150 lines
17 KiB
XML
150 lines
17 KiB
XML
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<document id="22FFCFB71FDCD0798ED754A6055CD00D" ID-DOI="10.11646/zootaxa.3700.3.7" ID-GBIF-Dataset="371f30c1-ff9d-4fde-9429-fc9d284caf03" ID-ISSN="1175-5326" ID-Zenodo-Dep="249376" ID-ZooBank="E3FA2E4B-DDF4-4FDB-8CF1-FFBA215D40CA" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460663374311" checkinUser="plazi" docAuthor="Massa, Bruno" docDate="2013" docId="03C187FAFFA0B346FF64D17B5ED5F9F0" docLanguage="en" docName="zt03700p475.pdf" docOrigin="Zootaxa 3700 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tmethis Fieber 1853" docType="treatment" docVersion="13" lastPageNumber="437" masterDocId="FFF8FF82FFA2B344FFF3D33D5D23FFDA" masterDocTitle="Pamphagidae (Orthoptera: Caelifera) of North Africa: key to genera and the annotated check-list of species" masterLastPageNumber="475" masterPageNumber="435" pageNumber="437" updateTime="1698347717602" updateUser="plazi">
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<mods:titleInfo id="70B558E5F22E933F597ED037ECDE68A4">
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<mods:title id="26044A3D5744029898442BB051CD3028">Pamphagidae (Orthoptera: Caelifera) of North Africa: key to genera and the annotated check-list of species</mods:title>
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<mods:name id="67A52B055D27AE9F0FA8BE17F62F37F0" type="personal">
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<mods:namePart id="D26ADE07408B0724E53AE1CAE5B91225">Massa, Bruno</mods:namePart>
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<mods:typeOfResource id="13FBBB75760D800A68021B53D982AED8">text</mods:typeOfResource>
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<mods:title id="BEC86FEC5CA8471203807476394DE1D0">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="BB2FEEC63B06B4465A432F9465C96580">
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<mods:date id="89646683084A3E745C01359F21DDF7C1">2013</mods:date>
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<mods:number id="9303E37E467475F8B33FA3E73AF1F8A9">3700</mods:number>
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</mods:detail>
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<mods:number id="4FF1809509EEF09F5EDF8A8745CC3FF2">3</mods:number>
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<mods:start id="2181E1F670094467E2BB47471C3D48D5">435</mods:start>
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<mods:end id="F7E15A16A025B3B49248E6FA1F86C61B">475</mods:end>
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<mods:classification id="F748064A0C0BC0D40402F0C6CE9AC29F">journal article</mods:classification>
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<mods:identifier id="D2D17A03760250D89D1887B1E1DC049C" type="DOI">10.11646/zootaxa.3700.3.7</mods:identifier>
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<mods:identifier id="320784420A97142565D4DB095905174E" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="30778F0BBC9B403AC6D550A1C026BCC0" type="Zenodo-Dep">249376</mods:identifier>
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</mods:mods>
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<treatment id="03C187FAFFA0B346FF64D17B5ED5F9F0" ID-DOI="http://doi.org/10.5281/zenodo.5697446" ID-GBIF-Taxon="119574872" ID-Zenodo-Dep="5697446" LSID="urn:lsid:plazi:treatment:03C187FAFFA0B346FF64D17B5ED5F9F0" httpUri="http://treatment.plazi.org/id/03C187FAFFA0B346FF64D17B5ED5F9F0" lastPageNumber="437" pageId="2" pageNumber="437">
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<subSubSection id="C3726567FFA0B346FF64D17B5ED5F9F0" pageId="2" pageNumber="437" type="nomenclature">
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<paragraph id="8BD736ECFFA0B346FF64D17B5E4AFDBA" blockId="2.[151,873,582,608]" box="[151,873,582,608]" pageId="2" pageNumber="437">
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<heading id="D09F8180FFA0B346FF64D17B5E4AFDBA" bold="true" box="[151,873,582,608]" fontSize="11" level="1" pageId="2" pageNumber="437" reason="1">
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<emphasis id="B91CEAFEFFA0B346FF64D17B5E4AFDBA" bold="true" box="[151,873,582,608]" pageId="2" pageNumber="437">
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<taxonomicName id="4C684D6FFFA0B346FF64D17B5CBBFDBA" ID-CoL="8P282" authority="Fieber, 1853" authorityName="Fieber" authorityYear="1853" box="[151,408,582,608]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="437" phylum="Arthropoda" rank="genus">
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<emphasis id="B91CEAFEFFA0B346FF64D17B5DDAFDBA" bold="true" box="[151,249,582,608]" italics="true" pageId="2" pageNumber="437">Tmethis</emphasis>
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Fieber, 1853
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</taxonomicName>
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(type-species:
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<emphasis id="B91CEAFEFFA0B346FDA3D17B5FC2FDBA" bold="true" box="[592,737,582,608]" italics="true" pageId="2" pageNumber="437">
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Gryllus
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<taxonomicName id="4C684D6FFFA0B346FD43D17B5FC2FDBA" authority="Fabricius" authorityName="Fabricius" box="[688,737,582,608]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="437" phylum="Arthropoda" rank="species" species="cisti">cisti</taxonomicName>
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</emphasis>
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Fabricius)
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD736ECFFA0B346FF64D1B05ED5F9F0" blockId="2.[151,1437,653,1578]" pageId="2" pageNumber="437">
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Fieber (1853) did not explain the origin of the name
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<taxonomicName id="4C684D6FFFA0B346FD0AD1B05E77FD7C" box="[761,852,653,678]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="437" phylum="Arthropoda" rank="genus">
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<emphasis id="B91CEAFEFFA0B346FD0AD1B05E77FD7C" box="[761,852,653,678]" italics="true" pageId="2" pageNumber="437">Tmethis</emphasis>
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</taxonomicName>
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; thank to my Greek colleague Haralabos Tsolakis, it was possible to trace a link with the verb "temno" (τέΜνω), which means "to cut", possibly referred to the particular shape of pronotum, with a distinct sulcus between prozona and metazona. I consider that
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<taxonomicName id="4C684D6FFFA0B346FAFBD1E85840FD34" box="[1288,1379,725,750]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="437" phylum="Arthropoda" rank="genus">
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<emphasis id="B91CEAFEFFA0B346FAFBD1E85840FD34" box="[1288,1379,725,750]" italics="true" pageId="2" pageNumber="437">Tmethis</emphasis>
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</taxonomicName>
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is of masculine gender, and consequently I will agree taxa with it. In his excellent revision of all Palaearctic
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<taxonomicName id="4C684D6FFFA0B346FF64D0205C07FCEC" box="[151,292,797,822]" class="Insecta" family="Pamphagidae" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="437" phylum="Arthropoda" rank="subFamily" subFamily="Thrinchinae">Thrinchinae</taxonomicName>
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, Uvarov (1943) pointed out the high variability of many species, mainly belonging to genera
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<taxonomicName id="4C684D6FFFA0B346FF64D07C5DD2FC80" box="[151,241,833,858]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="437" phylum="Arthropoda" rank="genus">
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<emphasis id="B91CEAFEFFA0B346FF64D07C5DD2FC80" box="[151,241,833,858]" italics="true" pageId="2" pageNumber="437">Tmethis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C684D6FFFA0B346FEDFD07C5F31FC83" authority="Uvarov" authorityName="Uvarov" box="[300,530,833,858]" class="Insecta" family="Pamphagidae" genus="Asiotmethis" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="437" phylum="Arthropoda" rank="genus">
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<emphasis id="B91CEAFEFFA0B346FEDFD07C5C91FC80" box="[300,434,833,858]" italics="true" pageId="2" pageNumber="437">Asiotmethis</emphasis>
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Uvarov
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</taxonomicName>
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, that may have wrinkled or smooth integuments; sculpturing of exposed parts (mainly pronotum) may be devoid of the usual tubercles and keel. Thus, taxa belonging to the genus
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<taxonomicName id="4C684D6FFFA0B346FAE4D0585851FCA4" box="[1303,1394,869,894]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="437" phylum="Arthropoda" rank="genus">
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<emphasis id="B91CEAFEFFA0B346FAE4D0585851FCA4" box="[1303,1394,869,894]" italics="true" pageId="2" pageNumber="437">Tmethis</emphasis>
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</taxonomicName>
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are much variable, not only in their general colour and in the length of wings, but also in their integuments and body structure, which may be more or less stout or slender. Some specimens appear squat and broad (
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<figureCitation id="13532A69FFA0B346FB10D090584CFC1C" box="[1251,1391,941,966]" captionStart="FIGURES 60 – 75. 60 – 71" captionStartId="31.[151,264,1945,1967]" captionTargetBox="[285,1297,196,1921]" captionTargetId="figure@31.[285,1297,193,1921]" captionTargetPageId="31" captionText="FIGURES 60 – 75. 60 – 71. Tegmina of Tuarega insignis from different North African localities to show the variability of vein branches, including topotype locality of T. ouarzazatensis (63, 65) and types of T. parisi (68 – 69) and T. sahara (70 – 71). 72. Lateral view of T. insignis. 73 – 75. Tmethis maroccanus and T. cisti from above to show variability of body stoutness." httpUri="https://zenodo.org/record/249382/files/figure.png" pageId="2" pageNumber="437">Figs. 73, 75</figureCitation>
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). In addition, the colour of hind tibiae varies from red to yellow and to purple or bluish-violet; it seems that this variation characterizes single populations. The arrangement of species within the genus was much unclear, in particular the separation of
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<taxonomicName id="4C684D6FFFA0B346FE16D7275F15FBEB" box="[485,566,1050,1073]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="437" phylum="Arthropoda" rank="species" species="cisti">
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<emphasis id="B91CEAFEFFA0B346FE16D7275F15FBEB" box="[485,566,1050,1073]" italics="true" pageId="2" pageNumber="437">T. cisti</emphasis>
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</taxonomicName>
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from
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<taxonomicName id="4C684D6FFFA0B346FD75D7275E66FBE8" box="[646,837,1049,1074]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="437" phylum="Arthropoda" rank="species" species="pulchripennis">
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<emphasis id="B91CEAFEFFA0B346FD75D7275E66FBE8" box="[646,837,1049,1074]" italics="true" pageId="2" pageNumber="437">T. pulchripennis</emphasis>
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</taxonomicName>
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. Reading Chopard (1943) it is evident that many populations of
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<taxonomicName id="4C684D6FFFA0B346FEB5D7035F22FB8C" box="[326,513,1085,1110]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="437" phylum="Arthropoda" rank="species" species="pulchripennis">
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<emphasis id="B91CEAFEFFA0B346FEB5D7035F22FB8C" box="[326,513,1085,1110]" italics="true" pageId="2" pageNumber="437">T. pulchripennis</emphasis>
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</taxonomicName>
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were actually
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<taxonomicName id="4C684D6FFFA0B346FD5CD7035FDEFB8F" box="[687,765,1086,1109]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="437" phylum="Arthropoda" rank="species" species="cisti">
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<emphasis id="B91CEAFEFFA0B346FD5CD7035FDEFB8F" box="[687,765,1086,1109]" italics="true" pageId="2" pageNumber="437">T. cisti</emphasis>
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</taxonomicName>
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(e.g., those living on
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<collectingCountry id="F37F767CFFA0B346FBF2D700596CFB8C" box="[1025,1103,1085,1110]" name="Tunisia" pageId="2" pageNumber="437">Djerba</collectingCountry>
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and Kerkenna Is.). The first author who defined characters to identify species was Uvarov (1943); he recognized
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<taxonomicName id="4C684D6FFFA0B346FB90D75F5829FBA3" box="[1123,1290,1122,1145]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="437" phylum="Arthropoda" rank="species" species="maroccanus">
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<emphasis id="B91CEAFEFFA0B346FB90D75F5829FBA3" box="[1123,1290,1122,1145]" italics="true" pageId="2" pageNumber="437">T. maroccanus</emphasis>
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</taxonomicName>
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from
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<taxonomicName id="4C684D6FFFA0B346FAA3D75F58BFFBA3" box="[1360,1436,1122,1145]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="437" phylum="Arthropoda" rank="species" species="cisti">
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<emphasis id="B91CEAFEFFA0B346FAA3D75F58BFFBA3" box="[1360,1436,1122,1145]" italics="true" pageId="2" pageNumber="437">T. cisti</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C684D6FFFA0B346FF34D7BB5CA2FB44" box="[199,385,1157,1182]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="437" phylum="Arthropoda" rank="species" species="pulchripennis">
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<emphasis id="B91CEAFEFFA0B346FF34D7BB5CA2FB44" box="[199,385,1157,1182]" italics="true" pageId="2" pageNumber="437">T. pulchripennis</emphasis>
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</taxonomicName>
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by the shape of the metazona of pronotum, which is broadly parabolic with convex margins in the former and triangular, with the sides straight or somewhat incurved in the other two species. Additionally, he identified
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<taxonomicName id="4C684D6FFFA0B346FEC3D7F35C5EFB3F" box="[304,381,1230,1253]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="437" phylum="Arthropoda" rank="species" species="cisti">
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<emphasis id="B91CEAFEFFA0B346FEC3D7F35C5EFB3F" box="[304,381,1230,1253]" italics="true" pageId="2" pageNumber="437">T. cisti</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C684D6FFFA0B346FE45D7F35F53FB3C" box="[438,624,1229,1254]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="437" phylum="Arthropoda" rank="species" species="pulchripennis">
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<emphasis id="B91CEAFEFFA0B346FE45D7F35F53FB3C" box="[438,624,1229,1254]" italics="true" pageId="2" pageNumber="437">T. pulchripennis</emphasis>
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</taxonomicName>
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by the shape of their male epiphallus, and by small differences on the lower valve of the ovipositor and length of spurs of hind tibiae. Thus, records of species in North African countries may be considered reliable only if criteria used by Uvarov (1943) were followed. However, La Greca (1969) considered much variable the shape of lower valve of the ovipositor (according to Uvarov (1943) in
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<taxonomicName id="4C684D6FFFA0B346FA74D6075C16FAAC" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="437" phylum="Arthropoda" rank="species" species="pulchripennis">
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<emphasis id="B91CEAFEFFA0B346FA74D6075C16FAAC" italics="true" pageId="2" pageNumber="437">T. pulchripennis</emphasis>
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</taxonomicName>
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with a distinct shoulder, which appears as a tooth in lateral view, while in
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<taxonomicName id="4C684D6FFFA0B346FB3FD663583EFAAF" box="[1228,1309,1374,1397]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="437" phylum="Arthropoda" rank="species" species="cisti">
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<emphasis id="B91CEAFEFFA0B346FB3FD663583EFAAF" box="[1228,1309,1374,1397]" italics="true" pageId="2" pageNumber="437">T. cisti</emphasis>
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</taxonomicName>
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it appears without a projection, not dentate), and identified some
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<taxonomicName id="4C684D6FFFA0B346FCCDD6BF5EDCFA40" box="[830,1023,1409,1434]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="437" phylum="Arthropoda" rank="species" species="pulchripennis">
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<emphasis id="B91CEAFEFFA0B346FCCDD6BF5EDCFA40" box="[830,1023,1409,1434]" italics="true" pageId="2" pageNumber="437">T. pulchripennis</emphasis>
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</taxonomicName>
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from
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<collectingCountry id="F37F767CFFA0B346FBA3D6BC59B5FA40" box="[1104,1174,1409,1434]" name="Libya" pageId="2" pageNumber="437">Libya</collectingCountry>
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, following criteria of Chopard (1943), based on the pronotum shape, that during this study resulted to be a very variable character. Tegmina length is also a variable character, as supposed by Jannone (1938). Eight subspecies or varieties have been described for
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<taxonomicName id="4C684D6FFFA0B346FE87D6D35CE1F9DF" box="[372,450,1518,1541]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="437" phylum="Arthropoda" rank="species" species="cisti">
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<emphasis id="B91CEAFEFFA0B346FE87D6D35CE1F9DF" box="[372,450,1518,1541]" italics="true" pageId="2" pageNumber="437">T. cisti</emphasis>
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</taxonomicName>
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, three for
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<taxonomicName id="4C684D6FFFA0B346FDCFD6D35FD4F9DC" box="[572,759,1517,1542]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="437" phylum="Arthropoda" rank="species" species="pulchripennis">
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<emphasis id="B91CEAFEFFA0B346FDCFD6D35FD4F9DC" box="[572,759,1517,1542]" italics="true" pageId="2" pageNumber="437">T. pulchripennis</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
and three for
|
|||
|
<taxonomicName id="4C684D6FFFA0B346FC6FD6D35967F9DF" box="[924,1092,1518,1541]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="437" phylum="Arthropoda" rank="species" species="maroccanus">
|
|||
|
<emphasis id="B91CEAFEFFA0B346FC6FD6D35967F9DF" box="[924,1092,1518,1541]" italics="true" pageId="2" pageNumber="437">T. maroccanus</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
, in some cases overlapping. I will examine each of them to establish if their taxonomic status is justified.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
</treatment>
|
|||
|
</document>
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