146 lines
18 KiB
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146 lines
18 KiB
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<document id="200F4A6CE99AAF080F2A71B55A806CF6" ID-DOI="10.5281/zenodo.10452458" ID-Zenodo-Dep="10452458" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1704223963807" checkinUser="julia" docAuthor="Löbl, Ivan" docDate="2023" docId="03CC2F59FF853F0EB0600DF3965AFA3F" docLanguage="en" docName="Vernate.42.135-142.pdf" docOrigin="Vernate 42" docTitle="Scaphisoma tienmuschanum Löbl 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="141" masterDocId="FFF55721FF803F08B00A09559475FF93" masterDocTitle="New species and records of Scaphidiinae (Coleoptera: Staphylinidae) from mainland China and Taiwan" masterLastPageNumber="142" masterPageNumber="135" pageNumber="140" updateTime="1704224970878" updateUser="ExternalLinkService">
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<mods:mods id="51CF390F1015E279D271AA830EDEE7DA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="382AA83A712FB74E9053812AF8E38A6A">
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<mods:title id="B63B555ADD2182809325E3B3CAF2E936">New species and records of Scaphidiinae (Coleoptera: Staphylinidae) from mainland China and Taiwan</mods:title>
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</mods:titleInfo>
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<mods:name id="10BB54E0D22247DC6FF1C6E321CA22D7" type="personal">
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<mods:role id="DDFB36B69B9317FE15375015D83B8C68">
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<mods:roleTerm id="98474268D5A62D3F67130C372CB308A5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B7B97FA0FDD885304CF549304524A8EB">Löbl, Ivan</mods:namePart>
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<mods:affiliation id="08D66AFBA2D6F1AAA95B463AEC0E08C9">Muséum d’histoire naturelle route de Malagnou 1 CH- 1208 Geneva, Switzerland</mods:affiliation>
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<mods:nameIdentifier id="8D798A43B8BC41B790503701379A8DF2" type="email">ivan.lobl@bluewin.ch</mods:nameIdentifier>
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<mods:typeOfResource id="95EF8E8990D22FB3358404842A2257AF">text</mods:typeOfResource>
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<mods:title id="0A4586701150EDBDF2F81B8939DEFE93">Vernate</mods:title>
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</mods:titleInfo>
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<mods:part id="2C3FE5473340DF72E05210B7AA5E752D">
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<mods:date id="17BD1DE8DEFAAF0343D1D4E3E9C7B23F">2023</mods:date>
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<mods:detail id="8820146C02936920EB164981F4F797E9" type="pubDate">
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<mods:number id="48CFD03D00069763B8CE9F4C21481AA6">2023-12-31</mods:number>
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</mods:detail>
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<mods:detail id="6F22E9F13F6A03E5202FC8DC2B66E254" type="volume">
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<mods:number id="BC1B1B04021AB7A85FD7449A020D9EDB">42</mods:number>
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</mods:detail>
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<mods:start id="C3D283594DADBAA710C60A4F92BCD9DF">135</mods:start>
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<mods:end id="AF6E1228A91153C99508CEB3C3977E41">142</mods:end>
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<mods:classification id="C6019FD72310B9A17BEB9FDAEB64C14E">journal article</mods:classification>
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<mods:identifier id="0191703BB2B962D0AB8B7CD377D16B09" type="DOI">10.5281/zenodo.10452458</mods:identifier>
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<mods:identifier id="0922D38B27562CB5251A59068C001564" type="Zenodo-Dep">10452458</mods:identifier>
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</mods:mods>
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<treatment id="03CC2F59FF853F0EB0600DF3965AFA3F" ID-DOI="http://doi.org/10.5281/zenodo.10452404" ID-Zenodo-Dep="10452404" LSID="urn:lsid:plazi:treatment:03CC2F59FF853F0EB0600DF3965AFA3F" httpUri="http://treatment.plazi.org/id/03CC2F59FF853F0EB0600DF3965AFA3F" lastPageId="6" lastPageNumber="141" pageId="5" pageNumber="140">
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<subSubSection id="C37FCDC4FF853F0DB0600DF395B3FB2F" box="[106,454,1190,1212]" pageId="5" pageNumber="140" type="nomenclature">
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<paragraph id="8BDA9E4FFF853F0DB0600DF395B3FB2F" blockId="5.[106,605,1190,1368]" box="[106,454,1190,1212]" pageId="5" pageNumber="140">
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<emphasis id="B911425DFF853F0DB0600DF395B3FB2F" bold="true" box="[106,454,1190,1212]" pageId="5" pageNumber="140">
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<taxonomicName id="4C65E5CCFF853F0DB0600DF3950DFB2F" authority="Löbl, 2023" authorityName="Löbl" authorityYear="2023" box="[106,376,1190,1212]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="140" phylum="Arthropoda" rank="species" species="tienmuschanum" status="sp. nov.">
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<emphasis id="B911425DFF853F0DB0600DF3950DFB2F" bold="true" box="[106,376,1190,1212]" italics="true" pageId="5" pageNumber="140">Scaphisoma tienmuschanum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A222FF26FF853F0DB1740DF295B3FB2F" box="[382,454,1191,1212]" pageId="5" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37FCDC4FF853F0DB0600D939510FACB" pageId="5" pageNumber="140" type="materials_examined">
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<paragraph id="8BDA9E4FFF853F0DB0600D939510FACB" blockId="5.[106,605,1190,1368]" pageId="5" pageNumber="140">
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<materialsCitation id="3B0D9412FF853F0DB0600D939510FACB" collectingDate="2017-06-24" collectionCode="NMPC" collectorName="J. Hajek & J. Ruzicka" country="China" elevation="430" latitude="30.3255" location="West Tianmushan Nat. Res." longLatPrecision="6" longitude="119.4406" municipality="Lin'an County" pageId="5" pageNumber="140" specimenCode="WTO3" specimenCount="1" specimenCount-male="1" stateProvince="ZHEJIANG PROV." typeStatus="holotype">
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<emphasis id="B911425DFF853F0DB0600D9394B7FB48" bold="true" box="[106,194,1222,1243]" pageId="5" pageNumber="140">
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<typeStatus id="54DE20EDFF853F0DB0600D9394B7FB48" box="[106,194,1222,1243]" pageId="5" pageNumber="140">Holotype</typeStatus>
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</emphasis>
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<specimenCount id="9D6355C6FF853F0DB0C60D939489FB4F" box="[204,252,1222,1244]" pageId="5" pageNumber="140" type="male">male</specimenCount>
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:
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<collectingCountry id="F372DEDFFF853F0DB1020D939525FB48" box="[264,336,1222,1243]" name="China" pageId="5" pageNumber="140">CHINA</collectingCountry>
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:
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<collectingRegion id="49A150ADFF853F0DB16A0D939662FB48" box="[352,535,1222,1243]" country="China" name="Zhejiang" pageId="5" pageNumber="140">ZHEJIANG PROV.</collectingRegion>
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<collectingMunicipality id="6BBE0435FF853F0DB22A0D9394DBFB69" pageId="5" pageNumber="140">Lin’an County</collectingMunicipality>
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,
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<location id="8EBAC894FF853F0DB0C90DB0959DFB69" LSID="urn:lsid:plazi:treatment:03CC2F59FF853F0EB0600DF3965AFA3F:8EBAC894FF853F0DB0C90DB0959DFB69" box="[195,488,1253,1275]" country="China" latitude="30.3255" longLatPrecision="6" longitude="119.4406" municipality="Lin'an County" name="West Tianmushan Nat. Res." pageId="5" pageNumber="140" stateProvince="ZHEJIANG PROV.">West Tianmushan Nat. Res.</location>
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<geoCoordinate id="EE51F888FF853F0DB1F00DB09628FB68" box="[506,605,1253,1275]" degrees="30.3255" direction="north" orientation="latitude" pageId="5" pageNumber="140" precision="5" value="30.3255">30.3255°N</geoCoordinate>
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<geoCoordinate id="EE51F888FF853F0DB0600C5194A2FA89" box="[106,215,1284,1306]" degrees="119.4406" direction="east" orientation="longitude" pageId="5" pageNumber="140" precision="5" value="119.4406">119.4406°E</geoCoordinate>
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,
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<collectorName id="2690FB99FF853F0DB0EC0C519542FA89" box="[230,311,1284,1306]" pageId="5" pageNumber="140">J. Hájek</collectorName>
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&
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<collectorName id="2690FB99FF853F0DB16A0C5195B1FA89" box="[352,452,1284,1306]" pageId="5" pageNumber="140">J. Růžička</collectorName>
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leg. / (
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<specimenCode id="DBC33634FF853F0DB21D0C519623FA89" box="[535,598,1284,1306]" pageId="5" pageNumber="140">WTO3</specimenCode>
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)
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<date id="FFDBB88FFF853F0DB0600C7694BEFAAA" box="[106,203,1315,1337]" pageId="5" pageNumber="140" value="2017-06-24">
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<collectingDate id="EF9F4167FF853F0DB0600C7694BEFAAA" box="[106,203,1315,1337]" pageId="5" pageNumber="140" value="2017-06-24">24.vi.2017</collectingDate>
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</date>
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<quantity id="4C9D33AAFF853F0DB0DB0C769578FAAA" box="[209,269,1315,1337]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="5" pageNumber="140" unit="m" value="430.0">
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<elevation id="0048797CFF853F0DB0DB0C769578FAAA" box="[209,269,1315,1337]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="5" pageNumber="140" unit="m" value="430.0">430 m</elevation>
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</quantity>
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, sift #02
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<location id="8EBAC894FF853F0DB16B0C76957CFACB" LSID="urn:lsid:plazi:treatment:03CC2F59FF853F0EB0600DF3965AFA3F:8EBAC894FF853F0DB16B0C76957CFACB" country="China" latitude="30.3255" longLatPrecision="6" longitude="119.4406" municipality="Lin'an County" name="large bamboo bushes above Chanyuan temple" pageId="5" pageNumber="140" stateProvince="ZHEJIANG PROV.">large bamboo bushes above Chanyuan temple</location>
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(
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<collectionCode id="ED74068AFF853F0DB11F0C17952EFACB" box="[277,347,1346,1368]" collectionName="Czech Republic, Prague, National Museum (Natural History)" pageId="5" pageNumber="140">NMPC</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37FCDC4FF853F0DB0600C2C97B2F99F" pageId="5" pageNumber="140" type="description">
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<paragraph id="8BDA9E4FFF853F0DB0600C2C9034FA1C" blockId="5.[106,606,1401,1641]" lastBlockId="5.[665,1165,158,1548]" pageId="5" pageNumber="140">
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<emphasis id="B911425DFF853F0DB0600C2C94AFFA1C" bold="true" box="[106,218,1401,1423]" pageId="5" pageNumber="140">Description</emphasis>
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. Length 1.35 mm, width 0.90 mm. Head and most of body dark reddish-brown, apex of abdomen light brown to yellowish, legs rufous, antennomeres I to IV yellowish, following antennomeres light brown. Length/width ratios of antennomeres: III 13/7: IV 24/5: V 29/6: VI 33/7: VII 39/12: VIII 33/10: IX 42/11: X 43/12: XI 43/12. Pronotum and elytra lacking microsculpture. Pronotum with lateral margins slightly arcuate; lateral margin carina concealed in dorsal view, impunctate; discal punctation sparse and very fine, visible at 50 times magnification. Minute tip of scutellum exposed. Elytron strongly narrowing apicad, with lateral margin arcuate, lateral margin stria concealed in dorsal view, impunctate; apical margin weakly rounded, lacking distinct crenulation; inner apical angle situated in level of outer angle; sutural margin not raised; sutural carina moderately deep, parallel, starting at each side of pronotal lobe, not extending along base; adsutural area flat, ca. 0.05 mm wide shortly posteriad of scutellar tip, with single, dense puncture row; punctation near bases rather fine, much coarser than pronotal punctation, becoming denser and coarser apicad, with puncture intervals mostly about two to three times as large as puncture diameters on apical half of disc. Hind wings fully developed. Hypomeron smooth. Mesanepisternum very finely and sparsely punctate. Mesoventrite with few fine punctures in middle, densely and rather coarsely punctate along lateral margins. Mesepimeron about 3 times as long as wide and as long as interval between its tip and mesocoxa. Metaventrite lacking microsculpture, nearly flat in middle, lacking impressions, with punctation rather coarse and conspicuously dense on apicomedian area, distinct at 40 times magnification, becoming sparse and very fine anteriad and laterad; antecoxal puncture rows absent; submesocoxal areas ca. 0.04 mm long, nearly as long as third of shortest interval between its margin and metacoxa; submesocoxal lines convex, finely punctate. Metanepisternum lacking microsculpture, convex, narrowing anteriad, suture not impressed, oblique, rounded at angles. Tibiae straight. Exposed tergites and ventrites with strigulate microsculpture, evanescent on lateral areas of ventrite I. Pygidium notched, with apical margin broadly concave. Ventrite I rather densely and finely punctate on basomedian area, very finely and sparsely punctate on prevailing surface; submetacoxal areas 0.04 mm long, nearly as long as half of shortest interval between its margin and apical margin of ventrite; submetacoxal lines convex, rather finely punctate.
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</paragraph>
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<paragraph id="8BDA9E4FFF853F0DB2930CCC97B2F99F" blockId="5.[665,1165,158,1548]" pageId="5" pageNumber="140">
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<emphasis id="B911425DFF853F0DB2930CCC96BDFA3D" box="[665,712,1433,1454]" italics="true" pageId="5" pageNumber="140">Male</emphasis>
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. Protarsomeres I to III slightly widened, with short tenent setae, protarsomere I narrower than apex of protibia. Mesotarsomeres I to III not widened. Aedeagus (
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<figureCitation id="135E82CAFF853F0DB2EE0CA39741F99F" box="[740,820,1526,1548]" captionStart="Figs 7, 8" captionStartId="6.[121,152,968,985]" captionTargetBox="[228,1095,144,949]" captionTargetId="figure-316@6.[168,1131,125,962]" captionTargetPageId="6" captionText="Figs 7, 8. Scaphisoma tienmuschanum sp. nov., aedeagus in dorsal and lateral views, scale = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.10452462" httpUri="https://zenodo.org/record/10452462/files/figure.png" pageId="5" pageNumber="140">Figs 7, 8</figureCitation>
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) 0.30 mm long.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37FCDC4FF853F0DB2930F66905BF9FB" pageId="5" pageNumber="140" type="etymology">
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<paragraph id="8BDA9E4FFF853F0DB2930F66905BF9FB" blockId="5.[665,1164,1587,1640]" pageId="5" pageNumber="140">
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<emphasis id="B911425DFF853F0DB2930F669777F9DB" bold="true" box="[665,770,1587,1608]" pageId="5" pageNumber="140">Etymology</emphasis>
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. The species epithet is the Latinized name of the natural park in which it was collected.
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</paragraph>
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</subSubSection>
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<caption id="DF1ACEC7FF863F0EB0730A9D9734FC4B" ID-DOI="http://doi.org/10.5281/zenodo.10452462" ID-Zenodo-Dep="10452462" box="[121,833,968,986]" httpUri="https://zenodo.org/record/10452462/files/figure.png" pageId="6" pageNumber="141" startId="6.[121,152,968,985]" targetBox="[228,1095,144,949]" targetPageId="6" targetType="figure">
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<paragraph id="8BDA9E4FFF863F0EB0730A9D9734FC4B" blockId="6.[121,833,968,986]" box="[121,833,968,986]" pageId="6" pageNumber="141">
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<emphasis id="B911425DFF863F0EB0730A9D94CEFC4A" bold="true" box="[121,187,968,985]" pageId="6" pageNumber="141">Figs 7, 8.</emphasis>
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<taxonomicName id="4C65E5CCFF863F0EB0B50A9D95F1FC4A" authority="Löbl, 2023" authorityName="Löbl" authorityYear="2023" box="[191,388,968,985]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="tienmuschanum" status="sp. nov.">
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<emphasis id="B911425DFF863F0EB0B50A9D95F1FC4A" box="[191,388,968,985]" italics="true" pageId="6" pageNumber="141">Scaphisoma tienmuschanum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A222FF26FF863F0EB1820A9D95B4FC4B" box="[392,449,968,984]" pageId="6" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
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, aedeagus in dorsal and lateral views, scale = 0.05 mm.
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</paragraph>
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</caption>
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<subSubSection id="C37FCDC4FF863F0EB0730D74965AFA3F" pageId="6" pageNumber="141" type="discussion">
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<paragraph id="8BDA9E4FFF863F0EB0730D74965AFA3F" blockId="6.[121,620,1057,1452]" pageId="6" pageNumber="141">
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<emphasis id="B911425DFF863F0EB0730D749494FBA4" bold="true" box="[121,225,1057,1079]" pageId="6" pageNumber="141">Comments</emphasis>
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. This new species would fall in the key to Chinese
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<taxonomicName id="4C65E5CCFF863F0EB0C10D15954DFBC6" baseAuthorityName="Lobl" baseAuthorityYear="2019" box="[203,312,1088,1109]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="genus">
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<emphasis id="B911425DFF863F0EB0C10D15954DFBC6" box="[203,312,1088,1109]" italics="true" pageId="6" pageNumber="141">Scaphisoma</emphasis>
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</taxonomicName>
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under couplet 64, to
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<taxonomicName id="4C65E5CCFF863F0EB2010D1494AAFBE6" authority="Lobl, 2000" authorityName="Lobl" authorityYear="2000" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="incisum">
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<emphasis id="B911425DFF863F0EB2010D149619FBC5" box="[523,620,1089,1110]" italics="true" pageId="6" pageNumber="141">S. incisum</emphasis>
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Löbl, 2000
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</taxonomicName>
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. The aedeagal characters suggest a relationship with
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<taxonomicName id="4C65E5CCFF863F0EB0D00D2A95B5FB07" authority="Lobl, 1980" authorityName="Lobl" authorityYear="1980" box="[218,448,1150,1172]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="taiwanum">
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<emphasis id="B911425DFF863F0EB0D00D2A9538FB07" box="[218,333,1151,1172]" italics="true" pageId="6" pageNumber="141">S. taiwanum</emphasis>
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<bibRefCitation id="EFF4E3BEFF863F0EB15D0D2B95B5FB07" author="LOBL, I." box="[343,448,1150,1172]" journalOrPublisher="Revue suisse de zoologie" pageId="6" pageNumber="141" pagination="91 - 123" part="87" refId="ref5386" refString="LOBL, I. (1980): Beitrag zur Kenntnis der Scaphidiidae (Coleoptera) Tai- wans. - Revue suisse de zoologie 87: 91 - 123." title="Beitrag zur Kenntnis der Scaphidiidae (Coleoptera) Tai- wans" type="journal article" year="1980">Löbl, 1980</bibRefCitation>
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</taxonomicName>
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.
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<taxonomicName id="4C65E5CCFF863F0EB1C70D2B9491FB20" authorityName="Löbl" authorityYear="2023" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="tienmuschanum">
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<emphasis id="B911425DFF863F0EB1C70D2B9491FB20" italics="true" pageId="6" pageNumber="141">Scaphisoma tienmuschanum</emphasis>
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</taxonomicName>
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may be easily distinguished from the latter species, as from other Chinese congeners, by the shape of the median lobe and the parameres. It has the apical process of the median lobe much more inflexed, nearly perpendicular to the axis of the basal bulb, and the parameres weakly narrowing apicad (in lateral view) and not bent mesally (in dorsal view). Besides, the body is smaller and the spines of the internal sac are notably longer in
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<taxonomicName id="4C65E5CCFF863F0EB0C50CC29507FA38" authorityName="Löbl" authorityYear="2023" box="[207,370,1430,1452]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="tienmuschanum">
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<emphasis id="B911425DFF863F0EB0C50CC29507FA38" box="[207,370,1430,1452]" italics="true" pageId="6" pageNumber="141">S. tienmuschanum</emphasis>
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</taxonomicName>
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than in
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<taxonomicName id="4C65E5CCFF863F0EB1B00CC2965FFA3F" authorityName="Lobl" authorityYear="1980" box="[442,554,1431,1452]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="taiwanum">
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<emphasis id="B911425DFF863F0EB1B00CC2965FFA3F" box="[442,554,1431,1452]" italics="true" pageId="6" pageNumber="141">S. taiwanum</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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