<documentid="3B994984B079798885A87CA78BB10C08"ID-CLB-Dataset="20642"ID-DOI="10.11646/zootaxa.5099.4.1"ID-GBIF-Dataset="1390c18f-21be-489e-a5d7-4114b1758cb7"ID-ISSN="1175-5326"ID-Zenodo-Dep="6091813"ID-ZooBank="D7FBA8FE-FC8E-4AA6-98EA-C7BC84B30F6C"IM.materialsCitations_approvedBy="valdenar"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="felipe"IM.treatmentCitations_approvedBy="valdenar"checkinTime="1644912017914"checkinUser="plazi"docAuthor="Shima, Hiroshi, Zhang, Wenxia & Tachi, Takuji"docDate="2022"docId="58177649FF93D922E4B27839FC3125EE"docLanguage="en"docName="zootaxa.5099.4.1.pdf"docOrigin="Zootaxa 5099 (4)"docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article"docStyleId="647186512141C8FC8976D5BCC54AEB7D"docStyleName="Zootaxa.2013-.journal_article"docStyleVersion="9"docTitle="Phyllomya aristalis"docType="treatment"docVersion="8"lastPageNumber="415"masterDocId="A42E0E31FF9ED92CE4257F0AFF8C2365"masterDocTitle="A systematic study of Old World Phyllomya Robineau-Desvoidy (Diptera: Tachinidae)"masterLastPageNumber="449"masterPageNumber="401"pageNumber="414"updateTime="1731602969078"updateUser="valdenar"zenodo-license-document="CLOSED">
<taxonomicNameid="17BEBCDCFF93D921E4B27839FD23242B"ID-CoL="4H7VY"authority="(Mesnil & Shima, 1978)"baseAuthorityName="Mesnil & Shima"baseAuthorityYear="1978"box="[151,687,1843,1870]"class="Insecta"family="Tachinidae"genus="Phyllomya"kingdom="Animalia"order="Diptera"pageId="13"pageNumber="414"phylum="Arthropoda"rank="species"species="aristalis">
<bibRefCitationid="B42FBAAEFF93D921E5AA783EFD2B242B"author="Mesnil, L. P. & Shima, H."box="[399,679,1844,1870]"pageId="13"pageNumber="414"pagination="312 - 328"refId="ref35270"refString="Mesnil, L. P. & Shima, H. (1978) New and little known Tachinidae from Japan (Diptera). Kontyu, 46, 312 - 328."type="journal article"year="1978">Mesnil & Shima, 1978</bibRefCitation>
<figureCitationid="4885DBDAFF93D921E4BA7852FF6B2417"box="[159,231,1880,1906]"captionStart="FIGURES 3–7"captionStartId="6.[151,264,1672,1696]"captionTargetBox="[161,1424,181,1640]"captionTargetId="figure-19@6.[151,1436,181,1648]"captionTargetPageId="6"captionText="FIGURES 3–7. Male terminalia of Phyllomya spp. 3. Epandrial structure of P. aristalis (Mesnil & Shima), ventral view; bs = bacilliform sclerite. 4. 6th tergite, 6th sternite and syntergosternite 7+8 of P. albipila Shima & Chao, lateral view. 5. 6th tergite and syntergosternite 7+8 of P. annularis (Villeneuve), dorsal view. 6. 5th sternite, 6th tergite and syntergosternite 7+8 of P. rufiventris Shima & Chao, lateral view. 7. Same, P. elegans Villeneuve. Scale bars: 0.1 mm."figureDoi="http://doi.org/10.5281/zenodo.6091821"httpUri="https://zenodo.org/record/6091821/files/figure.png"pageId="13"pageNumber="414">Figs 3</figureCitation>
,
<figureCitationid="4885DBDAFF93D921E4D17852FE9E2416"box="[244,274,1880,1907]"captionStart="FIGURES 12–20"captionStartId="11.[151,264,1886,1910]"captionTargetBox="[159,1430,188,1851]"captionTargetId="figure-18@11.[151,1436,181,1862]"captionTargetPageId="11"captionText="FIGURES 12–20. Epandrium, surstylus and cerci of Phyllomya spp., lateral view. 12. P. albipila Shima & Chao (after Shima & Chao 1992). 13. P. angusta Shima & Chao (after Shima & Chao 1992). 14. P. annularis (Villeneuve). 15. P. aristalis (Mesnil & Shima) (after Mesnil & Shima 1978). 16. P. elegans Villeneuve. 17. P. formosana Shima (after Shima 1988). 18. P. gibsonomyioides Crosskey. 19. P. gymnops (Villeneuve). 20. P. humilis Shima (after Shima 1988). Scale bars: 0.1 mm."figureDoi="http://doi.org/10.5281/zenodo.6091830"httpUri="https://zenodo.org/record/6091830/files/figure.png"pageId="13"pageNumber="414">15</figureCitation>
,
<figureCitationid="4885DBDAFF93D921E53B7852FEB02417"box="[286,316,1880,1906]"captionStart="FIGURES 21–29"captionStartId="12.[151,264,1898,1922]"captionTargetBox="[209,1379,191,1865]"captionTargetId="figure-19@12.[202,1386,181,1874]"captionTargetPageId="12"captionText="FIGURES 21–29. Epandrium, surstylus and cerci of Phyllomya spp., dorsal view. 21. P. albipila Shima & Chao (after Shima & Chao 1992). 22. P. angusta Shima & Chao (after Shima & Chao 1992). 23. P. annularis (Villeneuve). 24. P. aristalis (Mesnil & Shima) (after Mesnil & Shima 1978). 25. P. elegans Villeneuve. 26. P. formosana Shima (after Shima 1988). 27. P. gibsonomyioides Crosskey. 28. P. gymnops (Villeneuve). 29. P. humilis Shima (after Shima 1988). Scale bars: 0.1 mm."figureDoi="http://doi.org/10.5281/zenodo.6091838"httpUri="https://zenodo.org/record/6091838/files/figure.png"pageId="13"pageNumber="414">24</figureCitation>
,
<figureCitationid="4885DBDAFF93D921E56D7852FEEA2416"box="[328,358,1880,1907]"captionStart="FIGURES 55–58"captionStartId="20.[151,264,1581,1605]"captionTargetBox="[158,1426,184,1543]"captionTargetId="figure-20@20.[151,1436,181,1557]"captionTargetPageId="20"captionText="FIGURES 55–58. Hypandrium, pre- and postgonite and phallus of Phyllomya spp., lateral view. 55. P. albipila Shima & Chao (after Shima & Chao 1992). 56. P. angusta Shima & Chao (after Shima & Chao 1992). 57. P. annularis (Villeneuve). 58. P. aristalis (Mesnil & Shima) (after Mesnil & Shima 1978). Scale bar: 0.1 mm."figureDoi="http://doi.org/10.5281/zenodo.6091862"httpUri="https://zenodo.org/record/6091862/files/figure.png"pageId="13"pageNumber="414">58</figureCitation>
,
<figureCitationid="4885DBDAFF93D921E5577852FE1C2416"box="[370,400,1880,1907]"captionStart="FIGURES 73–78"captionStartId="30.[151,264,1300,1324]"captionTargetBox="[158,1436,280,1276]"captionTargetId="figure-55@30.[151,1436,272,1276]"captionTargetPageId="30"captionText="FIGURES 73–78. Male 5th sternite of Phyllomya spp., ventral view. 73. P. albipila Shima & Chao (after Shima & Chao 1992). 74. P. annularis (Villeneuve). 75. P. aristalis (Mesnil & Shima) (after Mesnil & Shima 1978). 76. P. elegans Villeneuve. 77. P. formosana Shima (after Shima 1988). 78. P. gibsonomyioides Crosskey. Scale bars: 0.1 mm."figureDoi="http://doi.org/10.5281/zenodo.6091870"httpUri="https://zenodo.org/record/6091870/files/figure.png"pageId="13"pageNumber="414">75</figureCitation>
,
<figureCitationid="4885DBDAFF93D921E5B97852FE442416"box="[412,456,1880,1907]"captionStart="FIGURES 101–111"captionStartId="40.[151,264,1894,1918]"captionTargetBox="[180,1404,181,1871]"captionTargetId="figure-19@40.[172,1416,181,1871]"captionTargetPageId="40"captionText="FIGURES 101–111. Adult habitus/male head of Phyllomya spp., left lateral view. 101. P. albipila Shima & Chao. 102. P. annularis (Villeneuve). 103–104. P. arctogena Shima & Tachi, sp. nov. 105. P. aristalis (Mesnil & Shima). 106–107. P. chaoi Shima & Zhang, sp. nov. 108. P. elegans Villeneuve. 109. P. formosana Shima. 110. P. gibsonomyioides Crosskey (female). 111. P. gymnops (Villeneuve)."figureDoi="http://doi.org/10.5281/zenodo.6091898"httpUri="https://zenodo.org/record/6091898/files/figure.png"pageId="13"pageNumber="414">105</figureCitation>
,
<figureCitationid="4885DBDAFF93D921E5F17852FDBB2417"box="[468,567,1880,1906]"captionStart="FIGURES 127–138"captionStartId="44.[151,264,1896,1920]"captionTargetBox="[295,1292,185,1872]"captionTargetId="figure-19@44.[295,1292,181,1872]"captionTargetPageId="44"captionText="FIGURES 127–138. First instar larva and cephaloskeleton of Phyllomya spp., lateral view. 127–128. P. annularis (Villeneuve). 129–130. P. arctogena Shima & Tachi, sp. nov. 131–132. P. aristalis (Mesnil & Shima). 133–134. P. chaoi Shima & Zhang, sp. nov. 135–136. P. gibsonomyioides Crosskey. 137–138. P. gymnops (Villeneuve).Abbreviations: T2 and T3 = thoracic segments 2 and 3. A1 to A7 = abdominal segments 1 to 7."figureDoi="http://doi.org/10.5281/zenodo.6091912"httpUri="https://zenodo.org/record/6091912/files/figure.png"pageId="13"pageNumber="414">131–132</figureCitation>
<treatmentCitationid="511FE14EFF93D921E4B27897FDB824D0"author="Kolomiets, N. G."box="[151,564,1948,1973]"page="91"pageId="13"pageNumber="414"year="1973">
<bibRefCitationid="B42FBAAEFF93D921E5477896FDB824D0"author="Kolomiets, N. G."box="[354,564,1948,1973]"pageId="13"pageNumber="414"pagination="85 - 95"refId="ref34991"refString="Kolomiets, N. G. (1973) Parasitic Diptera of the genus Myiostoma R. - D. (Diptera, Tachinidae) from the USSR fauna. Novye i Maloizvestnye Vidy Fauny Sibiri, 6, 85 - 95. [in Russian with English summary]"type="journal article"year="1973">Kolomiets, 1973: 91</bibRefCitation>
<bibRefCitationid="B42FBAAEFF93D921E01A7896FB7C24D0"author="Villeneuve, J."box="[1087,1264,1948,1973]"pageId="13"pageNumber="414"pagination="1 - 16"refId="ref36176"refString="Villeneuve, J. (1937) Myodaires superieurs de Chine. Bulletin du Musee Royal d'Histoire Naturelle de Belgique, 13, 1 - 16."type="journal article"year="1937">Villeneuve, 1937</bibRefCitation>
<treatmentCitationid="511FE14EFF93D921E4B278B1FD2624B6"author="Mesnil, L. P. & Shima, H."box="[151,682,1979,2003]"page="313"pageId="13"pageNumber="414"year="1978">
<taxonomicNameid="17BEBCDCFF93D921E4B278B1FD2624B6"ID-CoL="8C7X8"authority="Mesnil & Shima, 1978: 313"authorityName="Mesnil & Shima"authorityPageNumber="313"authorityYear="1978"box="[151,682,1979,2003]"class="Insecta"family="Tachinidae"genus="Gibsonomyia"higherTaxonomySource="GBIF"kingdom="Animalia"order="Diptera"pageId="13"pageNumber="414"phylum="Arthropoda"rank="species"species="aristalis">
<bibRefCitationid="B42FBAAEFF93D921E5A678B1FD2624B6"author="Mesnil, L. P. & Shima, H."box="[387,682,1979,2003]"pageId="13"pageNumber="414"pagination="312 - 328"refId="ref35270"refString="Mesnil, L. P. & Shima, H. (1978) New and little known Tachinidae from Japan (Diptera). Kontyu, 46, 312 - 328."type="journal article"year="1978">Mesnil & Shima, 1978: 313</bibRefCitation>
<treatmentCitationid="511FE14EFF90D922E4B27F9DFDF223CA"author="Mesnil, L. P. & Shima, H."box="[151,638,151,175]"page="486"pageId="14"pageNumber="415"year="1979">
<taxonomicNameid="17BEBCDCFF90D922E4B27F9DFDF223CA"ID-CoL="4H7VY"authority=": Mesnil & Shima 1979: 486"authorityName="Mesnil & Shima"authorityPageNumber="486"authorityYear="1979"box="[151,638,151,175]"class="Insecta"family="Tachinidae"genus="Phyllomya"kingdom="Animalia"order="Diptera"pageId="14"pageNumber="415"phylum="Arthropoda"rank="species"species="aristalis">
<bibRefCitationid="B42FBAAEFF90D922E54F7F9DFDF223CA"author="Mesnil, L. P. & Shima, H."box="[362,638,151,175]"pageId="14"pageNumber="415"pagination="476 - 486"refId="ref35303"refString="Mesnil, L. P. & Shima, H. (1979) New tribe, genera and species of Japanese and Oriental Tachinidae (Diptera), with note on synonymy. Kontyu, 47, 476 - 486."type="journal article"year="1979">Mesnil & Shima 1979: 486</bibRefCitation>
<bibRefCitationid="B42FBAAEFF90D922E0897F9DFAE023CB"author="Herting, B."box="[1196,1388,150,175]"pageId="14"pageNumber="415"pagination="1 - 228"refId="ref34902"refString="Herting, B. (1984) Catalogue of Palearctic Tachinidae (Diptera). Stuttgarter Beitrage zur Naturkunde, Serie A (Biologie), 369, 1 - 228."type="journal article"year="1984">Herting, 1984: 155</bibRefCitation>
<bibRefCitationid="B42FBAAEFF90D922E55F7FBFFD9923A8"author="Shima, H."box="[378,533,181,205]"pageId="14"pageNumber="415"pagination="1 - 37"refId="ref35628"refString="Shima, H. (1988) Some remarkable new species of Tachinidae (Diptera) from Japan and the Indo-Australian Region. Bulletin of the Kitakyushu Museum of Natural History, Kitakyushu, 8, 1 - 37."type="journal article"year="1988">Shima 1988:16</bibRefCitation>
</treatmentCitation>
(Japanese tachinid species);
<treatmentCitationid="511FE14EFF90D922E7647FBFFB4423A8"author="Herting, B. & Dely-Draskovits, A."box="[833,1224,181,205]"page="387"pageId="14"pageNumber="415"year="1993">
<bibRefCitationid="B42FBAAEFF90D922E7647FBFFB4423A8"author="Herting, B. & Dely-Draskovits, A."box="[833,1224,181,205]"pageId="14"pageNumber="415"refId="ref34934"refString="Herting, B. & Dely-Draskovits, A. (1993) Family Tachinidae. In: Soos, A. & Papp, L. (Eds.), Catalogue of Palaearctic Diptera. Vol. 13. Anthomyiidae - Tachinidae. Hungarian Natural History Museum, Budapest, pp. 118 - 540."type="book"year="1993">Herting & Dely-Draskovits 1993: 387</bibRefCitation>
<bibRefCitationid="B42FBAAEFF90D922E4E27FDEFEFE2389"author="Shima, H."box="[199,370,212,236]"pageId="14"pageNumber="415"pagination="483 - 495"refId="ref35698"refString="Shima, H. (2000) Tachinidae (Insecta, Diptera) of the Imperial Palace, Tokyo. Memoirs of the National Science Museum, Tokyo, 36, 483 - 495. [in Japanese with English summary]"type="journal article"year="2000">Shima 2000: 491</bibRefCitation>
</treatmentCitation>
(tachinids of Imperial Palace);
<treatmentCitationid="511FE14EFF90D922E68B7FDEFCEF2389"author="Richter, V. A."box="[686,867,212,236]"page="368"pageId="14"pageNumber="415"year="2004">
<bibRefCitationid="B42FBAAEFF90D922E68B7FDEFCEF2389"author="Richter, V. A."box="[686,867,212,236]"pageId="14"pageNumber="415"refId="ref35501"refString="Richter, V. A. (2004) Tachinidae. In: Lera, P. A. (Ed.), Key to insects of Russian Far East. Vol. 6. Dal'nauka, Vladivostok, pp. 148 - 398. [in Russian]"type="book"year="2004">Richter 2004: 368</bibRefCitation>
<bibRefCitationid="B42FBAAEFF90D922E08C7FDEFAD92389"author="Shima, H."box="[1193,1365,212,236]"pageId="14"pageNumber="415"pagination="279 - 289"refId="ref35788"refString="Shima, H. (2006) Tachinidae (Insecta, Diptera) of the Imperial Palace, Tokyo (Addenda and Corregenda). Memoirs of the National Science Museum, Tokyo, 43, 279 - 289. [in Japanese with English summary]"type="journal article"year="2006">Shima 2006: 287</bibRefCitation>
<bibRefCitationid="B42FBAAEFF90D922E5E27FF9FDF8226E"author="Shima, H."box="[455,628,243,267]"pageId="14"pageNumber="415"refId="ref35835"refString="Shima, H. (2014) Family Tachinidae. In: Nakamura, T., Saigusa, T. & Suwa, M. (Eds.), Catalog of the Insects of Japan. Vol. 8. The Entomological Society of Japan, Tsukuba, pp. 832 - 882."type="book"year="2014">Shima 2014: 836</bibRefCitation>
</treatmentCitation>
(Japanese catalog);
<treatmentCitationid="511FE14EFF90D922E7667FF9FBA4226F"author="Shima, H. & Hara, K."box="[835,1064,242,267]"page="5"pageId="14"pageNumber="415"year="2016">
<bibRefCitationid="B42FBAAEFF90D922E7667FF9FBA4226F"author="Shima, H. & Hara, K."box="[835,1064,242,267]"pageId="14"pageNumber="415"pagination="1 - 31"refId="ref35930"refString="Shima, H. & Hara, K. (2016) Tachinidae (Insecta, Diptera) of Saitama Prefecture. Bulletin of the Kyushu University Museum, 14, 1 - 31."type="journal article"year="2016">Shima & Hara 2016: 5</bibRefCitation>
Head almost holoptic in male, dichoptic in female; antennal pedicel with a long seta almost as long as total length of antenna; arista rather long plumose, total width including plumosity subequal in width to postpedicel; parafacial with dense, black hairs; palpus dark brown; 3 postsutural dorsocentral setae; 2, rarely 3, katepisternal setae; mid tibia with 2 anterodorsal setae; hind tibia with 3 preapical dorsal setae.
. Terminalia: Sixth tergite strongly concave mid-dorsally on anterior margin, with 2 pairs of strong setae; surstylus broad, with very weakly narrowed to truncated apex and dense, long hairs; cerci in dorsal view broad along basal 1/3 then narrowed to apex, completely fused along basal 1/4 with suture obliterated, narrowly separated along apical 1/3, in lateral view narrowed at basal 1/3 and weakly curved ventrally; pregonite very weakly and roundly produced posteriorly; basiphallus shorter than distiphallus, epiphallus narrow, shorter than basiphallus, dorsal sclerite of distiphallus flat, apical membranous portion very short, at most 1/4 as long as sclerite.
Terminalia: Sixth tergite 1.5 times as long as 7th tergite, concave mid-dorsally on anterior margin, with strong setae on posterior 1/4; 6th sternite with hairs on posterior 1/2 and strong marginal setae; 7th tergite broadly divided into 2 hemitergites, with a row of seta on posterior margin; 7th sternite slightly shorter than 6th sternite, with several short setae on posterior margin; 8th hemitergite fused with hypoproct on posteromedian portion and ventrally with 8th sternite.
<emphasisid="E2CA1B4DFF90D922E4E27C4DFDCF2004"bold="true"box="[199,579,839,865]"pageId="14"pageNumber="415">Description of first instar larva.</emphasis>
; cephaloskeleton rather broad in apical 1/3 and narrowed to a pointed apex, dorsal margin sharply convex in apical 1/2; 2nd thoracic segment with 4–5 rows of fine anterodorsal spinules; 7th abdominal segment with 6–7 posterodorsal rows of spinules that are almost continuous with posteroventral rows of spinules.
<emphasisid="E2CA1B4DFF90D922E4E27CDEFE62208B"bold="true"box="[199,494,980,1006]"pageId="14"pageNumber="415">Type material examined.</emphasis>
<materialsCitationid="60D6CD02FF90D922E5D07CDEFDF92774"ID-GBIF-Occurrence="3466636374"collectingDate="1969-07-14"collectionCode="BLKU"collectorName="Japan-Honshu & Shinhodaka & Hida & A. Nagatomi"pageId="14"pageNumber="415"specimenCount="1"specimenCount-male="1"typeStatus="holotype">
<taxonomicNameid="17BEBCDCFF90D922E6B87CDEFBD5208B"authority="Mesnil & Shima"authorityName="Mesnil & Shima"authorityYear="1978"box="[669,1113,980,1006]"class="Insecta"family="Tachinidae"genus="Gibsonomyia"higherTaxonomySource="GBIF"kingdom="Animalia"order="Diptera"pageId="14"pageNumber="415"phylum="Arthropoda"rank="species"species="aristalis">
<taxonomicNameid="17BEBCDCFF90D922E72A7CF2FC012774"authorityName="Mesnil & Shima"authorityYear="1978"box="[783,909,1015,1042]"class="Insecta"family="Tachinidae"genus="Gibsonomyia"higherTaxonomySource="GBIF"kingdom="Animalia"order="Diptera"pageId="14"pageNumber="415"phylum="Arthropoda"rank="species"species="aristalis">
<materialsCitationid="60D6CD02FF90D922E0557B11FF76273D"ID-GBIF-Occurrence="3466636496"collectionCode="BLKU"collectorName="Japan-Honshu & Shinhodaka & Hida & A. Nagatomi"pageId="14"pageNumber="415"specimenCount="1"specimenCount-female="1"typeStatus="paratype">
<materialsCitationid="60D6CD02FF90D922E4B27BE4FCD3266D"ID-GBIF-Occurrence="3466636431"box="[151,863,1262,1288]"collectingDate="1977-07-10"collectionCode="KUM"collectorName="K. Yamagishi & Honshu"country="Japan"location="Bonto"municipality="Akkesi"pageId="14"pageNumber="415"specimenCount="1"specimenCount-female="1">
<collectionCodeid="B6AF5F9AFF90D922E6007A3EFDF5262B"box="[549,633,1332,1358]"country="Japan"httpUri="http://grbio.org/cool/f678-ci92"name="National Science Museum (Natural History)"pageId="14"pageNumber="415">NSMT</collectionCode>
<materialsCitationid="60D6CD02FF90D922E7067A3FFC162617"ID-GBIF-Occurrence="3466636482"collectingDateMax="1997-05-22"collectingDateMin="1997-05-01"collectionCode="KUM, NSMT"collectorName="H. Shima & S. Shinonaga & Nambu"country="Japan"location="Imperial Palace"pageId="14"pageNumber="415"specimenCount="1"specimenCount-male="1"stateProvince="Tokyo">
<collectionCodeid="B6AF5F9AFF90D922E72F7A5DFCD12614"box="[778,861,1367,1393]"country="Japan"httpUri="http://grbio.org/cool/f678-ci92"name="National Science Museum (Natural History)"pageId="14"pageNumber="415">NSMT</collectionCode>
<materialsCitationid="60D6CD02FF90D922E7807A52FE1826F0"ID-GBIF-Occurrence="3466636376"collectingDate="1952-06-20"collectionCode="KUM"collectorName="K. Iwata & Kyushu"country="Japan"location="Sasayama"pageId="14"pageNumber="415"specimenCount="1"specimenCount-male="1"stateProvince="Hyogo">
<bibRefCitationid="B42FBAAEFF90D922E6417944FC9E250D"author="Crosskey, R. W."box="[612,786,1614,1640]"pageId="14"pageNumber="415"pagination="267 - 308"refId="ref34609"refString="Crosskey, R. W. (1974) British Tachinidae of Walker and Stephens (Diptera). Bulletin of the British Museum (Natural History), Entomology, 30, 267 - 308."type="journal article"year="1974">Crosskey, 1974</bibRefCitation>
</taxonomicName>
, but can be distinguished from it by the abdominal dorsum with narrow pruinose bands and hind tibia with 3 preapical dorsal setae.