174 lines
19 KiB
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174 lines
19 KiB
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<document id="2BC382C066D9F7315E47212B4211339A" ID-CLB-Dataset="31902" ID-DOI="10.11646/zootaxa.4329.2.1" ID-GBIF-Dataset="7e33ec7d-211b-4042-8d07-344364ee805d" ID-ISSN="1175-5326" ID-Zenodo-Dep="1001915" ID-ZooBank="8226E27D-E3A9-481A-B51B-558643Bbb03A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507106636447" checkinUser="plazi" docAuthor="Tarasov, Sergei" docDate="2017" docId="03C187E1AE57B63ADD9E44B4FE110453" docLanguage="en" docName="zootaxa.4329.2.1.pdf" docOrigin="Zootaxa 4329 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Panelus (Panelus) minutus Tarasov 2017, new combination" docType="treatment" docVersion="12" lastPageNumber="111" masterDocId="FFF8FF99AE5DB630DD09474FFFDB0270" masterDocTitle="A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data" masterLastPageNumber="149" masterPageNumber="101" pageNumber="111" updateTime="1698469310636" updateUser="ExternalLinkService">
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<mods:title id="964481E509A1C823C6C0D11486E58F56">A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data</mods:title>
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<mods:namePart id="1FB4D1DA9C7C13D49953A6D4B524A42F">Tarasov, Sergei</mods:namePart>
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<mods:title id="0EB771783AB740FFC1FC031434CCA752">Zootaxa</mods:title>
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<mods:date id="F6C411CEF90C7B710A031F16D22AF63B">2017</mods:date>
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<mods:number id="A973DC8E23C31D54A9B98DAEBB78E0C1">2017-10-04</mods:number>
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<mods:identifier id="97739A47ECAE4E1E51D56F20448E7762" type="DOI">10.11646/zootaxa.4329.2.1</mods:identifier>
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<treatment id="03C187E1AE57B63ADD9E44B4FE110453" ID-DOI="http://doi.org/10.5281/zenodo.6010796" ID-GBIF-Taxon="135106118" ID-Zenodo-Dep="6010796" LSID="urn:lsid:plazi:treatment:03C187E1AE57B63ADD9E44B4FE110453" httpUri="http://treatment.plazi.org/id/03C187E1AE57B63ADD9E44B4FE110453" lastPageNumber="111" pageId="10" pageNumber="111">
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<subSubSection id="C372657CAE57B63ADD9E44B4FC6B0665" box="[151,944,1019,1045]" pageId="10" pageNumber="111" type="nomenclature">
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<paragraph id="8BD736F7AE57B63ADD9E44B4FC6B0665" blockId="10.[151,944,1019,1045]" box="[151,944,1019,1045]" pageId="10" pageNumber="111">
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<heading id="D09F819BAE57B63ADD9E44B4FC6B0665" bold="true" box="[151,944,1019,1045]" fontSize="11" level="1" pageId="10" pageNumber="111" reason="1">
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<emphasis id="B91CEAE5AE57B63ADD9E44B4FC6B0665" bold="true" box="[151,944,1019,1045]" pageId="10" pageNumber="111">
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<taxonomicName id="4C684D74AE57B63ADD9E44B4FD080665" ID-CoL="7ZQ67" authority="(Boucomont, 1914)" authorityName="Tarasov" authorityYear="2017" baseAuthorityName="Boucomont" baseAuthorityYear="1914" box="[151,723,1019,1045]" class="Insecta" family="Scarabaeidae" genus="Panelus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="minutus" status="comb. nov." subGenus="Panelus">
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<emphasis id="B91CEAE5AE57B63ADD9E44B4FF220665" bold="true" box="[151,249,1019,1045]" italics="true" pageId="10" pageNumber="111">Panelus</emphasis>
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(
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<emphasis id="B91CEAE5AE57B63ADC0344B4FEB00665" bold="true" box="[266,363,1019,1045]" italics="true" pageId="10" pageNumber="111">Panelus</emphasis>
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)
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<emphasis id="B91CEAE5AE57B63ADC7544B4FE3B0665" bold="true" box="[380,480,1019,1045]" italics="true" pageId="10" pageNumber="111">minutus</emphasis>
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(
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<bibRefCitation id="EFF94B06AE57B63ADCE644B4FD100665" author="Boucomont" box="[495,715,1019,1045]" pageId="10" pageNumber="111" refString="Boucomont, A. (1914) Les Coprophages de l'Archipel Malais. Annales de la Societe Entomologique de France, 83, 238 - 350." type="journal article" year="1914">Boucomont, 1914</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="A22F579EAE57B63ADFD344B4FC6B0665" box="[730,944,1019,1045]" pageId="10" pageNumber="111">new combination</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C372657CAE57B63ADD9E430DFDF80606" pageId="10" pageNumber="111" type="reference_group">
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<paragraph id="8BD736F7AE57B63ADD9E430DFDF80606" blockId="10.[151,568,1090,1142]" pageId="10" pageNumber="111">
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<treatmentCitationGroup id="AB7811D9AE57B63ADD9E430DFDF80606" pageId="10" pageNumber="111">
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<taxonomicName id="4C684D74AE57B63ADD9E430DFDE30628" ID-CoL="7TG36" authority="Boucomont, 1914: 252" authorityName="Boucomont" authorityPageNumber="252" authorityYear="1914" box="[151,568,1090,1112]" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="minutus">
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<emphasis id="B91CEAE5AE57B63ADD9E430DFE920628" box="[151,329,1090,1112]" italics="true" pageId="10" pageNumber="111">Cassolus minutus</emphasis>
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<treatmentCitation id="0AC910E6AE57B63ADC46430CFDE30628" author="Boucomont" box="[335,568,1090,1112]" page="252" pageId="10" pageNumber="111" year="1914">
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<bibRefCitation id="EFF94B06AE57B63ADC46430CFDD30628" author="Boucomont" box="[335,520,1090,1112]" pageId="10" pageNumber="111" refString="Boucomont, A. (1914) Les Coprophages de l'Archipel Malais. Annales de la Societe Entomologique de France, 83, 238 - 350." type="journal article" year="1914">Boucomont, 1914</bibRefCitation>
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: 252
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</treatmentCitation>
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</taxonomicName>
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<emphasis id="B91CEAE5AE57B63ADD9E432FFE8A0606" box="[151,337,1120,1142]" italics="true" pageId="10" pageNumber="111">
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<taxonomicName id="4C684D74AE57B63ADD9E432FFE900606" ID-CoL="7TG36" authorityName="Boucomont" authorityYear="1914" box="[151,331,1120,1142]" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="minutus">Cassolus minutus</taxonomicName>
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;
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</emphasis>
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<treatmentCitation id="0AC910E6AE57B63ADC5E432FFDF80606" author="Balthasar" box="[343,547,1120,1142]" page="263" pageId="10" pageNumber="111" year="1963">
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<bibRefCitation id="EFF94B06AE57B63ADC5E432FFE280606" author="Balthasar" box="[343,499,1120,1142]" pageId="10" pageNumber="111" refString="Balthasar, V. (1963) Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Folume 1. Tschechoslowakische Akademie der Wissenschaften, Prague, 391 pp., XXIV pls." type="book" year="1963">Balthasar 1963</bibRefCitation>
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: 263
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</treatmentCitation>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C372657CAE57B63ADD9E43ECFE2806CB" box="[151,499,1187,1212]" pageId="10" pageNumber="111" type="materials_examined">
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<paragraph id="8BD736F7AE57B63ADD9E43ECFE2806CB" blockId="10.[151,1437,1187,1572]" box="[151,499,1187,1212]" pageId="10" pageNumber="111">
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<materialsCitation id="3B003CAAAE57B63ADD9E43ECFE2806CB" ID-GBIF-Occurrence="1658411522" box="[151,499,1187,1212]" location="Mt. Gedeh" pageId="10" pageNumber="111" specimenCount="1" stateProvince="Java" typeStatus="holotype">
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<emphasis id="B91CEAE5AE57B63ADD9E43ECFEF606CC" bold="true" box="[151,301,1187,1212]" pageId="10" pageNumber="111">
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<typeStatus id="54D38855AE57B63ADD9E43ECFF0906CC" box="[151,210,1187,1212]" pageId="10" pageNumber="111">Type</typeStatus>
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locality
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</emphasis>
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.
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<location id="8EB7602CAE57B63ADC3543EBFE6D06CC" LSID="urn:lsid:plazi:treatment:03C187E1AE57B63ADD9E44B4FE110453:8EB7602CAE57B63ADC3543EBFE6D06CC" box="[316,438,1187,1212]" name="Mt. Gedeh" pageId="10" pageNumber="111" stateProvince="Java">Mt. Gedeh</location>
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,
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<collectingRegion id="49ACF815AE57B63ADCC843EBFE2806CB" box="[449,499,1188,1211]" country="Indonesia" name="Jawa Timur" pageId="10" pageNumber="111">Java</collectingRegion>
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C372657CAE57B63ADDCE4388FE110453" pageId="10" pageNumber="111" type="discussion">
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<paragraph id="8BD736F7AE57B63ADDCE4388FE110453" blockId="10.[151,1437,1187,1572]" pageId="10" pageNumber="111">
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<emphasis id="B91CEAE5AE57B63ADDCE4388FE930690" bold="true" box="[199,328,1223,1248]" pageId="10" pageNumber="111">Comments</emphasis>
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. This species was originally described in
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<taxonomicName id="4C684D74AE57B63ADE154388FC5A0690" box="[796,897,1223,1248]" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="genus">
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<emphasis id="B91CEAE5AE57B63ADE154388FC5A0690" box="[796,897,1223,1248]" italics="true" pageId="10" pageNumber="111">Cassolus</emphasis>
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</taxonomicName>
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. Unfortunately, I was not able to locate the
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<typeStatus id="54D38855AE57B63AD8654387FA470690" box="[1388,1436,1224,1248]" pageId="10" pageNumber="111">type</typeStatus>
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specimen of this species in the collection of MNHN where it was presumably deposited. However, the original description of
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<bibRefCitation id="EFF94B06AE57B63ADC49425FFDCC0757" author="Boucomont" box="[320,535,1295,1319]" pageId="10" pageNumber="111" refString="Boucomont, A. (1914) Les Coprophages de l'Archipel Malais. Annales de la Societe Entomologique de France, 83, 238 - 350." type="journal article" year="1914">Boucomont (1914)</bibRefCitation>
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includes evidence allowing the taxonomic assessment of this species. In that paper, Boucomont gives two important notes: (1) the species
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<taxonomicName id="4C684D74AE57B63ADE89427BFBDB073B" authorityName="Boucomont" authorityYear="1914" box="[896,1024,1332,1355]" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="minutus">
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<emphasis id="B91CEAE5AE57B63ADE89427BFBDB073B" box="[896,1024,1332,1355]" italics="true" pageId="10" pageNumber="111">C. minutus</emphasis>
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</taxonomicName>
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is so similar to
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<taxonomicName id="4C684D74AE57B63AD9DB427CFEBF071F" authority="Waterhouse, 1874" authorityName="Waterhouse" authorityYear="1874" class="Insecta" family="Scarabaeidae" genus="Panelus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="parvulus">
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<emphasis id="B91CEAE5AE57B63AD9DB427CFA47073C" box="[1234,1436,1331,1356]" italics="true" pageId="10" pageNumber="111">Panelus parvulus</emphasis>
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Waterhouse, 1874
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</taxonomicName>
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that can be confused with it, and (2) both
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<taxonomicName id="4C684D74AE57B63ADE6D4217FBD0071F" authorityName="S. Tarasov" authorityYear="2017" baseAuthorityName="Lansberge" baseAuthorityYear="1885" box="[868,1035,1368,1391]" class="Insecta" family="Scarabaeidae" genus="Panelus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="sumatranus">
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<emphasis id="B91CEAE5AE57B63ADE6D4217FBD0071F" box="[868,1035,1368,1391]" italics="true" pageId="10" pageNumber="111">C. sumatranus</emphasis>
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</taxonomicName>
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[
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<emphasis id="B91CEAE5AE57B63AD9164217FB99071F" box="[1055,1090,1368,1391]" italics="true" pageId="10" pageNumber="111">i.e.</emphasis>
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,
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<taxonomicName id="4C684D74AE57B63AD95A4218FAE0071F" authorityName="S. Tarasov" authorityYear="2017" baseAuthorityName="Lansberge" baseAuthorityYear="1885" box="[1107,1339,1367,1392]" class="Insecta" family="Scarabaeidae" genus="Panelus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="sumatranus">
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<emphasis id="B91CEAE5AE57B63AD95A4218FAE0071F" box="[1107,1339,1367,1392]" italics="true" pageId="10" pageNumber="111">Panelus sumatranus</emphasis>
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</taxonomicName>
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] and
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<taxonomicName id="4C684D74AE57B63AD88B4217FF2B07E3" authorityName="Boucomont" authorityYear="1914" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="minutus">
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<emphasis id="B91CEAE5AE57B63AD88B4217FF2B07E3" italics="true" pageId="10" pageNumber="111">C. minutus</emphasis>
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</taxonomicName>
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remarkably resemble
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<taxonomicName id="4C684D74AE57B63ADCE54234FD9D07E4" authorityName="Lewis" authorityYear="1895" box="[492,582,1403,1428]" class="Insecta" family="Scarabaeidae" genus="Panelus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="genus">
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<emphasis id="B91CEAE5AE57B63ADCE54234FD9D07E4" box="[492,582,1403,1428]" italics="true" pageId="10" pageNumber="111">Panelus</emphasis>
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</taxonomicName>
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in their body shape, epistome, tarsi, and protibia. Additionally,
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<bibRefCitation id="EFF94B06AE57B63AD8134233FF3A07C7" author="Boucomont" pageId="10" pageNumber="111" refString="Boucomont, A. (1914) Les Coprophages de l'Archipel Malais. Annales de la Societe Entomologique de France, 83, 238 - 350." type="journal article" year="1914">Boucomont (1914)</bibRefCitation>
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mentioned that
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<taxonomicName id="4C684D74AE57B63ADC9142EFFDC907C7" authorityName="Boucomont" authorityYear="1914" box="[408,530,1440,1463]" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="minutus">
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<emphasis id="B91CEAE5AE57B63ADC9142EFFDC907C7" box="[408,530,1440,1463]" italics="true" pageId="10" pageNumber="111">C. minutus</emphasis>
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</taxonomicName>
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is significantly smaller in size than
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<taxonomicName id="4C684D74AE57B63ADEA042EFFB9607C7" authorityName="S. Tarasov" authorityYear="2017" baseAuthorityName="Lansberge" baseAuthorityYear="1885" box="[937,1101,1440,1463]" class="Insecta" family="Scarabaeidae" genus="Panelus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="sumatranus">
|
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|
|
<emphasis id="B91CEAE5AE57B63ADEA042EFFB9607C7" box="[937,1101,1440,1463]" italics="true" pageId="10" pageNumber="111">C. sumatranus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Boucomont’s findings of the close affiliations between
|
|||
|
|
<taxonomicName id="4C684D74AE57B63ADCCD428BFDB007AB" authorityName="S. Tarasov" authorityYear="2017" baseAuthorityName="Lansberge" baseAuthorityYear="1885" box="[452,619,1476,1499]" class="Insecta" family="Scarabaeidae" genus="Panelus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="sumatranus">
|
|||
|
|
<emphasis id="B91CEAE5AE57B63ADCCD428BFDB007AB" box="[452,619,1476,1499]" italics="true" pageId="10" pageNumber="111">C. sumatranus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and the members of
|
|||
|
|
<taxonomicName id="4C684D74AE57B63ADE6F428CFC1B07AC" authorityName="Lewis" authorityYear="1895" box="[870,960,1475,1500]" class="Insecta" family="Scarabaeidae" genus="Panelus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B91CEAE5AE57B63ADE6F428CFC1B07AC" box="[870,960,1475,1500]" italics="true" pageId="10" pageNumber="111">Panelus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
are supported by this study. I follow the mentioned evidences for similarity between
|
|||
|
|
<taxonomicName id="4C684D74AE57B63ADF9642A7FCC6078F" authorityName="Boucomont" authorityYear="1914" box="[671,797,1512,1535]" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="minutus">
|
|||
|
|
<emphasis id="B91CEAE5AE57B63ADF9642A7FCC6078F" box="[671,797,1512,1535]" italics="true" pageId="10" pageNumber="111">C. minutus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and the members of
|
|||
|
|
<emphasis id="B91CEAE5AE57B63AD92942A8FB5A0470" box="[1056,1153,1511,1536]" italics="true" pageId="10" pageNumber="111">
|
|||
|
|
<taxonomicName id="4C684D74AE57B63AD92942A8FBA60470" authorityName="Lewis" authorityYear="1895" box="[1056,1149,1511,1536]" class="Insecta" family="Scarabaeidae" genus="Panelus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="genus">Panelus</taxonomicName>
|
|||
|
|
,
|
|||
|
|
</emphasis>
|
|||
|
|
and thereby transfer
|
|||
|
|
<taxonomicName id="4C684D74AE57B63AD88B42A7FF2B0453" authorityName="Boucomont" authorityYear="1914" class="Insecta" family="Scarabaeidae" genus="Cassolus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="minutus">
|
|||
|
|
<emphasis id="B91CEAE5AE57B63AD88B42A7FF2B0453" italics="true" pageId="10" pageNumber="111">C. minutus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
to
|
|||
|
|
<taxonomicName id="4C684D74AE57B63ADC1A4144FEB60454" authorityName="Lewis" authorityYear="1895" box="[275,365,1547,1572]" class="Insecta" family="Scarabaeidae" genus="Panelus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B91CEAE5AE57B63ADC1A4144FEB60454" box="[275,365,1547,1572]" italics="true" pageId="10" pageNumber="111">Panelus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
as well.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|