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<document id="1D03F47DFC3225B7452120451FE5129B" ID-CLB-Dataset="31902" ID-DOI="10.11646/zootaxa.4329.2.1" ID-GBIF-Dataset="7e33ec7d-211b-4042-8d07-344364ee805d" ID-ISSN="1175-5326" ID-Zenodo-Dep="1001915" ID-ZooBank="8226E27D-E3A9-481A-B51B-558643Bbb03A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507106636447" checkinUser="plazi" docAuthor="Tarasov, Sergei" docDate="2017" docId="03C187E1AE7AB618DD9E4250FEAD0080" docLanguage="en" docName="zootaxa.4329.2.1.pdf" docOrigin="Zootaxa 4329 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parachorius solodovnikovi Tarasov 2017, new species" docType="treatment" docUuid="4E27796A-EA33-425A-B0FE-F9D100FD799D" docUuidSource="ZooBank" docVersion="12" lastPageNumber="141" masterDocId="FFF8FF99AE5DB630DD09474FFFDB0270" masterDocTitle="A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data" masterLastPageNumber="149" masterPageNumber="101" pageNumber="140" updateTime="1698469310636" updateUser="ExternalLinkService">
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<mods:title id="E1C9F4B5B20BBCBBD32402128D63E0EB">A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data</mods:title>
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<mods:namePart id="AB2CC486AE13595E3712BD7EDAAFE9CB">Tarasov, Sergei</mods:namePart>
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<mods:title id="4A058302E3B4824FD33B920221F9B9AE">Zootaxa</mods:title>
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<mods:date id="3C30104CCE8E22AEFAB085AD24A39839">2017</mods:date>
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<mods:number id="F4F5D3A5E5F575610663D39F6503DFBC">2017-10-04</mods:number>
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<mods:identifier id="1B895452F46359EA4E4FF2B529D4ADCF" type="DOI">10.11646/zootaxa.4329.2.1</mods:identifier>
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<treatment id="03C187E1AE7AB618DD9E4250FEAD0080" ID-DOI="http://doi.org/10.5281/zenodo.6010844" ID-GBIF-Taxon="135106126" ID-Zenodo-Dep="6010844" LSID="urn:lsid:zoobank.org:act:4E27796A-EA33-425A-B0FE-F9D100FD799D" httpUri="http://treatment.plazi.org/id/03C187E1AE7AB618DD9E4250FEAD0080" lastPageId="40" lastPageNumber="141" pageId="39" pageNumber="140">
<subSubSection id="C372657CAE7AB617DD9E4250FEE4072A" pageId="39" pageNumber="140" type="nomenclature">
<paragraph id="8BD736F7AE7AB617DD9E4250FD070749" blockId="39.[151,972,1311,1406]" box="[151,732,1311,1337]" pageId="39" pageNumber="140">
<heading id="D09F819BAE7AB617DD9E4250FD070749" bold="true" box="[151,732,1311,1337]" fontSize="11" level="1" pageId="39" pageNumber="140" reason="1">
<emphasis id="B91CEAE5AE7AB617DD9E4250FD070749" bold="true" box="[151,732,1311,1337]" pageId="39" pageNumber="140">
<taxonomicName id="4C684D74AE7AB617DD9E4250FD9E0749" ID-CoL="4CPV3" authority="Tarasov" authorityName="Tarasov" authorityYear="2017" box="[151,581,1311,1337]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="140" phylum="Arthropoda" rank="species" species="solodovnikovi" status="sp. nov.">
<emphasis id="B91CEAE5AE7AB617DD9E4250FE010749" bold="true" box="[151,474,1311,1337]" italics="true" pageId="39" pageNumber="140">Parachorius solodovnikovi</emphasis>
Tarasov
</taxonomicName>
<taxonomicNameLabel id="A22F579EAE7AB617DF444250FD070749" box="[589,732,1311,1337]" pageId="39" pageNumber="140" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD736F7AE7AB617DD9E420DFEE4072A" blockId="39.[151,972,1311,1406]" box="[151,319,1346,1371]" pageId="39" pageNumber="140">
(
<figureCitation id="13532A72AE7AB617DD96420DFF27072A" box="[159,252,1346,1371]" captionStart="FIGURE 23" captionStartId="40.[151,250,1887,1909]" captionTargetBox="[151,1423,840,1815]" captionTargetId="figure@40.[151,1436,840,1830]" captionTargetPageId="40" captionText="FIGURE 23. Morphology of Parachorius solodovnikovi. (A) aedeagus, lateral view (holotype). (B) aedeagus, lateral view (paratype, Thailand). (C) aedeagus, lateral view (paratype, Vietnam). (D) aedeagus, lateral view (E) habitus (male, holotype). (F) left metafemur (male, paratype, Vietnam) (G) left metafemur (male, holotype). (H) left protibia (male, holotype, original image flipped)." httpUri="https://zenodo.org/record/1001963/files/figure.png" pageId="39" pageNumber="140">Figs. 23</figureCitation>
,
<figureCitation id="13532A72AE7AB617DC01420DFEEC072A" box="[264,311,1346,1370]" captionStart="FIGURE 27" captionStartId="44.[151,250,1770,1792]" captionTargetBox="[151,1436,193,1748]" captionTargetId="figure@44.[151,1436,193,1748]" captionTargetPageId="44" captionText="FIGURE 27. Distribution maps of Parachorius species. (A) Parachorius peninsularis. (B) Parachorius pseudojavanus. (C) Parachorius schuelkei. (D) Parachorius semsanganus. (E) Parachorius solodovnikovi. (F) Parachorius thomsoni. Type locality — type locality of species (holotype or lectotype); Type. Collection data — localities of type specimens associated with species (paratypes or paralectotypes); Collection data — localities of other specimens examined in this study. Literature data — localities retrieved from literature." httpUri="https://zenodo.org/record/1001971/files/figure.png" pageId="39" pageNumber="140">27E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE7AB617DD9E422AFC17070E" box="[151,972,1381,1406]" pageId="39" pageNumber="140" type="description">
<paragraph id="8BD736F7AE7AB617DD9E422AFC17070E" blockId="39.[151,972,1311,1406]" box="[151,972,1381,1406]" pageId="39" pageNumber="140">
urn:lsid:zoobank.org:act:4
<accessionNumber id="943BAB14AE7AB617DCBA422AFDD6070E" box="[435,525,1381,1406]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/E27796" pageId="39" pageNumber="140" type="EnaNcbi">E27796</accessionNumber>
A-EA33-425A-B0FE-F9D100FD799D
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE7AB617DD9E42E2FD6007B6" box="[151,699,1453,1479]" pageId="39" pageNumber="140" type="materials_examined">
<paragraph id="8BD736F7AE7AB617DD9E42E2FD6007B6" blockId="39.[151,1436,1453,2019]" box="[151,699,1453,1479]" pageId="39" pageNumber="140">
<emphasis id="B91CEAE5AE7AB617DD9E42E2FEF607B6" bold="true" box="[151,301,1453,1478]" pageId="39" pageNumber="140">Type locality</emphasis>
. Vientiane, Houayang forest, Laos.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE7AB617DDCE429DFD3F047E" pageId="39" pageNumber="140" type="etymology">
<paragraph id="8BD736F7AE7AB617DDCE429DFD3F047E" blockId="39.[151,1436,1453,2019]" pageId="39" pageNumber="140">
<emphasis id="B91CEAE5AE7AB617DDCE429DFE95079B" bold="true" box="[199,334,1490,1515]" pageId="39" pageNumber="140">Etymology.</emphasis>
This species is named after the great entomologist and staphylinid specialist, Alexey Solodovnikov, who was also my MSc thesis advisor.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE7AB617DDCE4155FE6A0426" pageId="39" pageNumber="140" type="distribution">
<paragraph id="8BD736F7AE7AB617DDCE4155FE6A0426" blockId="39.[151,1436,1453,2019]" pageId="39" pageNumber="140">
<emphasis id="B91CEAE5AE7AB617DDCE4155FEBB0443" bold="true" box="[199,352,1562,1587]" pageId="39" pageNumber="140">Distribution.</emphasis>
The species is known only from few localities in
<collectingCountry id="F37F7667AE7AB617DE9A4155FC2E0443" box="[915,1013,1562,1587]" name="Vietnam" pageId="39" pageNumber="140">Vietnam</collectingCountry>
,
<collectingCountry id="F37F7667AE7AB617D9094154FBE20442" box="[1024,1081,1563,1586]" name="Laos" pageId="39" pageNumber="140">Laos</collectingCountry>
, and
<collectingCountry id="F37F7667AE7AB617D97C4155FB070443" box="[1141,1244,1562,1587]" name="Thailand" pageId="39" pageNumber="140">Thailand</collectingCountry>
. It is presumably widespread in Indochina.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE7AB617DDCE412DFBA60496" pageId="39" pageNumber="140" type="diagnosis">
<paragraph id="8BD736F7AE7AB617DDCE412DFBA60496" blockId="39.[151,1436,1453,2019]" pageId="39" pageNumber="140">
<emphasis id="B91CEAE5AE7AB617DDCE412DFE9A040B" bold="true" box="[199,321,1634,1659]" pageId="39" pageNumber="140">Diagnosis.</emphasis>
This species is most similar to
<taxonomicName id="4C684D74AE7AB617DFAD412CFCD8040B" authorityName="S. Tarasov" authorityYear="2017" baseAuthorityName="Sharp" baseAuthorityYear="1875" box="[676,771,1634,1659]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="140" phylum="Arthropoda" rank="species" species="nudus">
<emphasis id="B91CEAE5AE7AB617DFAD412CFCD8040B" box="[676,771,1634,1659]" italics="true" pageId="39" pageNumber="140">P. nudus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C684D74AE7AB617DE18412CFC79040B" authorityName="S. Tarasov" authorityYear="2017" box="[785,930,1634,1659]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="140" phylum="Arthropoda" rank="species" species="longipenis">
<emphasis id="B91CEAE5AE7AB617DE18412CFC79040B" box="[785,930,1634,1659]" italics="true" pageId="39" pageNumber="140">P. longipenis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C684D74AE7AB617DEE9412CFB5D040B" authorityName="Tarasov" authorityYear="2017" baseAuthorityName="Arrow" baseAuthorityYear="1907" box="[992,1158,1634,1659]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="140" phylum="Arthropoda" rank="species" species="peninsularis">
<emphasis id="B91CEAE5AE7AB617DEE9412CFB5D040B" box="[992,1158,1634,1659]" italics="true" pageId="39" pageNumber="140">P. peninsularis</emphasis>
</taxonomicName>
. From those species and all other
<taxonomicName id="4C684D74AE7AB617DDF541CAFE5304EE" box="[252,392,1669,1694]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="140" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEAE5AE7AB617DDF541CAFE5304EE" box="[252,392,1669,1694]" italics="true" pageId="39" pageNumber="140">Parachorius</emphasis>
</taxonomicName>
, it can be separated by the following combination of characters (1) protibia with apical spur reaching protarsomere 2, (2) first tooth approximately twice as wide as second tooth, (3) metatibia inner margin denticulate with relatively small teeth, (4) parameres with dorsally horn-like upper lobes.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE7AB618DDCE41BDFC3C0315" lastPageId="40" lastPageNumber="141" pageId="39" pageNumber="140" type="description">
<paragraph id="8BD736F7AE7AB617DDCE41BDFBEF057B" blockId="39.[151,1436,1453,2019]" box="[199,1076,1778,1803]" pageId="39" pageNumber="140">
<emphasis id="B91CEAE5AE7AB617DDCE41BDFE82057B" bold="true" box="[199,345,1778,1803]" pageId="39" pageNumber="140">Description.</emphasis>
<emphasis id="B91CEAE5AE7AB617DC5641BDFE44057B" box="[351,415,1778,1803]" italics="true" pageId="39" pageNumber="140">Body.</emphasis>
Body black, length 3.9–5.0 mm. Elytra uniformly colored.
</paragraph>
<paragraph id="8BD736F7AE7AB617DDCE405AFBAD055E" blockId="39.[151,1436,1453,2019]" box="[199,1142,1813,1838]" pageId="39" pageNumber="140">
<emphasis id="B91CEAE5AE7AB617DDCE405AFED0055E" box="[199,267,1813,1838]" italics="true" pageId="39" pageNumber="140">Head.</emphasis>
Clypeus with two teeth, outer margin of each tooth distinctly notched basally.
</paragraph>
<paragraph id="8BD736F7AE7AB617DDCE4074FB580506" blockId="39.[151,1436,1453,2019]" pageId="39" pageNumber="140">
<emphasis id="B91CEAE5AE7AB617DDCE4074FEE60522" box="[199,317,1851,1874]" italics="true" pageId="39" pageNumber="140">Pronotum.</emphasis>
Pronotum rounded laterally. Frontal angles rounded. Disc covered with punctures separated by 1–2 puncture diameter(s). Pronotum in anterior-posterior direction notably shorter than elytra.
</paragraph>
<paragraph id="8BD736F7AE7AB617DDCE40CDFD1B05EA" blockId="39.[151,1436,1453,2019]" box="[199,704,1922,1947]" pageId="39" pageNumber="140">
<emphasis id="B91CEAE5AE7AB617DDCE40CDFECE05EB" box="[199,277,1922,1947]" italics="true" pageId="39" pageNumber="140">Elytra.</emphasis>
Elytra covered with sparse punctures.
</paragraph>
<paragraph id="8BD736F7AE7AB617DDCE40E8FD9405CE" blockId="39.[151,1436,1453,2019]" box="[199,591,1957,1982]" pageId="39" pageNumber="140">
<emphasis id="B91CEAE5AE7AB617DDCE40E8FEC905CE" box="[199,274,1959,1982]" italics="true" pageId="39" pageNumber="140">Wings.</emphasis>
Wings normally developed.
</paragraph>
<paragraph id="8BD736F7AE7AB618DDCE4084FC67036D" blockId="39.[151,1436,1453,2019]" lastBlockId="40.[151,1437,151,752]" lastPageId="40" lastPageNumber="141" pageId="39" pageNumber="140">
<emphasis id="B91CEAE5AE7AB617DDCE4084FED80592" box="[199,259,1995,2018]" italics="true" pageId="39" pageNumber="140">Legs.</emphasis>
Profemur with almost straight or sinuate anterior margin. Protibia with apical spur reaching protarsomere 2. First tooth notably modified in males, approximately twice as wide as second tooth; first and second teeth distinctly separated from each other; first tooth distinctly separated from denticles located between first and second teeth. Metafemoral posterior margin not serrate, simple, not modified. Metatibia slightly sinuate, inner margin denticulate with relatively small teeth, produced in small angle apically.
</paragraph>
<paragraph id="8BD736F7AE75B618DDCE4666FBBD0330" blockId="40.[151,1437,151,752]" box="[199,1126,295,320]" pageId="40" pageNumber="141">
<emphasis id="B91CEAE5AE75B618DDCE4666FEB90330" box="[199,354,297,320]" italics="true" pageId="40" pageNumber="141">Metasternum.</emphasis>
Transverse basal ridge of metasternum reduced and extremely short.
</paragraph>
<paragraph id="8BD736F7AE75B618DDCE4603FC3C0315" blockId="40.[151,1437,151,752]" box="[199,999,332,357]" pageId="40" pageNumber="141">
<emphasis id="B91CEAE5AE75B618DDCE4603FEE00315" box="[199,315,332,357]" italics="true" pageId="40" pageNumber="141">Aedeagus.</emphasis>
Parameres symmetrical, with dorsally horn-like upper lobes.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE75B618DDCE4620FEAD0080" pageId="40" pageNumber="141" type="materials_examined">
<paragraph id="8BD736F7AE75B618DDCE4620FC6303F8" blockId="40.[151,1437,151,752]" box="[199,952,367,392]" pageId="40" pageNumber="141">
<emphasis id="B91CEAE5AE75B618DDCE4620FEE003F8" bold="true" box="[199,315,367,392]" pageId="40" pageNumber="141">
<typeStatus id="54D38855AE75B618DDCE4620FEED03F8" box="[199,310,367,392]" pageId="40" pageNumber="141" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male (
<figureCitation id="13532A72AE75B618DC824620FE2B03F8" box="[395,496,367,392]" captionStart="FIGURE 23" captionStartId="40.[151,250,1887,1909]" captionTargetBox="[151,1423,840,1815]" captionTargetId="figure@40.[151,1436,840,1830]" captionTargetPageId="40" captionText="FIGURE 23. Morphology of Parachorius solodovnikovi. (A) aedeagus, lateral view (holotype). (B) aedeagus, lateral view (paratype, Thailand). (C) aedeagus, lateral view (paratype, Vietnam). (D) aedeagus, lateral view (E) habitus (male, holotype). (F) left metafemur (male, paratype, Vietnam) (G) left metafemur (male, holotype). (H) left protibia (male, holotype, original image flipped)." httpUri="https://zenodo.org/record/1001963/files/figure.png" pageId="40" pageNumber="141">Fig. 23E</figureCitation>
). Profemur with sinuate anterior margin.
</paragraph>
<paragraph id="8BD736F7AE75B618DDCE46DBFEAD0080" blockId="40.[151,1437,151,752]" pageId="40" pageNumber="141">
<materialsCitation id="3B003CAAAE75B618DDCE46DBFC110384" ID-GBIF-Occurrence="1658411519" collectingDate="2008-05-24" collectionCode="ZMUC" collectorName="S Tarasov" country="Laos" county="Material" elevation="200" latitude="18.097666" location="Houayang forest" longLatPrecision="9" longitude="102.67578" municipality="Laos" pageId="40" pageNumber="141" specimenCount="2" stateProvince="Vientiane" typeStatus="holotype">
<emphasis id="B91CEAE5AE75B618DDCE46DBFDE903DD" bold="true" box="[199,562,404,429]" pageId="40" pageNumber="141">
<collectingCounty id="62B64E7BAE75B618DDCE46DBFEF403DD" box="[199,303,404,429]" pageId="40" pageNumber="141">Material</collectingCounty>
examined.
<typeStatus id="54D38855AE75B618DCCC46DBFDE903DD" box="[453,562,404,429]" pageId="40" pageNumber="141" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingMunicipality id="6BB3AC8DAE75B618DF4146DBFD5903DD" box="[584,642,404,429]" pageId="40" pageNumber="141">
<emphasis id="B91CEAE5AE75B618DF4146DBFD5903DD" bold="true" box="[584,642,404,429]" pageId="40" pageNumber="141">Laos</emphasis>
</collectingMunicipality>
:
<emphasis id="B91CEAE5AE75B618DF9146DBFD0A03DD" bold="true" box="[664,721,404,429]" pageId="40" pageNumber="141">male</emphasis>
, &quot;
<collectingCountry id="F37F7667AE75B618DFE646DBFCE603DC" box="[751,829,404,428]" name="Laos" pageId="40" pageNumber="141">LAOS</collectingCountry>
,
<collectingRegion id="49ACF815AE75B618DE5946DBFC1F03DD" box="[848,964,404,429]" country="Laos" name="Viangchan" pageId="40" pageNumber="141">Vientiane</collectingRegion>
[,]
<location id="8EB7602CAE75B618DEEA46DAFB7303DD" LSID="urn:lsid:plazi:treatment:03C187E1AE7AB618DD9E4250FEAD0080:8EB7602CAE75B618DEEA46DAFB7303DD" box="[995,1192,404,429]" country="Laos" county="Material" latitude="18.097666" longLatPrecision="9" longitude="102.67578" municipality="Laos" name="Houayang forest" pageId="40" pageNumber="141" stateProvince="Vientiane">Houayang forest</location>
, N
<geoCoordinate id="EE5C5030AE75B618D9D946DBFABC03DC" box="[1232,1383,404,429]" direction="east" orientation="longitude" pageId="40" pageNumber="141" precision="1" value="18.097666">18° 5.860' E</geoCoordinate>
102° 40.547' h=
<elevation id="0045D1C4AE75B618DC0746F8FE8A03A0" box="[270,337,439,464]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="40" pageNumber="141" unit="m" value="200.0">
<quantity id="4C909B12AE75B618DC0746F8FE8A03A0" box="[270,337,439,464]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="40" pageNumber="141" unit="m" value="200.0">200m</quantity>
</elevation>
,
<collectingMethod id="52294EE0AE75B618DC5446F8FE3B03BF" box="[349,480,439,464]" pageId="40" pageNumber="141">carrion trap</collectingMethod>
&amp;
<collectingMethod id="52294EE0AE75B618DF0C46F8FD5203BF" box="[517,649,439,464]" pageId="40" pageNumber="141">pitfall traps</collectingMethod>
, disturb forest 10.3-
<collectingDate id="EF92E9DFAE75B618DE6046F8FC0603A0" box="[873,989,439,464]" pageId="40" pageNumber="141" value="2008-05-24">
<date id="FFD61037AE75B618DE6046F8FC0603A0" box="[873,989,439,464]" pageId="40" pageNumber="141" value="2008-05-24">24.5.2008</date>
</collectingDate>
leg.
<collectorName id="269D5321AE75B618D91F46F7FB5E03BF" box="[1046,1157,440,464]" pageId="40" pageNumber="141">S Tarasov</collectorName>
/
<typeStatus id="54D38855AE75B618D99446F7FAEA03A0" box="[1181,1329,440,464]" pageId="40" pageNumber="141" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C684D74AE75B618D83046F8FE6D0384" authority="S. Tarasov" authorityName="S. Tarasov" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="141" phylum="Arthropoda" rank="species" species="solodovnikovi">Cassolus solodovnikovi S. Tarasov</taxonomicName>
det. 2010&quot;,
<geoCoordinate id="EE5C5030AE75B618DF4B4693FD6E0385" box="[578,693,476,501]" direction="north" orientation="latitude" pageId="40" pageNumber="141" precision="9" value="18.097666">18°5.86'N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE75B618DFCB4693FC800385" box="[706,859,476,501]" direction="east" orientation="longitude" pageId="40" pageNumber="141" precision="1" value="102.67578">102°40.547'E</geoCoordinate>
(
<collectionCode id="ED79AE32AE75B618DE624693FC190384" box="[875,962,476,500]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="40" pageNumber="141">ZMUC</collectionCode>
)
</materialsCitation>
.
<emphasis id="B91CEAE5AE75B618DEDC4693FB190385" bold="true" box="[981,1218,476,501]" pageId="40" pageNumber="141">Paratypes (5 males)</emphasis>
:
<materialsCitation id="3B003CAAAE75B618D9DB4693FD0B004C" ID-GBIF-Occurrence="1658411521" collectingDate="2007-09-30" collectingDateMax="2007-11-28" collectingDateMin="2007-09-30" collectorName="S. Tarasov" country="Laos" elevation="880" latitude="18.325867" location="Ban Vangheua" longLatPrecision="1" longitude="102.81153" municipality="National Park Phu Khao Khouay" pageId="40" pageNumber="141" specimenCount="1" specimenCount-male="1" stateProvince="Vientiane" typeStatus="paratype">
<emphasis id="B91CEAE5AE75B618D9DB4693FA4E0385" bold="true" box="[1234,1429,476,501]" pageId="40" pageNumber="141">
<specimenCount id="9D6EFD7EAE75B618D9DB4693FAF90385" box="[1234,1314,476,501]" pageId="40" pageNumber="141" type="male">1 male</specimenCount>
<typeStatus id="54D38855AE75B618D8234693FA4E0385" box="[1322,1429,476,501]" pageId="40" pageNumber="141" type="paratype">paratype</typeStatus>
</emphasis>
,
<collectingRegion id="49ACF815AE75B618DD9E46B0FED90068" box="[151,258,511,536]" country="Laos" name="Viangchan" pageId="40" pageNumber="141">Vientiane</collectingRegion>
province,
<collectingMunicipality id="6BB3AC8DAE75B618DC8946B0FCDA0068" box="[384,769,511,536]" pageId="40" pageNumber="141">National Park Phu Khao Khouay</collectingMunicipality>
,
<location id="8EB7602CAE75B618DE06454FFC630068" LSID="urn:lsid:plazi:treatment:03C187E1AE7AB618DD9E4250FEAD0080:8EB7602CAE75B618DE06454FFC630068" box="[783,952,511,536]" country="Laos" latitude="18.325867" longLatPrecision="1" longitude="102.81153" municipality="National Park Phu Khao Khouay" name="Ban Vangheua" pageId="40" pageNumber="141" stateProvince="Vientiane">Ban Vangheua</location>
,
<geoCoordinate id="EE5C5030AE75B618DECE46B0FB8C0068" box="[967,1111,511,536]" direction="north" orientation="latitude" pageId="40" pageNumber="141" precision="1" value="18.325867">18°19.552'N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE75B618D96F46B0FADA0068" box="[1126,1281,511,536]" direction="east" orientation="longitude" pageId="40" pageNumber="141" precision="1" value="102.81153">102°48.692'E</geoCoordinate>
,
<elevation id="0045D1C4AE75B618D819454FFA870067" box="[1296,1372,512,536]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="40" pageNumber="141" unit="m" value="880.0">
<quantity id="4C909B12AE75B618D819454FFA870067" box="[1296,1372,512,536]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="40" pageNumber="141" unit="m" value="880.0">880 m</quantity>
</elevation>
, pine forest, on
<collectingDate id="EF92E9DFAE75B618DC11456BFE03004D" box="[280,472,548,573]" pageId="40" pageNumber="141" value="2007-09-30" valueMax="2007-11-28" valueMin="2007-09-30">
<date id="FFD61037AE75B618DC11456BFE03004D" box="[280,472,548,573]" pageId="40" pageNumber="141" value="2007-09-30" valueMax="2007-11-28" valueMin="2007-09-30">30/ix–28/xi/2007</date>
</collectingDate>
(
<collectorName id="269D5321AE75B618DCF8456BFDAB004C" box="[497,624,548,572]" pageId="40" pageNumber="141">S. Tarasov</collectorName>
) (CST)
</materialsCitation>
;
<materialsCitation id="3B003CAAAE75B618DFED456BFB4E0010" ID-GBIF-Occurrence="1658411518" collectingDate="2007-08-30" collectingDateMax="2007-09-11" collectingDateMin="2007-08-30" collectorName="S. Tarasov" country="Thailand" elevation="200" latitude="18.097666" location="Houayang forest" longLatPrecision="9" longitude="102.67578" pageId="40" pageNumber="141" specimenCount="1" specimenCount-male="1" stateProvince="Vientiane" typeStatus="paratype">
<emphasis id="B91CEAE5AE75B618DFED456BFC6F004D" bold="true" box="[740,948,548,573]" pageId="40" pageNumber="141">
<specimenCount id="9D6EFD7EAE75B618DFED456BFCE0004D" box="[740,827,548,573]" pageId="40" pageNumber="141" type="male">1 male</specimenCount>
<typeStatus id="54D38855AE75B618DE40456BFC6F004D" box="[841,948,548,573]" pageId="40" pageNumber="141" type="paratype">paratype</typeStatus>
</emphasis>
,
<collectingRegion id="49ACF815AE75B618DEC2456BFBE2004D" box="[971,1081,548,573]" country="Laos" name="Viangchan" pageId="40" pageNumber="141">Vientiane</collectingRegion>
,
<location id="8EB7602CAE75B618D945456AFACA004D" LSID="urn:lsid:plazi:treatment:03C187E1AE7AB618DD9E4250FEAD0080:8EB7602CAE75B618D945456AFACA004D" box="[1100,1297,548,573]" country="Thailand" latitude="18.097666" longLatPrecision="9" longitude="102.67578" name="Houayang forest" pageId="40" pageNumber="141" stateProvince="Vientiane">Houayang forest</location>
,
<geoCoordinate id="EE5C5030AE75B618D82D456BFA4C004D" box="[1316,1431,548,573]" direction="north" orientation="latitude" pageId="40" pageNumber="141" precision="9" value="18.097666">18°5.86'N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE75B618DD9E4508FEE90010" box="[151,306,583,608]" direction="east" orientation="longitude" pageId="40" pageNumber="141" precision="1" value="102.67578">102°40.547'E</geoCoordinate>
,
<elevation id="0045D1C4AE75B618DC364507FE50002F" box="[319,395,584,608]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="40" pageNumber="141" unit="m" value="200.0">
<quantity id="4C909B12AE75B618DC364507FE50002F" box="[319,395,584,608]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="40" pageNumber="141" unit="m" value="200.0">200 m</quantity>
</elevation>
, disturb forest, on
<collectingDate id="EF92E9DFAE75B618DF6C4508FCEF0010" box="[613,820,583,608]" pageId="40" pageNumber="141" value="2007-08-30" valueMax="2007-09-11" valueMin="2007-08-30">
<date id="FFD61037AE75B618DF6C4508FCEF0010" box="[613,820,583,608]" pageId="40" pageNumber="141" value="2007-08-30" valueMax="2007-09-11" valueMin="2007-08-30">30/viii–11/ix/2007</date>
</collectingDate>
(
<collectorName id="269D5321AE75B618DE4E4507FC1B0010" box="[839,960,584,608]" pageId="40" pageNumber="141">S. Tarasov</collectorName>
) (CST);
<collectingCountry id="F37F7667AE75B618D9214508FB4E0010" box="[1064,1173,583,608]" name="Thailand" pageId="40" pageNumber="141">
<emphasis id="B91CEAE5AE75B618D9214508FB4E0010" bold="true" box="[1064,1173,583,608]" pageId="40" pageNumber="141">Thailand</emphasis>
</collectingCountry>
</materialsCitation>
:
<materialsCitation id="3B003CAAAE75B618D9AF4508FAA000F5" ID-GBIF-Occurrence="1658411536" collectingDate="2001-05-05" collectingDateMax="2001-05-12" collectingDateMin="2001-05-05" collectionCode="ISNB" country="Vietnam" latitude="17.50385" location="Na Haeo" longLatPrecision="1" longitude="100.939384" pageId="40" pageNumber="141" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B91CEAE5AE75B618D9AF4508FAB00010" bold="true" box="[1190,1387,583,608]" pageId="40" pageNumber="141">
<specimenCount id="9D6EFD7EAE75B618D9AF4508FB2D0010" box="[1190,1270,583,608]" pageId="40" pageNumber="141" type="male">1 male</specimenCount>
<typeStatus id="54D38855AE75B618D8094508FAB00010" box="[1280,1387,583,608]" pageId="40" pageNumber="141" type="paratype">paratype</typeStatus>
</emphasis>
,
<location id="8EB7602CAE75B618D8724507FF0900F4" LSID="urn:lsid:plazi:treatment:03C187E1AE7AB618DD9E4250FEAD0080:8EB7602CAE75B618D8724507FF0900F4" country="Vietnam" latitude="17.50385" longLatPrecision="1" longitude="100.939384" name="Na Haeo" pageId="40" pageNumber="141">Na Haeo</location>
(secondary forest),
<geoCoordinate id="EE5C5030AE75B618DCB14523FD9300F4" box="[440,584,620,644]" direction="north" orientation="latitude" pageId="40" pageNumber="141" precision="1" value="17.50385">17°30.231'N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE75B618DF5F4523FD2B00F5" box="[598,752,620,645]" direction="east" orientation="longitude" pageId="40" pageNumber="141" precision="1" value="100.939384">100°56.363'E</geoCoordinate>
, on
<collectingDate id="EF92E9DFAE75B618DE2A4523FC6B00F5" box="[803,944,620,645]" pageId="40" pageNumber="141" value="2001-05-05" valueMax="2001-05-12" valueMin="2001-05-05">
<date id="FFD61037AE75B618DE2A4523FC6B00F5" box="[803,944,620,645]" pageId="40" pageNumber="141" value="2001-05-05" valueMax="2001-05-12" valueMin="2001-05-05">5–12/v/2001</date>
</collectingDate>
(Constant, Grootaert) (
<collectionCode id="ED79AE32AE75B618D9B44523FAD900F4" box="[1213,1282,620,644]" pageId="40" pageNumber="141">ISNB</collectionCode>
);
<collectingCountry id="F37F7667AE75B618D81E4523FAA000F5" box="[1303,1403,620,645]" name="Vietnam" pageId="40" pageNumber="141">
<emphasis id="B91CEAE5AE75B618D81E4523FAA000F5" bold="true" box="[1303,1403,620,645]" pageId="40" pageNumber="141">Vietnam</emphasis>
</collectingCountry>
</materialsCitation>
:
<materialsCitation id="3B003CAAAE75B618D8874523FED800BC" ID-GBIF-Occurrence="1658411538" collectionCode="MNHN" collectorName="A. De Cooman" country="Vietnam" latitude="20.828333" location="Hoa-Binh" longLatPrecision="870" longitude="105.333336" municipality="Tonkin" pageId="40" pageNumber="141" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B91CEAE5AE75B618D8874523FE9800D8" bold="true" pageId="40" pageNumber="141">
<specimenCount id="9D6EFD7EAE75B618D8874523FF0A00D8" pageId="40" pageNumber="141" type="male">1 male</specimenCount>
<typeStatus id="54D38855AE75B618DDD145C0FE9800D8" box="[216,323,655,680]" pageId="40" pageNumber="141" type="paratype">paratype</typeStatus>
</emphasis>
,
<collectingMunicipality id="6BB3AC8DAE75B618DC5A45C0FE7900D8" box="[339,418,655,680]" pageId="40" pageNumber="141">Tonkin</collectingMunicipality>
[northern
<collectingCountry id="F37F7667AE75B618DF1345C0FDA400D8" box="[538,639,655,680]" name="Vietnam" pageId="40" pageNumber="141">Vietnam</collectingCountry>
], region of
<location id="8EB7602CAE75B618DFF645C0FCAB00D8" LSID="urn:lsid:plazi:treatment:03C187E1AE7AB618DD9E4250FEAD0080:8EB7602CAE75B618DFF645C0FCAB00D8" box="[767,880,655,680]" country="Vietnam" latitude="20.828333" longLatPrecision="870" longitude="105.333336" municipality="Tonkin" name="Hoa-Binh" pageId="40" pageNumber="141">Hoa-Binh</location>
,
<geoCoordinate id="EE5C5030AE75B618DE7445DFFC2A00D8" box="[893,1009,656,680]" direction="north" orientation="latitude" pageId="40" pageNumber="141" precision="92" value="20.828333">20°49.7'N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE75B618DEF745C0FBBC00D8" box="[1022,1127,655,680]" direction="east" orientation="longitude" pageId="40" pageNumber="141" precision="925" value="105.333336">105°20'E</geoCoordinate>
, on 1928, (
<collectorName id="269D5321AE75B618D9EE45DFFA4E00D8" box="[1255,1429,656,680]" pageId="40" pageNumber="141">A. De Cooman</collectorName>
) (
<collectionCode id="ED79AE32AE75B618DD9645FBFF2000BC" box="[159,251,692,716]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="40" pageNumber="141">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B003CAAAE75B618DC0445FBFEAD0080" ID-GBIF-Occurrence="1658411526" collectionCode="MNHN" collectorName="A. De Cooman" country="Vietnam" latitude="20.828333" location="Hoa-Binh" longLatPrecision="870" longitude="105.333336" municipality="Tonkin" pageId="40" pageNumber="141" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B91CEAE5AE75B618DC0445FBFE1600BD" bold="true" box="[269,461,692,717]" pageId="40" pageNumber="141">
<specimenCount id="9D6EFD7EAE75B618DC0445FBFE8700BD" box="[269,348,692,717]" pageId="40" pageNumber="141" type="male">1 male</specimenCount>
<typeStatus id="54D38855AE75B618DC6B45FBFE1600BD" box="[354,461,692,717]" pageId="40" pageNumber="141" type="paratype">paratype</typeStatus>
</emphasis>
,
<collectingMunicipality id="6BB3AC8DAE75B618DCD345FBFDF200BD" box="[474,553,692,717]" pageId="40" pageNumber="141">Tonkin</collectingMunicipality>
[northern
<collectingCountry id="F37F7667AE75B618DF9745FBFCD800BD" box="[670,771,692,717]" name="Vietnam" pageId="40" pageNumber="141">Vietnam</collectingCountry>
], region of
<location id="8EB7602CAE75B618DE7645FBFC2B00BD" LSID="urn:lsid:plazi:treatment:03C187E1AE7AB618DD9E4250FEAD0080:8EB7602CAE75B618DE7645FBFC2B00BD" box="[895,1008,692,717]" country="Vietnam" latitude="20.828333" longLatPrecision="870" longitude="105.333336" municipality="Tonkin" name="Hoa-Binh" pageId="40" pageNumber="141">Hoa-Binh</location>
,
<geoCoordinate id="EE5C5030AE75B618DEF245FBFBAB00BC" box="[1019,1136,692,716]" direction="north" orientation="latitude" pageId="40" pageNumber="141" precision="92" value="20.828333">20°49.7'N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE75B618D97345FBFB3F00BD" box="[1146,1252,692,717]" direction="east" orientation="longitude" pageId="40" pageNumber="141" precision="925" value="105.333336">105°20'E</geoCoordinate>
, on 1927 (
<collectorName id="269D5321AE75B618D85045FBFF230080" pageId="40" pageNumber="141">A. De Cooman</collectorName>
) (
<collectionCode id="ED79AE32AE75B618DC074597FEB10080" box="[270,362,728,752]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="40" pageNumber="141">MNHN</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue Copy permalink