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<document id="4B7E0B1B15BD2DF553329CD801140792" ID-DOI="10.5281/zenodo.294209" ID-GBIF-Dataset="79e02464-c04f-4443-895a-457c4782a91b" ID-ISSN="1175-5326" ID-Zenodo-Dep="294209" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460230905155" checkinUser="plazi" docAuthor="Tarasov, Sergey, Krikken, Jan &amp; Huijbregts, Johannes" docDate="2010" docId="03E0507D1D2C1F30FF08FF2ACB5AFCE0" docLanguage="en" docName="zt02490p068.pdf" docOrigin="Zootaxa 2490" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Onthophagus (Parascatonomus) alexeevi Tarasov, Krikken &amp; Huijbregts, sp. n." docType="treatment" docVersion="7" lastPageNumber="67" masterDocId="FFD928051D2E1F34FF9FFFBDC858FFCC" masterDocTitle="The Indochinese members of the Onthophagus (Parascatonomus) aurifex species group (Coleoptera: Scarabaeidae), with the description of a new species" masterLastPageNumber="68" masterPageNumber="63" pageNumber="65" updateTime="1698238958640" updateUser="plazi">
<mods:mods id="4F21863AF906F737892B5A65FC519943" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="528FCA35F7978341A0424F6AB7D579FC">
<mods:title id="9EC18E53E0220613742C3944750D910F">The Indochinese members of the Onthophagus (Parascatonomus) aurifex species group (Coleoptera: Scarabaeidae), with the description of a new species</mods:title>
</mods:titleInfo>
<mods:name id="9039D56ACD49EBC13ED6AC0D6B917C73" type="personal">
<mods:role id="A502AF48E35DEB4BA4EEA146D8C56AA4">
<mods:roleTerm id="3846617B06C84AE8CA53A8E60DA26358">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C1617C8A5FDE9E274C1740F9AFE22B8D">Tarasov, Sergey</mods:namePart>
</mods:name>
<mods:name id="55F932FFABED30823B226C4403233FB6" type="personal">
<mods:role id="FF8DDFB5AEC44140D7937CA655CD3507">
<mods:roleTerm id="2E73095F93182FD8DA772A209CBA311F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="35505944AB66BA2A907C7BBBDFF50D95">Krikken, Jan</mods:namePart>
</mods:name>
<mods:name id="4166EB87D0507830E5A0BBB65A182B2D" type="personal">
<mods:role id="B555178F8C8AD30FC71BACBF1308EEB0">
<mods:roleTerm id="25234F6261D2CFADEDF17A134E2F5FAD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="73B6FD01B58F271D39F868C75B6367A4">Huijbregts, Johannes</mods:namePart>
</mods:name>
<mods:typeOfResource id="BAF8551A5F0F519D423C24AC391BBE61">text</mods:typeOfResource>
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<mods:titleInfo id="78E18F192F2B9127EEE2DCFF2CBC3B77">
<mods:title id="C38603C5624661ABA5BFB457F4C83856">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C5930FCE8BE201562178802DD106CD93">
<mods:date id="EB3F70351977D05C61ACF72731AA5149">2010</mods:date>
<mods:detail id="A3D1B63C55E9C395FCFEE7A09D0A548B" type="volume">
<mods:number id="33DF4162B9807FC89B8E5F1332CCEE2A">2490</mods:number>
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<mods:start id="5B7347FF4A7CFE6F24F072E227825FE7">63</mods:start>
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<mods:classification id="CB5EC6D2D71B9359D8AB868214AC1243">journal article</mods:classification>
<mods:identifier id="782FA7469C2882CA113A13E25D8B919A" type="DOI">10.5281/zenodo.294209</mods:identifier>
<mods:identifier id="05B6BFD31173D2C88B84B713AF879EAA" type="GBIF-Dataset">79e02464-c04f-4443-895a-457c4782a91b</mods:identifier>
<mods:identifier id="B5727BDB6C7BEE8902DE137FD40164D4" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6C7CF88217CF05D19FDD4E80E3C47C29" type="Zenodo-Dep">294209</mods:identifier>
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<treatment id="03E0507D1D2C1F30FF08FF2ACB5AFCE0" ID-DOI="http://doi.org/10.5281/zenodo.6211453" ID-GBIF-Taxon="119387305" ID-Zenodo-Dep="6211453" LSID="urn:lsid:plazi:treatment:03E0507D1D2C1F30FF08FF2ACB5AFCE0" httpUri="http://treatment.plazi.org/id/03E0507D1D2C1F30FF08FF2ACB5AFCE0" lastPageId="4" lastPageNumber="67" pageId="2" pageNumber="65">
<subSubSection id="C353B2E01D2C1F36FF08FF2AC902FF18" pageId="2" pageNumber="65" type="nomenclature">
<paragraph id="8BF6E16B1D2C1F36FF08FF2ACC3AFF7D" blockId="2.[151,1122,151,212]" box="[151,1122,151,177]" pageId="2" pageNumber="65">
<heading id="D0BE56071D2C1F36FF08FF2ACC3AFF7D" bold="true" box="[151,1122,151,177]" fontSize="11" level="1" pageId="2" pageNumber="65" reason="1">
<emphasis id="B93D3D791D2C1F36FF08FF2ACC3AFF7D" bold="true" box="[151,1122,151,177]" pageId="2" pageNumber="65">
<taxonomicName id="4C499AE81D2C1F36FF08FF2ACC41FF7D" ID-CoL="49MRM" authority="Tarasov, Krikken &amp; Huijbregts" authorityName="Tarasov, Krikken &amp; Huijbregts" box="[151,1049,151,177]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="alexeevi" status="sp. nov." subGenus="Parascatonomus">
<emphasis id="B93D3D791D2C1F36FF08FF2ACAD1FF7D" bold="true" box="[151,649,151,177]" italics="true" pageId="2" pageNumber="65">Onthophagus (Parascatonomus) alexeevi</emphasis>
Tarasov, Krikken &amp; Huijbregts
</taxonomicName>
<taxonomicNameLabel id="A20E80021D2C1F36FB80FF2ACC3AFF7D" box="[1055,1122,151,177]" pageId="2" pageNumber="65" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF6E16B1D2C1F36FF08FF07C902FF18" blockId="2.[151,1122,151,212]" box="[151,346,186,212]" pageId="2" pageNumber="65">
(
<figureCitation id="1372FDEE1D2C1F36FF00FF07C8A9FF18" box="[159,241,186,212]" captionStart="FIGURES 1" captionStartId="2.[151,269,1368,1392]" captionTargetBox="[580,996,816,1331]" captionTargetId="figure@2.[577,1010,791,1344]" captionTargetPageId="2" captionText="FIGURES 1. Onthophagus (Parascatonomus) alexeevi, dorsal view, holotype." httpUri="https://zenodo.org/record/294210/files/figure.png" pageId="2" pageNumber="65">Figs. 1</figureCitation>
,
<figureCitation id="1372FDEE1D2C1F36FF60FF07C955FF18" box="[255,269,186,212]" captionStart="FIGURE 2" captionStartId="3.[151,255,1688,1712]" captionTargetBox="[166,1436,283,1627]" captionTargetId="figure@3.[151,1436,269,1667]" captionTargetPageId="3" captionText="FIGURE 2. Aedeagi of O. aurifex - group species. a – f, m — O. semifex (a – c, f, m — paratypes, d – e — Vietnam); g – l — O. alexeevi (g – j, l — paratypes, k — holotype); a – k — aedeagus lateral view; l – m — aedeagus dorsal view; a – e, g – h, k – m — aedeagus photographed in glycerin; f, g — aedeagus photographed after drying." httpUri="https://zenodo.org/record/294211/files/figure.png" pageId="2" pageNumber="65">2</figureCitation>
g–k, l)
</paragraph>
</subSubSection>
<subSubSection id="C353B2E01D2C1F36FF08FEB8CCBAFD13" pageId="2" pageNumber="65" type="materials_examined">
<paragraph id="8BF6E16B1D2C1F36FF08FEB8CC00FED3" blockId="2.[151,1437,261,735]" box="[151,1112,261,287]" pageId="2" pageNumber="65">
<emphasis id="B93D3D791D2C1F36FF08FEB8C9D3FED3" bold="true" box="[151,395,261,287]" pageId="2" pageNumber="65">Material examined.</emphasis>
<emphasis id="B93D3D791D2C1F36FE0DFEB8C9A6FED2" box="[402,510,261,286]" italics="true" pageId="2" pageNumber="65">
<typeStatus id="54F25FC91D2C1F36FE0DFEB8C9A6FED2" box="[402,510,261,286]" pageId="2" pageNumber="65" type="holotype">Holotype</typeStatus>
</emphasis>
(
<collectionCode id="ED5879AE1D2C1F36FD92FEB8CA30FED3" box="[525,616,261,287]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="2" pageNumber="65">ZMUC</collectionCode>
), male major bearing the following labels:
</paragraph>
<paragraph id="8BF6E16B1D2C1F36FF5AFE97C9A3FEA6" blockId="2.[151,1437,261,735]" pageId="2" pageNumber="65">
1. White, printed: S[outh]
<collectingCountry id="F35EA1FB1D2C1F36FD9CFE97CAFCFE88" box="[515,676,298,324]" name="Cambodia" pageId="2" pageNumber="65">CAMBODIA</collectingCountry>
Sianoukville h=
<quantity id="4CB14C8E1D2C1F36FCF4FE97CB83FE88" box="[875,987,298,324]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="2" pageNumber="65" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50–150m</quantity>
3 carrion traps disturb tropical forest 12–
<date id="FFF7C7AB1D2C1F36FF5EFEEDC91FFEA6" box="[193,327,336,362]" pageId="2" pageNumber="65" value="2008-05-16">16.05.2008</date>
leg. S. Tarasov
</paragraph>
<paragraph id="8BF6E16B1D2C1F36FF5AFEC8C9DFFE78" blockId="2.[151,1437,261,735]" pageId="2" pageNumber="65">
2. Orange, printed:
<typeStatus id="54F25FC91D2C1F36FE31FEC8CA06FE43" box="[430,606,373,399]" pageId="2" pageNumber="65" type="holotype">HOLOTYPUS</typeStatus>
<taxonomicName id="4C499AE81D2C1F36FDF9FEC8C94BFE78" authority="S. Tarasov, J. Krikken &amp; J. Huijbregts" authorityName="S. Tarasov, J. Krikken &amp; J. Huijbregts" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="alexeevi" subGenus="Parascatonomus">
<emphasis id="B93D3D791D2C1F36FDF9FEC8CC0AFE42" box="[614,1106,373,398]" italics="true" pageId="2" pageNumber="65">Onthophagus (Parascatonomus) alexeevi</emphasis>
S. Tarasov, J. Krikken &amp; J. Huijbregts
</taxonomicName>
det. 2009
</paragraph>
<paragraph id="8BF6E16B1D2C1F36FF5AFE7DCCBAFD13" blockId="2.[151,1437,261,735]" pageId="2" pageNumber="65">
<emphasis id="B93D3D791D2C1F36FF5AFE7DC918FE15" box="[197,320,448,473]" italics="true" pageId="2" pageNumber="65">
<typeStatus id="54F25FC91D2C1F36FF5AFE7DC918FE15" box="[197,320,448,473]" pageId="2" pageNumber="65" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D4F2AE21D2C1F36FECEFE7DC9A2FE16" box="[337,506,448,474]" pageId="2" pageNumber="65" type="generic">53 specimens</specimenCount>
, South
<collectingCountry id="F35EA1FB1D2C1F36FDC9FE7DCA8EFE16" box="[598,726,448,474]" name="Cambodia" pageId="2" pageNumber="65">Cambodia</collectingCountry>
, Sianoukville h=
<quantity id="4CB14C8E1D2C1F36FC3AFE7DCC42FE16" box="[933,1050,448,474]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="2" pageNumber="65" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50–150m</quantity>
, 3 carrion traps disturb tropical forest, 12–
<date id="FFF7C7AB1D2C1F36FE83FE58C9F2FE33" box="[284,426,485,511]" pageId="2" pageNumber="65" value="2008-05-16">16.05.2008</date>
leg. S. Tarasov (
<collectionCode id="ED5879AE1D2C1F36FD1FFE58CAB8FE33" box="[640,736,485,511]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="2" pageNumber="65">ZMUC</collectionCode>
); 1 ɗ &amp; 1 Ψ,
<collectingCountry id="F35EA1FB1D2C1F36FC15FE58CBA4FE33" box="[906,1020,485,511]" name="Thailand" pageId="2" pageNumber="65">Thailand</collectingCountry>
, Umphang river,
<geoCoordinate id="EE7D87AC1D2C1F36FB48FE58CD6BFE33" box="[1239,1331,485,511]" direction="north" orientation="latitude" pageId="2" pageNumber="65" precision="555" value="16.07">16.07N</geoCoordinate>
<geoCoordinate id="EE7D87AC1D2C1F36FAA3FE58CDCFFE33" box="[1340,1431,485,511]" direction="east" orientation="longitude" pageId="2" pageNumber="65" precision="555" value="99.0">99.00E</geoCoordinate>
, h
<quantity id="4CB14C8E1D2C1F36FF39FDB7C8A4FDE8" box="[166,252,522,548]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="65" unit="m" value="1000.0">1000m</quantity>
, 28/4–6/5.91 leg. D. Kral (
<collectionCode id="ED5879AE1D2C1F36FDDDFDB7CADEFDE8" box="[578,646,522,548]" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="2" pageNumber="65">ZISP</collectionCode>
); 2 ɗ, W
<collectingCountry id="F35EA1FB1D2C1F36FD64FDB7CB31FDE8" box="[763,873,522,548]" name="Thailand" pageId="2" pageNumber="65">Thailand</collectingCountry>
,
<quantity id="4CB14C8E1D2C1F36FCE9FDB7CBE6FDE8" box="[886,958,522,548]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="65" unit="m" value="300.0">300m</quantity>
, Thung Yai Wildlife Sanctuary,
<geoCoordinate id="EE7D87AC1D2C1F36FADCFDB7CDC4FDE8" box="[1347,1436,522,548]" direction="north" orientation="latitude" pageId="2" pageNumber="65" precision="555" value="15.28">15.28N</geoCoordinate>
<geoCoordinate id="EE7D87AC1D2C1F36FF08FD8DC8B5FD86" box="[151,237,560,586]" direction="east" orientation="longitude" pageId="2" pageNumber="65" precision="555" value="98.48">98.48E</geoCoordinate>
, Tak Province, Umphang District, Song Bae Stream, 18–
<date id="FFF7C7AB1D2C1F36FC17FD8DCC5AFD86" box="[904,1026,560,586]" pageId="2" pageNumber="65" value="1988-04-27">27.4.1988</date>
, M.J.D. Brendell, FIT B (
<collectionCode id="ED5879AE1D2C1F36FAAFFD8DCDD7FD86" LSID="urn:lsid:biocol.org:col:34992" box="[1328,1423,560,586]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="2" pageNumber="65">RMNH</collectionCode>
); 1 Ψ, same data and locality, but carrion trap (fish) (
<collectionCode id="ED5879AE1D2C1F36FD67FDE8CB0FFDA3" box="[760,855,597,623]" pageId="2" pageNumber="65">BMNH</collectionCode>
); 1ɗ &amp; 1Ψ same locality and date, but evergreen rain forest M.J.D. Brendell, (
<collectionCode id="ED5879AE1D2C1F36FE71FDC7CA15FD58" box="[494,589,634,660]" pageId="2" pageNumber="65">BMNH</collectionCode>
); 2 ɗ &amp; 1Ψ, W
<collectingCountry id="F35EA1FB1D2C1F36FC96FDC7CB2DFD58" box="[777,885,634,660]" name="Thailand" pageId="2" pageNumber="65">Thailand</collectingCountry>
,
<quantity id="4CB14C8E1D2C1F36FC1EFDC7CB9FFD58" box="[897,967,634,660]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="65" unit="m" value="300.0">300m</quantity>
, Thung Yai Wildlife Sanctuary,
<geoCoordinate id="EE7D87AC1D2C1F36FADBFDC7CDC4FD58" box="[1348,1436,634,660]" direction="north" orientation="latitude" pageId="2" pageNumber="65" precision="555" value="15.3">15.30N</geoCoordinate>
<geoCoordinate id="EE7D87AC1D2C1F36FF08FD1DC8B5FD76" box="[151,237,672,698]" direction="east" orientation="longitude" pageId="2" pageNumber="65" precision="555" value="98.48">98.48E</geoCoordinate>
, Tak Province, Umphang District, Mae Chan/Mae Klong confluence, 27.4–
<date id="FFF7C7AB1D2C1F36FBFBFD1DCC94FD76" box="[1124,1228,672,698]" pageId="2" pageNumber="65" value="1988-05-06">6.5.1988</date>
, M.J.D. Brendell, oak/bamboo forest (
<collectionCode id="ED5879AE1D2C1F36FE1EFD78C9B8FD13" box="[385,480,709,735]" pageId="2" pageNumber="65">BMNH</collectionCode>
); 1ɗ same data and locality, but edge of Karen clearing (
<collectionCode id="ED5879AE1D2C1F36FBE8FD78CC8EFD13" box="[1143,1238,709,735]" pageId="2" pageNumber="65">BMNH</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="DF36B1E31D2C1F36FF08FAE5CBB9FABC" box="[151,993,1368,1392]" httpUri="https://zenodo.org/record/294210/files/figure.png" pageId="2" pageNumber="65" targetBox="[580,996,816,1331]" targetPageId="2">
<paragraph id="8BF6E16B1D2C1F36FF08FAE5CBB9FABC" blockId="2.[151,993,1368,1392]" box="[151,993,1368,1392]" pageId="2" pageNumber="65">
<emphasis id="B93D3D791D2C1F36FF08FAE5C970FABC" bold="true" box="[151,296,1368,1392]" pageId="2" pageNumber="65">FIGURES 1.</emphasis>
<taxonomicName id="4C499AE81D2C1F36FEB1FAE4CAB0FABC" box="[302,744,1369,1392]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="alexeevi" subGenus="Parascatonomus">
<emphasis id="B93D3D791D2C1F36FEB1FAE4CAB0FABC" box="[302,744,1369,1392]" italics="true" pageId="2" pageNumber="65">Onthophagus (Parascatonomus) alexeevi</emphasis>
</taxonomicName>
, dorsal view, holotype.
</paragraph>
</caption>
<subSubSection id="C353B2E01D2C1F36FF5AFA25CDC0F82F" pageId="2" pageNumber="65" type="description">
<paragraph id="8BF6E16B1D2C1F36FF5AFA25CB8DFA14" blockId="2.[151,1437,1432,2019]" pageId="2" pageNumber="65">
<emphasis id="B93D3D791D2C1F36FF5AFA25C906FA7E" bold="true" box="[197,350,1432,1458]" pageId="2" pageNumber="65">Description.</emphasis>
Head and pronotum cupreous or with greenish metallic luster, elytra black, matt, underside black with metallic cupreous or/and green luster. Length 9.0–
<quantity id="4CB14C8E1D2C1F36FCF8FA03CB88FA14" box="[871,976,1470,1496]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="2" pageNumber="65" unit="mm" value="11.5">11.5 mm</quantity>
.
</paragraph>
<paragraph id="8BF6E16B1D2C1F36FF5AFA5ECCE4F984" blockId="2.[151,1437,1432,2019]" pageId="2" pageNumber="65">
MALE major (
<figureCitation id="1372FDEE1D2C1F36FEEBFA5EC9E1FA31" box="[372,441,1507,1533]" captionStart="FIGURES 1" captionStartId="2.[151,269,1368,1392]" captionTargetBox="[580,996,816,1331]" captionTargetId="figure@2.[577,1010,791,1344]" captionTargetPageId="2" captionText="FIGURES 1. Onthophagus (Parascatonomus) alexeevi, dorsal view, holotype." httpUri="https://zenodo.org/record/294210/files/figure.png" pageId="2" pageNumber="65">Fig. 1</figureCitation>
). Head rounded; clypeal margin anteriorly distinctly reflexed, apex with short tooth, tip rounded; frontoclypeal ridge slightly expressed as smooth arcuate strip; clypeus densely, transversely rugulate; vertex simple with slightly protruding plate; eyes completely divided by canthus.
</paragraph>
<paragraph id="8BF6E16B1D2C1F36FF5AF9EECDC4F911" blockId="2.[151,1437,1432,2019]" pageId="2" pageNumber="65">Pronotum convex, with slightly steeper anterior declivity in major specimens; anterior and lateral sides of pronotum marginate, base marginate only laterally and medially angulate; anterolateral pronotal angle slightly obtuse; pronotal disc covered with small, simple, sparse punctures (separated by 1–2 puncture diameters) becoming denser and granulose laterally; anterior part of propleura with ridge reaching prosternal lateral side.</paragraph>
<paragraph id="8BF6E16B1D2C1F36FF5AF954CCC0F8CF" blockId="2.[151,1437,1432,2019]" box="[197,1176,1769,1795]" pageId="2" pageNumber="65">Elytra with short, yellow, fine setae; sparsely covered with small, simple punctures.</paragraph>
<paragraph id="8BF6E16B1D2C1F36FF5AF8B3CA8CF8E4" blockId="2.[151,1437,1432,2019]" box="[197,724,1806,1832]" pageId="2" pageNumber="65">Metasternum anteriorly raised, prow-shaped.</paragraph>
<paragraph id="8BF6E16B1D2C1F36FF5AF88ECD46F8BF" blockId="2.[151,1437,1432,2019]" pageId="2" pageNumber="65">Protibia with 4 outer teeth, apico-internal angle with small tooth; apical spur modified in males: small, acuminate, slightly bent downward and inward; mesotibial and metatibial apex bearing short setae.</paragraph>
<paragraph id="8BF6E16B1D2C1F36FF5AF8C3CB01F854" blockId="2.[151,1437,1432,2019]" box="[197,857,1918,1944]" pageId="2" pageNumber="65">
Aedeagus as in
<figureCitation id="1372FDEE1D2C1F36FEE2F8C3C995F854" box="[381,461,1918,1944]" captionStart="FIGURE 2" captionStartId="3.[151,255,1688,1712]" captionTargetBox="[166,1436,283,1627]" captionTargetId="figure@3.[151,1436,269,1667]" captionTargetPageId="3" captionText="FIGURE 2. Aedeagi of O. aurifex - group species. a – f, m — O. semifex (a – c, f, m — paratypes, d – e — Vietnam); g – l — O. alexeevi (g – j, l — paratypes, k — holotype); a – k — aedeagus lateral view; l – m — aedeagus dorsal view; a – e, g – h, k – m — aedeagus photographed in glycerin; f, g — aedeagus photographed after drying." httpUri="https://zenodo.org/record/294211/files/figure.png" pageId="2" pageNumber="65">
Figs.
<quantity id="4CB14C8E1D2C1F36FE20F8C3C995F854" box="[447,461,1918,1944]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="2" pageNumber="65" unit="g" value="2.0">2</quantity>
</figureCitation>
g–k, l (
<figureCitation id="1372FDEE1D2C1F36FDBFF8C3CA3EF854" box="[544,614,1918,1944]" captionStart="FIGURE 2" captionStartId="3.[151,255,1688,1712]" captionTargetBox="[166,1436,283,1627]" captionTargetId="figure@3.[151,1436,269,1667]" captionTargetPageId="3" captionText="FIGURE 2. Aedeagi of O. aurifex - group species. a – f, m — O. semifex (a – c, f, m — paratypes, d – e — Vietnam); g – l — O. alexeevi (g – j, l — paratypes, k — holotype); a – k — aedeagus lateral view; l – m — aedeagus dorsal view; a – e, g – h, k – m — aedeagus photographed in glycerin; f, g — aedeagus photographed after drying." httpUri="https://zenodo.org/record/294211/files/figure.png" pageId="2" pageNumber="65">
Fig.
<quantity id="4CB14C8E1D2C1F36FDC8F8C3CA3EF854" box="[599,614,1918,1944]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="2" pageNumber="65" unit="g" value="2.0">2</quantity>
</figureCitation>
g is of the
<typeStatus id="54F25FC91D2C1F36FD7EF8C3CB15F854" box="[737,845,1918,1944]" pageId="2" pageNumber="65" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8BF6E16B1D2C1F36FF5AF81ECC6BF871" blockId="2.[151,1437,1432,2019]" box="[197,1075,1955,1981]" pageId="2" pageNumber="65">FEMALE. Head and pronotum armature do not differ from those of males.</paragraph>
<paragraph id="8BF6E16B1D2C1F36FF5AF874CDC0F82F" blockId="2.[151,1437,1432,2019]" box="[197,1432,1993,2019]" pageId="2" pageNumber="65">VARIATION. Slightly expressed in minor specimens by flatter pronotum with slightly denser punctation.</paragraph>
</subSubSection>
<subSubSection id="C353B2E01D2D1F30FF5AFF25CD58FDEB" lastPageId="4" lastPageNumber="67" pageId="3" pageNumber="66" type="materials_examined">
<paragraph id="8BF6E16B1D2D1F37FF5AFF25CD1FFF1B" blockId="3.[151,1437,152,215]" pageId="3" pageNumber="66">
<typeStatus id="54F25FC91D2D1F37FF5AFF25C93FFF7E" box="[197,359,152,178]" pageId="3" pageNumber="66" type="holotype">HOLOTYPE</typeStatus>
. The body coloration and length of the
<typeStatus id="54F25FC91D2D1F37FCDCFF25CBF4FF7E" box="[835,940,152,178]" pageId="3" pageNumber="66" type="holotype">holotype</typeStatus>
as follows; head and pronotum cupreous with greenish metallic luster, underside black with metallic cupreous and green luster. Length
<quantity id="4CB14C8E1D2D1F37FB7AFF00CD1AFF1B" box="[1253,1346,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="3" pageNumber="66" unit="mm" value="9.7">9.7 mm</quantity>
.
</paragraph>
<caption id="DF36B1E31D2D1F37FF08F925CC70F93C" httpUri="https://zenodo.org/record/294211/files/figure.png" pageId="3" pageNumber="66" targetBox="[166,1436,283,1627]" targetPageId="3">
<paragraph id="8BF6E16B1D2D1F37FF08F925CC70F93C" blockId="3.[151,1436,1688,1776]" pageId="3" pageNumber="66">
<emphasis id="B93D3D791D2D1F37FF08F925C941F97C" bold="true" box="[151,281,1688,1712]" pageId="3" pageNumber="66">FIGURE 2.</emphasis>
Aedeagi of
<emphasis id="B93D3D791D2D1F37FE02F924CA4AF97C" box="[413,530,1688,1712]" italics="true" pageId="3" pageNumber="66">
<taxonomicName id="4C499AE81D2D1F37FE02F924CA51F97C" box="[413,521,1689,1712]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="aurifex">O. aurifex</taxonomicName>
<emphasis id="B93D3D791D2D1F37FD96F925CA4AF97C" bold="true" box="[521,530,1688,1712]" italics="true" pageId="3" pageNumber="66">-</emphasis>
</emphasis>
group species. a–f, m —
<taxonomicName id="4C499AE81D2D1F37FC81F924CBD7F97C" box="[798,911,1689,1712]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="semifex">
<emphasis id="B93D3D791D2D1F37FC81F924CBD7F97C" box="[798,911,1689,1712]" italics="true" pageId="3" pageNumber="66">O. semifex</emphasis>
</taxonomicName>
(a–c, f, m — paratypes, d–e — Vietnam); g–l —
<taxonomicName id="4C499AE81D2D1F37FF08F904C956F91C" box="[151,270,1721,1744]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="alexeevi">
<emphasis id="B93D3D791D2D1F37FF08F904C956F91C" box="[151,270,1721,1744]" italics="true" pageId="3" pageNumber="66">O. alexeevi</emphasis>
</taxonomicName>
(g–j, l — paratypes, k — holotype); a–k — aedeagus lateral view; l–m — aedeagus dorsal view; a–e, g–h, k– m — aedeagus photographed in glycerin; f, g — aedeagus photographed after drying.
</paragraph>
</caption>
<paragraph id="8BF6E16B1D2D1F30FF5AF8A5CC25FE10" blockId="3.[151,1436,1816,2029]" lastBlockId="4.[151,1437,152,812]" lastPageId="4" lastPageNumber="67" pageId="3" pageNumber="66">
<emphasis id="B93D3D791D2D1F37FF5AF8A5C981F8FE" bold="true" box="[197,473,1816,1842]" pageId="3" pageNumber="66">Differential diagnosis.</emphasis>
The new species is most similar to three other species of
<taxonomicName id="4C499AE81D2D1F37FB16F8A4CD5AF8FE" box="[1161,1282,1817,1842]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="aurifex">
<emphasis id="B93D3D791D2D1F37FB16F8A4CD5AF8FE" box="[1161,1282,1817,1842]" italics="true" pageId="3" pageNumber="66">
<collectingCountry id="F35EA1FB1D2D1F37FB16F8A4CCFAF8FE" box="[1161,1186,1817,1842]" name="Iceland" pageId="3" pageNumber="66">O</collectingCountry>
. aurifex
</emphasis>
</taxonomicName>
complex:
<emphasis id="B93D3D791D2D1F37FAE0F8A4C9FEF89B" italics="true" pageId="3" pageNumber="66">
<taxonomicName id="4C499AE81D2D1F37FAE0F8A4C8AAF89B" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="semifex">
<collectingCountry id="F35EA1FB1D2D1F37FAE0F8A4CDCFF8FE" box="[1407,1431,1817,1842]" name="Iceland" pageId="3" pageNumber="66">O</collectingCountry>
. semifex
</taxonomicName>
,
<taxonomicName id="4C499AE81D2D1F37FF61F883C9FEF89B" box="[254,422,1854,1879]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="semiaureus">
<collectingCountry id="F35EA1FB1D2D1F37FF61F883C94EF89B" box="[254,278,1854,1879]" name="Iceland" pageId="3" pageNumber="66">O</collectingCountry>
. semiaureus
</taxonomicName>
</emphasis>
and
<taxonomicName id="4C499AE81D2D1F37FE7EF883CA00F89B" box="[481,600,1854,1879]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="aurifex">
<emphasis id="B93D3D791D2D1F37FE7EF883CA00F89B" box="[481,600,1854,1879]" italics="true" pageId="3" pageNumber="66">
<collectingCountry id="F35EA1FB1D2D1F37FE7EF883C9A1F89B" box="[481,505,1854,1879]" name="Iceland" pageId="3" pageNumber="66">O</collectingCountry>
. aurifex
</emphasis>
</taxonomicName>
, which also have the upper side of the forebody shiny metallic. This is in contrast to the other species of the
<taxonomicName id="4C499AE81D2D1F37FDC3F8D9CA8EF8B1" box="[604,726,1892,1917]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="aurifex">
<emphasis id="B93D3D791D2D1F37FDC3F8D9CA8EF8B1" box="[604,726,1892,1917]" italics="true" pageId="3" pageNumber="66">
<collectingCountry id="F35EA1FB1D2D1F37FDC3F8D9CA2DF8B1" box="[604,629,1892,1917]" name="Iceland" pageId="3" pageNumber="66">O</collectingCountry>
. aurifex
</emphasis>
</taxonomicName>
group belonging to
<taxonomicName id="4C499AE81D2D1F37FC54F8D9CC37F8B1" box="[971,1135,1892,1917]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="sarawacus">
<emphasis id="B93D3D791D2D1F37FC54F8D9CC37F8B1" box="[971,1135,1892,1917]" italics="true" pageId="3" pageNumber="66">
<collectingCountry id="F35EA1FB1D2D1F37FC54F8D9CBBCF8B1" box="[971,996,1892,1917]" name="Iceland" pageId="3" pageNumber="66">O</collectingCountry>
. sarawacus
</emphasis>
</taxonomicName>
complex, which have an entirely black dorsal surface. The new species can only be reliably separated from these three species using the aedeagus. Among these three species, the aedeagal shape of
<taxonomicName id="4C499AE81D2D1F37FC2DF813CC65F80B" box="[946,1085,1966,1991]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="alexeevi">
<emphasis id="B93D3D791D2D1F37FC2DF813CC65F80B" box="[946,1085,1966,1991]" italics="true" pageId="3" pageNumber="66">
<collectingCountry id="F35EA1FB1D2D1F37FC2DF813CB93F80B" box="[946,971,1966,1991]" name="Iceland" pageId="3" pageNumber="66">O</collectingCountry>
. alexeevi
</emphasis>
</taxonomicName>
is most similar to that of
<taxonomicName id="4C499AE81D2D1F37FAE0F813C8A8F821" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="semifex">
<emphasis id="B93D3D791D2D1F37FAE0F813C8A8F821" italics="true" pageId="3" pageNumber="66">
<collectingCountry id="F35EA1FB1D2D1F37FAE0F813CDC0F80B" box="[1407,1432,1966,1991]" name="Iceland" pageId="3" pageNumber="66">O</collectingCountry>
. semifex
</emphasis>
</taxonomicName>
but differs in the sclerotized anterior part of parameral apex, which is thinner and without the lower tooth-like short appendage (
<figureCitation id="1372FDEE1D2A1F30FE7EFF25CA6AFF7E" box="[481,562,152,178]" captionStart="FIGURE 2" captionStartId="3.[151,255,1688,1712]" captionTargetBox="[166,1436,283,1627]" captionTargetId="figure@3.[151,1436,269,1667]" captionTargetPageId="3" captionText="FIGURE 2. Aedeagi of O. aurifex - group species. a – f, m — O. semifex (a – c, f, m — paratypes, d – e — Vietnam); g – l — O. alexeevi (g – j, l — paratypes, k — holotype); a – k — aedeagus lateral view; l – m — aedeagus dorsal view; a – e, g – h, k – m — aedeagus photographed in glycerin; f, g — aedeagus photographed after drying." httpUri="https://zenodo.org/record/294211/files/figure.png" pageId="4" pageNumber="67">
Figs.
<quantity id="4CB14C8E1D2A1F30FDBBFF25CA6AFF7E" box="[548,562,152,178]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="4" pageNumber="67" unit="g" value="2.0">2</quantity>
</figureCitation>
g–k, in
<figureCitation id="1372FDEE1D2A1F30FD0AFF25CABFFF7E" box="[661,743,152,178]" captionStart="FIGURE 2" captionStartId="3.[151,255,1688,1712]" captionTargetBox="[166,1436,283,1627]" captionTargetId="figure@3.[151,1436,269,1667]" captionTargetPageId="3" captionText="FIGURE 2. Aedeagi of O. aurifex - group species. a – f, m — O. semifex (a – c, f, m — paratypes, d – e — Vietnam); g – l — O. alexeevi (g – j, l — paratypes, k — holotype); a – k — aedeagus lateral view; l – m — aedeagus dorsal view; a – e, g – h, k – m — aedeagus photographed in glycerin; f, g — aedeagus photographed after drying." httpUri="https://zenodo.org/record/294211/files/figure.png" pageId="4" pageNumber="67">
Figs.
<quantity id="4CB14C8E1D2A1F30FD48FF25CABFFF7E" box="[727,743,152,178]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="4" pageNumber="67" unit="g" value="2.0">2</quantity>
</figureCitation>
g, j indicated with circle) present in
<taxonomicName id="4C499AE81D2A1F30FB0EFF25CD55FF7D" box="[1169,1293,152,177]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="semifex">
<emphasis id="B93D3D791D2A1F30FB0EFF25CD55FF7D" box="[1169,1293,152,177]" italics="true" pageId="4" pageNumber="67">
<collectingCountry id="F35EA1FB1D2A1F30FB0EFF25CCF1FF7D" box="[1169,1193,152,177]" name="Iceland" pageId="4" pageNumber="67">O</collectingCountry>
. semifex
</emphasis>
</taxonomicName>
(
<figureCitation id="1372FDEE1D2A1F30FA82FF25CD36FF7E" box="[1309,1390,152,178]" captionStart="FIGURE 2" captionStartId="3.[151,255,1688,1712]" captionTargetBox="[166,1436,283,1627]" captionTargetId="figure@3.[151,1436,269,1667]" captionTargetPageId="3" captionText="FIGURE 2. Aedeagi of O. aurifex - group species. a – f, m — O. semifex (a – c, f, m — paratypes, d – e — Vietnam); g – l — O. alexeevi (g – j, l — paratypes, k — holotype); a – k — aedeagus lateral view; l – m — aedeagus dorsal view; a – e, g – h, k – m — aedeagus photographed in glycerin; f, g — aedeagus photographed after drying." httpUri="https://zenodo.org/record/294211/files/figure.png" pageId="4" pageNumber="67">Figs. 2</figureCitation>
a–f, in
<figureCitation id="1372FDEE1D2A1F30FF28FF00C954FF1B" box="[183,268,189,215]" captionStart="FIGURE 2" captionStartId="3.[151,255,1688,1712]" captionTargetBox="[166,1436,283,1627]" captionTargetId="figure@3.[151,1436,269,1667]" captionTargetPageId="3" captionText="FIGURE 2. Aedeagi of O. aurifex - group species. a – f, m — O. semifex (a – c, f, m — paratypes, d – e — Vietnam); g – l — O. alexeevi (g – j, l — paratypes, k — holotype); a – k — aedeagus lateral view; l – m — aedeagus dorsal view; a – e, g – h, k – m — aedeagus photographed in glycerin; f, g — aedeagus photographed after drying." httpUri="https://zenodo.org/record/294211/files/figure.png" pageId="4" pageNumber="67">Figs. 2</figureCitation>
a, f indicated with circle).
<taxonomicName id="4C499AE81D2A1F30FDCDFF00CB3AFF1A" box="[594,866,189,214]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="alexeevi">
<emphasis id="B93D3D791D2A1F30FDCDFF00CB3AFF1A" box="[594,866,189,214]" italics="true" pageId="4" pageNumber="67">Onthophagus alexeevi</emphasis>
</taxonomicName>
also has the membranous surface of paramere preapically excavated (
<figureCitation id="1372FDEE1D2A1F30FE3AFF5FC9B2FF30" box="[421,490,226,252]" captionStart="FIGURE 2" captionStartId="3.[151,255,1688,1712]" captionTargetBox="[166,1436,283,1627]" captionTargetId="figure@3.[151,1436,269,1667]" captionTargetPageId="3" captionText="FIGURE 2. Aedeagi of O. aurifex - group species. a – f, m — O. semifex (a – c, f, m — paratypes, d – e — Vietnam); g – l — O. alexeevi (g – j, l — paratypes, k — holotype); a – k — aedeagus lateral view; l – m — aedeagus dorsal view; a – e, g – h, k – m — aedeagus photographed in glycerin; f, g — aedeagus photographed after drying." httpUri="https://zenodo.org/record/294211/files/figure.png" pageId="4" pageNumber="67">Fig. 2</figureCitation>
l indicated with arrow, see also
<figureCitation id="1372FDEE1D2A1F30FCC5FF5FCBC7FF30" box="[858,927,226,252]" captionStart="FIGURE 2" captionStartId="3.[151,255,1688,1712]" captionTargetBox="[166,1436,283,1627]" captionTargetId="figure@3.[151,1436,269,1667]" captionTargetPageId="3" captionText="FIGURE 2. Aedeagi of O. aurifex - group species. a – f, m — O. semifex (a – c, f, m — paratypes, d – e — Vietnam); g – l — O. alexeevi (g – j, l — paratypes, k — holotype); a – k — aedeagus lateral view; l – m — aedeagus dorsal view; a – e, g – h, k – m — aedeagus photographed in glycerin; f, g — aedeagus photographed after drying." httpUri="https://zenodo.org/record/294211/files/figure.png" pageId="4" pageNumber="67">Fig. 2</figureCitation>
j) while this structure is not excavated in
<taxonomicName id="4C499AE81D2A1F30FAE0FF5FC8B7FEED" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="semifex">
<emphasis id="B93D3D791D2A1F30FAE0FF5FC8B7FEED" italics="true" pageId="4" pageNumber="67">
<collectingCountry id="F35EA1FB1D2A1F30FAE0FF5FCDCFFF37" box="[1407,1431,226,251]" name="Iceland" pageId="4" pageNumber="67">O</collectingCountry>
. semifex
</emphasis>
</taxonomicName>
(
<figureCitation id="1372FDEE1D2A1F30FF60FEB5C91CFEEE" box="[255,324,264,290]" captionStart="FIGURE 2" captionStartId="3.[151,255,1688,1712]" captionTargetBox="[166,1436,283,1627]" captionTargetId="figure@3.[151,1436,269,1667]" captionTargetPageId="3" captionText="FIGURE 2. Aedeagi of O. aurifex - group species. a – f, m — O. semifex (a – c, f, m — paratypes, d – e — Vietnam); g – l — O. alexeevi (g – j, l — paratypes, k — holotype); a – k — aedeagus lateral view; l – m — aedeagus dorsal view; a – e, g – h, k – m — aedeagus photographed in glycerin; f, g — aedeagus photographed after drying." httpUri="https://zenodo.org/record/294211/files/figure.png" pageId="4" pageNumber="67">
Fig.
<quantity id="4CB14C8E1D2A1F30FEA9FEB5C91CFEEE" box="[310,324,264,290]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="67" unit="m" value="2.0">2</quantity>
</figureCitation>
m indicated with arrow, see also
<figureCitation id="1372FDEE1D2A1F30FD5BFEB5CB52FEEE" box="[708,778,264,290]" captionStart="FIGURE 2" captionStartId="3.[151,255,1688,1712]" captionTargetBox="[166,1436,283,1627]" captionTargetId="figure@3.[151,1436,269,1667]" captionTargetPageId="3" captionText="FIGURE 2. Aedeagi of O. aurifex - group species. a – f, m — O. semifex (a – c, f, m — paratypes, d – e — Vietnam); g – l — O. alexeevi (g – j, l — paratypes, k — holotype); a – k — aedeagus lateral view; l – m — aedeagus dorsal view; a – e, g – h, k – m — aedeagus photographed in glycerin; f, g — aedeagus photographed after drying." httpUri="https://zenodo.org/record/294211/files/figure.png" pageId="4" pageNumber="67">Fig. 2</figureCitation>
f). Although the morphological differences between two species are subtle, there is robust evidence of their specific separation. First, the differences in aedeagal shape are stable between two species in the large series of examined material. Second, these two species seems to be sympatric (
<figureCitation id="1372FDEE1D2A1F30FEBAFEC5C929FE5E" box="[293,369,376,402]" captionStart="FIGURE 3" captionStartId="4.[151,255,1867,1891]" captionTargetBox="[155,1428,888,1822]" captionTargetId="figure@4.[151,1436,869,1846]" captionTargetPageId="4" captionText="FIGURE 3. Distribution map of the species in the O. aurifex complex." httpUri="https://zenodo.org/record/294212/files/figure.png" pageId="4" pageNumber="67">Fig. 3</figureCitation>
) and thus the stable differences in aedeagal shape are not a product of geographical variation, which indicated the specific independence of two species. Third, both species are possibly ecologically isolated as they have not been so far collected at the same place (
<figureCitation id="1372FDEE1D2A1F30FBB9FE7FCC35FE10" box="[1062,1133,450,476]" captionStart="FIGURE 3" captionStartId="4.[151,255,1867,1891]" captionTargetBox="[155,1428,888,1822]" captionTargetId="figure@4.[151,1436,869,1846]" captionTargetPageId="4" captionText="FIGURE 3. Distribution map of the species in the O. aurifex complex." httpUri="https://zenodo.org/record/294212/files/figure.png" pageId="4" pageNumber="67">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BF6E16B1D2A1F30FF5AFE55CD58FDEB" blockId="4.[151,1437,152,812]" pageId="4" pageNumber="67">
The new species differs from
<taxonomicName id="4C499AE81D2A1F30FDACFE55CABAFDCD" box="[563,738,488,513]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="semiaureus">
<emphasis id="B93D3D791D2A1F30FDACFE55CABAFDCD" box="[563,738,488,513]" italics="true" pageId="4" pageNumber="67">
<collectingCountry id="F35EA1FB1D2A1F30FDACFE55CA14FDCD" box="[563,588,488,513]" name="Iceland" pageId="4" pageNumber="67">O</collectingCountry>
. semiaureus
</emphasis>
</taxonomicName>
and
<taxonomicName id="4C499AE81D2A1F30FC81FE55CBC2FDCD" box="[798,922,488,513]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="aurifex">
<emphasis id="B93D3D791D2A1F30FC81FE55CBC2FDCD" box="[798,922,488,513]" italics="true" pageId="4" pageNumber="67">
<collectingCountry id="F35EA1FB1D2A1F30FC81FE55CB6FFDCD" box="[798,823,488,513]" name="Iceland" pageId="4" pageNumber="67">O</collectingCountry>
. aurifex
</emphasis>
</taxonomicName>
by the different shape and location of the sclerotized anterior part of the parameral apex (for details, see also
<bibRefCitation id="EFD89C9A1D2A1F30FC34FDB0CCABFDEB" author="Krikken" box="[939,1267,525,551]" pageId="4" pageNumber="68" refString="Krikken, J. &amp; Huijbregts, J. (2008) Distinguishing the Sundaland species in the Onthophagus (Parascatonomus) aurifex group (Coleoptera: Scarabaeidae: Scarabaeinae). Tijdschrift voor Entomologie, 151, 173 - 185. Masumoto, K. (2002) New record of Onthophagus semiaureus Lansberge from Thailand. Elytra, 30, 172." type="journal article" year="2008">Krikken &amp; Huijbregts 2008</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C353B2E01D2A1F30FF5AFD8FCD1CFD70" pageId="4" pageNumber="67" type="etymology">
<paragraph id="8BF6E16B1D2A1F30FF5AFD8FCD1CFD70" blockId="4.[151,1437,152,812]" pageId="4" pageNumber="67">
<emphasis id="B93D3D791D2A1F30FF5AFD8FC90EFD80" bold="true" box="[197,342,562,588]" pageId="4" pageNumber="67">Etymology.</emphasis>
The new species is named after Dr. Sergey Alexeev, Kaluga Regional Biological Center, Russia—thanks to whom, one of us (S.T.) became an entomologist. S.T. is much obliged to Dr. Sergey Alexeev for invaluable support and personal help at the beginning of his career, as well as for an interesting and enjoyable time in both expeditions and laboratory work at the Kaluga Regional Biological Center.
</paragraph>
</subSubSection>
<subSubSection id="C353B2E01D2A1F30FF5AFD7ACB5AFCE0" pageId="4" pageNumber="67" type="distribution">
<paragraph id="8BF6E16B1D2A1F30FF5AFD7ACB5AFCE0" blockId="4.[151,1437,152,812]" pageId="4" pageNumber="67">
<emphasis id="B93D3D791D2A1F30FF5AFD7AC9A5FD2D" bold="true" box="[197,509,711,737]" pageId="4" pageNumber="67">Distribution and ecology.</emphasis>
The new species is known only from West
<collectingCountry id="F35EA1FB1D2A1F30FC66FD75CC39FD2E" box="[1017,1121,712,738]" name="Thailand" pageId="4" pageNumber="67">Thailand</collectingCountry>
and South
<collectingCountry id="F35EA1FB1D2A1F30FB7BFD75CD04FD2E" box="[1252,1372,712,738]" name="Cambodia" pageId="4" pageNumber="67">Cambodia</collectingCountry>
(
<figureCitation id="1372FDEE1D2A1F30FAF2FD75C8FFFCCB" captionStart="FIGURE 3" captionStartId="4.[151,255,1867,1891]" captionTargetBox="[155,1428,888,1822]" captionTargetId="figure@4.[151,1436,869,1846]" captionTargetPageId="4" captionText="FIGURE 3. Distribution map of the species in the O. aurifex complex." httpUri="https://zenodo.org/record/294212/files/figure.png" pageId="4" pageNumber="67">Fig. 3</figureCitation>
), but is probably widespread in Indochina. All Cambodian specimens were collected in disturbed tropical forest by means of pitfall traps baited with dead fish.
</paragraph>
</subSubSection>
</treatment>
</document>
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