<documentid="157369008D807BDAE6E24DA25C96C572"ID-CLB-Dataset="48132"ID-DOI="10.5281/zenodo.207073"ID-GBIF-Dataset="a3815507-6100-464e-aaf6-60d6820a1eec"ID-ISSN="1175-5326"ID-Zenodo-Dep="207073"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.tables_requiresApprovalFor="existingObjects,plazi"IM.taxonomicNames_approvedBy="felipe"checkinTime="1460228776049"checkinUser="plazi"docAuthor="Torres-Carvajal, Omar, Etheridge, Richard & Queiroz, Kevin De"docDate="2011"docId="038287FDFFA5605969838AFAD0E0F84E"docLanguage="en"docName="zt02752p044.pdf"docOrigin="Zootaxa 2752"docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article"docStyleId="890A69B780ED73D6DB8551B71C8AC79E"docStyleName="Zootaxa.2009-2012.journal_article"docStyleVersion="4"docTitle="Enyalioides touzeti Torres-Carvajal et al. 2008"docType="treatment"docVersion="11"lastPageNumber="31"masterDocId="FFBBFF85FFBB604769148B6ED174FF93"masterDocTitle="A systematic revision of Neotropical lizards in the clade Hoplocercinae (Squamata: Iguania)"masterLastPageNumber="44"masterPageNumber="1"pageNumber="31"updateTime="1720202091178"updateUser="ExternalLinkService">
<mods:titleid="4A829628333429F8F95B9A80BA0CB617">A systematic revision of Neotropical lizards in the clade Hoplocercinae (Squamata: Iguania)</mods:title>
<taxonomicNameid="4C2B4D68FFA5605969838AFAD3A4FE3E"ID-CoL="6FGTM"authority="Torres-Carvajal et al. 2008"authorityName="Torres-Carvajal et al."authorityYear="2008"box="[151,720,403,430]"class="Reptilia"family="Hoplocercidae"genus="Enyalioides"kingdom="Animalia"order="Squamata"pageId="30"pageNumber="31"phylum="Chordata"rank="species"species="touzeti">
<bibRefCitationid="EFBA4B1AFFA5605968978AFDD3A4FE3E"author="Torres-Carvajal"box="[387,720,403,430]"pageId="30"pageNumber="41"refString="Torres-Carvajal, O., Almendariz, A., Valencia, J., Yanez-Munoz, M. & Reyes, J. (2008) A new species of Enyalioides (Iguanidae: Hoplocercinae) from southwestern Ecuador. Papeis Avulsos de Zoologia, 48, 227 - 235."type="journal article"year="2008">
<paragraphid="8B9436EBFFA5605969838AB4D3B3FE63"blockId="30.[151,777,474,527]"box="[151,711,474,496]"pageId="30"pageNumber="31">Proposed standard English name: Touzet’s woodlizards</paragraph>
<paragraphid="8B9436EBFFA5605969838A97D27DFD9C"blockId="30.[151,777,474,527]"box="[151,777,505,527]"pageId="30"pageNumber="31">Proposed standard Spanish name: lagartijas de palo de Touzet</paragraph>
<taxonomicNameid="4C2B4D68FFA5605969838941D3C9FDD5"ID-CoL="6FGTM"authority="Torres-Carvajal et al. (2008:228)"authorityName="Torres-Carvajal et al."authorityPageNumber="228"authorityYear="2008"box="[151,701,559,582]"class="Reptilia"family="Hoplocercidae"genus="Enyalioides"kingdom="Animalia"order="Squamata"pageId="30"pageNumber="31"phylum="Chordata"rank="species"species="touzeti">
<bibRefCitationid="EFBA4B1AFFA5605968708941D3FDFDD5"author="Torres-Carvajal"box="[356,649,559,582]"pageId="30"pageNumber="41"refString="Torres-Carvajal, O., Almendariz, A., Valencia, J., Yanez-Munoz, M. & Reyes, J. (2008) A new species of Enyalioides (Iguanidae: Hoplocercinae) from southwestern Ecuador. Papeis Avulsos de Zoologia, 48, 227 - 235."type="journal article"year="2008">
by having the following combination of characters: ventrals keeled; dorsals keeled; flank scales homogeneous in size; vertebral scales in neck region in adult males more than twice as high as vertebrals in pelvic region; gular region in males extensively covered by a dark gular patch; and dorsum and limbs without projecting scales.
(1) dorsal head scales conical or multicarinate, strongly projecting dorsally; (2) posterior superciliaries not enlarged relative to adjacent scales; (3) scales on lateral edge of skull roof just posterior to superciliaries more projecting than adjacent scales; the projection is more pronounced in adults; (4) one or two enlarged pretympanic scales present; (5) gular scales conical or multicarinate, strongly projecting ventrally; (6) dorsal neck scales homogeneous in size, mostly conical or cycloid and slightly imbricate; lateral neck scales similar in size to dorsal neck scales, sometimes with a few enlarged conical scales; (7) vertebrals larger than adjacent dorsals, forming distinct raised middorsal crest that extends onto tail as a pair of crests; (8) nuchal region with continuous and single middorsal crest; (9) dorsals small, prominently keeled, imbricate, and homogeneous in size; (10) longitudinal row of raised, enlarged scales between dorsals and flank scales present and continuous; (11) scales on flanks homogeneous in size, smaller than dorsals; (12) ventrals keeled; (13) fore limb scales keeled dorsally and ventrally; (14) hind limb scales keeled dorsally and ventrally; scattered enlarged scales absent; dorsal scales of pes homogeneous in size; (15) caudals heterogeneous, increasing in size posteriorly on each segment (6–8 scales in lateral view), not modified as conspicuous spines (
<figureCitationid="13102A6EFFA560596B6B8FBFD3B6FB7A"box="[639,706,1233,1257]"captionStart="FIGURE 3"captionStartId="12.[151,250,1517,1540]"captionTargetBox="[280,1305,321,1491]"captionTargetId="figure@12.[274,1314,306,1505]"captionTargetPageId="12"captionText="FIGURE 3. Caudal segments from the anterior third of the tail of ten species of Hoplocercinae in lateral view. (A) Enyalioides cofanorum, (B) E. heterolepis, (C) E. laticeps, (D) E. microlepis, (E) E. oshaughnessyi, (F) E. palpebralis, (G) E. praestabilis, (H) E. rubrigularis, (I) E. touzeti, and (J) Morunasaurus annularis. Scale bars = 5 mm. Illustration by D. Paucar."httpUri="https://zenodo.org/record/207076/files/figure.png"pageId="30"pageNumber="31">Fig. 3</figureCitation>
); (16) tail compressed laterally. Meristic and morphometric characters are presented in
<tableCitationid="C6A90350FFA56059688C8F9BD09FFA9E"box="[408,491,1269,1293]"captionStart="TABLE 1"captionStartId="6.[151,235,1304,1325]"captionTargetBox="[151,1436,1377,1892]"captionTargetPageId="6"captionText="TABLE 1. Summary of counts and measurements (mm) for species of Hoplocercinae. Range (first line) and mean ± standard"httpUri="http://table.plazi.org/id/DF546663FFBD604169838E76D4E8FABD"pageId="30"pageNumber="31"tableUuid="DF546663FFBD604169838E76D4E8FABD">Table 1</tableCitation>
<emphasisid="B95FEAF9FFA5605969D18E76D0E1FAA2"bold="true"box="[197,405,1304,1329]"pageId="30"pageNumber="31">Coloration in life</emphasis>
(
<figureCitationid="13102A6EFFA5605968BC8E77D09BFAA2"box="[424,495,1305,1329]"captionStart="FIGURE 4"captionStartId="14.[151,250,1897,1920]"captionTargetBox="[204,1383,190,1874]"captionTargetId="figure@14.[204,1384,184,1876]"captionTargetPageId="14"captionText="FIGURE 4. Photographs of live lizards representing twelve species of Hoplocercinae."httpUri="https://zenodo.org/record/207077/files/figure.png"pageId="30"pageNumber="31">Fig. 4</figureCitation>
). Dorsal background olive green or light brown in adults (reddish brown in juveniles), normally with yellow spots in males and large dark blotches (some with background color in the center) in females; supralabials and infralabials normally yellow; light stripe extending from mouth commisure to tympanum in some specimens; light stripe between tympanum and scapular region in some specimens; chin and gular regions in males dark; venter light brown or whitish cream, with dark irregular marks in some specimens; iris reddish brown (
<bibRefCitationid="EFBA4B1AFFA5605969F98EA3D355FA76"author="Torres-Carvajal"box="[237,545,1485,1509]"pageId="30"pageNumber="41"refString="Torres-Carvajal, O., Almendariz, A., Valencia, J., Yanez-Munoz, M. & Reyes, J. (2008) A new species of Enyalioides (Iguanidae: Hoplocercinae) from southwestern Ecuador. Papeis Avulsos de Zoologia, 48, 227 - 235."type="journal article"year="2008">
high during the day, or sleeping in a vertical position on tree fern branches at night (
<bibRefCitationid="EFBA4B1AFFA56059698A8D57D0A6F9C2"author="Torres-Carvajal"box="[158,466,1593,1617]"pageId="30"pageNumber="41"refString="Torres-Carvajal, O., Almendariz, A., Valencia, J., Yanez-Munoz, M. & Reyes, J. (2008) A new species of Enyalioides (Iguanidae: Hoplocercinae) from southwestern Ecuador. Papeis Avulsos de Zoologia, 48, 227 - 235."type="journal article"year="2008">
<figureCitationid="13102A6EFFA5605968508DEFD0E3F90A"box="[324,407,1665,1689]"captionStart="FIGURE 10"captionStartId="28.[151,250,1666,1689]"captionTargetBox="[161,1426,207,1630]"captionTargetId="figure@28.[151,1436,193,1645]"captionTargetPageId="28"captionText="FIGURE 10. Distribution of Enyalioides praestabilis (dots) and E. touzeti (triangles)."httpUri="https://zenodo.org/record/207087/files/figure.png"pageId="30"pageNumber="31">Fig. 10</figureCitation>
). This species is not known to occur in sympatry with other hoplocercines, although
<figureCitationid="13102A6EFFA560596A0C8DCBD211F92E"box="[792,869,1701,1725]"captionStart="FIGURE 6"captionStartId="20.[151,250,1959,1982]"captionTargetBox="[159,1429,530,1933]"captionTargetId="figure@20.[151,1436,517,1938]"captionTargetPageId="20"captionText="FIGURE 6. Distribution of Enyalioides heterolepis (triangles) and E. laticeps (dots)."httpUri="https://zenodo.org/record/207083/files/figure.png"pageId="30"pageNumber="31">Figs. 6</figureCitation>
and
<figureCitationid="13102A6EFFA560596A888DC8D2D8F92E"box="[924,940,1702,1725]"captionStart="FIGURE 7"captionStartId="23.[151,250,1564,1587]"captionTargetBox="[197,1383,204,1538]"captionTargetId="figure@23.[189,1403,169,1564]"captionTargetPageId="23"captionText="FIGURE 7. Distribution of Enyalioides microlepis (dots) and E. oshaughnessyi (triangles). A, Alto Tambo; B, Bilsa Ecological Reserve (see Remarks under E. oshaughnessyi)."httpUri="https://zenodo.org/record/207084/files/figure.png"pageId="30"pageNumber="31">7</figureCitation>
as a separate species, which suggests that this could be a case of fixed morphological differences evolving prior to the evolution of reciprocal monophyly of mtDNA alleles. Alternatively,
