152 lines
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152 lines
16 KiB
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<document id="A7CCED87F744FC310C29EEAFA693290F" ID-DOI="0.1649/0010-065X-69.mo4.27" ID-ISSN="1938-4394" ID-Zenodo-Dep="10086147" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623095935056" checkinUser="carolina" docAuthor="Johnston, M. Andrew, Fleming, David, Franz, Nico M. & Smith, Aaron D." docDate="2015" docId="B31FCE62BB6DFFBEFD0AFD6FFCF30A81" docLanguage="en" docName="TheColeopBull.69.mo14.27-54.pdf.imf" docOrigin="The Coleopterists Bulletin (mo 14) 69" docStyle="DocumentStyle:4CAA047AD53AEBA766D4E826C2986B94.5:TheColeopBull.2010-2016.monograph" docStyleId="4CAA047AD53AEBA766D4E826C2986B94" docStyleName="TheColeopBull.2010-2016.monograph" docStyleVersion="5" docTitle="Eleodes (Litheleodes) extricatus" docType="treatment" docVersion="4" lastPageNumber="46" masterDocId="4F26B61ABB7EFFADFFC8FFB9FFAD0C40" masterDocTitle="Amphidorini Leconte (Coleoptera: Tenebrionidae) of Arizona: Keys and Species Accounts" masterLastPageNumber="54" masterPageNumber="27" pageNumber="46" updateTime="1699918030632" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="A0BE0CB24B31890AF435DDBF1CB0DD34">
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<mods:title id="25A57E8BA96ED8FB14982FF9CEB1F780">Amphidorini Leconte (Coleoptera: Tenebrionidae) of Arizona: Keys and Species Accounts</mods:title>
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</mods:titleInfo>
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<mods:name id="39699056452ACAD91E6AF3FB8AF70342" type="personal">
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<mods:roleTerm id="722F16CC3E75DDE4DBF8FA5638845EC9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8D91DCF60831C280DFE230BCCE7C0F46">Johnston, M. Andrew</mods:namePart>
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<mods:affiliation id="AEA1EE3497A888E1E0A672ECEA036DE0">School of Life Sciences, Arizona State University PO Box 874501, Tempe, AZ 85287 - 4501, U. S. A.</mods:affiliation>
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<mods:nameIdentifier id="30B4A91C21E31E65D80DC7DB2B9FEB4A" type="email">ajohnston@asu.edu,</mods:nameIdentifier>
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<mods:name id="27B26180BD0D287E36F38C3B48A69266" type="personal">
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</mods:role>
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<mods:namePart id="0DB0855329478D9B2FBCBBC27BBE5E73">Fleming, David</mods:namePart>
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<mods:affiliation id="7FE61DBEAA34332706A9972046034655">School of Life Sciences, Arizona State University PO Box 874501, Tempe, AZ 85287 - 4501, U. S. A.</mods:affiliation>
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<mods:nameIdentifier id="485E5312AB21030BFD5FA8A36274FCEA" type="email">david.fleming.1@asu.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name id="4C1465CB1FDA50CEB38DFA31F723E4F8" type="personal">
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<mods:roleTerm id="8F85E1F0C5CACF29A3235A18436AEA93">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9494136FC40E38BC3B214339BFC7A4F5">Franz, Nico M.</mods:namePart>
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<mods:affiliation id="F1F07E4F188C92BF8EBD6F4631A1B7C9">School of Life Sciences, Arizona State University PO Box 874501, Tempe, AZ 85287 - 4501, U. S. A.</mods:affiliation>
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<mods:nameIdentifier id="CD3EEB092A3ED3F5E62E75B5CFC937C7" type="email">nico.franz@asu.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name id="5A1599CD415012A7ECBE320210B3924A" type="personal">
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<mods:roleTerm id="C80BCF0FE5D54FDE32939C5765E04AAB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2014E8F38F450A6E94885AF86A36064A">Smith, Aaron D.</mods:namePart>
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<mods:affiliation id="AB9D231AFDF18B72CEFCF8EF16DE3A47">Department of Biological Sciences, Northern Arizona University Flagstaff, AZ 86011 - 5640, U. S. A.</mods:affiliation>
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<mods:nameIdentifier id="2B868B523D1289D134CAC9B6ADC58FBE" type="email">aaron.smith@nau.edu</mods:nameIdentifier>
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<mods:typeOfResource id="BF9E6EB55FC218797586E3E890D2C63D">text</mods:typeOfResource>
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<mods:titleInfo id="B3521777F37FE6E8945B6293ED37D47A">
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<mods:title id="4835DD2E727C43ADDA20809CDF06E1C4">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="2ACA6D8BBFEBA4340D4CB73255A3DFDC">
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<mods:date id="118BF38B0E652C7A81C529E171CC836E">2015</mods:date>
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<mods:detail id="DF4E432B40758ECB8B22D5A627749869" type="series">
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<mods:title id="5D508DCD6EACF1765BCA4E08B45C4F16">mo 14</mods:title>
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</mods:detail>
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<mods:detail id="E8CC92C8B5F159630C329E0C985E3C99" type="pubDate">
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<mods:number id="95503A683F9939D8EC7F960657191ACC">2015-12-18</mods:number>
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</mods:detail>
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<mods:detail id="794048ED62FCBB1A7B364256DCDC18B2" type="volume">
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<mods:number id="56333E4F369C22AD5BADE298FAECB9F5">69</mods:number>
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</mods:detail>
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<mods:extent id="932E86D4E50489686F6A7BD3118CD93A" unit="page">
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<mods:start id="8AD5D7C0A004B499C5B50BABA86F5E27">27</mods:start>
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<mods:end id="841AF2AB1AEC78ED6CEB75AD68659D7E">54</mods:end>
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</mods:extent>
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<mods:classification id="481811470ADD86E951757F2DA37B2699">journal article</mods:classification>
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<mods:identifier id="2BFDEA974A63E20A3E5E91585ADAF72B" type="DOI">0.1649/0010-065X-69.mo4.27</mods:identifier>
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<mods:identifier id="924A99F4E3EC8CF9C3EF845F7C19AB25" type="ISSN">1938-4394</mods:identifier>
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<mods:identifier id="5ACBAF5A74CC265240D669FAE1D943FE" type="Zenodo-Dep">10086147</mods:identifier>
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</mods:mods>
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<treatment id="B31FCE62BB6DFFBEFD0AFD6FFCF30A81" LSID="urn:lsid:plazi:treatment:B31FCE62BB6DFFBEFD0AFD6FFCF30A81" httpUri="http://treatment.plazi.org/id/B31FCE62BB6DFFBEFD0AFD6FFCF30A81" lastPageNumber="46" pageId="19" pageNumber="46">
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<subSubSection id="73AC2CFFBB6DFFBEFD0AFD6FFBD20EAB" box="[706,1151,726,747]" pageId="19" pageNumber="46" type="nomenclature">
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<paragraph id="3B097F74BB6DFFBEFD0AFD6FFBD20EAB" blockId="19.[706,1151,726,774]" box="[706,1151,726,747]" pageId="19" pageNumber="46">
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<heading id="6041C818BB6DFFBEFD0AFD6FFBD20EAB" box="[706,1151,726,747]" centered="true" fontSize="10" level="2" pageId="19" pageNumber="46" reason="2">
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<taxonomicName id="FCB604F7BB6DFFBEFD0AFD6FFBD20EAB" authority="(Say, 1823)" baseAuthorityName="Say" baseAuthorityYear="1823" box="[706,1151,726,747]" class="Insecta" family="Tenebrionidae" genus="Eleodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="46" phylum="Arthropoda" rank="species" species="extricatus" subGenus="Litheleodes">
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<emphasis id="09C2A366BB6DFFBEFD0AFD6FFBD20EAB" bold="true" box="[706,1151,726,747]" pageId="19" pageNumber="46">
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<emphasis id="09C2A366BB6DFFBEFD0AFD6FFCA30EAB" bold="true" box="[706,782,726,747]" italics="true" pageId="19" pageNumber="46">Eleodes</emphasis>
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(
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<emphasis id="09C2A366BB6DFFBEFCE8FD6FFC220EAB" bold="true" box="[800,911,726,747]" italics="true" pageId="19" pageNumber="46">Litheleodes</emphasis>
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)
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<emphasis id="09C2A366BB6DFFBEFC69FD6FFBAC0EAB" bold="true" box="[929,1025,726,747]" italics="true" pageId="19" pageNumber="46">extricatus</emphasis>
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(Say, 1823)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="73AC2CFFBB6DFFBEFCADFD48FC710F46" box="[869,988,752,774]" pageId="19" pageNumber="46" type="description">
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<paragraph id="3B097F74BB6DFFBEFCADFD48FC710F46" blockId="19.[706,1151,726,774]" box="[869,988,752,774]" pageId="19" pageNumber="46">
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(
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<figureCitation id="A38D63F1BB6DFFBEFCA4FD48FC780F46" box="[876,981,752,774]" captionStart="Fig" captionStartId="8.[197,229,1644,1663]" captionTargetBox="[173,1193,193,1621]" captionTargetId="figure-1@8.[172,1194,192,1621]" captionTargetPageId="8" captionText="Fig. 6. Eleodes species, dorsal habitus. A) E. eschscholtzi, male, B) E. eschscholtzi, female; C) E. caudiferus, D) E. spinipes macrurus, male. Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.10120680" httpUri="https://zenodo.org/record/10120680/files/figure.png" pageId="19" pageNumber="46">Fig. 6C–D</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="73AC2CFFBB6DFFBEFD65FCA2FBCE0F25" pageId="19" pageNumber="46" type="reference_group">
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<paragraph id="3B097F74BB6DFFBEFD65FCA2FC8C0F0B" blockId="19.[685,1171,795,870]" pageId="19" pageNumber="46">
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<taxonomicName id="FCB604F7BB6DFFBEFD65FCA2FB3E0F70" authority="Blaisdell, 1909" authorityName="Blaisdell" authorityYear="1909" box="[685,1171,795,816]" class="Insecta" family="Tenebrionidae" genus="Eleodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="46" phylum="Arthropoda" rank="subSpecies" species="extricatus" subSpecies="arizonensis">
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<emphasis id="09C2A366BB6DFFBEFD65FCA2FC460F70" box="[685,1003,795,816]" italics="true" pageId="19" pageNumber="46">Eleodes extricatus arizonensis</emphasis>
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<bibRefCitation id="5F270285BB6DFFBEFC3EFCA2FB3E0F70" author="Blaisdell" box="[1014,1171,795,816]" pageId="19" pageNumber="46" pagination="1 - 524" refId="ref12543" refString="Blaisdell, F. E. 1909. A monographic revision of the Coleoptera belonging to the Tenebrionidae tribe Eleodiini inhabiting the United States, Lower California, and adjacent islands. Bulletin of the United States National Museum 63: 1 - 524." type="journal article" year="1909">Blaisdell, 1909</bibRefCitation>
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</taxonomicName>
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(
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<figureCitation id="A38D63F1BB6DFFBEFD06FC8FFCB70F0B" box="[718,794,822,843]" captionStart="Fig" captionStartId="8.[197,229,1644,1663]" captionTargetBox="[173,1193,193,1621]" captionTargetId="figure-1@8.[172,1194,192,1621]" captionTargetPageId="8" captionText="Fig. 6. Eleodes species, dorsal habitus. A) E. eschscholtzi, male, B) E. eschscholtzi, female; C) E. caudiferus, D) E. spinipes macrurus, male. Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.10120680" httpUri="https://zenodo.org/record/10120680/files/figure.png" pageId="19" pageNumber="46">Fig. 6D</figureCitation>
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)
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</paragraph>
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<paragraph id="3B097F74BB6DFFBEFD65FCE8FBCE0F25" blockId="19.[685,1171,795,870]" box="[685,1123,848,870]" pageId="19" pageNumber="46">
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<taxonomicName id="FCB604F7BB6DFFBEFD65FCE8FBCE0F25" authority="Haldeman, 1852" authorityName="Haldeman" authorityYear="1852" box="[685,1123,848,870]" class="Insecta" family="Tenebrionidae" genus="Eleodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="46" phylum="Arthropoda" rank="subSpecies" species="extricatus" subSpecies="cognatus">
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<emphasis id="09C2A366BB6DFFBEFD65FCE8FC160F26" box="[685,955,849,870]" italics="true" pageId="19" pageNumber="46">Eleodes extricatus cognatus</emphasis>
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Haldeman, 1852
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="73AC2CFFBB6DFFBEFD0DFCC2FBFD08F1" pageId="19" pageNumber="46" type="diagnosis">
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<paragraph id="3B097F74BB6DFFBEFD0DFCC2FBFD08F1" blockId="19.[685,1173,891,1729]" pageId="19" pageNumber="46">
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<emphasis id="09C2A366BB6DFFBEFD0DFCC2FC830FD0" bold="true" box="[709,814,891,912]" pageId="19" pageNumber="46">Diagnosis.</emphasis>
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Body oblong-ovate. Pronotum subquadrate, gently arcuate laterally, anterior angles always produced forward, angles ranging from acute and projected anterolaterally to broadly rounded, posterior margin not constricted. Prosternum strongly produced behind procoxae, forming posteriorly directled triangular wedge. Elytra asperately to muricately punctured, more or less striate. Profemora feebly to strongly spined in males, sinuate in females. Female genitalia with coxite with inner lobe projected and triangular, gonostyle large, flattened, positioned distally on outer lobe of coxite.
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</paragraph>
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</subSubSection>
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<subSubSection id="73AC2CFFBB6DFFBEFD0DFB0EFCE20976" pageId="19" pageNumber="46" type="distribution">
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<paragraph id="3B097F74BB6DFFBEFD0DFB0EFCE20976" blockId="19.[685,1173,891,1729]" pageId="19" pageNumber="46">
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<emphasis id="09C2A366BB6DFFBEFD0DFB0EFCED088C" bold="true" box="[709,832,1207,1228]" pageId="19" pageNumber="46">Distribution.</emphasis>
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Mohave, Coconino, Navajo, Apache, Yavapai, Gila, Graham, Greenlee, Maricopa, Pinal, Pima, Santa Cruz, and Cochise Counties, Arizona. Texas west to
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<collectingRegion id="F972B196BB6DFFBEFC53FABFFC4D095B" box="[923,992,1286,1307]" country="Mexico" name="Sonora" pageId="19" pageNumber="46">Sonora</collectingRegion>
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,
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<collectingCountry id="43A13FE4BB6DFFBEFC26FABFFB95095B" box="[1006,1080,1286,1307]" name="Mexico" pageId="19" pageNumber="46">Mexico</collectingCountry>
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, north to
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<collectingRegion id="F972B196BB6DFFBEFD65FA98FD580976" box="[685,757,1313,1334]" country="Canada" name="Alberta" pageId="19" pageNumber="46">Alberta</collectingRegion>
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,
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<collectingCountry id="43A13FE4BB6DFFBEFCCAFA98FCE60976" box="[770,843,1313,1334]" name="Canada" pageId="19" pageNumber="46">Canada</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="73AC2CFFBB6DFFBEFD0DFA85FCF30A81" pageId="19" pageNumber="46" type="discussion">
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<paragraph id="3B097F74BB6DFFBEFD0DFA85FCF30A81" blockId="19.[685,1173,891,1729]" pageId="19" pageNumber="46">
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<emphasis id="09C2A366BB6DFFBEFD0DFA85FC840911" bold="true" box="[709,809,1340,1361]" pageId="19" pageNumber="46">Remarks.</emphasis>
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This widespread and variable species is challenging to characterize. It can most easily be recognized by having roughly sculptured elytra, males with spined profemora, and distinctive female genitalia (the produced inner lobe of the coxite separating it from similar species in the subgenus
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<taxonomicName id="FCB604F7BB6DFFBEFD65FA63FC3909AF" authority="Blaisdell" authorityName="Blaisdell" box="[685,916,1498,1519]" class="Insecta" family="Tenebrionidae" genus="Eleodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Melaneleodes">
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<emphasis id="09C2A366BB6DFFBEFD65FA63FC9E09AF" box="[685,819,1498,1519]" italics="true" pageId="19" pageNumber="46">Melaneleodes</emphasis>
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Blaisdell
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</taxonomicName>
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). Found across the state at mid– to high–elevation, three subspecies are known from
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<collectingRegion id="F972B196BB6DFFBEFD16F9B7FC840A63" box="[734,809,1550,1571]" country="United States of America" name="Arizona" pageId="19" pageNumber="46">Arizona</collectingRegion>
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.
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<taxonomicName id="FCB604F7BB6DFFBEFCF9F9B7FB9C0A63" authority="extricatus" box="[817,1073,1550,1571]" class="Insecta" family="Tenebrionidae" genus="Eleodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="46" phylum="Arthropoda" rank="subSpecies" species="extricatus" subSpecies="extricatus">
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<emphasis id="09C2A366BB6DFFBEFCF9F9B7FB9C0A63" box="[817,1073,1550,1571]" italics="true" pageId="19" pageNumber="46">Eleodes extricatus extricatus</emphasis>
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</taxonomicName>
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is coarsely to scabrously punctured and from the central and western mountains.
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<taxonomicName id="FCB604F7BB6DFFBEFCB8F9FAFB390A18" authorityName="Blaisdell" authorityYear="1909" box="[880,1172,1603,1624]" class="Insecta" family="Tenebrionidae" genus="Eleodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="46" phylum="Arthropoda" rank="subSpecies" species="extricatus" subSpecies="arizonensis">
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<emphasis id="09C2A366BB6DFFBEFCB8F9FAFB390A18" box="[880,1172,1603,1624]" italics="true" pageId="19" pageNumber="46">Eleodes extricatus arizonensis</emphasis>
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</taxonomicName>
|
|||
|
|
has elytral striae more or less sulcate and is from the southeastern “sky islands”.
|
|||
|
|
<taxonomicName id="FCB604F7BB6DFFBEFC2AF9C1FCA50AE7" authorityName="Haldeman" authorityYear="1852" class="Insecta" family="Tenebrionidae" genus="Eleodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="46" phylum="Arthropoda" rank="subSpecies" species="extricatus" subSpecies="cognatus">
|
|||
|
|
<emphasis id="09C2A366BB6DFFBEFC2AF9C1FCA50AE7" italics="true" pageId="19" pageNumber="46">Eleodes extricatus cognatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is more robust and less scabrous and widely distributed.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|