treatments-xml/data/03/D5/FE/03D5FE7E7049BF2DFD72FD83FCB4FD69.xml

<document id="1F98DD000F2FF290F3BD8C81DCFFEC56" ID-DOI="10.5852/ejt.2018.401" ID-GBIF-Dataset="49d45069-6150-4b6e-873c-fc8058025a39" ID-ISSN="2118-9773" ID-Zenodo-Dep="1167581" ID-ZooBank="9C9DB157-AAA5-40B7-BA0B-9A57779382C1" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1517911480053" checkinUser="plazi" docAuthor="Salnitska, Maria &amp; Solodovnikov, Alexey" docDate="2018" docId="03D5FE7E7049BF2DFD72FD83FCB4FD69" docLanguage="en" docName="ejt-401_salnitska_solodovnikov.pdf.imf" docOrigin="European Journal of Taxonomy 401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Quedius mutilatus Eppelsheim A. Smetana 1888" docType="treatment" docVersion="17" lastPageNumber="4" masterDocId="FFEC8606704BBF2EFFF0FFB0FFBBFFEB" masterDocTitle="Taxonomy of the poorly known Quedius mutilatus group of wingless montane species from Middle Asia (Coleoptera: Staphylinidae: Staphylinini)" masterLastPageNumber="17" masterPageNumber="1" originalUpdateDomain="Boston" originalUpdateTime="1547222236327" originalUpdateUser="yanell" pageNumber="3" updateTime="1734634589678" updateUser="guilherme">
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<mods:title id="176B6ECBC6797CA4141759633327BF81">Taxonomy of the poorly known Quedius mutilatus group of wingless montane species from Middle Asia (Coleoptera: Staphylinidae: Staphylinini)</mods:title>
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<treatment id="03D5FE7E7049BF2DFD72FD83FCB4FD69" ID-DOI="http://doi.org/10.5281/zenodo.5695862" ID-GBIF-Taxon="141125765" ID-Zenodo-Dep="5695862" LSID="urn:lsid:plazi:treatment:03D5FE7E7049BF2DFD72FD83FCB4FD69" httpUri="http://treatment.plazi.org/id/03D5FE7E7049BF2DFD72FD83FCB4FD69" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" scope_family="Staphylinidae" scope_order="Coleoptera" scope_subFamily="Staphylininae" scope_subGenus="Microsaurus" scope_subTribe="Quediina" scope_tribe="Staphylinini">
<subSubSection id="C3661CE37049BF2CFD72FD83FC0AFDA6" box="[642,945,563,590]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BC34F687049BF2CFD72FD83FC0AFDA6" blockId="2.[642,945,563,590]" box="[642,945,563,590]" pageId="2" pageNumber="3">
<heading id="D08BF8047049BF2CFD72FD83FC0AFDA6" box="[642,945,563,590]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
<emphasis id="B908937A7049BF2CFD72FD83FC0AFDA6" bold="true" box="[642,945,563,590]" pageId="2" pageNumber="3">
<taxonomicName id="4C7C34EB7049BF2CFD72FD83FCDBFDA5" ID-CoL="4R2SL" authority="Eppelsheim, 1888" authorityName="Eppelsheim A. Smetana" authorityYear="1888" box="[642,864,563,590]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mutilatus">
<emphasis id="B908937A7049BF2CFD72FD83FCDBFDA5" bold="true" box="[642,864,563,590]" italics="true" pageId="2" pageNumber="3">Quedius mutilatus</emphasis>
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3661CE37049BF2CFF4DFDC4FCF3FC10" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="8BC34F687049BF2CFF4DFDC4FE8DFD65" blockId="2.[189,310,628,654]" box="[189,310,628,654]" pageId="2" pageNumber="3">
<heading id="D08BF8047049BF2CFF4DFDC4FE8DFD65" bold="true" box="[189,310,628,654]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B908937A7049BF2CFF4DFDC4FE8DFD65" bold="true" box="[189,310,628,654]" pageId="2" pageNumber="3">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BC34F687049BF2CFF4DFD13FCF3FC10" blockId="2.[189,1399,675,1020]" pageId="2" pageNumber="3">
The 
<taxonomicName id="4C7C34EB7049BF2CFF06FD13FE6BFD56" authorityName="Eppelsheim A. Smetana" authorityYear="1888" box="[246,464,675,701]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mutilatus">
<emphasis id="B908937A7049BF2CFF06FD13FE6BFD56" box="[246,464,675,701]" italics="true" pageId="2" pageNumber="3">Quedius mutilatus</emphasis>
</taxonomicName>
group of species can be recognized as follows: relatively large 
<taxonomicName id="4C7C34EB7049BF2CFB12FD14FACCFD56" authorityName="Dejean" authorityYear="1833" box="[1250,1399,676,701]" class="Insecta" family="Staphylinidae" genus="Microsaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B908937A7049BF2CFB12FD14FACCFD56" box="[1250,1399,676,701]" italics="true" pageId="2" pageNumber="3">Microsaurus</emphasis>
</taxonomicName>
(body length 
<quantity id="4C84E28D7049BF2CFEA9FD76FE5BFD0A" box="[345,480,710,737]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" metricValueMax="11.6" metricValueMin="8.0" pageId="2" pageNumber="3" unit="mm" value="9.8" valueMax="11.6" valueMin="8.0">8–11.6 mm</quantity>
) with brown to dark brown, dorso-ventrally flattened body, with notably small eyes and pronounced signs of brachyptery such as short elytra and absent palisade fringe on abdominal tergite VII; head with posterior frontal puncture situated very close to neck; pronotum with distinct anterior and posterior angles, normally with three punctures in dorsal rows with anteriormost (smaller) puncture located latero-anteriorly at pronotal anterior margin, and with one to two sublateral punctures situated slightly anterior to large lateral puncture; scutellum impunctate; aedeagus robust, symmetrical, with apical portion of median lobe slightly curved towards paramere, with characteristic tooth near apex (in lateral view), with paramere widest shortly before apex (in parameral view) and with four distinct groups of sensory peg setae: two apical and two lateral.
</paragraph>
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<subSubSection id="C3661CE37049BF2CFF4DFB92FE2EFB59" pageId="2" pageNumber="3" type="taxon_list">
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<heading id="D08BF8047049BF2CFF4DFB92FED0FBD7" bold="true" box="[189,363,1058,1084]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B908937A7049BF2CFF4DFB92FED0FBD7" bold="true" box="[189,363,1058,1084]" pageId="2" pageNumber="3">List of species</emphasis>
</heading>
</paragraph>
<paragraph id="8BC34F687049BF2CFF4DFBE1FE2EFB59" blockId="2.[189,1399,1105,1202]" pageId="2" pageNumber="3">
<taxonomicName id="4C7C34EB7049BF2CFF4DFBE1FDC1FB80" authority="Eppelsheim, 1888" authorityName="Eppelsheim A. Smetana" authorityYear="1888" box="[189,634,1105,1131]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mutilatus">
<emphasis id="B908937A7049BF2CFF4DFBE1FE2DFB80" box="[189,406,1105,1131]" italics="true" pageId="2" pageNumber="3">Quedius mutilatus</emphasis>
<bibRefCitation id="EFED32997049BF2CFE50FBE1FDC1FB80" author="Eppelsheim E." box="[416,634,1105,1131]" journalOrPublisher="Deutsche Entomologische Zeitung" pageId="2" pageNumber="3" pagination="49 - 67" part="32" refId="ref6461" refString="Eppelsheim E. 1888. Neue Staphylinen Central-Asiens. Deutsche Entomologische Zeitung 32: 49 - 67." title="Neue Staphylinen Central-Asiens" type="journal article" year="1888">Eppelsheim, 1888</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="4C7C34EB7049BF2CFD79FBE1FC47FB87" authority="Smetana, 1995" authorityName="Smetana" authorityYear="1995" box="[649,1020,1105,1132]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="kalabi">
<emphasis id="B908937A7049BF2CFD79FBE1FC85FB80" box="[649,830,1105,1131]" italics="true" pageId="2" pageNumber="3">Quedius kalabi</emphasis>
<bibRefCitation id="EFED32997049BF2CFCB8FBE1FC47FB87" author="Smetana A." box="[840,1020,1105,1132]" journalOrPublisher="Elytra (Tokyo)" pageId="2" pageNumber="3" pagination="77 - 88" part="23" refId="ref6582" refString="Smetana A. 1995. Taxonomic and faunistic contributions to the knowledge of Palearctic Quediina (Coleoptera, Staphylinidae, Staphylinini). Elytra (Tokyo) 23: 77 - 88." title="Taxonomic and faunistic contributions to the knowledge of Palearctic Quediina (Coleoptera, Staphylinidae, Staphylinini)" type="journal article" year="1995">Smetana, 1995</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="4C7C34EB7049BF2CFBFBFBE1FACCFB80" authority="Smetana, 2014" authorityName="Smetana" authorityYear="2014" box="[1035,1399,1105,1131]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="equus">
<emphasis id="B908937A7049BF2CFBFBFBE1FB00FB80" box="[1035,1211,1105,1131]" italics="true" pageId="2" pageNumber="3">Quedius equus</emphasis>
<bibRefCitation id="EFED32997049BF2CFB35FBE1FACCFB80" author="Smetana A." box="[1221,1399,1105,1131]" journalOrPublisher="Folia Heyrovskyana, Series A" pageId="2" pageNumber="3" pagination="1 - 39" part="21" refId="ref6707" refString="Smetana A. 2014. Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 44. Genus Quedius Stephens, 1829. Subgenus Microsaurus Dejean, 1833. Section 22. Folia Heyrovskyana, Series A 21 (1 - 4): 1 - 39." title="Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 44. Genus Quedius Stephens, 1829. Subgenus Microsaurus Dejean, 1833. Section 22" type="journal article" year="2014">Smetana, 2014</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C7C34EB7049BF2CFF1EFBC5FE77FB64" authority="Salnitska &amp; Solodovnikov, 2018" authorityName="Salnitska &amp; Solodovnikov" authorityYear="2018" box="[238,460,1141,1167]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="kungeicus" status="sp. nov.">
<emphasis id="B908937A7049BF2CFF1EFBC5FE77FB64" box="[238,460,1141,1167]" italics="true" pageId="2" pageNumber="3">Quedius kungeicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B2E017049BF2CFE23FBC5FD8BFB64" box="[467,560,1141,1167]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
; all species can be reliably separated from each other by the structure of the aedeagus only.
</paragraph>
</subSubSection>
<subSubSection id="C3661CE37049BF2CFF4DFB69FE86F968" pageId="2" pageNumber="3" type="distribution">
<paragraph id="8BC34F687049BF2CFF4DFB69FEEDFB18" blockId="2.[189,342,1241,1267]" box="[189,342,1241,1267]" pageId="2" pageNumber="3">
<heading id="D08BF8047049BF2CFF4DFB69FEEDFB18" bold="true" box="[189,342,1241,1267]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B908937A7049BF2CFF4DFB69FEEDFB18" bold="true" box="[189,342,1241,1267]" pageId="2" pageNumber="3">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BC34F687049BF2CFF4DFAB8FE86F968" blockId="2.[189,1399,1288,1667]" pageId="2" pageNumber="3">
All species of the group seem to be allopatric (
<figureCitation id="134753ED7049BF2CFD29FAB8FCA5FAC9" box="[729,798,1288,1314]" captionStart="Fig.1" captionStartId="4.[189,232,1930,1956]" captionTargetBox="[221,1375,624,1881]" captionTargetId="figure@4.[208,1379,622,1889]" captionTargetPageId="4" captionText="Fig.1.DistributionandaedeagusvariabilityoftheQuediusmutilatusgroupintheTien-ShanMountains. Numbers are specified in Table 1. Empty symbols = typespcimens (for Q. kalabi precise locality of the holotype unknown);? = ambiguously labelled or undetermined material. Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/1167583/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
): 
<taxonomicName id="4C7C34EB7049BF2CFCC6FAB8FB53FAC9" authority="Eppelsheim, 1888" authorityName="Eppelsheim A. Smetana" authorityYear="1888" box="[822,1256,1288,1314]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mutilatus">
<emphasis id="B908937A7049BF2CFCC6FAB8FBB5FAC9" box="[822,1038,1288,1314]" italics="true" pageId="2" pageNumber="3">Quedius mutilatus</emphasis>
<bibRefCitation id="EFED32997049BF2CFBE4FAB8FB53FAC9" author="Eppelsheim E." box="[1044,1256,1288,1314]" journalOrPublisher="Deutsche Entomologische Zeitung" pageId="2" pageNumber="3" pagination="49 - 67" part="32" refId="ref6461" refString="Eppelsheim E. 1888. Neue Staphylinen Central-Asiens. Deutsche Entomologische Zeitung 32: 49 - 67." title="Neue Staphylinen Central-Asiens" type="journal article" year="1888">Eppelsheim, 1888</bibRefCitation>
</taxonomicName>
is restricted to the central part of Terskei Alatau, south of Issyk-Kul Lake; it is replaced eastwardly by 
<taxonomicName id="4C7C34EB7049BF2CFAF6FA9CFECAFA81" authority="Smetana, 1995" authorityName="Smetana" authorityYear="1995" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="kalabi">
<emphasis id="B908937A7049BF2CFAF6FA9CFACDFAAE" box="[1286,1398,1323,1350]" italics="true" pageId="2" pageNumber="3">Q. kalabi</emphasis>
<bibRefCitation id="EFED32997049BF2CFF4DFAFFFECAFA81" author="Smetana A." box="[189,369,1359,1386]" journalOrPublisher="Elytra (Tokyo)" pageId="2" pageNumber="3" pagination="77 - 88" part="23" refId="ref6582" refString="Smetana A. 1995. Taxonomic and faunistic contributions to the knowledge of Palearctic Quediina (Coleoptera, Staphylinidae, Staphylinini). Elytra (Tokyo) 23: 77 - 88." title="Taxonomic and faunistic contributions to the knowledge of Palearctic Quediina (Coleoptera, Staphylinidae, Staphylinini)" type="journal article" year="1995">Smetana, 1995</bibRefCitation>
</taxonomicName>
and then 
<taxonomicName id="4C7C34EB7049BF2CFE01FAFFFC9BFA82" authority="Smetana, 2014" authorityName="Smetana" authorityYear="2014" box="[497,800,1359,1385]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="equus">
<emphasis id="B908937A7049BF2CFE01FAFFFDE5FA82" box="[497,606,1359,1385]" italics="true" pageId="2" pageNumber="3">Q. equus</emphasis>
<bibRefCitation id="EFED32997049BF2CFD99FAFFFC9BFA82" author="Smetana A." box="[617,800,1359,1385]" journalOrPublisher="Folia Heyrovskyana, Series A" pageId="2" pageNumber="3" pagination="1 - 39" part="21" refId="ref6707" refString="Smetana A. 2014. Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 44. Genus Quedius Stephens, 1829. Subgenus Microsaurus Dejean, 1833. Section 22. Folia Heyrovskyana, Series A 21 (1 - 4): 1 - 39." title="Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 44. Genus Quedius Stephens, 1829. Subgenus Microsaurus Dejean, 1833. Section 22" type="journal article" year="2014">Smetana, 2014</bibRefCitation>
</taxonomicName>
, the latter probably being distributed throughout NE Terskei Alatau to 
<collectingRegion id="49B8818A7049BF2CFE4FFAC2FD9CFA67" box="[447,551,1394,1420]" country="China" name="Xinjiang Uygur" pageId="2" pageNumber="3">Xinjiang</collectingRegion>
Province in 
<collectingCountry id="F36B0FF87049BF2CFD4FFAC2FCB1FA67" box="[703,778,1394,1420]" name="China" pageId="2" pageNumber="3">China</collectingCountry>
. 
<taxonomicName id="4C7C34EB7049BF2CFCE5FAC2FC4CFA67" authority="Salnitska &amp; Solodovnikov, 2018" authorityName="Salnitska &amp; Solodovnikov" authorityYear="2018" box="[789,1015,1394,1420]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="kungeicus" status="sp. nov.">
<emphasis id="B908937A7049BF2CFCE5FAC2FC4CFA67" box="[789,1015,1394,1420]" italics="true" pageId="2" pageNumber="3">Quedius kungeicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B2E017049BF2CFC0EFAC3FBE1FA66" box="[1022,1114,1395,1421]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
is known from the 
<typeStatus id="54C7F1CA7049BF2CFAB3FAC3FACDFA66" box="[1347,1398,1395,1421]" pageId="2" pageNumber="3">type</typeStatus>
locality in eastern Kungei Alatau only. There is a single female from the high altitudes of Atbashi Mountain in Central Tien-Shan, which presumably belongs to the 
<taxonomicName id="4C7C34EB7049BF2CFC2FFA09FBCFFA38" authorityName="Eppelsheim A. Smetana" authorityYear="1888" box="[991,1140,1465,1491]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mutilatus">
<emphasis id="B908937A7049BF2CFC2FFA09FBCFFA38" box="[991,1140,1465,1491]" italics="true" pageId="2" pageNumber="3">Q. mutilatus</emphasis>
</taxonomicName>
group, but cannot be further identified without associated males. Three ambiguous specimens from Toksanbai in Dzungarian Alatau, damaged by dermestids, are presumably mislabelled and in fact originate from Terskei Alatau, because a single fully preserved male among them displays an aedeagus of the 
<taxonomicName id="4C7C34EB7049BF2CFB61F993FAB9F9D6" authorityName="Smetana" authorityYear="1995" box="[1169,1282,1571,1597]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="kalabi">
<emphasis id="B908937A7049BF2CFB61F993FAB9F9D6" box="[1169,1282,1571,1597]" italics="true" pageId="2" pageNumber="3">Q. kalabi</emphasis>
</taxonomicName>
<typeStatus id="54C7F1CA7049BF2CFAFDF993FAFFF9D6" box="[1293,1348,1571,1597]" pageId="2" pageNumber="3">type</typeStatus>
. All specimens of the 
<taxonomicName id="4C7C34EB7049BF2CFE66F9F6FD96F98B" authorityName="Eppelsheim A. Smetana" authorityYear="1888" box="[406,557,1606,1632]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mutilatus">
<emphasis id="B908937A7049BF2CFE66F9F6FD96F98B" box="[406,557,1606,1632]" italics="true" pageId="2" pageNumber="3">Q. mutilatus</emphasis>
</taxonomicName>
group with recorded bionomic data indicate a specialization to high elevations.
</paragraph>
</subSubSection>
<subSubSection id="C3661CE37049BF2DFF4DF91BFCB4FD69" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="discussion">
<paragraph id="8BC34F687049BF2CFF4DF91BFEEBF92E" blockId="2.[189,336,1707,1733]" box="[189,336,1707,1733]" pageId="2" pageNumber="3">
<heading id="D08BF8047049BF2CFF4DF91BFEEBF92E" bold="true" box="[189,336,1707,1733]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B908937A7049BF2CFF4DF91BFEEBF92E" bold="true" box="[189,336,1707,1733]" pageId="2" pageNumber="3">Systematics</emphasis>
</heading>
</paragraph>
<paragraph id="8BC34F687049BF2DFF4DF96AFBC9FE11" blockId="2.[189,1399,1753,2027]" lastBlockId="3.[189,1399,267,506]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
Some authors affiliated 
<taxonomicName id="4C7C34EB7049BF2CFE3EF96AFDE6F918" authorityName="Eppelsheim A. Smetana" authorityYear="1888" box="[462,605,1753,1780]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mutilatus">
<emphasis id="B908937A7049BF2CFE3EF96AFDE6F918" box="[462,605,1753,1780]" italics="true" pageId="2" pageNumber="3">Q. mutilatus</emphasis>
</taxonomicName>
with 
<taxonomicName id="4C7C34EB7049BF2CFD6CF96AFC64F91F" authority="Reitter, 1887" authorityName="Reitter" authorityYear="1887" box="[668,991,1753,1780]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="przewalskii">
<emphasis id="B908937A7049BF2CFD6CF96AFCFFF918" box="[668,836,1753,1780]" italics="true" pageId="2" pageNumber="3">Q. przewalskii</emphasis>
<bibRefCitation id="EFED32997049BF2CFCB9F969FC64F91F" author="Reitter E." box="[841,991,1753,1780]" journalOrPublisher="Horae Societatis Entomologicae Rossicae" pageId="2" pageNumber="3" pagination="201 - 234" part="21" refId="ref6545" refString="Reitter E. 1887. Insecta in itinere Cl. N. Przewalskii in Asia Centrali novisime lecta. VI. Clavicornia, Lamellicornia et Serricornia. Horae Societatis Entomologicae Rossicae 21: 201 - 234." title="Insecta in itinere Cl. N. Przewalskii in Asia Centrali novisime lecta. VI. Clavicornia, Lamellicornia et Serricornia" type="journal article" year="1887">Reitter, 1887</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFED32997049BF2CFC1BF969FB3AF91F" author="Reitter E." box="[1003,1153,1753,1780]" journalOrPublisher="Horae Societatis Entomologicae Rossicae" pageId="2" pageNumber="3" pagination="201 - 234" part="21" refId="ref6545" refString="Reitter E. 1887. Insecta in itinere Cl. N. Przewalskii in Asia Centrali novisime lecta. VI. Clavicornia, Lamellicornia et Serricornia. Horae Societatis Entomologicae Rossicae 21: 201 - 234." title="Insecta in itinere Cl. N. Przewalskii in Asia Centrali novisime lecta. VI. Clavicornia, Lamellicornia et Serricornia" type="journal article" year="1887">Reitter 1887</bibRefCitation>
; 
<bibRefCitation id="EFED32997049BF2CFB7BF969FAACF91F" author="Bohac J." box="[1163,1303,1753,1780]" journalOrPublisher="Entomologicheskoe Obozrenie" pageId="2" pageNumber="3" pagination="549 - 557" part="67" refId="ref6389" refString="Bohac J. 1988. New and little known Staphylininae (Coleoptera, Staphylinidae). Entomologicheskoe Obozrenie 67: 549 - 557. [In Russian.]" title="New and little known Staphylininae (Coleoptera, Staphylinidae)" type="journal article" year="1988">Boháč 1988</bibRefCitation>
) or with the 
<taxonomicName id="4C7C34EB7049BF2CFF18F94DFE28F8FC" authorityName="Reitter" authorityYear="1887" box="[232,403,1789,1815]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="przewalskii">
<emphasis id="B908937A7049BF2CFF18F94DFE28F8FC" box="[232,403,1789,1815]" italics="true" pageId="2" pageNumber="3">Q. przewalskii</emphasis>
</taxonomicName>
group (
<bibRefCitation id="EFED32997049BF2CFE1EF94DFD27F8FC" author="Smetana A." box="[494,668,1789,1815]" journalOrPublisher="Folia Heyrovskyana, Series A" pageId="2" pageNumber="3" pagination="1 - 39" part="21" refId="ref6707" refString="Smetana A. 2014. Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 44. Genus Quedius Stephens, 1829. Subgenus Microsaurus Dejean, 1833. Section 22. Folia Heyrovskyana, Series A 21 (1 - 4): 1 - 39." title="Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 44. Genus Quedius Stephens, 1829. Subgenus Microsaurus Dejean, 1833. Section 22" type="journal article" year="2014">Smetana 2014</bibRefCitation>
); the latter, however, lacks a clear definition (
<bibRefCitation id="EFED32997049BF2CFB41F94DFEE6F8D0" author="Solodovnikov A. &amp; Hansen A. K." journalOrPublisher="Zootaxa" pageId="2" pageNumber="3" pagination="475 - 490" part="4170" publicationUrl="https://doi.org/10.11646/zootaxa.4170.3.3" refId="ref6763" refString="Solodovnikov A. &amp; Hansen A. K. 2016. Review of subterranean Quedius, with description of the first hypogean species from the Russian Far East (Coleoptera: Staphylinidae: Staphylinini). Zootaxa 4170 (3): 475 - 490. https: // doi. org / 10.11646 / zootaxa. 4170.3.3" title="Review of subterranean Quedius, with description of the first hypogean species from the Russian Far East (Coleoptera: Staphylinidae: Staphylinini)" type="journal article" year="2016">Solodovnikov &amp; Hansen 2016</bibRefCitation>
). For example, 
<bibRefCitation id="EFED32997049BF2CFDE3F890FD75F8D1" author="Smetana A." box="[531,718,1824,1850]" journalOrPublisher="Folia Heyrovskyana, Series A" pageId="2" pageNumber="3" pagination="1 - 39" part="21" refId="ref6707" refString="Smetana A. 2014. Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 44. Genus Quedius Stephens, 1829. Subgenus Microsaurus Dejean, 1833. Section 22. Folia Heyrovskyana, Series A 21 (1 - 4): 1 - 39." title="Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 44. Genus Quedius Stephens, 1829. Subgenus Microsaurus Dejean, 1833. Section 22" type="journal article" year="2014">Smetana (2014)</bibRefCitation>
affiliated 
<taxonomicName id="4C7C34EB7049BF2CFCB6F891FC63F8D1" authorityName="Eppelsheim A. Smetana" authorityYear="1888" box="[838,984,1824,1851]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mutilatus">
<emphasis id="B908937A7049BF2CFCB6F891FC63F8D1" box="[838,984,1824,1851]" italics="true" pageId="2" pageNumber="3">Q. mutilatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7C34EB7049BF2CFBE2F891FBC1F8D1" authorityName="Smetana" authorityYear="2014" box="[1042,1146,1825,1851]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="equus">
<emphasis id="B908937A7049BF2CFBE2F891FBC1F8D1" box="[1042,1146,1825,1851]" italics="true" pageId="2" pageNumber="3">Q. equus</emphasis>
</taxonomicName>
to the 
<taxonomicName id="4C7C34EB7049BF2CFB3CF891FACCF8D1" authorityName="Reitter" authorityYear="1887" box="[1228,1399,1824,1851]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="przewalskii">
<emphasis id="B908937A7049BF2CFB3CF891FACCF8D1" box="[1228,1399,1824,1851]" italics="true" pageId="2" pageNumber="3">Q. przewalskii</emphasis>
</taxonomicName>
group, contrary to his (e.g., 
<bibRefCitation id="EFED32997049BF2CFDE6F8F4FD79F8B5" author="Smetana A." box="[534,706,1860,1886]" journalOrPublisher="Elytra (Tokyo)" pageId="2" pageNumber="3" pagination="193 - 216" part="29" refId="ref6654" refString="Smetana A. 2001. Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 20. Genus Quedius Stephens, 1829. Subgenus Microsaurus Dejean, 1833. Section 12. Elytra (Tokyo) 29 (1): 193 - 216." title="Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 20. Genus Quedius Stephens, 1829. Subgenus Microsaurus Dejean, 1833. Section 12" type="journal article" year="2001">Smetana 2001</bibRefCitation>
) earlier opinion of the composition of this group. Based on body chaetotaxy and the shape of the aedeagus, it could be related to 
<taxonomicName id="4C7C34EB7049BF2CFBCCF8D7FEBFF84E" authority="(Thomson, 1867)" authorityName="Thomson" authorityYear="1867" baseAuthorityName="Thomson" baseAuthorityYear="1867" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="B908937A7049BF2CFBCCF8D7FB57F86A" box="[1084,1260,1895,1921]" italics="true" pageId="2" pageNumber="3">Q. puncticollis</emphasis>
(Thomson, 1867)
</taxonomicName>
, as proposed by 
<bibRefCitation id="EFED32997049BF2CFE39F83AFDC6F84F" author="Coiffait H." box="[457,637,1930,1956]" journalOrPublisher="Nouvelle Revue d'Entomologie, Supplement" pageId="2" pageNumber="3" pagination="1 - 364" part="8" refId="ref6418" refString="Coiffait H. 1978. Coleopteres Staphylinidae de la region palearctique occidentale. III. Sous famille Staphylininae, tribu Quediini; sous famille Paederinae, tribu Pinophilini. Nouvelle Revue d'Entomologie, Supplement 8 (4): 1 - 364." title="Coleopteres Staphylinidae de la region palearctique occidentale. III. Sous famille Staphylininae, tribu Quediini; sous famille Paederinae, tribu Pinophilini" type="journal article" year="1978">Coiffait (1978)</bibRefCitation>
. The 
<taxonomicName id="4C7C34EB7049BF2CFD4EF83BFCD0F84F" baseAuthorityName="Thomson" baseAuthorityYear="1867" box="[702,875,1930,1957]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="B908937A7049BF2CFD4EF83BFCD0F84F" box="[702,875,1930,1957]" italics="true" pageId="2" pageNumber="3">Q. puncticollis</emphasis>
</taxonomicName>
group of 
<bibRefCitation id="EFED32997049BF2CFC2FF83AFB2BF84F" author="Coiffait H." box="[991,1168,1930,1956]" journalOrPublisher="Nouvelle Revue d'Entomologie, Supplement" pageId="2" pageNumber="3" pagination="1 - 364" part="8" refId="ref6418" refString="Coiffait H. 1978. Coleopteres Staphylinidae de la region palearctique occidentale. III. Sous famille Staphylininae, tribu Quediini; sous famille Paederinae, tribu Pinophilini. Nouvelle Revue d'Entomologie, Supplement 8 (4): 1 - 364." title="Coleopteres Staphylinidae de la region palearctique occidentale. III. Sous famille Staphylininae, tribu Quediini; sous famille Paederinae, tribu Pinophilini" type="journal article" year="1978">Coiffait (1978)</bibRefCitation>
is phylogenetically heterogeneous, based on the diverse morphology and scattered distributions of species. We establish the 
<taxonomicName id="4C7C34EB7049BF2CFF18F861FEC2F800" authorityName="Eppelsheim A. Smetana" authorityYear="1888" box="[232,377,2001,2027]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mutilatus">
<emphasis id="B908937A7049BF2CFF18F861FEC2F800" box="[232,377,2001,2027]" italics="true" pageId="2" pageNumber="3">Q. mutilatus</emphasis>
</taxonomicName>
group here for 
<taxonomicName id="4C7C34EB7049BF2CFDDEF861FD04F800" authorityName="Eppelsheim A. Smetana" authorityYear="1888" box="[558,703,2001,2027]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mutilatus">
<emphasis id="B908937A7049BF2CFDDEF861FD04F800" box="[558,703,2001,2027]" italics="true" pageId="2" pageNumber="3">Q. mutilatus</emphasis>
</taxonomicName>
and related species from Tien-Shan. Members of the group markedly differ from 
<taxonomicName id="4C7C34EB7048BF2DFE4EFEBCFDD7FECE" baseAuthorityName="Thomson" baseAuthorityYear="1867" box="[446,620,267,294]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="B908937A7048BF2DFE4EFEBCFDD7FECE" box="[446,620,267,294]" italics="true" pageId="3" pageNumber="4">Q. puncticollis</emphasis>
</taxonomicName>
and related species in a microphthalmous, brachypterous habitus, with weakly pigmented coloration and in the shape of the aedeagus, with a pointed apex of the median lobe and sensory peg setae arranged in apical and lateral groups. The 
<taxonomicName id="4C7C34EB7048BF2DFC0FFEE3FB29FE87" authorityName="Eppelsheim A. Smetana" authorityYear="1888" box="[1023,1170,338,365]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="mutilatus">
<emphasis id="B908937A7048BF2DFC0FFEE3FB29FE87" box="[1023,1170,338,365]" italics="true" pageId="3" pageNumber="4">Q. mutilatus</emphasis>
</taxonomicName>
group differs from the 
<taxonomicName id="4C7C34EB7048BF2DFF1AFEC6FE2EFE64" authorityName="Reitter" authorityYear="1887" box="[234,405,373,400]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="przewalskii">
<emphasis id="B908937A7048BF2DFF1AFEC6FE2EFE64" box="[234,405,373,400]" italics="true" pageId="3" pageNumber="4">Q. przewalskii</emphasis>
</taxonomicName>
group, in the more restricted sense of 
<bibRefCitation id="EFED32997048BF2DFC93FEC6FB99FE7B" author="Smetana A." box="[867,1058,374,400]" journalOrPublisher="Elytra (Tokyo)" pageId="3" pageNumber="4" pagination="193 - 216" part="29" refId="ref6654" refString="Smetana A. 2001. Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 20. Genus Quedius Stephens, 1829. Subgenus Microsaurus Dejean, 1833. Section 12. Elytra (Tokyo) 29 (1): 193 - 216." title="Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 20. Genus Quedius Stephens, 1829. Subgenus Microsaurus Dejean, 1833. Section 12" type="journal article" year="2001">Smetana (2001)</bibRefCitation>
, in having three (contrary to one or two) punctures in the dorsal rows of the pronotum and in the structure of the aedeagus, which is symmetrical, with a median lobe having a pointed apex and a more basally positioned ventral tooth, and with wide parameres having two groups of sensory peg setae (apical and lateral).
</paragraph>
<paragraph id="8BC34F687048BF2DFF4DFD91FCB4FD69" blockId="3.[189,1399,545,642]" pageId="3" pageNumber="4">
All species of this group are individually treated below. In addition to the description of a new species, a redescription is provided for 
<taxonomicName id="4C7C34EB7048BF2DFDD9FDF5FD00FDB5" authorityName="Eppelsheim A. Smetana" authorityYear="1888" box="[553,699,580,607]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="mutilatus">
<emphasis id="B908937A7048BF2DFDD9FDF5FD00FDB5" box="[553,699,580,607]" italics="true" pageId="3" pageNumber="4">Q. mutilatus</emphasis>
</taxonomicName>
, the most commonly cited species of the group with which other, later described species have been compared.
</paragraph>
</subSubSection>
</treatment>
</document>