/5279300/files/figure.png" pageId="8" pageNumber="1643">Figs 18-19</figureCitation>
) large, and elongate, with a distinctly developed apical lamella. Female genitalia as figured (
<figureCitation id="130DB4D9C349EC27FBA019E6FF11FDB3" captionStart="Figs" captionStartId="20.[178,220,1499,1519]" captionTargetBox="[198,1064,239,1458]" captionTargetId="figure-0@20.[178,1086,219,1505]" captionTargetPageId="20" captionText="Figs. 22-25. Female genitalia. Hemisternite and gonocoxa. (22) Psychristus dentatus nov.sp., (Holotype); (23) P. brunneus nov.sp. (Paratype, Laos, Ban Khoun Ngeun); (24) P. discretus ANDREWES (India, Bomdila); (25) P. shibatai ITO (Taiwan, Mt. Kwena)." figureDoi="http://doi.org/10.5281/zenodo.5279302" httpUri="https://zenodo.org/record/5279302/files/figure.png" pageId="8" pageNumber="1643">Fig. 24</figureCitation>
). For ratios representing various body proportions see table 1 and 2.
</paragraph>
<paragraph id="8B89A85CC349EC27FF061A21FD6AFCAE" blockId="8.[178,1085,549,776]" pageId="8" pageNumber="1643">
C o m p a r i s o n s: Within the
<taxonomicName id="4C36D3DFC349EC27FDAD1A22FD4AFD9D" authorityName="ANDREWES. Habitus" authorityYear="1930" box="[537,660,549,571]" class="Insecta" family="Carabidae" genus="Psychristus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1643" phylum="Arthropoda" rank="species" species="discretus">
<emphasis id="B942744EC349EC27FDAD1A22FD4AFD9D" box="[537,660,549,571]" italics="true" pageId="8" pageNumber="1643">P. discretus</emphasis>
</taxonomicName>
group the species (
<figureCitation id="130DB4D9C349EC27FCD31A21FC79FD9C" box="[871,935,549,571]" captionStart="Figs 7-9" captionStartId="16.[178,218,1480,1500]" captionTargetBox="[183,1077,232,1449]" captionTargetId="figure-0@16.[174,1081,219,1617]" captionTargetPageId="16" captionText="Figs 7-9. Psychristus discretus ANDREWES. Habitus, head and pronotum. (7) Nepal, Basantapur; (8) Vietnam, Sa Pa; (9) India, Bomdila." figureDoi="http://doi.org/10.5281/zenodo.5279290" httpUri="https://zenodo.org/record/5279290/files/figure.png" pageId="8" pageNumber="1643">Fig. 7</figureCitation>
) is in general appearance most similar to
<taxonomicName id="4C36D3DFC349EC27FE7C1A47FDE8FDFE" authorityName="ITO. Habitus" authorityYear="1985" box="[456,566,578,600]" class="Insecta" family="Carabidae" genus="Psychristus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1643" phylum="Arthropoda" rank="species" species="shibatai">
<emphasis id="B942744EC349EC27FE7C1A47FDE8FDFE" box="[456,566,578,600]" italics="true" pageId="8" pageNumber="1643">P. shibatai</emphasis>
</taxonomicName>
(
<figureCitation id="130DB4D9C349EC27FDF71A46FD59FDFE" box="[579,647,578,600]" captionStart="Figs 10-12" captionStartId="17.[178,218,1470,1490]" captionTargetBox="[190,1072,217,1421]" captionTargetId="figure-0@17.[178,1085,159,1557]" captionTargetPageId="17" captionText="Figs 10-12. Psychristus shibatai ITO. Habitus, head and pronotum. (10-11) Paratype, Taiwan, Wushe; (12) Female, Mt. Kwena." figureDoi="http://doi.org/10.5281/zenodo.5279294" httpUri="https://zenodo.org/record/5279294/files/figure.png" pageId="8" pageNumber="1643">Fig.10</figureCitation>
) from which it can be separated usually by the larger size; darker, blackish brown body colour; the pronotum (
<figureCitation id="130DB4D9C349EC27FCEA1A5BFC6AFDD3" box="[862,948,607,629]" captionStart="Figs 7-9" captionStartId="16.[178,218,1480,1500]" captionTargetBox="[183,1077,232,1449]" captionTargetId="figure-0@16.[174,1081,219,1617]" captionTargetPageId="16" captionText="Figs 7-9. Psychristus discretus ANDREWES. Habitus, head and pronotum. (7) Nepal, Basantapur; (8) Vietnam, Sa Pa; (9) India, Bomdila." figureDoi="http://doi.org/10.5281/zenodo.5279290" httpUri="https://zenodo.org/record/5279290/files/figure.png" pageId="8" pageNumber="1643">Figs 7-9</figureCitation>
) with medial third of median line usually impunctate and anterior angles more projecting forward; the elytra, which are longer relative to the pronotum (EL/PL&gt; 2.99), with humeral tooth very small and not projecting, and dorsal surface strongly convex; the large, elongated median lobe of aedoeagus with developed apical lamella (
<figureCitation id="130DB4D9C349EC27FCBA1AD0FCA1FD4C" box="[782,895,724,746]" captionStart="Figs 18-21" captionStartId="19.[202,242,1272,1292]" captionTargetBox="[256,1458,411,1190]" captionTargetId="figure-32@19.[229,1539,397,1270]" captionTargetPageId="19" captionText="Figs 18-21. Median lobe of aedoagus, lateral and dorsal aspect. (18-19) Psychristus discretus ANDREWES (India, Bomdila); (20-21) P. hibatai ITO (Paratype, Taiwan, Wushe)." figureDoi="http://doi.org/10.5281/zenodo.5279300" httpUri="https://zenodo.org/record/5279300/files/figure.png" pageId="8" pageNumber="1643">Figs 18-19</figureCitation>
) and the different shape of hemisternite (
<figureCitation id="130DB4D9C349EC27FE2C1AF6FE3CFCAE" box="[408,482,754,776]" captionStart="Figs" captionStartId="20.[178,220,1499,1519]" captionTargetBox="[198,1064,239,1458]" captionTargetId="figure-0@20.[178,1086,219,1505]" captionTargetPageId="20" captionText="Figs. 22-25. Female genitalia. Hemisternite and gonocoxa. (22) Psychristus dentatus nov.sp., (Holotype); (23) P. brunneus nov.sp. (Paratype, Laos, Ban Khoun Ngeun); (24) P. discretus ANDREWES (India, Bomdila); (25) P. shibatai ITO (Taiwan, Mt. Kwena)." figureDoi="http://doi.org/10.5281/zenodo.5279302" httpUri="https://zenodo.org/record/5279302/files/figure.png" pageId="8" pageNumber="1643">Fig. 24</figureCitation>
) of female genitalia.
</paragraph>
<paragraph id="8B89A85CC349EC27FF061B1CFC24FC8B" blockId="8.[178,1018,791,813]" box="[178,1018,791,813]" pageId="8" pageNumber="1643">
For differentiation from
<taxonomicName id="4C36D3DFC349EC27FE1E1B1CFDFAFC8B" authorityName="Jaeger &amp; H" authorityYear="2009" box="[426,548,791,813]" class="Insecta" family="Carabidae" genus="Psychristus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1643" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="B942744EC349EC27FE1E1B1CFDFAFC8B" box="[426,548,791,813]" italics="true" pageId="8" pageNumber="1643">P. brunneus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C36D3DFC349EC27FDE11B1CFD19FC8B" authorityName="Jaeger &amp; H" authorityYear="2009" box="[597,711,791,813]" class="Insecta" family="Carabidae" genus="Psychristus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1643" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="B942744EC349EC27FDE11B1CFD19FC8B" box="[597,711,791,813]" italics="true" pageId="8" pageNumber="1643">P. dentatus</emphasis>
</taxonomicName>
see these species and the key.
</paragraph>
<paragraph id="8B89A85CC349EC27FF061B39FD22FC61" blockId="8.[178,1085,828,967]" pageId="8" pageNumber="1643">
D i s t r i b u t i o n: The species occurs from northwest
<collectingCountry id="F321E8CCC349EC27FCA31B38FC95FCF4" box="[791,843,828,850]" name="India" pageId="8" pageNumber="1643">India</collectingCountry>
(
<collectingRegion id="49F266BEC349EC27FCE91B38FC31FCF4" box="[861,1007,828,850]" country="India" name="Uttar Pradesh" pageId="8" pageNumber="1643">Uttar Pradesh</collectingRegion>
) in the West, across
<collectingCountry id="F321E8CCC349EC27FEF81B5EFE55FCD6" box="[332,395,858,880]" name="Nepal" pageId="8" pageNumber="1643">Nepal</collectingCountry>
, north
<collectingCountry id="F321E8CCC349EC27FE501B5EFDC6FCD6" box="[484,536,858,880]" name="India" pageId="8" pageNumber="1643">India</collectingCountry>
(
<collectingRegion id="49F266BEC349EC27FD9B1B5EFD1EFCD6" box="[559,704,858,880]" country="India" name="West Bengal" pageId="8" pageNumber="1643">West Bengal</collectingRegion>
,
<collectingRegion id="49F266BEC349EC27FD601B5EFCFFFCD6" box="[724,801,858,880]" country="India" name="Sikkim" pageId="8" pageNumber="1643">Sikkim</collectingRegion>
),
<collectingCountry id="F321E8CCC349EC27FC881B5EFC56FCD6" box="[828,904,858,880]" name="Bhutan" pageId="8" pageNumber="1643">Bhutan</collectingCountry>
, northeast
<collectingCountry id="F321E8CCC349EC27FBBD1B5EFBE3FCD6" box="[1033,1085,858,880]" name="India" pageId="8" pageNumber="1643">India</collectingCountry>
(
<collectingRegion id="49F266BEC349EC27FF0C1B73FEA0FC2B" box="[184,382,887,909]" country="India" name="Arunachal Pradesh" pageId="8" pageNumber="1643">Arunachal Pradesh</collectingRegion>
),
<collectingCountry id="F321E8CCC349EC27FE3B1B7CFE2FFC2B" box="[399,497,888,909]" name="Myanmar" pageId="8" pageNumber="1643">Myanmar</collectingCountry>
to northwest
<collectingCountry id="F321E8CCC349EC27FDC91B73FD0AFC2B" box="[637,724,887,909]" name="Vietnam" pageId="8" pageNumber="1643">Vietnam</collectingCountry>
in the East. It has been collected at elevations of about
<quantity id="4CCE05B9C349EC27FE321B90FDCAFC0C" box="[390,532,916,938]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.8" metricValueMin="1.5" pageId="8" pageNumber="1643" unit="m" value="2150.0" valueMax="2800.0" valueMin="1500.0">1500-2800 m</quantity>
.
<figureCitation id="130DB4D9C349EC27FD971B90FD57FC0C" box="[547,649,916,938]" captionStart="Figs 26-27" captionStartId="21.[178,218,1483,1503]" captionTargetBox="[178,1058,219,1459]" captionTargetId="figure-33@21.[178,1085,219,1460]" captionTargetPageId="21" captionText="Figs 26-27. Distribution. (26) P. dentatus nov.sp. (open circles), P. brunneus nov.sp. (filled circles) and P. shibatai ITO (filled squares); (27) P. discretus ANDREWES." figureDoi="http://doi.org/10.5281/zenodo.5279304" httpUri="https://zenodo.org/record/5279304/files/figure.png" pageId="8" pageNumber="1643">Figure 27</figureCitation>
shows the known distribution, based on revised records (see
<bibRefCitation id="EFA7D5ADC349EC27FE351BB6FDF2FC61" author="JAEGER B" box="[385,556,945,967]" pageId="8" pageNumber="1643" pagination="63 - 93" refId="ref8994" refString="JAEGER B. (1997): Revision der Himalaya-Arten der Gattung Psychristus ANDREWES 1930 (Col., Carabidae) - Linzer biologische Beitrage 29 (1): 63 - 93." type="journal article" year="1997">JAEGER 1997: 72</bibRefCitation>
and records below).
</paragraph>
<paragraph id="8B89A85CC349EC27FF061BD3FD4BFBD9" blockId="8.[178,1085,983,1151]" pageId="8" pageNumber="1643">
LINDROTH (1939) described the species also as
<taxonomicName id="4C36D3DFC349EC27FD1A1BD3FC7BFC4B" authorityName="LINDROTH" authorityYear="1939" box="[686,933,983,1005]" class="Insecta" family="Carabidae" genus="Bradycellus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1643" phylum="Arthropoda" rank="species" species="ponderosus">
<emphasis id="B942744EC349EC27FD1A1BD3FC7BFC4B" box="[686,933,983,1005]" italics="true" pageId="8" pageNumber="1643">Bradycellus ponderosus</emphasis>
</taxonomicName>
from Karelia. However, due to zoogeographical reasons an occurrence of the species in this northeast European region seems to be impossible.
<bibRefCitation id="EFA7D5ADC349EC27FDDA1C16FCE1FB81" author="JAEGER B" box="[622,831,1041,1063]" pageId="8" pageNumber="1643" pagination="955 - 973" refId="ref9028" refString="JAEGER B. (1999): Nachtrag zur Acupalpus - Fauna des Nahen und Mittleren Ostens und neue Erkenntnisse zur Verbreitung und Synonymie einiger Arten der Gattungen Acupalpus, Anthracus, Bradycellus und Psychristus (Coleoptera, Carabidae). - Linzer biologische Beitrage 31 (2): 955 - 973." type="journal article" year="1999">JAEGER (1999: 968)</bibRefCitation>
suspected that the
<typeStatus id="548D16FEC349EC27FBA51C16FBE3FB81" box="[1041,1085,1042,1063]" pageId="8" pageNumber="1643">type</typeStatus>
species was mislabelled and originated from collections from former
<collectingCountry id="F321E8CCC349EC27FC041C2AFBE6FBE2" box="[944,1080,1070,1092]" name="India" pageId="8" pageNumber="1643">British India</collectingCountry>
, probably from a large series from
<collectingCountry id="F321E8CCC349EC27FD961C48FDB9FBC7" box="[546,615,1100,1121]" name="Myanmar" pageId="8" pageNumber="1643">Burma</collectingCountry>
collected by R. Malaise in 1934 during the Swedish expedition to
<collectingCountry id="F321E8CCC349EC27FE2D1C6DFDC6FBD9" box="[409,536,1129,1151]" name="India" pageId="8" pageNumber="1643">British India</collectingCountry>
and
<collectingCountry id="F321E8CCC349EC27FDFD1C6EFD4FFBD9" box="[585,657,1130,1151]" name="Myanmar" pageId="8" pageNumber="1643">Burma</collectingCountry>
.
</paragraph>
<paragraph id="8B89A85CC349EC27FF061C8AFBD1FB64" blockId="8.[178,1085,1166,1218]" pageId="8" pageNumber="1643">
Since the revision of the species (
<bibRefCitation id="EFA7D5ADC349EC27FDAD1C8AFD11FB02" author="JAEGER B" box="[537,719,1166,1188]" pageId="8" pageNumber="1643" pagination="63 - 93" refId="ref8994" refString="JAEGER B. (1997): Revision der Himalaya-Arten der Gattung Psychristus ANDREWES 1930 (Col., Carabidae) - Linzer biologische Beitrage 29 (1): 63 - 93." type="journal article" year="1997">JAEGER 1997: 72</bibRefCitation>
) the following additional material, including the first record for the Indian state of
<collectingRegion id="49F266BEC349EC27FD271CA8FC88FB64" box="[659,854,1196,1218]" country="India" name="Arunachal Pradesh" pageId="8" pageNumber="1643">Arunachal Pradesh</collectingRegion>
, has been studied:
</paragraph>
<paragraph id="8B89A85CC349EC27FF061CCBFEE1F98A" blockId="8.[178,1086,1231,1580]" pageId="8" pageNumber="1643">
<materialsCitation id="3B5EA201C349EC27FF061CCBFCE6FAB0" collectingDate="1997-05" collectingDateMax="2005-06" collectingDateMin="1997-05" collectionCode="LF" county="Kaski District" elevation="2100" latitude="29.858055" location="Far-Western Region" longLatPrecision="20" longitude="80.90945" municipality="Western Region" pageId="8" pageNumber="1643" specimenCount="1">
N e p a l:
<location id="8EE9FE87C349EC27FE961CCBFE03FB45" LSID="urn:lsid:plazi:treatment:039F194AC346EC26FF061AC1FC36FEE2:8EE9FE87C349EC27FE961CCBFE03FB45" box="[290,477,1231,1251]" county="Kaski District" latitude="29.858055" longLatPrecision="20" longitude="80.90945" municipality="Western Region" name="Far-Western Region" pageId="8" pageNumber="1643">Far-Western Region</location>
:
<location id="8EE9FE87C349EC27FE581CCBFD52FB45" LSID="urn:lsid:plazi:treatment:039F194AC346EC26FF061AC1FC36FEE2:8EE9FE87C349EC27FE581CCBFD52FB45" box="[492,652,1231,1251]" county="Kaski District" latitude="29.858055" longLatPrecision="20" longitude="80.90945" municipality="Western Region" name="Darchula District" pageId="8" pageNumber="1643">Darchula District</location>
:
<locationDeviation id="9A3BB17EC349EC27FD2E1CCBFCE8FB45" box="[666,822,1231,1251]" location="Batar" pageId="8" pageNumber="1643">
<quantity id="4CCE05B9C349EC27FD2E1CCBFD11FB45" box="[666,719,1231,1251]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="1643" unit="km" value="1.0">1 km</quantity>
NE Batar
</locationDeviation>
, Chamliya Khola,
<quantity id="4CCE05B9C349EC27FC5A1CCBFBE7FB45" box="[1006,1081,1231,1251]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="8" pageNumber="1643" unit="m" value="2100.0">
<elevation id="001B4F6FC349EC27FC5A1CCBFBE7FB45" box="[1006,1081,1231,1251]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="8" pageNumber="1643" unit="m" value="2100.0">2100 m</elevation>
</quantity>
,
<geoCoordinate id="EE02CE9BC349EC27FF6B1CEDFE9BFB5B" box="[223,325,1257,1277]" degrees="29" direction="north" minutes="51" orientation="latitude" pageId="8" pageNumber="1643" precision="15" seconds="29" value="29.858055">29°51'29&quot;N</geoCoordinate>
<geoCoordinate id="EE02CE9BC349EC27FEFA1CEDFE6AFB5B" box="[334,436,1257,1277]" degrees="80" direction="east" minutes="54" orientation="longitude" pageId="8" pageNumber="1643" precision="15" seconds="34" value="80.90945">80°54'34&quot;E</geoCoordinate>
,
<collectionCode id="ED273099C349EC27FE0B1CEDFE05FB5B" box="[447,475,1257,1277]" pageId="8" pageNumber="1643">LF</collectionCode>
,
<date id="FF888E9CC349EC27FE521CEDFDE7FB5B" box="[486,569,1257,1277]" pageId="8" pageNumber="1643" value="2005-06">
<collectingDate id="EFCC7774C349EC27FE521CEDFDE7FB5B" box="[486,569,1257,1277]" pageId="8" pageNumber="1643" value="2005-06">VI. 2005</collectingDate>
</date>
, Weigel (1 - cWEIG). -
<collectingMunicipality id="6BED3226C349EC27FCA91CEDFC6EFB5B" box="[797,944,1257,1277]" pageId="8" pageNumber="1643">Western Region</collectingMunicipality>
:
<collectingCounty id="62E8D0D0C349EC27FC0F1CEDFBE4FB5B" box="[955,1082,1257,1277]" pageId="8" pageNumber="1643">Kaski District</collectingCounty>
: Annapurna, Krapa Danda,
<quantity id="4CCE05B9C349EC27FE7F1D06FDCEFAB0" box="[459,528,1282,1302]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="1643" unit="m" value="2500.0">
<elevation id="001B4F6FC349EC27FE7F1D06FDCEFAB0" box="[459,528,1282,1302]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="1643" unit="m" value="2500.0">2500 m</elevation>
</quantity>
,
<date id="FF888E9CC349EC27FDAE1D06FDBAFAB0" box="[538,612,1282,1302]" pageId="8" pageNumber="1643" value="1997-05">
<collectingDate id="EFCC7774C349EC27FDAE1D06FDBAFAB0" box="[538,612,1282,1302]" pageId="8" pageNumber="1643" value="1997-05">V. 1997</collectingDate>
</date>
, Schmidt (1 – cSCHM)
</materialsCitation>
;
<materialsCitation id="3B5EA201C349EC27FCF51D06FBFBFAEE" collectingDate="1973-04" collectingDateMax="1996-05" collectingDateMin="1973-04" collectionCode="SMNS" collectorName="Jager &amp; Cherakapti &amp; Jiri &amp; Martens" county="Dolakha District" elevation="2350" location="Lamjung Himal" municipality="Central Region" pageId="8" pageNumber="1643" specimenCount="1">
<location id="8EE9FE87C349EC27FCF51D06FC0EFAB0" LSID="urn:lsid:plazi:treatment:039F194AC346EC26FF061AC1FC36FEE2:8EE9FE87C349EC27FCF51D06FC0EFAB0" box="[833,976,1282,1302]" county="Dolakha District" municipality="Central Region" name="Lamjung Himal" pageId="8" pageNumber="1643">Lamjung Himal</location>
, unt.
<location id="8EE9FE87C349EC27FBB41D06FEC4FA89" LSID="urn:lsid:plazi:treatment:039F194AC346EC26FF061AC1FC36FEE2:8EE9FE87C349EC27FBB41D06FEC4FA89" county="Dolakha District" municipality="Central Region" name="Taunja Danda" pageId="8" pageNumber="1643">Taunja Danda</location>
, oberh. Hogo-Kh.,W-Hang,
<quantity id="4CCE05B9C349EC27FD961D1FFDB5FA89" box="[546,619,1307,1327]" metricMagnitude="3" metricUnit="m" metricValue="2.35" pageId="8" pageNumber="1643" unit="m" value="2350.0">
<elevation id="001B4F6FC349EC27FD961D1FFDB5FA89" box="[546,619,1307,1327]" metricMagnitude="3" metricUnit="m" metricValue="2.35" pageId="8" pageNumber="1643" unit="m" value="2350.0">2350 m</elevation>
</quantity>
,
<date id="FF888E9CC349EC27FDCE1D1FFD16FA89" box="[634,712,1307,1327]" pageId="8" pageNumber="1643" value="1996-05">
<collectingDate id="EFCC7774C349EC27FDCE1D1FFD16FA89" box="[634,712,1307,1327]" pageId="8" pageNumber="1643" value="1996-05">V. 1996</collectingDate>
</date>
,
<collectorName id="26C3CD8AC349EC27FD621D1FFCDAFA89" box="[726,772,1307,1327]" pageId="8" pageNumber="1643">Jäger</collectorName>
(17 – cSCHM, cJAE). -
<collectingMunicipality id="6BED3226C349EC27FC491D1FFEC0FAEE" pageId="8" pageNumber="1643">Central Region</collectingMunicipality>
:
<collectingCounty id="62E8D0D0C349EC27FE9F1D30FE61FAEE" box="[299,447,1332,1352]" pageId="8" pageNumber="1643">Dolakha District</collectingCounty>
:
<collectorName id="26C3CD8AC349EC27FE731D30FDF6FAEE" box="[455,552,1332,1352]" pageId="8" pageNumber="1643">Cherakapti</collectorName>
, nördl.
<collectorName id="26C3CD8AC349EC27FDDF1D30FD58FAEE" box="[619,646,1332,1352]" pageId="8" pageNumber="1643">Jiri</collectorName>
,
<quantity id="4CCE05B9C349EC27FD251D30FCD7FAEE" box="[657,777,1332,1352]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="8" pageNumber="1643" unit="m" value="1650.0" valueMax="1800.0" valueMin="1500.0">
<elevation id="001B4F6FC349EC27FD251D30FCD7FAEE" box="[657,777,1332,1352]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="8" pageNumber="1643" unit="m" value="1650.0" valueMax="1800.0" valueMin="1500.0">1500-1800 m</elevation>
</quantity>
,
<date id="FF888E9CC349EC27FCA61D30FCBDFAEE" box="[786,867,1332,1352]" pageId="8" pageNumber="1643" value="1973-04">
<collectingDate id="EFCC7774C349EC27FCA61D30FCBDFAEE" box="[786,867,1332,1352]" pageId="8" pageNumber="1643" value="1973-04">IV. 1973</collectingDate>
</date>
,
<collectorName id="26C3CD8AC349EC27FCD81D30FC6AFAEE" box="[876,948,1332,1352]" pageId="8" pageNumber="1643">Martens</collectorName>
(1 -
<collectionCode id="ED273099C349EC27FC541D30FBFEFAEE" box="[992,1056,1332,1352]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="8" pageNumber="1643">SMNS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B5EA201C349EC27FB9B1D30FDBAFADD" collectingDate="2000-06" collectionCode="VI, SMNS, V" collectorName="E Ting Sang La &amp; Schawaller" elevation="2600" location="Khare Kola" pageId="8" pageNumber="1643" specimenCount="1">
<collectorName id="26C3CD8AC349EC27FB9B1D30FE84FAC4" pageId="8" pageNumber="1643">E Ting Sang La</collectorName>
,
<quantity id="4CCE05B9C349EC27FED11D4AFE74FAC4" box="[357,426,1358,1378]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="8" pageNumber="1643" unit="m" value="2600.0">
<elevation id="001B4F6FC349EC27FED11D4AFE74FAC4" box="[357,426,1358,1378]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="8" pageNumber="1643" unit="m" value="2600.0">2600 m</elevation>
</quantity>
, 627,
<date id="FF888E9CC349EC27FE561D4AFDEDFAC4" box="[482,563,1358,1378]" pageId="8" pageNumber="1643" value="2000-06">
<collectingDate id="EFCC7774C349EC27FE561D4AFDEDFAC4" box="[482,563,1358,1378]" pageId="8" pageNumber="1643" value="2000-06">
<collectionCode id="ED273099C349EC27FE561D4AFE22FAC4" box="[482,508,1358,1378]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="8" pageNumber="1643">VI</collectionCode>
. 2000
</collectingDate>
</date>
,
<collectorName id="26C3CD8AC349EC27FD891D4AFD40FAC4" box="[573,670,1358,1378]" pageId="8" pageNumber="1643">Schawaller</collectorName>
(4 -
<collectionCode id="ED273099C349EC27FD781D4AFCD5FAC4" box="[716,779,1358,1378]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="8" pageNumber="1643">SMNS</collectionCode>
), N slope of
<location id="8EE9FE87C349EC27FC341D4AFC34FAC4" LSID="urn:lsid:plazi:treatment:039F194AC346EC26FF061AC1FC36FEE2:8EE9FE87C349EC27FC341D4AFC34FAC4" box="[896,1002,1358,1378]" name="Khare Kola" pageId="8" pageNumber="1643">Khare Kola</location>
,
<quantity id="4CCE05B9C349EC27FC471D4AFBE7FAC4" box="[1011,1081,1358,1378]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="1643" unit="m" value="2200.0">
<elevation id="001B4F6FC349EC27FC471D4AFBE7FAC4" box="[1011,1081,1358,1378]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="1643" unit="m" value="2200.0">2200 m</elevation>
</quantity>
, 612,
<collectionCode id="ED273099C349EC27FEBA1D63FEC1FADD" box="[270,287,1383,1403]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="1643" type="Museum">V</collectionCode>
./
<date id="FF888E9CC349EC27FE9E1D63FEA3FADD" box="[298,381,1383,1403]" pageId="8" pageNumber="1643" value="2000-06">
<collectingDate id="EFCC7774C349EC27FE9E1D63FEA3FADD" box="[298,381,1383,1403]" pageId="8" pageNumber="1643" value="2000-06">
<collectionCode id="ED273099C349EC27FE9E1D63FE9DFADD" box="[298,323,1383,1403]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="8" pageNumber="1643">VI</collectionCode>
. 2000
</collectingDate>
</date>
,
<collectorName id="26C3CD8AC349EC27FE3D1D63FE34FADD" box="[393,490,1383,1403]" pageId="8" pageNumber="1643">Schawaller</collectorName>
(1 -
<collectionCode id="ED273099C349EC27FDAA1D63FD80FADD" box="[542,606,1383,1403]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="8" pageNumber="1643">SMNS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B5EA201C349EC27FDC51D63FEFAFA08" collectingDate="2000-05" collectingDateMax="2000-06" collectingDateMin="2000-05" collectionCode="V, SMNS, VI" collectorName="Serukapti &amp; Schawaller" elevation="2400" location="Sindhupalchok District" municipality="Dolangsa" pageId="8" pageNumber="1643" specimenCount="1">
<collectorName id="26C3CD8AC349EC27FDC51D63FD18FADD" box="[625,710,1383,1403]" pageId="8" pageNumber="1643">Serukapti</collectorName>
,
<quantity id="4CCE05B9C349EC27FD661D63FCC4FADD" box="[722,794,1383,1403]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="8" pageNumber="1643" unit="m" value="2400.0">
<elevation id="001B4F6FC349EC27FD661D63FCC4FADD" box="[722,794,1383,1403]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="8" pageNumber="1643" unit="m" value="2400.0">2400 m</elevation>
</quantity>
, 605,
<date id="FF888E9CC349EC27FCE31D63FC7AFADD" box="[855,932,1383,1403]" pageId="8" pageNumber="1643" value="2000-05">
<collectingDate id="EFCC7774C349EC27FCE31D63FC7AFADD" box="[855,932,1383,1403]" pageId="8" pageNumber="1643" value="2000-05">
<collectionCode id="ED273099C349EC27FCE31D63FCB4FADD" box="[855,874,1383,1403]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="1643" type="Museum">V</collectionCode>
. 2000
</collectingDate>
</date>
,
<collectorName id="26C3CD8AC349EC27FC041D63FBCFFADD" box="[944,1041,1383,1403]" pageId="8" pageNumber="1643">Schawaller</collectorName>
(5 -
<collectionCode id="ED273099C349EC27FF6B1D84FEC1FA32" box="[223,287,1408,1428]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="8" pageNumber="1643">SMNS</collectionCode>
). -
<location id="8EE9FE87C349EC27FE891D84FDD5FA32" LSID="urn:lsid:plazi:treatment:039F194AC346EC26FF061AC1FC36FEE2:8EE9FE87C349EC27FE891D84FDD5FA32" box="[317,523,1408,1428]" municipality="Dolangsa" name="Sindhupalchok District" pageId="8" pageNumber="1643">Sindhupalchok District</location>
: SW below
<collectingMunicipality id="6BED3226C349EC27FDCE1D84FD10FA32" box="[634,718,1408,1428]" pageId="8" pageNumber="1643">Dolangsa</collectingMunicipality>
,
<quantity id="4CCE05B9C349EC27FD6D1D84FCC1FA32" box="[729,799,1408,1428]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="8" pageNumber="1643" unit="m" value="2300.0">
<elevation id="001B4F6FC349EC27FD6D1D84FCC1FA32" box="[729,799,1408,1428]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="8" pageNumber="1643" unit="m" value="2300.0">2300 m</elevation>
</quantity>
, 631,
<date id="FF888E9CC349EC27FCEC1D84FC74FA32" box="[856,938,1408,1428]" pageId="8" pageNumber="1643" value="2000-06">
<collectingDate id="EFCC7774C349EC27FCEC1D84FC74FA32" box="[856,938,1408,1428]" pageId="8" pageNumber="1643" value="2000-06">
<collectionCode id="ED273099C349EC27FCEC1D84FCACFA32" box="[856,882,1408,1428]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="8" pageNumber="1643">VI</collectionCode>
. 2000
</collectingDate>
</date>
,
<collectorName id="26C3CD8AC349EC27FC001D84FBCBFA32" box="[948,1045,1408,1428]" pageId="8" pageNumber="1643">Schawaller</collectorName>
(1 -
<collectionCode id="ED273099C349EC27FF6B1D9EFEC1FA08" box="[223,287,1434,1454]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="8" pageNumber="1643">SMNS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B5EA201C349EC27FE851D9EFEE5F98A" collectingDate="1978-06" collectingDateMax="2000-06" collectingDateMin="1978-06" collectionCode="MHNG, SMNS, NHMB" collectorName="Pokhare NE Barahbise &amp; Lobl &amp; Smetana &amp; Phulchoki &amp; Quercus &amp; Martens &amp; Ausobsky &amp; Junbesi &amp; Schawaller &amp; Basantapur" county="Terhathum District" elevation="2700" latitude="27.11" location="Lalitpur District" longLatPrecision="743" longitude="87.27" municipality="Solukhumbu District" pageId="8" pageNumber="1643" specimenCount="1">
<collectorName id="26C3CD8AC349EC27FE851D9EFDDCFA08" box="[305,514,1434,1454]" pageId="8" pageNumber="1643">Pokhare NE Barahbise</collectorName>
,
<quantity id="4CCE05B9C349EC27FDBB1D9EFD89FA08" box="[527,599,1434,1454]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="8" pageNumber="1643" unit="m" value="2700.0">
<elevation id="001B4F6FC349EC27FDBB1D9EFD89FA08" box="[527,599,1434,1454]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="8" pageNumber="1643" unit="m" value="2700.0">2700 m</elevation>
</quantity>
,
<date id="FF888E9CC349EC27FDD01D9EFD6FFA08" box="[612,689,1434,1454]" pageId="8" pageNumber="1643" value="1981-05">
<collectingDate id="EFCC7774C349EC27FDD01D9EFD6FFA08" box="[612,689,1434,1454]" pageId="8" pageNumber="1643" value="1981-05">V. 1981</collectingDate>
</date>
,
<collectorName id="26C3CD8AC349EC27FD0A1D9EFD39FA08" box="[702,743,1434,1454]" pageId="8" pageNumber="1643">Löbl</collectorName>
&amp;
<collectorName id="26C3CD8AC349EC27FCBF1D9EFC88FA08" box="[779,854,1434,1454]" pageId="8" pageNumber="1643">Smetana</collectorName>
(1 -
<collectionCode id="ED273099C349EC27FC391D9EFC0BFA08" box="[909,981,1434,1454]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="8" pageNumber="1643" type="Museum">MHNG</collectionCode>
). -
<location id="8EE9FE87C349EC27FC431D9EFEFDFA61" LSID="urn:lsid:plazi:treatment:039F194AC346EC26FF061AC1FC36FEE2:8EE9FE87C349EC27FC431D9EFEFDFA61" county="Terhathum District" latitude="27.11" longLatPrecision="743" longitude="87.27" municipality="Solukhumbu District" name="Lalitpur District" pageId="8" pageNumber="1643">Lalitpur District</location>
: 106 KTM
<collectorName id="26C3CD8AC349EC27FE3C1DB7FE3EFA61" box="[392,480,1459,1479]" pageId="8" pageNumber="1643">Phulchoki</collectorName>
<quantity id="4CCE05B9C349EC27FE511DB7FD82FA61" box="[485,604,1459,1479]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="8" pageNumber="1643" unit="m" value="2150.0" valueMax="2300.0" valueMin="2000.0">
<elevation id="001B4F6FC349EC27FE511DB7FD82FA61" box="[485,604,1459,1479]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="8" pageNumber="1643" unit="m" value="2150.0" valueMax="2300.0" valueMin="2000.0">2000-2300 m</elevation>
</quantity>
,
<collectorName id="26C3CD8AC349EC27FDD11DB7FD6EFA61" box="[613,688,1459,1479]" pageId="8" pageNumber="1643">Quercus</collectorName>
,
<date id="FF888E9CC349EC27FD0C1DB7FCD9FA61" box="[696,775,1459,1479]" pageId="8" pageNumber="1643" value="1980-03">
<collectingDate id="EFCC7774C349EC27FD0C1DB7FCD9FA61" box="[696,775,1459,1479]" pageId="8" pageNumber="1643" value="1980-03">III. 1980</collectingDate>
</date>
,
<collectorName id="26C3CD8AC349EC27FCBB1DB7FC88FA61" box="[783,854,1459,1479]" pageId="8" pageNumber="1643">Martens</collectorName>
&amp;
<collectorName id="26C3CD8AC349EC27FCC71DB7FC14FA61" box="[883,970,1459,1479]" pageId="8" pageNumber="1643">Ausobsky</collectorName>
(2 -
<collectionCode id="ED273099C349EC27FC401DB7FBEAFA61" box="[1012,1076,1459,1479]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="8" pageNumber="1643">SMNS</collectionCode>
). -
<location id="8EE9FE87C349EC27FF581DC8FEABFA46" LSID="urn:lsid:plazi:treatment:039F194AC346EC26FF061AC1FC36FEE2:8EE9FE87C349EC27FF581DC8FEABFA46" box="[236,373,1484,1504]" county="Terhathum District" latitude="27.11" longLatPrecision="743" longitude="87.27" municipality="Solukhumbu District" name="Eastern Region" pageId="8" pageNumber="1643">Eastern Region</location>
:
<collectingMunicipality id="6BED3226C349EC27FEC91DC8FDE8FA46" box="[381,566,1484,1504]" pageId="8" pageNumber="1643">Solukhumbu District</collectingMunicipality>
:
<collectorName id="26C3CD8AC349EC27FD8B1DC8FD5CFA46" box="[575,642,1484,1504]" pageId="8" pageNumber="1643">Junbesi</collectorName>
,
<quantity id="4CCE05B9C349EC27FD3F1DC8FD11FA46" box="[651,719,1484,1504]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="8" pageNumber="1643" unit="m" value="2700.0">
<elevation id="001B4F6FC349EC27FD3F1DC8FD11FA46" box="[651,719,1484,1504]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="8" pageNumber="1643" unit="m" value="2700.0">2700 m</elevation>
</quantity>
, 514,
<date id="FF888E9CC349EC27FCB11DC8FC90FA46" box="[773,846,1484,1504]" pageId="8" pageNumber="1643" value="1997-05">
<collectingDate id="EFCC7774C349EC27FCB11DC8FC90FA46" box="[773,846,1484,1504]" pageId="8" pageNumber="1643" value="1997-05">V. 1997</collectingDate>
</date>
,
<collectorName id="26C3CD8AC349EC27FCE31DC8FC66FA46" box="[855,952,1484,1504]" pageId="8" pageNumber="1643">Schawaller</collectorName>
(1 -
<collectionCode id="ED273099C349EC27FC571DC8FBFDFA46" box="[995,1059,1484,1504]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="8" pageNumber="1643">SMNS</collectionCode>
). –
<collectingCounty id="62E8D0D0C349EC27FF6B1DE1FE55FA5F" box="[223,395,1509,1529]" pageId="8" pageNumber="1643">Terhathum District</collectingCounty>
:
<collectorName id="26C3CD8AC349EC27FE231DE1FE27FA5F" box="[407,505,1509,1529]" pageId="8" pageNumber="1643">Basantapur</collectorName>
env.,
<geoCoordinate id="EE02CE9BC349EC27FD811DE1FDA8FA5F" box="[565,630,1509,1529]" degrees="27.11" direction="north" orientation="latitude" pageId="8" pageNumber="1643" precision="555" value="27.11">27.11N</geoCoordinate>
<geoCoordinate id="EE02CE9BC349EC27FDCB1DE1FD1EFA5F" box="[639,704,1509,1529]" degrees="87.27" direction="east" orientation="longitude" pageId="8" pageNumber="1643" precision="555" value="87.27">87.27E</geoCoordinate>
,
<quantity id="4CCE05B9C349EC27FD781DE1FCCAFA5F" box="[716,788,1509,1529]" metricMagnitude="3" metricUnit="m" metricValue="2.19" pageId="8" pageNumber="1643" unit="m" value="2190.0">
<elevation id="001B4F6FC349EC27FD781DE1FCCAFA5F" box="[716,788,1509,1529]" metricMagnitude="3" metricUnit="m" metricValue="2.19" pageId="8" pageNumber="1643" unit="m" value="2190.0">2190 m</elevation>
</quantity>
,
<date id="FF888E9CC349EC27FC951DE1FCAAFA5F" box="[801,884,1509,1529]" pageId="8" pageNumber="1643" value="2000-06">
<collectingDate id="EFCC7774C349EC27FC951DE1FCAAFA5F" box="[801,884,1509,1529]" pageId="8" pageNumber="1643" value="2000-06">VI. 2000</collectingDate>
</date>
, Farkač (5 –
<collectionCode id="ED273099C349EC27FC411DE1FBE4FA5F" box="[1013,1082,1509,1529]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="8" pageNumber="1643">NHMB</collectionCode>
, cJAE). - Region/District not traced: Syabnu [Sic! Probably meaning Syabru?],
<date id="FF888E9CC349EC27FC3A1DFBFC01F9B5" box="[910,991,1535,1555]" pageId="8" pageNumber="1643" value="1978-06">
<collectingDate id="EFCC7774C349EC27FC3A1DFBFC01F9B5" box="[910,991,1535,1555]" pageId="8" pageNumber="1643" value="1978-06">VI. 1978</collectingDate>
</date>
, Bhakta (1 –
<collectionCode id="ED273099C349EC27FF5B1E1CFEEBF98A" box="[239,309,1560,1580]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="8" pageNumber="1643">NHMB</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B89A85CC348EC26FF0618DCFCFAFF4A" blockId="9.[178,1085,216,324]" box="[178,804,216,236]" pageId="9" pageNumber="1644">
B h u t a n: Timphu Umg.,
<quantity id="4CCE05B9C348EC26FE0018DCFE26FF4A" box="[436,504,216,236]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="1644" unit="m" value="2500.0">2500 m</quantity>
,
<date id="FF888E9CC348EC26FDB518DCFD89FF4A" box="[513,599,216,236]" pageId="9" pageNumber="1644" value="1988-07">VII. 1988</date>
, Holzschuh (1 - NME).
</paragraph>
<paragraph id="8B89A85CC348EC26FF0618FCFBE3FEAA" blockId="9.[178,1085,216,324]" box="[178,1085,248,268]" pageId="9" pageNumber="1644">
I n d i a:
<collectingRegion id="49F266BEC348EC26FEA218FCFE1FFEAA" box="[278,449,248,268]" country="India" name="Arunachal Pradesh" pageId="9" pageNumber="1644">Arunachal Pradesh</collectingRegion>
: W of Bomdila,
<quantity id="4CCE05B9C348EC26FDE918FCFD7BFEAA" box="[605,677,248,268]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="9" pageNumber="1644" unit="m" value="2600.0">2600 m</quantity>
,
<geoCoordinate id="EE02CE9BC348EC26FD0518FCFD27FEAA" box="[689,761,248,268]" degrees="27" direction="north" minutes="16" orientation="latitude" pageId="9" pageNumber="1644" precision="925" value="27.266666">27°16'N</geoCoordinate>
<geoCoordinate id="EE02CE9BC348EC26FCB518FCFC97FEAA" box="[769,841,248,268]" degrees="92" direction="east" minutes="24" orientation="longitude" pageId="9" pageNumber="1644" precision="925" value="92.4">92°24'E</geoCoordinate>
,
<date id="FF888E9CC348EC26FCE118FCFC7FFEAA" box="[853,929,248,268]" pageId="9" pageNumber="1644" value="2004-05">V. 2004</date>
, Dembický (16 -
</paragraph>
<paragraph id="8B89A85CC348EC26FF6B1914FEB7FE82" blockId="9.[178,1085,216,324]" box="[223,361,272,292]" pageId="9" pageNumber="1644">NHMB, cJAE).</paragraph>
<paragraph id="8B89A85CC348EC26FF061934FC36FEE2" blockId="9.[178,1085,216,324]" box="[178,1000,304,324]" pageId="9" pageNumber="1644">
V i e t n a m: Sa Pa (
<collectingRegion id="49F266BEC348EC26FECB1934FE16FEE2" box="[383,456,304,324]" country="Vietnam" name="Lao Cai" pageId="9" pageNumber="1644">Lao Cai</collectingRegion>
),
<geoCoordinate id="EE02CE9BC348EC26FE631934FDC1FEE2" box="[471,543,304,324]" degrees="22" direction="north" minutes="20" orientation="latitude" pageId="9" pageNumber="1644" precision="925" value="22.333334">22°20'N</geoCoordinate>
<geoCoordinate id="EE02CE9BC348EC26FD901934FDABFEE2" box="[548,629,304,324]" degrees="103" direction="east" minutes="50" orientation="longitude" pageId="9" pageNumber="1644" precision="925" value="103.833336">103°50'E</geoCoordinate>
,
<date id="FF888E9CC348EC26FDCA1934FD34FEE2" box="[638,746,304,324]" pageId="9" pageNumber="1644" value="1991-05" valueMax="1991-06-31" valueMin="1991-05-00">V.-VI. 1991</date>
, Jendek (3 - NHMW, cJAE).
</paragraph>
</subSubSection>
</treatment>
</document>
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