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<document id="002BE8C5F35598ECA9723605979C08BD" ID="10.11646/zootaxa.4150.3.3" ID-DOI="10.11646/zootaxa.4150.3.3" ID-GBIF-Dataset="e98edafd-7489-488d-ab8a-276a4a4926a4" ID-ISSN="1175-5326" ID-Zenodo-Dep="265618" ID-ZooBank="79657D7A-324F-4539-A80F-E11BB4189753" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1470984303137" checkinUser="plazi" docAuthor="Patitucci, Luciano Damián, Mulieri, Pablo Ricardo &amp; Mariluis, Juan Carlos" docDate="2016" docId="03F65F6EF056FFC9FF6CF968FECE2B14" docLanguage="en" docName="zootaxa.4150.3.3.pdf" docOrigin="Zootaxa 4150 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helina chilensis Malloch 1934" docType="treatment" docVersion="7" lastPageNumber="294" masterDocId="FFCF2716F05CFFC4FFFBFF88FFAE2F04" masterDocTitle="Taxonomic review of the species of Helina R. - D. (Diptera: Muscidae) from Andean-Patagonian forests" masterLastPageNumber="313" masterPageNumber="281" pageNumber="291" updateTime="1698381830647" updateUser="plazi">
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<mods:title id="DE7A860FCDAFF863CABEAB90944D7E6C">Taxonomic review of the species of Helina R. - D. (Diptera: Muscidae) from Andean-Patagonian forests</mods:title>
</mods:titleInfo>
<mods:name id="DCC87A3EDD097C7FF7E442BD52D4401C" type="personal">
<mods:role id="2A849A4E4691F915A8558FA0600DDCB0">
<mods:roleTerm id="025A75970B66C4529940D3336E27CAA2">Author</mods:roleTerm>
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<mods:namePart id="210033036BF0332BA8DAE506F83B84C3">Patitucci, Luciano Damián</mods:namePart>
</mods:name>
<mods:name id="7506206680EFBC9D215A26BE37721122" type="personal">
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<mods:namePart id="50AF557C688118CDE05EE2353132F48E">Mulieri, Pablo Ricardo</mods:namePart>
</mods:name>
<mods:name id="FF666EE0589C5103466CF5F932E0B487" type="personal">
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<mods:roleTerm id="47E3C36413F82D474C22574A0F21BF5F">Author</mods:roleTerm>
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<mods:namePart id="0A0504E0DB9E8029C6644A68AD0CC38C">Mariluis, Juan Carlos</mods:namePart>
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<mods:title id="7CF54B1C96A593DED0D79D27CEDD7A3F">Zootaxa</mods:title>
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<mods:date id="1D65420DEEE82C5164FD57218A60DFFB">2016</mods:date>
<mods:detail id="D5C7DA72333964A9F5EF60C9DD24C5D3" type="volume">
<mods:number id="72AE6308E4B236FC6DB5344A8989FAD4">4150</mods:number>
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<mods:detail id="86D7A9600ABF127E1E01266C68A483C1" type="issue">
<mods:number id="AD209FD340669EB05C2638482D055C97">3</mods:number>
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<mods:classification id="DBDE99E640A09CE6C934422A5AF513C6">journal article</mods:classification>
<mods:identifier id="D6CAB035C5B0E5077C164E120F874C83" type="DOI">10.11646/zootaxa.4150.3.3</mods:identifier>
<mods:identifier id="BE359045111250D5DEA870167F82F4BD" type="GBIF-Dataset">e98edafd-7489-488d-ab8a-276a4a4926a4</mods:identifier>
<mods:identifier id="D18258307556D7F81C2A070A8896C17E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="060FB3296B42495DB0AA1A5C4C39A663" type="Zenodo-Dep">265618</mods:identifier>
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<treatment id="03F65F6EF056FFC9FF6CF968FECE2B14" ID-DOI="http://doi.org/10.5281/zenodo.6088853" ID-GBIF-Taxon="124329283" ID-Zenodo-Dep="6088853" LSID="urn:lsid:plazi:treatment:03F65F6EF056FFC9FF6CF968FECE2B14" httpUri="http://treatment.plazi.org/id/03F65F6EF056FFC9FF6CF968FECE2B14" lastPageId="13" lastPageNumber="294" pageId="10" pageNumber="291">
<subSubSection id="C345BDF3F056FFCEFF6CF968FE882818" pageId="10" pageNumber="291" type="nomenclature">
<paragraph id="8BE0EE78F056FFCEFF6CF968FDBC29FE" blockId="10.[151,530,1760,1820]" box="[151,530,1760,1786]" pageId="10" pageNumber="291">
<heading id="D0A85914F056FFCEFF6CF968FDBC29FE" bold="true" box="[151,530,1760,1786]" fontSize="11" level="1" pageId="10" pageNumber="291" reason="1">
<taxonomicName id="4C5F95FBF056FFCEFF6CF968FDBC29FE" authority="Malloch, 1934" authorityName="Malloch" authorityYear="1934" box="[151,530,1760,1786]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="291" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="B92B326AF056FFCEFF6CF968FDBC29FE" bold="true" box="[151,530,1760,1786]" pageId="10" pageNumber="291">
<emphasis id="B92B326AF056FFCEFF6CF968FEF529FE" bold="true" box="[151,347,1760,1786]" italics="true" pageId="10" pageNumber="291">Helina chilensis</emphasis>
<bibRefCitation id="EFCE9389F056FFCEFE99F968FDBC29FE" author="Malloch" box="[354,530,1760,1786]" pageId="10" pageNumber="291" refString="Malloch, J. R. (1934) Muscidae. In: Diptera of Patagonia and South Chile. Part 7, Fascicle 2. British Museum (Natural History), London, pp. 171 - 346." type="book chapter" year="1934">Malloch, 1934</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE0EE78F056FFCEFF6CF88BFE882818" blockId="10.[151,530,1760,1820]" box="[151,294,1795,1820]" pageId="10" pageNumber="291">
(
<figureCitation id="1364F2FDF056FFCEFF64F88BFF432818" box="[159,237,1795,1820]" captionStart="FIGURE 5" captionStartId="12.[151,250,1410,1432]" captionTargetBox="[189,1399,193,1386]" captionTargetId="figure@12.[187,1399,193,1389]" captionTargetPageId="12" captionText="FIGURE 5. Helina chilensis Malloch. A – H. Male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate and surstyli, posterior view. G. Cercal plate and surstyli, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). I – J. Female, apical portion of ovipositor: I. Dorsal view. J. Ventral view (scalebar: 0.5 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; epiph, epiphallus; epiprct, epiproct; gon, gonopodo; hyprct, hypoproct; par, paramere; st 8, sternite 8; tg 8, tergite 8.)" httpUri="https://zenodo.org/record/265623/files/figure.png" pageId="10" pageNumber="291">Figs. 5</figureCitation>
,
<figureCitation id="1364F2FDF056FFCEFF01F88CFEA62818" box="[250,264,1796,1820]" captionStart="FIGURE 9" captionStartId="18.[151,250,1937,1959]" captionTargetBox="[168,1423,829,1910]" captionTargetId="figure@18.[163,1423,829,1916]" captionTargetPageId="18" captionText="FIGURE 9. Geographical distribution. A. Helina araucana sp. nov. B. Helina australis Carvalho &amp; Pont. C. Helina bigoti Malloch. D. Helina chilensis Malloch. E. Helina connexa Malloch. F. Helina discolor (Stein). G. Helina dorada sp. nov. (Green area: Andean-Patagonian forests; orange circle: new records; blue triangle: previous distribution.)" httpUri="https://zenodo.org/record/265627/files/figure.png" pageId="10" pageNumber="291">9</figureCitation>
D)
</paragraph>
</subSubSection>
<subSubSection id="C345BDF3F056FFCFFF6CF8C3FACA2FEF" lastPageId="11" lastPageNumber="292" pageId="10" pageNumber="291" type="reference_group">
<paragraph id="8BE0EE78F056FFCEFF6CF8C3FDD02884" blockId="10.[151,715,1867,1920]" pageId="10" pageNumber="291">
<taxonomicName id="4C5F95FBF056FFCEFF6CF8C3FDB42865" authority="Malloch, 1934" authorityName="Malloch" authorityYear="1934" box="[151,538,1867,1889]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="291" phylum="Arthropoda" rank="species" species="fulvocalyptrata">
<emphasis id="B92B326AF056FFCEFF6CF8C3FED32865" box="[151,381,1867,1889]" italics="true" pageId="10" pageNumber="291">Helina fulvocalyptrata</emphasis>
<bibRefCitation id="EFCE9389F056FFCEFE78F8C3FDB42865" author="Malloch" box="[387,538,1867,1889]" pageId="10" pageNumber="291" refString="Malloch, J. R. (1934) Muscidae. In: Diptera of Patagonia and South Chile. Part 7, Fascicle 2. British Museum (Natural History), London, pp. 171 - 346." type="book chapter" year="1934">Malloch, 1934</bibRefCitation>
</taxonomicName>
.
<emphasis id="B92B326AF056FFCEFDD8F8C3FD6B2865" bold="true" box="[547,709,1867,1889]" pageId="10" pageNumber="291">New synonymy</emphasis>
.
<taxonomicName id="4C5F95FBF056FFCEFF6CF8E2FE632884" authority="Malloch, 1934" authorityName="Malloch" authorityYear="1934" box="[151,461,1898,1920]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="291" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B92B326AF056FFCEFF6CF8E2FE812884" box="[151,303,1898,1920]" italics="true" pageId="10" pageNumber="291">Helina simplex</emphasis>
<bibRefCitation id="EFCE9389F056FFCEFECDF8E2FE632884" author="Malloch" box="[310,461,1898,1920]" pageId="10" pageNumber="291" refString="Malloch, J. R. (1934) Muscidae. In: Diptera of Patagonia and South Chile. Part 7, Fascicle 2. British Museum (Natural History), London, pp. 171 - 346." type="book chapter" year="1934">Malloch, 1934</bibRefCitation>
</taxonomicName>
.
<emphasis id="B92B326AF056FFCEFE2DF8E2FDD62884" bold="true" box="[470,632,1898,1920]" pageId="10" pageNumber="291">New synonymy</emphasis>
.
</paragraph>
<paragraph id="8BE0EE78F056FFCEFF6CF825FB4628E5" blockId="10.[151,1309,1964,2017]" pageId="10" pageNumber="291">
For a complete list of references, see the catalog by
<bibRefCitation id="EFCE9389F056FFCEFD5EF824FC2928C6" box="[677,903,1964,1986]" pageId="10" pageNumber="291" refString="Carvalho, C. J. B. de, Couri, M. S., Pont, A. C., Pamplona, D. &amp; Lopes, S. M. (2005) A Catalogue of the Muscidae (Diptera) of the Neotropical Region. Zootaxa, 860, 1 - 282. http: // dx. doi. org / 10.11646 / zootaxa. 860.1.1" type="journal article">
Carvalho
<emphasis id="B92B326AF056FFCEFCF3F825FC9228C6" box="[776,828,1964,1986]" italics="true" pageId="10" pageNumber="291">et al.</emphasis>
(2005)
</bibRefCitation>
. Information subsequent to this catalog:
<taxonomicName id="4C5F95FBF056FFCEFF6CF843FE9528E5" box="[151,315,1995,2017]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="291" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="B92B326AF056FFCEFF6CF843FE9528E5" box="[151,315,1995,2017]" italics="true" pageId="10" pageNumber="291">Helina chilensis</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFCE9389F056FFCEFEB3F843FDCF28E5" author="Lowerberg-Neto" box="[328,609,1995,2017]" pageId="10" pageNumber="291" refString="Lowerberg-Neto, P., Carvalho, C. J. B. de &amp; Hawkins, B. A. (2011) Tropical niche conservatism as a historical narrative hypothesis for the Neotropics: a case study using the fly family Muscidae. Journal of Biogeography, 38, 1936 - 1947. http: // dx. doi. org / 10.1111 / j. 1365 - 2699.2011.02540. x" type="journal article" year="2011">
Löwerberg-Neto
<emphasis id="B92B326AF056FFCEFE0DF844FD8D28E5" box="[502,547,1995,2017]" italics="true" pageId="10" pageNumber="291">et al</emphasis>
. 2011
</bibRefCitation>
(biogeography);
<bibRefCitation id="EFCE9389F056FFCEFCEAF843FBC128E5" author="Lowerberg-Neto" box="[785,1135,1995,2017]" pageId="10" pageNumber="291" refString="Lowerberg-Neto, P. &amp; Carvalho, C. J. B. de (2013) Muscidae (Insecta: Diptera) of Latin America and the Caribbean: geographic distribution and check-list by country. Zootaxa, 3650 (1), 01 - 147. http: // dx. doi. org / 10.11646 / zootaxa. 3650.1.1" type="journal article" year="2013">Löwerberg-Neto &amp; Carvalho 2013</bibRefCitation>
(checklist).
</paragraph>
<paragraph id="8BE0EE78F057FFCFFF6CFF10FCB82FC8" blockId="11.[151,1436,152,235]" pageId="11" pageNumber="292">
<taxonomicName id="4C5F95FBF057FFCFFF6CFF10FED02FAA" box="[151,382,152,174]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="species" species="fulvocalyptrata">
<emphasis id="B92B326AF057FFCFFF6CFF10FED02FAA" box="[151,382,152,174]" italics="true" pageId="11" pageNumber="292">Helina fulvocalyptrata</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFCE9389F057FFCFFE77FF11FDCC2FAA" author="Pont" box="[396,610,152,174]" pageId="11" pageNumber="292" refString="Pont, A. C. &amp; Werner, D. (2006) The Types of Fanniidae and Muscidae (Diptera) in the Museum fur Naturkunde, Humboldt- Universitat zu Berlin, Germany. Mitteilungen aus dem Zoologischen Museum in Berlin, 82, 3 - 139. http: // dx. doi. org / 10.1002 / mmnz. 200600001" type="journal article" year="2006">Pont &amp; Werner 2006</bibRefCitation>
(
<typeStatus id="54E450DAF057FFCFFD8BFF10FD332FAA" box="[624,669,152,174]" pageId="11" pageNumber="292">type</typeStatus>
specimens);
<bibRefCitation id="EFCE9389F057FFCFFCDEFF10FBEC2FAA" author="Lowerberg-Neto" box="[805,1090,152,174]" pageId="11" pageNumber="292" refString="Lowerberg-Neto, P., Carvalho, C. J. B. de &amp; Hawkins, B. A. (2011) Tropical niche conservatism as a historical narrative hypothesis for the Neotropics: a case study using the fly family Muscidae. Journal of Biogeography, 38, 1936 - 1947. http: // dx. doi. org / 10.1111 / j. 1365 - 2699.2011.02540. x" type="journal article" year="2011">
Löwerberg-Neto
<emphasis id="B92B326AF057FFCFFC2FFF11FBAC2FAA" box="[980,1026,152,174]" italics="true" pageId="11" pageNumber="292">et al</emphasis>
. 2011
</bibRefCitation>
(biogeography);
<bibRefCitation id="EFCE9389F057FFCFFB0FFF10FED92FC8" author="Lowerberg-Neto" pageId="11" pageNumber="292" refString="Lowerberg-Neto, P. &amp; Carvalho, C. J. B. de (2013) Muscidae (Insecta: Diptera) of Latin America and the Caribbean: geographic distribution and check-list by country. Zootaxa, 3650 (1), 01 - 147. http: // dx. doi. org / 10.11646 / zootaxa. 3650.1.1" type="journal article" year="2013">Löwerberg-Neto &amp; Carvalho 2013</bibRefCitation>
(checklist);
<bibRefCitation id="EFCE9389F057FFCFFE0EFF3FFDF32FC8" author="Pont" box="[501,605,182,204]" pageId="11" pageNumber="292" refString="Pont, A. C. (2013). The Fanniidae and Muscidae (Diptera) described by Paul Stein (1852 - 1921). Journal of Zoological Systematics and Evolutionary Research, 89, 31 - 166. http: // dx. doi. org / 10.1002 / zoos. 201300004" type="journal article" year="2013">Pont 2013</bibRefCitation>
(
<typeStatus id="54E450DAF057FFCFFD91FF3EFD392FC8" box="[618,663,182,204]" pageId="11" pageNumber="292">type</typeStatus>
specimens).
</paragraph>
<paragraph id="8BE0EE78F057FFCFFF6CFF5DFACA2FEF" blockId="11.[151,1436,152,235]" box="[151,1380,213,235]" pageId="11" pageNumber="292">
<taxonomicName id="4C5F95FBF057FFCFFF6CFF5DFE812FEF" box="[151,303,213,235]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B92B326AF057FFCFFF6CFF5DFE812FEF" box="[151,303,213,235]" italics="true" pageId="11" pageNumber="292">Helina simplex</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFCE9389F057FFCFFEC6FF5DFDF82FEF" author="Lowerberg-Neto" box="[317,598,213,235]" pageId="11" pageNumber="292" refString="Lowerberg-Neto, P., Carvalho, C. J. B. de &amp; Hawkins, B. A. (2011) Tropical niche conservatism as a historical narrative hypothesis for the Neotropics: a case study using the fly family Muscidae. Journal of Biogeography, 38, 1936 - 1947. http: // dx. doi. org / 10.1111 / j. 1365 - 2699.2011.02540. x" type="journal article" year="2011">
Löwerberg-Neto
<emphasis id="B92B326AF057FFCFFE10FF5EFDB62FEF" box="[491,536,213,235]" italics="true" pageId="11" pageNumber="292">et al</emphasis>
. 2011
</bibRefCitation>
(biogeography);
<bibRefCitation id="EFCE9389F057FFCFFCFEFF5DFBCA2FEF" author="Lowerberg-Neto" box="[773,1124,213,235]" pageId="11" pageNumber="292" refString="Lowerberg-Neto, P. &amp; Carvalho, C. J. B. de (2013) Muscidae (Insecta: Diptera) of Latin America and the Caribbean: geographic distribution and check-list by country. Zootaxa, 3650 (1), 01 - 147. http: // dx. doi. org / 10.11646 / zootaxa. 3650.1.1" type="journal article" year="2013">Löwerberg-Neto &amp; Carvalho 2013</bibRefCitation>
(checklist, new records).
</paragraph>
</subSubSection>
<subSubSection id="C345BDF3F057FFCFFF6CFE9FFDA029AB" pageId="11" pageNumber="292" type="description">
<paragraph id="8BE0EE78F057FFCFFF6CFE9FFBE02E2B" blockId="11.[151,1437,279,1999]" box="[151,1102,279,304]" pageId="11" pageNumber="292">
<emphasis id="B92B326AF057FFCFFF6CFE9FFE932E34" bold="true" box="[151,317,279,304]" pageId="11" pageNumber="292">Redescription</emphasis>
.
<emphasis id="B92B326AF057FFCFFEB0FE9FFE292E34" bold="true" box="[331,391,279,304]" italics="true" pageId="11" pageNumber="292">Male</emphasis>
(
<figureCitation id="1364F2FDF057FFCFFE6EFE9FFE762E34" box="[405,472,279,304]" captionStart="FIGURE 5" captionStartId="12.[151,250,1410,1432]" captionTargetBox="[189,1399,193,1386]" captionTargetId="figure@12.[187,1399,193,1389]" captionTargetPageId="12" captionText="FIGURE 5. Helina chilensis Malloch. A – H. Male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate and surstyli, posterior view. G. Cercal plate and surstyli, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). I – J. Female, apical portion of ovipositor: I. Dorsal view. J. Ventral view (scalebar: 0.5 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; epiph, epiphallus; epiprct, epiproct; gon, gonopodo; hyprct, hypoproct; par, paramere; st 8, sternite 8; tg 8, tergite 8.)" httpUri="https://zenodo.org/record/265623/files/figure.png" pageId="11" pageNumber="292">Fig. 5</figureCitation>
A). Length. Body:
<quantity id="4CA7439DF057FFCFFD57FE9FFCFF2E2B" box="[684,849,279,304]" metricMagnitude="-3" metricUnit="m" metricValue="6.615" metricValueMax="7.67" metricValueMin="5.56" pageId="11" pageNumber="292" unit="mm" value="6.615" valueMax="7.67" valueMin="5.56">5.56–7.67 mm</quantity>
, wing:
<quantity id="4CA7439DF057FFCFFC58FE9FFBE72E2B" box="[931,1097,279,304]" metricMagnitude="-3" metricUnit="m" metricValue="5.995" metricValueMax="6.76" metricValueMin="5.23" pageId="11" pageNumber="292" unit="mm" value="5.995" valueMax="6.76" valueMin="5.23">5.23–6.76 mm</quantity>
.
</paragraph>
<paragraph id="8BE0EE78F057FFCFFF3CFEB4FC102EC4" blockId="11.[151,1437,279,1999]" pageId="11" pageNumber="292">
<emphasis id="B92B326AF057FFCFFF3CFEB4FEAA2E51" box="[199,260,316,341]" italics="true" pageId="11" pageNumber="292">Head</emphasis>
(
<figureCitation id="1364F2FDF057FFCFFEEDFEB4FEF32E51" box="[278,349,316,341]" captionStart="FIGURE 5" captionStartId="12.[151,250,1410,1432]" captionTargetBox="[189,1399,193,1386]" captionTargetId="figure@12.[187,1399,193,1389]" captionTargetPageId="12" captionText="FIGURE 5. Helina chilensis Malloch. A – H. Male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate and surstyli, posterior view. G. Cercal plate and surstyli, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). I – J. Female, apical portion of ovipositor: I. Dorsal view. J. Ventral view (scalebar: 0.5 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; epiph, epiphallus; epiprct, epiproct; gon, gonopodo; hyprct, hypoproct; par, paramere; st 8, sternite 8; tg 8, tergite 8.)" httpUri="https://zenodo.org/record/265623/files/figure.png" pageId="11" pageNumber="292">Fig. 5</figureCitation>
B). Black. Holoptic, the shortest distance between eyes is
<quantity id="4CA7439DF057FFCFFBFFFEB4FB032E50" box="[1028,1197,316,341]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" metricValueMax="3.2" metricValueMin="1.6" pageId="11" pageNumber="292" unit="mm" value="0.24" valueMax="0.32" valueMin="0.16">0.16–0.32 mm</quantity>
. 6–8 pairs of frontal setae. Eye hairs long and sparse. Lunule pale brown; fronto-orbital plate, parafacial, gena, postgena, and occiput black with silver pollinosity. Postocular setae proclinate. Parafrontal plate bare. Antenna dark brown; arista plumose, two or fourth times as long as its basal diameter. Palpus black.
</paragraph>
<paragraph id="8BE0EE78F057FFCFFF3CFE44FE8D2D93" blockId="11.[151,1437,279,1999]" pageId="11" pageNumber="292">
<emphasis id="B92B326AF057FFCFFF3CFE44FEB62EE1" box="[199,280,460,485]" italics="true" pageId="11" pageNumber="292">Thorax</emphasis>
(
<figureCitation id="1364F2FDF057FFCFFED3FE44FEC52EE1" box="[296,363,460,485]" captionStart="FIGURE 5" captionStartId="12.[151,250,1410,1432]" captionTargetBox="[189,1399,193,1386]" captionTargetId="figure@12.[187,1399,193,1389]" captionTargetPageId="12" captionText="FIGURE 5. Helina chilensis Malloch. A – H. Male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate and surstyli, posterior view. G. Cercal plate and surstyli, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). I – J. Female, apical portion of ovipositor: I. Dorsal view. J. Ventral view (scalebar: 0.5 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; epiph, epiphallus; epiprct, epiproct; gon, gonopodo; hyprct, hypoproct; par, paramere; st 8, sternite 8; tg 8, tergite 8.)" httpUri="https://zenodo.org/record/265623/files/figure.png" pageId="11" pageNumber="292">Fig. 5</figureCitation>
C). Scutum black with three white-grey pollinose vittae; scutellum; anepisternum, anepimeron, katepimeron, katepisternum, proepisternum, proepimeron and meron black with grey pollinosity; anterior and posterior spiracles black. Chaetotaxy: acrostichal setae 1-2+1; dorsocentral setae 2–4; humeral setae 3–5; notopleural setae 2, posterior seta smaller. Prealar strong, one third-one fourth of the length of the anterior supraalar postsutural seta. Scutellum with thinner and short basal and subapical setae. Anepisternum with a series of 4–6 strong setae.
</paragraph>
<paragraph id="8BE0EE78F057FFCFFF3CFD2DFDEA2C03" blockId="11.[151,1437,279,1999]" pageId="11" pageNumber="292">
<emphasis id="B92B326AF057FFCFFF3CFD2DFF512DB8" box="[199,255,677,700]" italics="true" pageId="11" pageNumber="292">Wing</emphasis>
. Infuscate, brown; costal spine two or three time as long as its costal bristles; the transverse cross-vein dm-cu slightly curved; vein R 4+5 and vein M divergent apically; cross vein dm-cu in basal half of cell r4+5. Both calypters yellow with yellow margins.
</paragraph>
<paragraph id="8BE0EE78F057FFCFFF3CFC9CFD282B6B" blockId="11.[151,1437,279,1999]" pageId="11" pageNumber="292">
<emphasis id="B92B326AF057FFCFFF3CFC9CFF522C2F" box="[199,252,788,811]" italics="true" pageId="11" pageNumber="292">Legs</emphasis>
. Coxa and trochanter black; fore femur black (some specimens yellow apically), fore tibia black; mid femur black at base with apical third yellow (some specimens with apical middle yellow), mid tibia black or darkbrown; hind femur black at base with apical third yellow (some specimens with apical two-third or more yellow), hind tibia dark-brown. Fore femur with rows of dorsal, posterodorsal, and posteroventral setae; fore tibia with one posterior setae, and 3 preapical setae. Mid femur with 4–6 setae in the basal third on ventral surface, and 3 preapical setae on posterodorsal to posterior surface; mid tibia with 3–4 posterior setae, 1–2 posteroventral seta, 1– 2 anterodorsal setae (some speciemens without setae) and 4 apical setae (anterodorsal, posteroventral, ventral, and anteroventral). Hind femur with a anterodorsal row, and 4–5 setae in the apical middle of anteroventral surface; hind tibia with 3–5 anteroventral setae, 3–6 anterodorsal setae, and 6–7 posterodorsal setae, calcar absent. Fore claws and pulvilli longer than mid and hind.
</paragraph>
<paragraph id="8BE0EE78F057FFCFFF3CFBF2FC902BB3" blockId="11.[151,1437,279,1999]" pageId="11" pageNumber="292">
<emphasis id="B92B326AF057FFCFFF3CFBF2FE9F2B97" box="[199,305,1146,1171]" italics="true" pageId="11" pageNumber="292">Abdomen</emphasis>
(
<figureCitation id="1364F2FDF057FFCFFEC4FBF2FE2C2B97" box="[319,386,1146,1171]" captionStart="FIGURE 5" captionStartId="12.[151,250,1410,1432]" captionTargetBox="[189,1399,193,1386]" captionTargetId="figure@12.[187,1399,193,1389]" captionTargetPageId="12" captionText="FIGURE 5. Helina chilensis Malloch. A – H. Male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate and surstyli, posterior view. G. Cercal plate and surstyli, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). I – J. Female, apical portion of ovipositor: I. Dorsal view. J. Ventral view (scalebar: 0.5 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; epiph, epiphallus; epiprct, epiproct; gon, gonopodo; hyprct, hypoproct; par, paramere; st 8, sternite 8; tg 8, tergite 8.)" httpUri="https://zenodo.org/record/265623/files/figure.png" pageId="11" pageNumber="292">Fig. 5</figureCitation>
D). Black with brown pollinosity. Sternite 5 longer than wide, with several strong setae on the apical third; posterior margin with two projections (
<figureCitation id="1364F2FDF057FFCFFD21FB16FCB32BB3" box="[730,797,1182,1207]" captionStart="FIGURE 5" captionStartId="12.[151,250,1410,1432]" captionTargetBox="[189,1399,193,1386]" captionTargetId="figure@12.[187,1399,193,1389]" captionTargetPageId="12" captionText="FIGURE 5. Helina chilensis Malloch. A – H. Male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate and surstyli, posterior view. G. Cercal plate and surstyli, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). I – J. Female, apical portion of ovipositor: I. Dorsal view. J. Ventral view (scalebar: 0.5 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; epiph, epiphallus; epiprct, epiproct; gon, gonopodo; hyprct, hypoproct; par, paramere; st 8, sternite 8; tg 8, tergite 8.)" httpUri="https://zenodo.org/record/265623/files/figure.png" pageId="11" pageNumber="292">Fig. 5</figureCitation>
E).
</paragraph>
<paragraph id="8BE0EE78F057FFCFFF3CFB4AFD682A6F" blockId="11.[151,1437,279,1999]" pageId="11" pageNumber="292">
<taxonomicName id="4C5F95FBF057FFCFFF3CFB4AFEEF2BDF" box="[199,321,1218,1243]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="292" phylum="Tracheophyta" rank="genus">
<emphasis id="B92B326AF057FFCFFF3CFB4AFEEF2BDF" box="[199,321,1218,1243]" italics="true" pageId="11" pageNumber="292">Terminalia</emphasis>
</taxonomicName>
. Cercal plate longer than wide, with a distal incision; with two straight protuberances at the apex (
<figureCitation id="1364F2FDF057FFCFFF64FB6EFF4C2BFB" box="[159,226,1254,1279]" captionStart="FIGURE 5" captionStartId="12.[151,250,1410,1432]" captionTargetBox="[189,1399,193,1386]" captionTargetId="figure@12.[187,1399,193,1389]" captionTargetPageId="12" captionText="FIGURE 5. Helina chilensis Malloch. A – H. Male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate and surstyli, posterior view. G. Cercal plate and surstyli, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). I – J. Female, apical portion of ovipositor: I. Dorsal view. J. Ventral view (scalebar: 0.5 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; epiph, epiphallus; epiprct, epiproct; gon, gonopodo; hyprct, hypoproct; par, paramere; st 8, sternite 8; tg 8, tergite 8.)" httpUri="https://zenodo.org/record/265623/files/figure.png" pageId="11" pageNumber="292">Fig. 5</figureCitation>
F). Surstylus curving forwards basally and inwards apically, with a wide margin in posterior view, and with strong setae on inner surface (
<figureCitation id="1364F2FDF057FFCFFE08FA82FDED2A27" box="[499,579,1290,1315]" captionStart="FIGURE 5" captionStartId="12.[151,250,1410,1432]" captionTargetBox="[189,1399,193,1386]" captionTargetId="figure@12.[187,1399,193,1389]" captionTargetPageId="12" captionText="FIGURE 5. Helina chilensis Malloch. A – H. Male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate and surstyli, posterior view. G. Cercal plate and surstyli, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). I – J. Female, apical portion of ovipositor: I. Dorsal view. J. Ventral view (scalebar: 0.5 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; epiph, epiphallus; epiprct, epiproct; gon, gonopodo; hyprct, hypoproct; par, paramere; st 8, sternite 8; tg 8, tergite 8.)" httpUri="https://zenodo.org/record/265623/files/figure.png" pageId="11" pageNumber="292">Figs. 5</figureCitation>
F–G). Aedeagus with aedeagal apodeme straight and strongly sclerotized; epiphalus slightly sclerotized, paramere slightly bent downward with setula; gonopod with short setulae; and distiphallus tubular, slightly sclerotized (
<figureCitation id="1364F2FDF057FFCFFDA4FADAFD0F2A6F" box="[607,673,1362,1387]" captionStart="FIGURE 5" captionStartId="12.[151,250,1410,1432]" captionTargetBox="[189,1399,193,1386]" captionTargetId="figure@12.[187,1399,193,1389]" captionTargetPageId="12" captionText="FIGURE 5. Helina chilensis Malloch. A – H. Male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate and surstyli, posterior view. G. Cercal plate and surstyli, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). I – J. Female, apical portion of ovipositor: I. Dorsal view. J. Ventral view (scalebar: 0.5 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; epiph, epiphallus; epiprct, epiproct; gon, gonopodo; hyprct, hypoproct; par, paramere; st 8, sternite 8; tg 8, tergite 8.)" httpUri="https://zenodo.org/record/265623/files/figure.png" pageId="11" pageNumber="292">Fig. 5</figureCitation>
H).
</paragraph>
<paragraph id="8BE0EE78F057FFCFFF3CFAFEFCDB2A8B" blockId="11.[151,1437,279,1999]" box="[199,885,1398,1423]" pageId="11" pageNumber="292">
<emphasis id="B92B326AF057FFCFFF3CFAFEFEB02A8B" bold="true" box="[199,286,1398,1423]" italics="true" pageId="11" pageNumber="292">Female</emphasis>
. Length. Body:
<quantity id="4CA7439DF057FFCFFE2EFAFEFDD72A8B" box="[469,633,1398,1423]" metricMagnitude="-3" metricUnit="m" metricValue="6.58" metricValueMax="7.05" metricValueMin="6.11" pageId="11" pageNumber="292" unit="mm" value="6.58" valueMax="7.05" valueMin="6.11">6.11–7.05 mm</quantity>
, wing:
<quantity id="4CA7439DF057FFCFFD30FAFEFCDF2A8B" box="[715,881,1398,1423]" metricMagnitude="-3" metricUnit="m" metricValue="5.91" metricValueMax="6.02" metricValueMin="5.8" pageId="11" pageNumber="292" unit="mm" value="5.91" valueMax="6.02" valueMin="5.8">5.80–6.02 mm</quantity>
.
</paragraph>
<paragraph id="8BE0EE78F057FFCFFF3CFA12FDA029AB" blockId="11.[151,1437,279,1999]" pageId="11" pageNumber="292">
Differs from male as follows:
<emphasis id="B92B326AF057FFCFFDDFFA12FDCE2AB7" box="[548,608,1434,1459]" italics="true" pageId="11" pageNumber="292">Head</emphasis>
, dichoptic, the shortest distance between eyes is
<quantity id="4CA7439DF057FFCFFB63FA12FAEF2AB6" box="[1176,1345,1434,1459]" metricMagnitude="-4" metricUnit="m" metricValue="8.7" metricValueMax="8.9" metricValueMin="8.5" pageId="11" pageNumber="292" unit="mm" value="0.87" valueMax="0.89" valueMin="0.85">0.85–0.89 mm</quantity>
; frontoorbital plate setulose with 2 reclinate orbital setae. Parafrontal plate setulose.
<emphasis id="B92B326AF057FFCFFC0EFA48FB842AD3" box="[1013,1066,1472,1495]" italics="true" pageId="11" pageNumber="292">Legs</emphasis>
. Mid femur with 4–5 setae in the basal third on ventral surface; with a row of anterior setae in the basal middle, and 3 preapical setae on anterodorsal to posterior surface. Hind femur with only an anterodorsal row setae, and 3 long and curved setae in the basal third on anteroventral surface, hind tibia with only 3 anteroventral setae and 4 anterodorsal setae.
<taxonomicName id="4C5F95FBF057FFCFFB54F9A2FA882947" box="[1199,1318,1578,1603]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="292" phylum="Tracheophyta" rank="genus">
<emphasis id="B92B326AF057FFCFFB54F9A2FA882947" box="[1199,1318,1578,1603]" italics="true" pageId="11" pageNumber="292">Terminalia</emphasis>
</taxonomicName>
. Tergite 8 with 2 parallel sclerotized plates, fused and with setulae on distal margin; epiproct with proximal margin curved, and several strong spines, cercus digitiform with strong spine (
<figureCitation id="1364F2FDF057FFCFFCB4F9FAFC3F298F" box="[847,913,1650,1675]" captionStart="FIGURE 5" captionStartId="12.[151,250,1410,1432]" captionTargetBox="[189,1399,193,1386]" captionTargetId="figure@12.[187,1399,193,1389]" captionTargetPageId="12" captionText="FIGURE 5. Helina chilensis Malloch. A – H. Male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate and surstyli, posterior view. G. Cercal plate and surstyli, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). I – J. Female, apical portion of ovipositor: I. Dorsal view. J. Ventral view (scalebar: 0.5 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; epiph, epiphallus; epiprct, epiproct; gon, gonopodo; hyprct, hypoproct; par, paramere; st 8, sternite 8; tg 8, tergite 8.)" httpUri="https://zenodo.org/record/265623/files/figure.png" pageId="11" pageNumber="292">
Fig.
<date id="FFE1C8B8F057FFCFFC79F9FAFC3F298F" box="[898,913,1650,1675]" pageId="11" pageNumber="292">5</date>
</figureCitation>
I). Hypoproct rounded, sclerotized, with strong setulae on distal margin (
<figureCitation id="1364F2FDF057FFCFFE4BF91EFE5A29AB" box="[432,500,1686,1711]" captionStart="FIGURE 5" captionStartId="12.[151,250,1410,1432]" captionTargetBox="[189,1399,193,1386]" captionTargetId="figure@12.[187,1399,193,1389]" captionTargetPageId="12" captionText="FIGURE 5. Helina chilensis Malloch. A – H. Male: A. Lateral view (scale bar: 1 mm). B. Head, frontal view. C. Thorax, dorsal view. D. Abdomen, dorsal view (scale bars: 0.5 mm). E. Sternite 5 (scale bar: 0.2 mm). F. Cercal plate and surstyli, posterior view. G. Cercal plate and surstyli, lateral view (scale bar: 0.1 mm). H. Phallic complex, lateral view (scale bar: 0.1 mm). I – J. Female, apical portion of ovipositor: I. Dorsal view. J. Ventral view (scalebar: 0.5 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; epiph, epiphallus; epiprct, epiproct; gon, gonopodo; hyprct, hypoproct; par, paramere; st 8, sternite 8; tg 8, tergite 8.)" httpUri="https://zenodo.org/record/265623/files/figure.png" pageId="11" pageNumber="292">Fig. 5</figureCitation>
J).
</paragraph>
</subSubSection>
<subSubSection id="C345BDF3F057FFC9FF3CF932FAC52EA8" lastPageId="13" lastPageNumber="294" pageId="11" pageNumber="292" type="materials_examined">
<paragraph id="8BE0EE78F057FFCFFF3CF932FB7B28CB" blockId="11.[151,1437,279,1999]" pageId="11" pageNumber="292">
<emphasis id="B92B326AF057FFCFFF3CF932FE5E29D7" bold="true" box="[199,496,1722,1747]" pageId="11" pageNumber="292">Type material examined</emphasis>
.
<emphasis id="B92B326AF057FFCFFDF8F932FDDE29D7" bold="true" box="[515,624,1722,1747]" pageId="11" pageNumber="292">Holotype</emphasis>
1 male, pinned, without left wing, labels: “
<taxonomicName id="4C5F95FBF057FFCFFB83F932FB6B29D7" box="[1144,1221,1722,1747]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="genus">Helina</taxonomicName>
/
<taxonomicName id="4C5F95FBF057FFCFFB1FF932FAE829D7" box="[1252,1350,1722,1747]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="species" species="chilensis">chilensis</taxonomicName>
/ Type [handwritten] / det. JRMalloch [printed]” on white paper, black frame; “Perales /
<date id="FFE1C8B8F057FFCFFBDAF956FBFF29F3" box="[1057,1105,1758,1783]" pageId="11" pageNumber="292" value="1925-01">I-25</date>
” handwritten on white paper; “Type No.[printed] / 49855 [handwritten] / U.S.N.M. [printed]” on red paper (USNM).
<emphasis id="B92B326AF057FFCFFB88F88AFB4E281F" bold="true" box="[1139,1248,1794,1819]" pageId="11" pageNumber="292">Holotype</emphasis>
, 1 male, pinned, damaged, “
<taxonomicName id="4C5F95FBF057FFCFFEEEF8AEFECC283B" box="[277,354,1830,1855]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="genus">Helina</taxonomicName>
/
<taxonomicName id="4C5F95FBF057FFCFFE80F8AEFE7A283B" box="[379,468,1830,1855]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="species" species="simplex">simplex</taxonomicName>
/ Type [handwritten] / det. JRMalloch [printed]” on white paper, black frame; “Type No. [printed] / 49854 [handwritten]” on red paper (USNM).
<emphasis id="B92B326AF057FFCFFCC5F8C2FC052867" bold="true" box="[830,939,1866,1891]" pageId="11" pageNumber="292">Holotype</emphasis>
1 male, pinned, without fore and mind legs, labels: “
<taxonomicName id="4C5F95FBF057FFCFFF0FF8E6FEEF2883" box="[244,321,1902,1927]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="genus">Helina</taxonomicName>
/
<taxonomicName id="4C5F95FBF057FFCFFEA5F8E6FDA92883" box="[350,519,1902,1927]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="292" phylum="Arthropoda" rank="species" species="fulvocalyptrata">fulvocalyptrata</taxonomicName>
/ Type [handwritten] / det. JRMalloch [printed]” on white paper, black frame; “Argentina: / Terr. Rio Negro. / F. &amp; M. Edwards. / B. M.
<date id="FFE1C8B8F057FFCFFCD1F81AFC2528AF" box="[810,907,1938,1963]" pageId="11" pageNumber="292" value="1927" valueMax="1963">1927-63</date>
.” printed, on white paper; “L Correntoso. /
<date id="FFE1C8B8F057FFCFFA8CF81BFEBD28CB" pageId="11" pageNumber="292" value="1926-11-18" valueMax="1926-11-25" valueMin="1926-11-18">18- 25.xi.1926</date>
.” printed, on white paper; “Holo- / type” printed, on white paper, red frame (BMNH).
</paragraph>
<caption id="DF20BEF0F050FFC8FF6CFA0AFE382935" httpUri="https://zenodo.org/record/265623/files/figure.png" pageId="12" pageNumber="293" targetBox="[189,1399,193,1386]" targetPageId="12">
<paragraph id="8BE0EE78F050FFC8FF6CFA0AFE382935" blockId="12.[151,1436,1410,1585]" pageId="12" pageNumber="293">
<emphasis id="B92B326AF050FFC8FF6CFA0AFEBC2A9C" bold="true" box="[151,274,1410,1432]" pageId="12" pageNumber="293">FIGURE 5.</emphasis>
<taxonomicName id="4C5F95FBF050FFC8FEE3FA0AFD982A9C" authority="Malloch. A" authorityName="Malloch. A" box="[280,566,1410,1432]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="293" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="B92B326AF050FFC8FEE3FA0AFE132A9C" box="[280,445,1410,1432]" italics="true" pageId="12" pageNumber="293">Helina chilensis</emphasis>
Malloch.
<emphasis id="B92B326AF050FFC8FDD8FA0AFD982A9C" bold="true" box="[547,566,1410,1432]" pageId="12" pageNumber="293">A</emphasis>
</taxonomicName>
<emphasis id="B92B326AF050FFC8FDCDFA0AFDF32A9C" bold="true" box="[566,605,1410,1432]" pageId="12" pageNumber="293">–H.</emphasis>
Male:
<emphasis id="B92B326AF050FFC8FD58FA0AFD122A9C" bold="true" box="[675,700,1410,1432]" pageId="12" pageNumber="293">A.</emphasis>
Lateral view (scale bar: 1 mm).
<emphasis id="B92B326AF050FFC8FBFEFA0AFBB22A9C" bold="true" box="[1029,1052,1410,1432]" pageId="12" pageNumber="293">B.</emphasis>
Head, frontal view.
<emphasis id="B92B326AF050FFC8FB11FA0AFAAC2A9C" bold="true" box="[1258,1282,1410,1432]" pageId="12" pageNumber="293">C.</emphasis>
Thorax, dorsal view.
<emphasis id="B92B326AF050FFC8FF2FFA29FF432AB3" bold="true" box="[212,237,1441,1463]" pageId="12" pageNumber="293">D.</emphasis>
Abdomen, dorsal view (scale bars: 0.5 mm).
<emphasis id="B92B326AF050FFC8FD38FA29FD742AB3" bold="true" box="[707,730,1441,1463]" pageId="12" pageNumber="293">E.</emphasis>
Sternite 5 (scale bar: 0.2 mm).
<emphasis id="B92B326AF050FFC8FBDAFA29FB9C2AB3" bold="true" box="[1057,1074,1441,1463]" pageId="12" pageNumber="293">F.</emphasis>
Cercal plate and surstyli, posterior view.
<emphasis id="B92B326AF050FFC8FF2FFA37FF462AD1" bold="true" box="[212,232,1471,1493]" pageId="12" pageNumber="293">G.</emphasis>
Cercal plate and surstyli, lateral view (scale bar: 0.1 mm).
<emphasis id="B92B326AF050FFC8FCB0FA37FCC82AD1" bold="true" box="[843,870,1471,1493]" pageId="12" pageNumber="293">H.</emphasis>
Phallic complex, lateral view (scale bar: 0.1 mm).
<emphasis id="B92B326AF050FFC8FA89FA37FA322AD1" bold="true" box="[1394,1436,1471,1493]" pageId="12" pageNumber="293">I–J.</emphasis>
Female, apical portion of ovipositor:
<emphasis id="B92B326AF050FFC8FDE2FA56FD862AF0" bold="true" box="[537,552,1502,1524]" pageId="12" pageNumber="293">I.</emphasis>
Dorsal view.
<emphasis id="B92B326AF050FFC8FD40FA56FD602AF0" bold="true" box="[699,718,1502,1524]" pageId="12" pageNumber="293">J.</emphasis>
Ventral view (scalebar: 0.5 mm). (Abbreviations: aed apod, aedeagal apodeme; distiph, distiphallus; epiph, epiphallus; epiprct, epiproct; gon, gonopodo; hyprct, hypoproct; par, paramere; st 8, sternite 8; tg 8, tergite 8.)
</paragraph>
</caption>
<paragraph id="8BE0EE78F050FFC9FF3CF9D0FAC52EA8" blockId="12.[151,1436,1624,2009]" lastBlockId="13.[151,1437,151,1040]" lastPageId="13" lastPageNumber="294" pageId="12" pageNumber="293">
<materialsCitation id="3B37E425F050FFC8FF3CF9D0FC9A2991" ID-GBIF-Occurrence="1291551069" collectingDate="2011-01-15" collectionCode="MACN" collectorName="PN Lago Puelo &amp; Intendencia &amp; Mulieri &amp; Patitucci" country="Argentina" location="Other" pageId="12" pageNumber="293" specimenCount="1" specimenCount-male="1" stateProvince="Chubut">
<emphasis id="B92B326AF050FFC8FF3CF9D0FC832975" bold="true" box="[199,813,1624,1649]" pageId="12" pageNumber="293">
<location id="8E80B8A3F050FFC8FF3CF9D0FEA12975" LSID="urn:lsid:plazi:treatment:03F65F6EF056FFC9FF6CF968FECE2B14:8E80B8A3F050FFC8FF3CF9D0FEA12975" box="[199,271,1624,1649]" country="Argentina" name="Other" pageId="12" pageNumber="293" stateProvince="Chubut">Other</location>
material examined.
<collectingCountry id="F348AEE8F050FFC8FDEAF9D0FD6F2975" box="[529,705,1624,1649]" name="Argentina" pageId="12" pageNumber="293">ARGENTINA</collectingCountry>
:
<collectingRegion id="499B209AF050FFC8FD2AF9D0FC832975" box="[721,813,1624,1649]" country="Argentina" name="Chubut" pageId="12" pageNumber="293">Chubut</collectingRegion>
</emphasis>
:
<specimenCount id="9D5925F1F050FFC8FCBBF9D1FC3D2975" box="[832,915,1624,1649]" pageId="12" pageNumber="293" type="male">1 male</specimenCount>
,
<collectorName id="26AA8BAEF050FFC8FC58F9D1FBF52975" box="[931,1115,1624,1649]" pageId="12" pageNumber="293">PN Lago Puelo</collectorName>
,
<collectorName id="26AA8BAEF050FFC8FB91F9D0FB5E2975" box="[1130,1264,1624,1649]" pageId="12" pageNumber="293">Intendencia</collectorName>
, -42.085077, - 71.614662,
<date id="FFE1C8B8F050FFC8FEDAF9F4FE3D2991" box="[289,403,1660,1685]" pageId="12" pageNumber="293" value="2011-01-15">
<collectingDate id="EFA53150F050FFC8FEDAF9F4FE3D2991" box="[289,403,1660,1685]" pageId="12" pageNumber="293" value="2011-01-15">15-I-2011</collectingDate>
</date>
,
<collectorName id="26AA8BAEF050FFC8FE58F9F4FE592991" box="[419,503,1660,1685]" pageId="12" pageNumber="293">Mulieri</collectorName>
&amp;
<collectorName id="26AA8BAEF050FFC8FDDDF9F4FD262991" box="[550,648,1660,1685]" pageId="12" pageNumber="293">Patitucci</collectorName>
leg. (
<collectionCode id="ED4E76BDF050FFC8FD2AF9F5FC822991" box="[721,812,1661,1685]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="12" pageNumber="293">MACN</collectionCode>
)
</materialsCitation>
; 6 females, PN Los Alerces, Cabaña La Cascada, - 42.888499, -71.592376,
<date id="FFE1C8B8F050FFC8FE50F929FDAC29BD" box="[427,514,1697,1721]" pageId="12" pageNumber="293" value="2013-02">II-2013</date>
, Patitucci leg. (MACN); 1 male, PN Los Alerces, Puerto Mermoud, -42.723187, - 71.748741,
<date id="FFE1C8B8F050FFC8FEDEF94DFED029D9" box="[293,382,1733,1757]" pageId="12" pageNumber="293" value="2014-10">X-2014</date>
, Mulieri leg. (MACN);
<materialsCitation id="3B37E425F050FFC8FD4AF94DFD3F2805" ID-GBIF-Occurrence="1291551064" collectingDate="1993-01-10" collectionCode="FAUBA" collectorName="Malargue &amp; Valle Hermoso &amp; Medan" country="Argentina" location="Mendoza" pageId="12" pageNumber="293" specimenCount="3" specimenCount-male="3" stateProvince="Mendoza">
<emphasis id="B92B326AF050FFC8FD4AF94DFCB129DA" bold="true" box="[689,799,1733,1758]" pageId="12" pageNumber="293">
<collectingRegion id="499B209AF050FFC8FD4AF94DFCB129DA" box="[689,799,1733,1758]" country="Argentina" name="Mendoza" pageId="12" pageNumber="293">Mendoza</collectingRegion>
</emphasis>
:
<specimenCount id="9D5925F1F050FFC8FCC3F94DFC3429DA" box="[824,922,1733,1758]" pageId="12" pageNumber="293" type="male">3 males</specimenCount>
,
<collectorName id="26AA8BAEF050FFC8FC54F94DFBB329DA" box="[943,1053,1733,1758]" pageId="12" pageNumber="293">Malargüe</collectorName>
,
<collectorName id="26AA8BAEF050FFC8FBC9F94DFB4829D9" box="[1074,1254,1733,1758]" pageId="12" pageNumber="293">Valle Hermoso</collectorName>
, -35.066667, - 70.216667,
<date id="FFE1C8B8F050FFC8FEE7F961FE202805" box="[284,398,1769,1793]" pageId="12" pageNumber="293" value="1993-01-10">
<collectingDate id="EFA53150F050FFC8FEE7F961FE202805" box="[284,398,1769,1793]" pageId="12" pageNumber="293" value="1993-01-10">10-I-1993</collectingDate>
</date>
,
<collectorName id="26AA8BAEF050FFC8FE62F960FE492805" box="[409,487,1768,1793]" pageId="12" pageNumber="293">Medan</collectorName>
leg. (
<collectionCode id="ED4E76BDF050FFC8FDDDF961FD242805" box="[550,650,1769,1793]" pageId="12" pageNumber="293">FAUBA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B37E425F050FFC8FD67F960FD2D2821" ID-GBIF-Occurrence="1291551056" collectingDate="2013-01" collectionCode="MACN" collectorName="Batea Mahuida &amp; Mulieri &amp; Patitucci" country="Argentina" location="Neuquen" pageId="12" pageNumber="293" specimenCount="4" specimenCount-female="4" stateProvince="Neuquen">
<emphasis id="B92B326AF050FFC8FD67F960FCA92805" bold="true" box="[668,775,1768,1793]" pageId="12" pageNumber="293">
<collectingRegion id="499B209AF050FFC8FD67F960FCA92805" box="[668,775,1768,1793]" country="Argentina" name="Neuquen" pageId="12" pageNumber="293">Neuquén</collectingRegion>
</emphasis>
:
<specimenCount id="9D5925F1F050FFC8FCEDF961FC2B2805" box="[790,901,1768,1793]" pageId="12" pageNumber="293" type="female">4 females</specimenCount>
, APN
<collectorName id="26AA8BAEF050FFC8FC35F961FBD22805" box="[974,1148,1768,1793]" pageId="12" pageNumber="293">Batea Mahuida</collectorName>
, -38.845457, -71.093002,
<date id="FFE1C8B8F050FFC8FF6CF885FF4A2821" box="[151,228,1805,1829]" pageId="12" pageNumber="293" value="2013-01">
<collectingDate id="EFA53150F050FFC8FF6CF885FF4A2821" box="[151,228,1805,1829]" pageId="12" pageNumber="293" value="2013-01">I-2013</collectingDate>
</date>
,
<collectorName id="26AA8BAEF050FFC8FF0FF885FEE62822" box="[244,328,1805,1830]" pageId="12" pageNumber="293">Mulieri</collectorName>
&amp;
<collectorName id="26AA8BAEF050FFC8FE8DF885FE762822" box="[374,472,1805,1830]" pageId="12" pageNumber="293">Patitucci</collectorName>
leg. (
<collectionCode id="ED4E76BDF050FFC8FDDAF885FDD52821" box="[545,635,1805,1829]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="12" pageNumber="293">MACN</collectionCode>
)
</materialsCitation>
, 2 males, APN Batea Mahuida, -38.837593, -71.097928,
<date id="FFE1C8B8F050FFC8FAD6F885FA392821" box="[1325,1431,1805,1829]" pageId="12" pageNumber="293" value="2013-12">XII-2013</date>
, Mulieri &amp; Patitucci leg. (MACN), 4 females, Cerro Chapelco, -40.197050, -71.298453, I/
<date id="FFE1C8B8F050FFC8FB48F8B9FAA7284D" box="[1203,1289,1841,1865]" pageId="12" pageNumber="293" value="1983-02">II-1983</date>
, Gentili leg. (MACN); 2 females, 4 males, Cerro Chapelco, -40.197050, -71.298453,
<date id="FFE1C8B8F050FFC8FC35F8DDFB8D2869" box="[974,1059,1877,1901]" pageId="12" pageNumber="293" value="2011-02">II-2011</date>
, Mariluis leg. (MACN); 2 males, Copahue, -37.820278, -71.101111,
<date id="FFE1C8B8F050FFC8FDD1F8F1FDD92895" box="[554,631,1913,1937]" pageId="12" pageNumber="293" value="2002-01">I-2002</date>
, Medan &amp; Alvarez leg., (FAUBA), 1 female, 1 male, PN Lanín, Hua Hum, -40.116265, -71.662723,
<date id="FFE1C8B8F050FFC8FE05F815FDFD28B1" box="[510,595,1949,1973]" pageId="12" pageNumber="293" value="2011-02">II-2011</date>
, Patitucci leg. (MACN); 3 males, PN Lanín, Lago Tromen, -39.563419, - 71.418872,
<date id="FFE1C8B8F050FFC8FEDBF849FE2428DD" box="[288,394,1985,2009]" pageId="12" pageNumber="293" value="1998-12">XII-1998</date>
, Medan &amp; Basili leg. (FAUBA), 1 female, PN Lanín, Mirador Bandurrias, -40.159804, - 71.371661,
<date id="FFE1C8B8F051FFC9FEE5FF10FEDD2FB4" box="[286,371,152,176]" pageId="13" pageNumber="294" value="2011-02">II-2011</date>
, Mulieri &amp; Patitucci leg. (MACN), 2 females, 3 males, PN Lanín, Ñorquinco, -39.146931, - 71.232717,
<date id="FFE1C8B8F051FFC9FED9FF34FEC12FD0" box="[290,367,188,212]" pageId="13" pageNumber="294" value="2013-01">I-2013</date>
, Olea, Mulieri &amp; Patitucci leg. (MACN), 4 females, 2 males, PN Lanín, Ruca Choroi, - 39.229641, -71.177417,
<date id="FFE1C8B8F051FFC9FE4CFF68FD872FFC" box="[439,553,224,248]" pageId="13" pageNumber="294" value="2013-01-10">10-I-2013</date>
, Olea, Mulieri &amp; Patitucci leg. (MACN), 5 females, 2 males, PN Lanín, Seccional Bandurrias, -40.144240, -71.342986,
<date id="FFE1C8B8F051FFC9FD55FE8CFCAD2E18" box="[686,771,260,284]" pageId="13" pageNumber="294" value="2011-02">II-2011</date>
, Mulieri &amp; Patitucci leg. (MACN).
<materialsCitation id="3B37E425F051FFC9FB6AFE8CFCA72E44" ID-GBIF-Occurrence="1291551063" collectingDate="1962-01" collectionCode="MACN" collectorName="Andes" country="Argentina" location="San Martin" pageId="13" pageNumber="294" specimenCount="1" specimenCount-female="1" stateProvince="San Martin">
<specimenCount id="9D5925F1F051FFC9FB6AFE8CFB5A2E19" box="[1169,1268,260,285]" pageId="13" pageNumber="294" type="female">1 female</specimenCount>
,
<collectingRegion id="499B209AF051FFC9FB05FE8CFAD52E18" box="[1278,1403,260,284]" country="Peru" name="San Martin" pageId="13" pageNumber="294">San Martín</collectingRegion>
de los
<collectorName id="26AA8BAEF051FFC9FF38FEAFFEA32E44" box="[195,269,295,320]" pageId="13" pageNumber="294">Andes</collectorName>
, -40.162242, -71.356484,
<date id="FFE1C8B8F051FFC9FDB9FEAFFD3E2E44" box="[578,656,295,320]" pageId="13" pageNumber="294" value="1962-01">
<collectingDate id="EFA53150F051FFC9FDB9FEAFFD3E2E44" box="[578,656,295,320]" pageId="13" pageNumber="294" value="1962-01">I-1962</collectingDate>
</date>
, (
<collectionCode id="ED4E76BDF051FFC9FD5CFEA0FCAF2E44" box="[679,769,296,320]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="13" pageNumber="294">MACN</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B37E425F051FFC9FCE3FEAFFDD62E60" ID-GBIF-Occurrence="1291551066" collectingDate="1962-01-08" collectionCode="MACN" collectorName="PN Nahuel Huapi &amp; Llao Llao" country="Argentina" location="Rio Negro" pageId="13" pageNumber="294" specimenCount="3" specimenCount-female="3" stateProvince="Rio Negro">
<emphasis id="B92B326AF051FFC9FCE3FEAFFC382E44" bold="true" box="[792,918,295,320]" pageId="13" pageNumber="294">
<collectingRegion id="499B209AF051FFC9FCE3FEAFFC382E44" box="[792,918,295,320]" country="Argentina" name="Rio Negro" pageId="13" pageNumber="294">Río Negro</collectingRegion>
</emphasis>
:
<specimenCount id="9D5925F1F051FFC9FC52FEA0FBB32E44" box="[937,1053,295,320]" pageId="13" pageNumber="294" type="female">3 females</specimenCount>
,
<collectorName id="26AA8BAEF051FFC9FBD7FEA0FB502E44" box="[1068,1278,295,320]" pageId="13" pageNumber="294">PN Nahuel Huapí</collectorName>
,
<collectorName id="26AA8BAEF051FFC9FAF5FEAFFA2D2E44" box="[1294,1411,295,320]" pageId="13" pageNumber="294">Llao Llao</collectorName>
, - 41.046, -71.572897,
<date id="FFE1C8B8F051FFC9FE6FFEC4FE572E61" box="[404,505,332,357]" pageId="13" pageNumber="294" value="1962-01-08">
<collectingDate id="EFA53150F051FFC9FE6FFEC4FE572E61" box="[404,505,332,357]" pageId="13" pageNumber="294" value="1962-01-08">8-I-1962</collectingDate>
</date>
, (
<collectionCode id="ED4E76BDF051FFC9FDEDFEC4FDDF2E60" box="[534,625,332,356]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="13" pageNumber="294">MACN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B37E425F051FFC9FD76FEC4FC1F2E61" ID-GBIF-Occurrence="1291551065" box="[653,945,332,357]" country="Argentina" location="Santa Cruz" pageId="13" pageNumber="294" specimenCount="2" specimenCount-female="2" stateProvince="Santa Cruz">
<emphasis id="B92B326AF051FFC9FD76FEC4FCB12E61" bold="true" box="[653,799,332,357]" pageId="13" pageNumber="294">
<collectingRegion id="499B209AF051FFC9FD76FEC4FCB12E61" box="[653,799,332,357]" country="Argentina" name="Santa Cruz" pageId="13" pageNumber="294">Santa Cruz</collectingRegion>
</emphasis>
:
<specimenCount id="9D5925F1F051FFC9FCC3FEC4FC1F2E61" box="[824,945,332,357]" pageId="13" pageNumber="294" type="female">2 females</specimenCount>
</materialsCitation>
,
<materialsCitation id="3B37E425F051FFC9FC3DFEC4FB9D2E8C" ID-GBIF-Occurrence="1291551059" collectingDate="1994-12" collectionCode="MACN" collectorName="PN Los Glaciares &amp; Mariluis" country="Argentina" location="Rio Mitre" pageId="13" pageNumber="294" specimenCount="4" specimenCount-male="4" stateProvince="Magallanes">
<specimenCount id="9D5925F1F051FFC9FC3DFEC4FB872E61" box="[966,1065,332,357]" pageId="13" pageNumber="294" type="male">4 males</specimenCount>
,
<collectorName id="26AA8BAEF051FFC9FBC5FEC5FAB52E61" box="[1086,1307,332,357]" pageId="13" pageNumber="294">PN Los Glaciares</collectorName>
, Península
<collectingRegion id="499B209AF051FFC9FF6CFEE7FEB52E8C" box="[151,283,367,392]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="13" pageNumber="294">Magallanes</collectingRegion>
,
<location id="8E80B8A3F051FFC9FEDDFEF8FE382E8C" LSID="urn:lsid:plazi:treatment:03F65F6EF056FFC9FF6CF968FECE2B14:8E80B8A3F051FFC9FEDDFEF8FE382E8C" box="[294,406,367,392]" country="Argentina" name="Rio Mitre" pageId="13" pageNumber="294" stateProvince="Magallanes">Río Mitre</location>
, -50.418967, -72.742651,
<date id="FFE1C8B8F051FFC9FD46FEF8FC862E8C" box="[701,808,368,392]" pageId="13" pageNumber="294" value="1994-12">
<collectingDate id="EFA53150F051FFC9FD46FEF8FC862E8C" box="[701,808,368,392]" pageId="13" pageNumber="294" value="1994-12">XII-1994</collectingDate>
</date>
,
<collectorName id="26AA8BAEF051FFC9FCC8FEE7FC3C2E8C" box="[819,914,367,392]" pageId="13" pageNumber="294">Mariluis</collectorName>
leg. (
<collectionCode id="ED4E76BDF051FFC9FC2BFEF8FB852E8C" box="[976,1067,368,392]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="13" pageNumber="294">MACN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B37E425F051FFC9FBC6FEE7FF5F2EA9" ID-GBIF-Occurrence="1291551061" country="Chile" location="Bio-Bio" pageId="13" pageNumber="294" specimenCount="3" specimenCount-female="3" stateProvince="Bio-Bio">
<emphasis id="B92B326AF051FFC9FBC6FEE7FAD12E8C" bold="true" box="[1085,1407,367,392]" pageId="13" pageNumber="294">
<collectingCountry id="F348AEE8F051FFC9FBC6FEE7FB352E8C" box="[1085,1179,367,392]" name="Chile" pageId="13" pageNumber="294">CHILE</collectingCountry>
: Región de
<collectingRegion id="499B209AF051FFC9FADEFEE7FAD12E8C" box="[1317,1407,367,392]" country="Chile" name="Biobio" pageId="13" pageNumber="294">Bio-Bio</collectingRegion>
</emphasis>
:
<specimenCount id="9D5925F1F051FFC9FA75FEF8FF5F2EA9" pageId="13" pageNumber="294" type="female">3 females</specimenCount>
</materialsCitation>
, 1 male, Concepción, Parque Botánico Hualpen, -36.8300, -73.0500,
<date id="FFE1C8B8F051FFC9FC00FE1CFBFC2EA8" box="[1019,1106,404,428]" pageId="13" pageNumber="294" value="1970-02">II-1970</date>
, Cekalovic leg. (MNRJ).
</paragraph>
</subSubSection>
<subSubSection id="C345BDF3F051FFC9FF3CFE3FFC5E2EF1" pageId="13" pageNumber="294" type="distribution">
<paragraph id="8BE0EE78F051FFC9FF3CFE3FFC5E2EF1" blockId="13.[151,1437,151,1040]" pageId="13" pageNumber="294">
<emphasis id="B92B326AF051FFC9FF3CFE3FFEF72ED4" bold="true" box="[199,345,439,464]" pageId="13" pageNumber="294">Distribution</emphasis>
(
<figureCitation id="1364F2FDF051FFC9FE90FE3FFE1F2ED4" box="[363,433,439,464]" captionStart="FIGURE 9" captionStartId="18.[151,250,1937,1959]" captionTargetBox="[168,1423,829,1910]" captionTargetId="figure@18.[163,1423,829,1916]" captionTargetPageId="18" captionText="FIGURE 9. Geographical distribution. A. Helina araucana sp. nov. B. Helina australis Carvalho &amp; Pont. C. Helina bigoti Malloch. D. Helina chilensis Malloch. E. Helina connexa Malloch. F. Helina discolor (Stein). G. Helina dorada sp. nov. (Green area: Andean-Patagonian forests; orange circle: new records; blue triangle: previous distribution.)" httpUri="https://zenodo.org/record/265627/files/figure.png" pageId="13" pageNumber="294">Fig. 9</figureCitation>
D).
<collectingCountry id="F348AEE8F051FFC9FE25FE30FD282ED4" box="[478,646,440,464]" name="Argentina" pageId="13" pageNumber="294">ARGENTINA</collectingCountry>
:
<collectingRegion id="499B209AF051FFC9FD68FE3FFD482ED4" box="[659,742,439,464]" country="Argentina" name="Chubut" pageId="13" pageNumber="294">Chubut</collectingRegion>
(new record),
<collectingRegion id="499B209AF051FFC9FC69FE3FFC542ED4" box="[914,1018,439,464]" country="Argentina" name="Mendoza" pageId="13" pageNumber="294">Mendoza</collectingRegion>
(new record),
<collectingRegion id="499B209AF051FFC9FB5CFE30FAA12ED4" box="[1191,1295,440,464]" country="Argentina" name="Neuquen" pageId="13" pageNumber="294">Neuquén</collectingRegion>
,
<collectingRegion id="499B209AF051FFC9FAE6FE30FA362ED4" box="[1309,1432,440,464]" country="Argentina" name="Rio Negro" pageId="13" pageNumber="294">Río Negro</collectingRegion>
,
<collectingRegion id="499B209AF051FFC9FF6CFE54FEBD2EF0" box="[151,275,476,500]" country="Argentina" name="Santa Cruz" pageId="13" pageNumber="294">Santa Cruz</collectingRegion>
(new rercord).
<collectingCountry id="F348AEE8F051FFC9FE3AFE54FDB62EF0" box="[449,536,476,500]" name="Chile" pageId="13" pageNumber="294">CHILE</collectingCountry>
: Región de
<collectingRegion id="499B209AF051FFC9FD60FE54FD562EF1" box="[667,760,476,501]" country="Chile" name="Biobio" pageId="13" pageNumber="294">Bio-Bio</collectingRegion>
,
<collectingRegion id="499B209AF051FFC9FCF8FE54FC422EF1" box="[771,1004,476,501]" country="Chile" name="Los Lagos" pageId="13" pageNumber="294">Región de los Lagos</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C345BDF3F051FFC9FF3CFE77FCD72CA1" pageId="13" pageNumber="294" type="discussion">
<paragraph id="8BE0EE78F051FFC9FF3CFE77FCD72CA1" blockId="13.[151,1437,151,1040]" pageId="13" pageNumber="294">
<emphasis id="B92B326AF051FFC9FF3CFE77FE9A2D1C" bold="true" box="[199,308,511,536]" pageId="13" pageNumber="294">Remarks</emphasis>
.
<bibRefCitation id="EFCE9389F051FFC9FEBCFE77FE792D1C" author="Stein" box="[327,471,511,536]" pageId="13" pageNumber="294" refString="Stein, P. (1904) Die Amerikanischen Anthomyiden des Koniglichen Museums fur Naturkunde zu Berlin und des Ungarischen National-Museums zu Budapest. Annales Historico-Naturales Musei Nationalis Hungarici, 2, 414 - 495." type="journal article" year="1904">Stein (1904)</bibRefCitation>
described
<taxonomicName id="4C5F95FBF051FFC9FDA0FE77FCDE2D1C" box="[603,880,511,536]" class="Insecta" family="Muscidae" genus="Spilogaster" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="294" phylum="Arthropoda" rank="species" species="fulvisquama">
<emphasis id="B92B326AF051FFC9FDA0FE77FCDE2D1C" box="[603,880,511,536]" italics="true" pageId="13" pageNumber="294">Spilogaster fulvisquama</emphasis>
</taxonomicName>
based on a single female from an unspecified location of
<collectingCountry id="F348AEE8F051FFC9FEE6FDACFEF32D39" box="[285,349,548,573]" name="Chile" pageId="13" pageNumber="294">Chile</collectingCountry>
. Later,
<bibRefCitation id="EFCE9389F051FFC9FE4EFDACFDC92D38" author="Malloch" box="[437,615,548,573]" pageId="13" pageNumber="294" refString="Malloch, J. R. (1934) Muscidae. In: Diptera of Patagonia and South Chile. Part 7, Fascicle 2. British Museum (Natural History), London, pp. 171 - 346." type="book chapter" year="1934">Malloch (1934)</bibRefCitation>
described
<taxonomicName id="4C5F95FBF051FFC9FD1DFDADFC192D39" box="[742,951,548,573]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="294" phylum="Arthropoda" rank="species" species="fulvocalyptrata">
<emphasis id="B92B326AF051FFC9FD1DFDADFC192D39" box="[742,951,548,573]" italics="true" pageId="13" pageNumber="294">H. fulvocalyptrata</emphasis>
</taxonomicName>
with specimens from several locations of
<collectingCountry id="F348AEE8F051FFC9FF6CFDCFFEA82D64" box="[151,262,583,608]" name="Argentina" pageId="13" pageNumber="294">Argentina</collectingCountry>
and
<collectingCountry id="F348AEE8F051FFC9FEB8FDCFFE2C2D64" box="[323,386,583,608]" name="Chile" pageId="13" pageNumber="294">Chile</collectingCountry>
, and suggested that it was similar to
<emphasis id="B92B326AF051FFC9FCCCFDC0FC492D64" box="[823,999,583,608]" italics="true" pageId="13" pageNumber="294">
<taxonomicName id="4C5F95FBF051FFC9FCCCFDC0FC4D2D64" box="[823,995,583,608]" class="Insecta" family="Muscidae" genus="Spilogaster" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="294" phylum="Arthropoda" rank="species" species="fulvisquama">S. fulvisquama</taxonomicName>
.
</emphasis>
More recently,
<bibRefCitation id="EFCE9389F051FFC9FB5BFDC0FA862D64" author="Pont" box="[1184,1320,584,608]" pageId="13" pageNumber="294" refString="Pont, A. C. (1972) Family Muscidae. In: A Catalogue of the Diptera of the Americas South of the United States, Museu de Zoologia, Universidade de Sao Paulo, 97, 111 pp." type="book" year="1972">Pont (1972)</bibRefCitation>
explicitly established the synonym. Also,
<bibRefCitation id="EFCE9389F051FFC9FDF6FDE4FD6E2D80" author="Malloch" box="[525,704,620,645]" pageId="13" pageNumber="294" refString="Malloch, J. R. (1934) Muscidae. In: Diptera of Patagonia and South Chile. Part 7, Fascicle 2. British Museum (Natural History), London, pp. 171 - 346." type="book chapter" year="1934">Malloch (1934)</bibRefCitation>
described
<taxonomicName id="4C5F95FBF051FFC9FCB9FDE5FC642D81" box="[834,970,620,645]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="294" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="B92B326AF051FFC9FCB9FDE5FC642D81" box="[834,970,620,645]" italics="true" pageId="13" pageNumber="294">H. chilensis</emphasis>
</taxonomicName>
from one male specimen from Perales,
<collectingCountry id="F348AEE8F051FFC9FF6CFD07FF792DAC" box="[151,215,655,680]" name="Chile" pageId="13" pageNumber="294">Chile</collectingCountry>
, and
<taxonomicName id="4C5F95FBF051FFC9FEE8FD19FE252DAC" box="[275,395,655,680]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="294" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B92B326AF051FFC9FEE8FD19FE252DAC" box="[275,395,655,680]" italics="true" pageId="13" pageNumber="294">H. simplex</emphasis>
</taxonomicName>
from a single male specimen without locality. In our samplings, we collected the following combinations from different localities: male specimens of
<taxonomicName id="4C5F95FBF051FFC9FCD8FD3DFC5F2DC9" box="[803,1009,692,717]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="294" phylum="Arthropoda" rank="species" species="fulvocalyptrata">
<emphasis id="B92B326AF051FFC9FCD8FD3DFC5F2DC9" box="[803,1009,692,717]" italics="true" pageId="13" pageNumber="294">H. fulvocalyptrata</emphasis>
</taxonomicName>
with female specimens of
<taxonomicName id="4C5F95FBF051FFC9FAE7FD3DFA3A2DC9" box="[1308,1428,692,717]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="294" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B92B326AF051FFC9FAE7FD3DFA3A2DC9" box="[1308,1428,692,717]" italics="true" pageId="13" pageNumber="294">H. simplex</emphasis>
</taxonomicName>
; male specimens of
<taxonomicName id="4C5F95FBF051FFC9FE87FD51FDA82DF4" box="[380,518,727,752]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="294" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="B92B326AF051FFC9FE87FD51FDA82DF4" box="[380,518,727,752]" italics="true" pageId="13" pageNumber="294">H. chilensis</emphasis>
</taxonomicName>
with female specimens of
<taxonomicName id="4C5F95FBF051FFC9FCB0FD51FC692DF4" box="[843,967,727,752]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="294" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B92B326AF051FFC9FCB0FD51FC692DF4" box="[843,967,727,752]" italics="true" pageId="13" pageNumber="294">H. simplex</emphasis>
</taxonomicName>
, and male and female specimens of
<taxonomicName id="4C5F95FBF051FFC9FA7AFD51FF422C11" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="294" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B92B326AF051FFC9FA7AFD51FF422C11" italics="true" pageId="13" pageNumber="294">H. simplex</emphasis>
</taxonomicName>
. We observed that the variation in the coloration of legs and in the number of setae in the mid and hind tibia of males was not correlated with differences in
<taxonomicName id="4C5F95FBF051FFC9FD54FC97FC8F2C3C" box="[687,801,799,824]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="13" pageNumber="310" phylum="Tracheophyta" rank="genus">terminalia</taxonomicName>
structures, which are similar in all specimens. Based on these observations, we concluded that these are the same biological entity. The three names available were published by
<bibRefCitation id="EFCE9389F051FFC9FECEFCEFFE452C84" author="Malloch" box="[309,491,871,896]" pageId="13" pageNumber="294" refString="Malloch, J. R. (1934) Muscidae. In: Diptera of Patagonia and South Chile. Part 7, Fascicle 2. British Museum (Natural History), London, pp. 171 - 346." type="book chapter" year="1934">Malloch (1934)</bibRefCitation>
; we chose
<taxonomicName id="4C5F95FBF051FFC9FD8AFCE1FD572C84" box="[625,761,871,896]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="294" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="B92B326AF051FFC9FD8AFCE1FD572C84" box="[625,761,871,896]" italics="true" pageId="13" pageNumber="294">H. chilensis</emphasis>
</taxonomicName>
because it was first mentioned (
<bibRefCitation id="EFCE9389F051FFC9FB81FCEFFAF52C84" author="Malloch" box="[1146,1371,871,896]" pageId="13" pageNumber="294" refString="Malloch, J. R. (1934) Muscidae. In: Diptera of Patagonia and South Chile. Part 7, Fascicle 2. British Museum (Natural History), London, pp. 171 - 346." type="book chapter" year="1934">Malloch 1934: 176</bibRefCitation>
), and provide a redescripton with details of female and male
<taxonomicName id="4C5F95FBF051FFC9FD04FC04FCDA2CA1" box="[767,884,908,933]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="13" pageNumber="310" phylum="Tracheophyta" rank="genus">terminalia</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C345BDF3F051FFC9FF3CFC27FECE2B14" pageId="13" pageNumber="294" type="biology_ecology">
<paragraph id="8BE0EE78F051FFC9FF3CFC27FECE2B14" blockId="13.[151,1437,151,1040]" pageId="13" pageNumber="294">
<emphasis id="B92B326AF051FFC9FF3CFC27FEB12CCC" bold="true" box="[199,287,943,968]" pageId="13" pageNumber="294">Biology</emphasis>
. Specimens collected by the authors were captured with a baited trap, a Malaise trap, and a hand net over vegetation. Several of the specimens studied in this work were collected on
<taxonomicName id="4C5F95FBF051FFC9FBC6FC5DFB4E2CE8" box="[1085,1248,981,1004]" class="Magnoliopsida" family="Rhamnaceae" genus="Discaria" kingdom="Plantae" order="Rosales" pageId="13" pageNumber="294" phylum="Tracheophyta" rank="species" species="nana">
<emphasis id="B92B326AF051FFC9FBC6FC5DFB4E2CE8" box="[1085,1248,981,1004]" italics="true" pageId="13" pageNumber="294">Discaria nana</emphasis>
</taxonomicName>
with a hand net over
<quantity id="4CA7439DF051FFC9FF2BFC70FE8A2B0B" box="[208,292,1016,1040]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="294" unit="m" value="2000.0">2000 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
</treatment>
</document>
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