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<mods:title id="E1C5F533B20899476E9E1D2B668B4E6D">Taxonomic notes on Sarju Ghauri, 1977 (Hemiptera: Heteroptera: Pentatomidae) with description of a new species from India</mods:title>
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<mods:affiliation id="DD4BAE0421423958670FC3D3E4997256">National Bureau of Agricultural Insect Resources, Bangalore, 560024, India. &amp; https: // orcid. org / 0000 - 0001 - 7306 - 082 X</mods:affiliation>
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<taxonomicName id="4C5C7088FFC2FFA7BE99F965FE75F95D" authority="Ghauri, 1977" authorityName="Ghauri" authorityYear="1977" box="[151,390,1734,1761]" class="Insecta" family="Pentatomidae" genus="Sarju" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="genus">
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<emphasis id="B928D719FFC2FFA7BE99F965FF29F95C" bold="true" box="[151,218,1735,1761]" italics="true" pageId="1" pageNumber="284">Sarju</emphasis>
<bibRefCitation id="EFCD76FAFFC2FFA7BEEFF964FE75F95D" author="Ghauri, M. S. K." box="[225,390,1734,1760]" pageId="1" pageNumber="284" pagination="9 - 27" refId="ref12389" refString="Ghauri, M. S. K. (1977) Sarju - a new genus of Halyini (Heteroptera, Pentatomidae, Pentatominae) with new species. Tu ¨ rkiye Bitki Koruma Dergisi, 1 (1), 9 - 27." type="journal article" year="1977">Ghauri, 1977</bibRefCitation>
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<treatmentCitation id="0AFD2D1AFFC2FFA7BE99F8A9FE5AF89E" author="Ghauri, M. S. K." box="[151,425,1803,1827]" page="11" pageId="1" pageNumber="284" year="1977">
<taxonomicName id="4C5C7088FFC2FFA7BE99F8A9FE5AF89E" authority="Ghauri, 1977: 10 - 11" authorityName="Ghauri" authorityPageNumber="10 - 11" authorityYear="1977" box="[151,425,1803,1827]" class="Insecta" family="Pentatomidae" genus="Sarju" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="genus">
<emphasis id="B928D719FFC2FFA7BE99F8A9FF3DF89F" box="[151,206,1803,1826]" italics="true" pageId="1" pageNumber="284">Sarju</emphasis>
<bibRefCitation id="EFCD76FAFFC2FFA7BEDAF8A9FE5AF89E" author="Ghauri, M. S. K." box="[212,425,1803,1827]" pageId="1" pageNumber="284" pagination="9 - 27" refId="ref12389" refString="Ghauri, M. S. K. (1977) Sarju - a new genus of Halyini (Heteroptera, Pentatomidae, Pentatominae) with new species. Tu ¨ rkiye Bitki Koruma Dergisi, 1 (1), 9 - 27." type="journal article" year="1977">Ghauri, 1977: 1011</bibRefCitation>
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;
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<typeStatus id="54E7B5A9FFC2FFA7BFBDF8AEFE16F899" box="[435,485,1804,1828]" pageId="1" pageNumber="284">Type</typeStatus>
species:
<taxonomicName id="4C5C7088FFC2FFA7BC4CF8A9FC74F89E" authority="Westwood, 1837" authorityName="Westwood" authorityYear="1837" box="[578,903,1803,1827]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="1" pageNumber="284" phylum="Chordata" rank="species" species="obscura">
<emphasis id="B928D719FFC2FFA7BC4CF8A9FD26F89E" box="[578,725,1803,1827]" italics="true" pageId="1" pageNumber="284">Halys obscura</emphasis>
<bibRefCitation id="EFCD76FAFFC2FFA7BCD5F8A9FC74F89E" author="Westwood, J. O." box="[731,903,1803,1827]" pageId="1" pageNumber="284" pagination="1 - 46" refId="ref13046" refString="Westwood, J. O. (1837) s. n. In: Hope, F. W. (Eds.), A Catalogue of Hemiptera in the collection of the Rev. F. W. Hope, M. A. with short Latin diagnoses of the new species. Pt. 1. Printed by J. C. Bridgewater, London, pp. 1 - 46." type="book chapter" year="1837">Westwood, 1837</bibRefCitation>
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, by original designation.
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<treatmentCitation id="0AFD2D1AFFC2FFA7BE99F888FE8FF8FF" author="Rider, D. A." box="[151,380,1834,1858]" page="311" pageId="1" pageNumber="284" year="2006">
<taxonomicName id="4C5C7088FFC2FFA7BE99F888FE8FF8FF" authority="Rider, 2006: 311" authorityName="Rider" authorityPageNumber="311" authorityYear="2006" box="[151,380,1834,1858]" class="Insecta" family="Pentatomidae" genus="Sarju" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="genus">
<emphasis id="B928D719FFC2FFA7BE99F888FF3DF8FC" box="[151,206,1834,1857]" italics="true" pageId="1" pageNumber="284">Sarju</emphasis>
<bibRefCitation id="EFCD76FAFFC2FFA7BEDAF888FE8FF8FF" author="Rider, D. A." box="[212,380,1834,1858]" pageId="1" pageNumber="284" refId="ref12695" refString="Rider, D. A. (2006) Family Pentatomidae. In: Aukema, B. &amp; Rieger, Ch. (Eds.), Catalogue of the Heteroptera of the Palaearctic Region. Vol. 5. The Netherlands Entomological Society, Amsterdam, pp. 233 - 402." type="book" year="2006">Rider, 2006: 311</bibRefCitation>
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(Palearctic catalog).
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<paragraph id="8BE30B0BFFC2FFA4BE99F8D2FD6EFED8" blockId="1.[151,1437,1904,2039]" lastBlockId="2.[151,1437,151,2014]" lastPageId="2" lastPageNumber="285" pageId="1" pageNumber="284">
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Redescription.
<emphasis id="B928D719FFC2FFA7BF44F8D2FE44F837" bold="true" box="[330,439,1904,1930]" italics="true" pageId="1" pageNumber="284">Structure</emphasis>
.
</emphasis>
<emphasis id="B928D719FFC2FFA7BFCAF8D2FDF2F837" box="[452,513,1904,1930]" italics="true" pageId="1" pageNumber="284">Head</emphasis>
(
<figureCitation id="1367178EFFC2FFA7BC00F8D2FD83F837" box="[526,624,1904,1930]" captionStart="FIGURES 38" captionStartId="5.[151,264,1565,1589]" captionTargetBox="[156,1428,667,1437]" captionTargetId="figure-176@5.[151,1436,633,1541]" captionTargetPageId="5" captionText="FIGURES 38. Sarju brevirostrata sp. nov. 3, Holotype (headdorsal); 4, Paratype (headdorsal); 5, Holotype—label; 6, pronotum; 7, External scent efferent system; 8, Genital capsule (dorsal). Scale: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4664142" httpUri="https://zenodo.org/record/4664142/files/figure.png" pageId="1" pageNumber="284">Figs 3, 4</figureCitation>
,
<figureCitation id="1367178EFFC2FFA7BC74F8D2FD6AF837" box="[634,665,1904,1930]" captionStart="FIGURES 2227" captionStartId="10.[151,264,1926,1950]" captionTargetBox="[187,1401,1156,1867]" captionTargetId="figure-440@10.[170,1416,1136,1902]" captionTargetPageId="10" captionText="FIGURES 2227. Sarju farida Ghauri. 22, head (dorsal); 23, pronotum; 24, external scent efferent system; 25, Genital capsule (dorsal); 26, Genital capsule (ventral); 27, Genital capsule (caudal). Scale: 0.5 mm. Lettering: p—peritreme;tl—triangular lobelike projection." figureDoi="http://doi.org/10.5281/zenodo.4664154" httpUri="https://zenodo.org/record/4664154/files/figure.png" pageId="1" pageNumber="284">22</figureCitation>
,
<figureCitation id="1367178EFFC2FFA7BCADF8D2FD33F837" box="[675,704,1904,1930]" captionStart="FIGURES 3944" captionStartId="14.[151,264,1651,1675]" captionTargetBox="[171,1428,768,1584]" captionTargetId="figure-267@14.[151,1436,740,1628]" captionTargetPageId="14" captionText="FIGURES 3944. Sarju nigricollis (Westwood). 39, head (dorsal); 40, pronotum; 41, external scent efferent system; 42, Genital capsule (dorsal); 43, Genital capsule (ventral); 44, Genital capsule (caudal). Scale: 0.5 mm. Lettering: p—peritreme." figureDoi="http://doi.org/10.5281/zenodo.4664160" httpUri="https://zenodo.org/record/4664160/files/figure.png" pageId="1" pageNumber="284">39</figureCitation>
) nearly parallel to body axis, usually longer than wide (sometimes as long as wide); lateral margins with one or two pairs of teeth, first pair subapical to mandibular plates, always present, rounded or quadrangular and the second pair in front of compound eyes, sometimes indistinct or absent. Mandibular plates variable, straight or obliquely bend downwards at apex, meeting or not meeting in front of clypeus, sometimes partly covering apex of clypeus, apex of mandibular plates rounded or acuminate, longer than clypeus, sometimes nearly equal to clypeus, but not shorter; clypeus open or closed apically; dorsal surface of head usually more or less flattened, sometimes apical region bend downwards. Compound eyes rounded and protruding out. Antennae five segmented, slender with second segment usually bowed and thickened at apex, sometimes thin and straight. Anterior apex of bacculae blunt, quadrangular, not rounded; length of labium variable, reaching meta coxae to middle of fourth abdominal segment.
</paragraph>
<paragraph id="8BE30B0BFFC1FFA4BEC9FECDFAFAFE10" blockId="2.[151,1437,151,2014]" pageId="2" pageNumber="285">
<emphasis id="B928D719FFC1FFA4BEC9FECDFEC5FE35" box="[199,310,367,392]" italics="true" pageId="2" pageNumber="285">Pronotum</emphasis>
(
<figureCitation id="1367178EFFC1FFA4BF48FECDFE7CFE34" box="[326,399,367,393]" captionStart="FIGURES 38" captionStartId="5.[151,264,1565,1589]" captionTargetBox="[156,1428,667,1437]" captionTargetId="figure-176@5.[151,1436,633,1541]" captionTargetPageId="5" captionText="FIGURES 38. Sarju brevirostrata sp. nov. 3, Holotype (headdorsal); 4, Paratype (headdorsal); 5, Holotype—label; 6, pronotum; 7, External scent efferent system; 8, Genital capsule (dorsal). Scale: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4664142" httpUri="https://zenodo.org/record/4664142/files/figure.png" pageId="2" pageNumber="285">Figs 6</figureCitation>
,
<figureCitation id="1367178EFFC1FFA4BF92FECDFE49FE34" box="[412,442,367,393]" captionStart="FIGURES 2227" captionStartId="10.[151,264,1926,1950]" captionTargetBox="[187,1401,1156,1867]" captionTargetId="figure-440@10.[170,1416,1136,1902]" captionTargetPageId="10" captionText="FIGURES 2227. Sarju farida Ghauri. 22, head (dorsal); 23, pronotum; 24, external scent efferent system; 25, Genital capsule (dorsal); 26, Genital capsule (ventral); 27, Genital capsule (caudal). Scale: 0.5 mm. Lettering: p—peritreme;tl—triangular lobelike projection." figureDoi="http://doi.org/10.5281/zenodo.4664154" httpUri="https://zenodo.org/record/4664154/files/figure.png" pageId="2" pageNumber="285">23</figureCitation>
,
<figureCitation id="1367178EFFC1FFA4BFC9FECDFE17FE34" box="[455,484,367,393]" captionStart="FIGURES 3944" captionStartId="14.[151,264,1651,1675]" captionTargetBox="[171,1428,768,1584]" captionTargetId="figure-267@14.[151,1436,740,1628]" captionTargetPageId="14" captionText="FIGURES 3944. Sarju nigricollis (Westwood). 39, head (dorsal); 40, pronotum; 41, external scent efferent system; 42, Genital capsule (dorsal); 43, Genital capsule (ventral); 44, Genital capsule (caudal). Scale: 0.5 mm. Lettering: p—peritreme." figureDoi="http://doi.org/10.5281/zenodo.4664160" httpUri="https://zenodo.org/record/4664160/files/figure.png" pageId="2" pageNumber="285">40</figureCitation>
). Anterior pronotal margin concave, anterolateral angles with small tooth, anterolateral margins concave (or bisinuate by
<bibRefCitation id="EFCD76FAFFC1FFA4BC4CFE31FD20FE10" author="Ghauri, M. S. K." box="[578,723,403,429]" pageId="2" pageNumber="285" pagination="9 - 27" refId="ref12389" refString="Ghauri, M. S. K. (1977) Sarju - a new genus of Halyini (Heteroptera, Pentatomidae, Pentatominae) with new species. Tu ¨ rkiye Bitki Koruma Dergisi, 1 (1), 9 - 27." type="journal article" year="1977">Ghauri 1977</bibRefCitation>
). Humeri variable, mostly short blunt or nodulose.
</paragraph>
<paragraph id="8BE30B0BFFC1FFA4BEC9FE15FA8AFE48" blockId="2.[151,1437,151,2014]" pageId="2" pageNumber="285">
<emphasis id="B928D719FFC1FFA4BEC9FE15FEC6FE6C" box="[199,309,439,465]" italics="true" pageId="2" pageNumber="285">Scutellum</emphasis>
(
<figureCitation id="1367178EFFC1FFA4BF4BFE15FE7EFE6C" box="[325,397,439,465]" captionStart="FIGURES 12" captionStartId="4.[151,264,1627,1651]" captionTargetBox="[183,1411,534,1581]" captionTargetId="figure-180@4.[156,1431,520,1604]" captionTargetPageId="4" captionText="FIGURES 12. Sarju brevirostrata sp. nov.—Habitus. 1, dorsal; 2, ventral. Scale: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4664138" httpUri="https://zenodo.org/record/4664138/files/figure.png" pageId="2" pageNumber="285">Figs 1</figureCitation>
,
<figureCitation id="1367178EFFC1FFA4BF92FE15FE17FE6C" box="[412,484,439,465]" captionStart="FIGURES 1821" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[199,1400,211,1909]" captionTargetId="figure-17@9.[179,1408,181,1920]" captionTargetPageId="9" captionText="FIGURES 1821. 1819. Sarju farida Ghauri—Habitus. 18, dorsal; 19, ventral. 2021. Sarju nigricollis (Westwood)—Habitus. 20, dorsal; 21, ventral. Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4664152" httpUri="https://zenodo.org/record/4664152/files/figure.png" pageId="2" pageNumber="285">18, 20</figureCitation>
) subtriangular, longer than wide at base, abruptly narrowing in posterior third, apex rounded; apex of scutellum much shorter than anterodistal angles of corium; disc of scutellum more or less flat.
</paragraph>
<paragraph id="8BE30B0BFFC1FFA4BEC9FE5DFB7DFD80" blockId="2.[151,1437,151,2014]" pageId="2" pageNumber="285">
<emphasis id="B928D719FFC1FFA4BEC9FE5DFECEFDA4" box="[199,317,511,537]" italics="true" pageId="2" pageNumber="285">Hemelytra</emphasis>
. Clavus narrowly triangular, corium much surpassing apex of scutellum. Membrane translucent, broadly rounded apically, usually exceeding apex of abdomen, without reticulate venation.
</paragraph>
<paragraph id="8BE30B0BFFC1FFA4BEC9FDE5FB49FD14" blockId="2.[151,1437,151,2014]" pageId="2" pageNumber="285">
<emphasis id="B928D719FFC1FFA4BEC9FDE5FDD8FDDD" box="[199,555,583,609]" italics="true" pageId="2" pageNumber="285">Thoracic pleurites and sternum</emphasis>
. External scent efferent system typical of
<taxonomicName id="4C5C7088FFC1FFA4BA02FDE5FB93FDDC" authorityName="Amyot &amp; Serville" authorityYear="1843" box="[1036,1120,583,609]" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="285" phylum="Arthropoda" rank="tribe" tribe="Halyini">Halyini</taxonomicName>
<typeStatus id="54E7B5A9FFC1FFA4BA66FDE5FB6BFDDC" box="[1128,1176,583,609]" pageId="2" pageNumber="285">type</typeStatus>
(
<bibRefCitation id="EFCD76FAFFC1FFA4BAA6FDE5FADDFDDC" author="Salini, S." box="[1192,1326,583,609]" pageId="2" pageNumber="285" pagination="351 - 375" refId="ref12853" refString="Salini, S. (2019) Revision of the genus Halys (Hemiptera: Pentatomidae) with description of a new species from India. Zootaxa, 4586 (2), 351 - 375. https: // doi. org / 10.11646 / zootaxa. 4586.2.9" type="journal article" year="2019">Salini 2019</bibRefCitation>
, Fig. 91); peritreme spout-shaped (crescent-shaped) usually reaching middle of metapleuron with well developed evaporatorium (
<figureCitation id="1367178EFFC1FFA4BED5FD2DFED7FD14" box="[219,292,655,681]" captionStart="FIGURES 38" captionStartId="5.[151,264,1565,1589]" captionTargetBox="[156,1428,667,1437]" captionTargetId="figure-176@5.[151,1436,633,1541]" captionTargetPageId="5" captionText="FIGURES 38. Sarju brevirostrata sp. nov. 3, Holotype (headdorsal); 4, Paratype (headdorsal); 5, Holotype—label; 6, pronotum; 7, External scent efferent system; 8, Genital capsule (dorsal). Scale: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4664142" httpUri="https://zenodo.org/record/4664142/files/figure.png" pageId="2" pageNumber="285">Figs 7</figureCitation>
,
<figureCitation id="1367178EFFC1FFA4BF3EFD2DFEBDFD14" box="[304,334,655,681]" captionStart="FIGURES 2227" captionStartId="10.[151,264,1926,1950]" captionTargetBox="[187,1401,1156,1867]" captionTargetId="figure-440@10.[170,1416,1136,1902]" captionTargetPageId="10" captionText="FIGURES 2227. Sarju farida Ghauri. 22, head (dorsal); 23, pronotum; 24, external scent efferent system; 25, Genital capsule (dorsal); 26, Genital capsule (ventral); 27, Genital capsule (caudal). Scale: 0.5 mm. Lettering: p—peritreme;tl—triangular lobelike projection." figureDoi="http://doi.org/10.5281/zenodo.4664154" httpUri="https://zenodo.org/record/4664154/files/figure.png" pageId="2" pageNumber="285">24</figureCitation>
,
<figureCitation id="1367178EFFC1FFA4BF54FD2DFE8AFD14" box="[346,377,655,681]" captionStart="FIGURES 3944" captionStartId="14.[151,264,1651,1675]" captionTargetBox="[171,1428,768,1584]" captionTargetId="figure-267@14.[151,1436,740,1628]" captionTargetPageId="14" captionText="FIGURES 3944. Sarju nigricollis (Westwood). 39, head (dorsal); 40, pronotum; 41, external scent efferent system; 42, Genital capsule (dorsal); 43, Genital capsule (ventral); 44, Genital capsule (caudal). Scale: 0.5 mm. Lettering: p—peritreme." figureDoi="http://doi.org/10.5281/zenodo.4664160" httpUri="https://zenodo.org/record/4664160/files/figure.png" pageId="2" pageNumber="285">41</figureCitation>
). Metathoracic spiracle long, narrow and distinctly visible in ventral view.
</paragraph>
<paragraph id="8BE30B0BFFC1FFA4BEC9FD11FDAFFCA8" blockId="2.[151,1437,151,2014]" pageId="2" pageNumber="285">
<emphasis id="B928D719FFC1FFA4BEC9FD11FF0FFD71" box="[199,252,691,716]" italics="true" pageId="2" pageNumber="285">Legs</emphasis>
. Slender. Femora unarmed, sparsely pubescent, cylindrical, rounded in cross section. All tibiae slender than respective femora. All tibiae pubescent, dorsally with narrow, median, longitudinal groove. All tarsal segments dorsally regularly rounded, not grooved.
</paragraph>
<paragraph id="8BE30B0BFFC1FFA4BEC9FCBDFDF0FC18" blockId="2.[151,1437,151,2014]" pageId="2" pageNumber="285">
<emphasis id="B928D719FFC1FFA4BEC9FCBDFE45FC84" box="[199,438,799,825]" italics="true" pageId="2" pageNumber="285">Pregenital abdomen.</emphasis>
Narrower than pronotum across humeri. Connexivum broad and exposed most of its width. Posterolateral angles of each ventrite angulate or with minute spinous process. Abdominal venter convex medially, with median longitudinal groove, shallow, sometimes absent. Ventrite III without spine or tubercle, usually with slight groove medially.
</paragraph>
<paragraph id="8BE30B0BFFC1FFA4BEC9FC0DFB9BFAE8" blockId="2.[151,1437,151,2014]" pageId="2" pageNumber="285">
<emphasis id="B928D719FFC1FFA4BEC9FC0DFE81FC74" box="[199,370,943,969]" italics="true" pageId="2" pageNumber="285">Male genitalia.</emphasis>
Genital capsule quadrangular dorsal rim variable (
<figureCitation id="1367178EFFC1FFA4BDAEFC0DFC14FC74" box="[928,999,943,969]" captionStart="FIGURES 38" captionStartId="5.[151,264,1565,1589]" captionTargetBox="[156,1428,667,1437]" captionTargetId="figure-176@5.[151,1436,633,1541]" captionTargetPageId="5" captionText="FIGURES 38. Sarju brevirostrata sp. nov. 3, Holotype (headdorsal); 4, Paratype (headdorsal); 5, Holotype—label; 6, pronotum; 7, External scent efferent system; 8, Genital capsule (dorsal). Scale: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4664142" httpUri="https://zenodo.org/record/4664142/files/figure.png" pageId="2" pageNumber="285">Figs 8</figureCitation>
,
<figureCitation id="1367178EFFC1FFA4BDFCFC0CFBE3FC74" box="[1010,1040,942,969]" captionStart="FIGURES 2227" captionStartId="10.[151,264,1926,1950]" captionTargetBox="[187,1401,1156,1867]" captionTargetId="figure-440@10.[170,1416,1136,1902]" captionTargetPageId="10" captionText="FIGURES 2227. Sarju farida Ghauri. 22, head (dorsal); 23, pronotum; 24, external scent efferent system; 25, Genital capsule (dorsal); 26, Genital capsule (ventral); 27, Genital capsule (caudal). Scale: 0.5 mm. Lettering: p—peritreme;tl—triangular lobelike projection." figureDoi="http://doi.org/10.5281/zenodo.4664154" httpUri="https://zenodo.org/record/4664154/files/figure.png" pageId="2" pageNumber="285">25</figureCitation>
,
<figureCitation id="1367178EFFC1FFA4BA15FC0DFBC8FC74" box="[1051,1083,943,969]" captionStart="FIGURES 3944" captionStartId="14.[151,264,1651,1675]" captionTargetBox="[171,1428,768,1584]" captionTargetId="figure-267@14.[151,1436,740,1628]" captionTargetPageId="14" captionText="FIGURES 3944. Sarju nigricollis (Westwood). 39, head (dorsal); 40, pronotum; 41, external scent efferent system; 42, Genital capsule (dorsal); 43, Genital capsule (ventral); 44, Genital capsule (caudal). Scale: 0.5 mm. Lettering: p—peritreme." figureDoi="http://doi.org/10.5281/zenodo.4664160" httpUri="https://zenodo.org/record/4664160/files/figure.png" pageId="2" pageNumber="285">42</figureCitation>
); dorsal sclerite lacking; ventral rim with deep or shallow median excavation cup-shaped or broad U-shaped (
<figureCitation id="1367178EFFC1FFA4BA07FC71FBBCFC50" box="[1033,1103,979,1005]" captionStart="FIGURES 914" captionStartId="6.[151,264,1536,1560]" captionTargetBox="[184,1422,633,1489]" captionTargetId="figure-216@6.[151,1436,633,1513]" captionTargetPageId="6" captionText="FIGURES 914. Sarju brevirostrata sp. nov. 9, Genital capsule (ventral); 10, Genital capsule (caudal); 1113, paramere (different planes); 14, articulatory apparatus. Scale: 0.25 mm. Lettering: cp—capitate process; mb—mushroom shaped bodies." figureDoi="http://doi.org/10.5281/zenodo.4664144" httpUri="https://zenodo.org/record/4664144/files/figure.png" pageId="2" pageNumber="285">Fig. 9</figureCitation>
) or V-shaped or arcuate with (
<figureCitation id="1367178EFFC1FFA4BE91FC55FF0BFBAC" box="[159,248,1015,1041]" captionStart="FIGURES 2227" captionStartId="10.[151,264,1926,1950]" captionTargetBox="[187,1401,1156,1867]" captionTargetId="figure-440@10.[170,1416,1136,1902]" captionTargetPageId="10" captionText="FIGURES 2227. Sarju farida Ghauri. 22, head (dorsal); 23, pronotum; 24, external scent efferent system; 25, Genital capsule (dorsal); 26, Genital capsule (ventral); 27, Genital capsule (caudal). Scale: 0.5 mm. Lettering: p—peritreme;tl—triangular lobelike projection." figureDoi="http://doi.org/10.5281/zenodo.4664154" httpUri="https://zenodo.org/record/4664154/files/figure.png" pageId="2" pageNumber="285">Figs 26</figureCitation>
,
<figureCitation id="1367178EFFC1FFA4BF08FC55FED0FBAC" box="[262,291,1015,1041]" captionStart="FIGURES 3944" captionStartId="14.[151,264,1651,1675]" captionTargetBox="[171,1428,768,1584]" captionTargetId="figure-267@14.[151,1436,740,1628]" captionTargetPageId="14" captionText="FIGURES 3944. Sarju nigricollis (Westwood). 39, head (dorsal); 40, pronotum; 41, external scent efferent system; 42, Genital capsule (dorsal); 43, Genital capsule (ventral); 44, Genital capsule (caudal). Scale: 0.5 mm. Lettering: p—peritreme." figureDoi="http://doi.org/10.5281/zenodo.4664160" httpUri="https://zenodo.org/record/4664160/files/figure.png" pageId="2" pageNumber="285">43</figureCitation>
) or without sinuate margin; caudal angles rounded or broad and truncate, produced or not produced medially. Paramere (
<figureCitation id="1367178EFFC1FFA4BF71FBB9FE0CFB88" box="[383,511,1051,1077]" captionStart="FIGURES 914" captionStartId="6.[151,264,1536,1560]" captionTargetBox="[184,1422,633,1489]" captionTargetId="figure-216@6.[151,1436,633,1513]" captionTargetPageId="6" captionText="FIGURES 914. Sarju brevirostrata sp. nov. 9, Genital capsule (ventral); 10, Genital capsule (caudal); 1113, paramere (different planes); 14, articulatory apparatus. Scale: 0.25 mm. Lettering: cp—capitate process; mb—mushroom shaped bodies." figureDoi="http://doi.org/10.5281/zenodo.4664144" httpUri="https://zenodo.org/record/4664144/files/figure.png" pageId="2" pageNumber="285">Figs 1113</figureCitation>
,
<figureCitation id="1367178EFFC1FFA4BC05FBB9FDA7FB88" box="[523,596,1051,1077]" captionStart="FIGURES 2833" captionStartId="11.[151,264,1472,1496]" captionTargetBox="[168,1405,436,1436]" captionTargetId="figure-90@11.[151,1436,417,1449]" captionTargetPageId="11" captionText="FIGURES 2833. Sarju farida Ghauri. 2829, paramere; 30, articulatory apparatus; 31, phallus (ventral); 32, phallus (dorsal); 33, phallus (lateral). Scale: 0.25 mm. Lettering: cp—capitate process; dcp—dorsal conjunctival processes; mcp—median conjunctival processes; pa—processes of aedeagus; sp—setose process by Ghauri, 1977; vcp—ventral conjunctival processes; vt—ventral tubercle." figureDoi="http://doi.org/10.5281/zenodo.4664156" httpUri="https://zenodo.org/record/4664156/files/figure.png" pageId="2" pageNumber="285">2829</figureCitation>
,
<figureCitation id="1367178EFFC1FFA4BC51FBB8FD54FB88" box="[607,679,1050,1077]" captionStart="FIGURES 4550" captionStartId="15.[151,264,1840,1864]" captionTargetBox="[170,1436,777,1786]" captionTargetId="figure-274@15.[151,1436,777,1817]" captionTargetPageId="15" captionText="FIGURES 4550. Sarju nigricollis (Westwood). 4546, paramere (different planes); 47, articulatory apparatus; 48, phallus (ventral); 49, phallus (dorsal); 50, phallus (lateral). Scale: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4664162" httpUri="https://zenodo.org/record/4664162/files/figure.png" pageId="2" pageNumber="285">4546</figureCitation>
) with broad and thick crown, parameral crown with transverse narrow ridge, inner lobe variable, produced into elongate finger-like process or rounded apex or abruptly narrow or gradually narrowed towards apex; stem short. A short process bearing setae (setose process by
<bibRefCitation id="EFCD76FAFFC1FFA4BAC7FBC1FAAEFBC0" author="Ghauri, M. S. K." box="[1225,1373,1123,1149]" pageId="2" pageNumber="285" pagination="9 - 27" refId="ref12389" refString="Ghauri, M. S. K. (1977) Sarju - a new genus of Halyini (Heteroptera, Pentatomidae, Pentatominae) with new species. Tu ¨ rkiye Bitki Koruma Dergisi, 1 (1), 9 - 27." type="journal article" year="1977">Ghauri, 1977</bibRefCitation>
) near posterior end on inner side, small, thumb-shaped or conical or apically rounded. Phallus with phallotheca sclerotized and constricted towards base (proximal end), a short angular process (ventral tubercle, vt) just before proximal end of phallotheca ventromedially (
<figureCitation id="1367178EFFC1FFA4BC2FFB6DFD85FB54" box="[545,630,1231,1257]" captionStart="FIGURES 1517" captionStartId="7.[151,264,1112,1136]" captionTargetBox="[166,1418,340,1073]" captionTargetId="figure-77@7.[151,1436,308,1088]" captionTargetPageId="7" captionText="FIGURES 1517. Sarju brevirostrata sp. nov. 15, phallus (ventral); 16, phallus (dorsal); 17, phallus (lateral). Scale: 0.1 mm. Lettering: ae—aedeagus; dcp—dorsal conjunctival processes; mcp—median conjunctival processes; pa—processes of aedeagus; vcp—ventral conjunctival processes; vt—ventral tubercle." figureDoi="http://doi.org/10.5281/zenodo.4664150" httpUri="https://zenodo.org/record/4664150/files/figure.png" pageId="2" pageNumber="285">Figs 17</figureCitation>
,
<figureCitation id="1367178EFFC1FFA4BC8FFB6DFD6CFB54" box="[641,671,1231,1257]" captionStart="FIGURES 2833" captionStartId="11.[151,264,1472,1496]" captionTargetBox="[168,1405,436,1436]" captionTargetId="figure-90@11.[151,1436,417,1449]" captionTargetPageId="11" captionText="FIGURES 2833. Sarju farida Ghauri. 2829, paramere; 30, articulatory apparatus; 31, phallus (ventral); 32, phallus (dorsal); 33, phallus (lateral). Scale: 0.25 mm. Lettering: cp—capitate process; dcp—dorsal conjunctival processes; mcp—median conjunctival processes; pa—processes of aedeagus; sp—setose process by Ghauri, 1977; vcp—ventral conjunctival processes; vt—ventral tubercle." figureDoi="http://doi.org/10.5281/zenodo.4664156" httpUri="https://zenodo.org/record/4664156/files/figure.png" pageId="2" pageNumber="285">33</figureCitation>
,
<figureCitation id="1367178EFFC1FFA4BCA4FB6CFD39FB54" box="[682,714,1230,1257]" captionStart="FIGURES 4550" captionStartId="15.[151,264,1840,1864]" captionTargetBox="[170,1436,777,1786]" captionTargetId="figure-274@15.[151,1436,777,1817]" captionTargetPageId="15" captionText="FIGURES 4550. Sarju nigricollis (Westwood). 4546, paramere (different planes); 47, articulatory apparatus; 48, phallus (ventral); 49, phallus (dorsal); 50, phallus (lateral). Scale: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4664162" httpUri="https://zenodo.org/record/4664162/files/figure.png" pageId="2" pageNumber="285">50</figureCitation>
); three pairs of conjunctival processes-dorsal pair of conjunctival processes (dcp) short, membranous or semi sclerotised, median conjunctival processes (mcp) membranous, trilobate or with three lobes, ventral pair (vcp) and processes of aedeagus (=penial lobes) (pa) semisclerotised or sclerotised; aedeagus (ae) moderately elongate, tubular, usually shorter than processes of aedeagus.
</paragraph>
<paragraph id="8BE30B0BFFC1FFA4BEC9FAFDFEA2FA58" blockId="2.[151,1437,151,2014]" pageId="2" pageNumber="285">
<emphasis id="B928D719FFC1FFA4BEC9FAFDFE7FFAC4" box="[199,396,1375,1401]" italics="true" pageId="2" pageNumber="285">Female genitalia.</emphasis>
Terminalia (
<figureCitation id="1367178EFFC1FFA4BC16FAFDFD9DFAC4" box="[536,622,1375,1401]" captionStart="FIGURES 3438" captionStartId="12.[151,264,1821,1845]" captionTargetBox="[182,1429,799,1793]" captionTargetId="figure-250@12.[151,1436,777,1797]" captionTargetPageId="12" captionText="FIGURES 3438. Sarju farida Ghauri. 34, terminalia (dorsal); 35, spermatheca; 36, apical receptacle; 37, terminalia (ventral); 38, valvifers VIII. 3435: Scale: 0.5 mm; 3638: Scale: 0.25 mm. Lettering: lt IX—laterotergite IX; lt VIII—laterotergite VIII; rs—ring sclerite; sso—sclerite guarding spermathecal opening; vlf VIII—valvifers VIII." figureDoi="http://doi.org/10.5281/zenodo.4664158" httpUri="https://zenodo.org/record/4664158/files/figure.png" pageId="2" pageNumber="285">Figs 34</figureCitation>
,
<figureCitation id="1367178EFFC1FFA4BC77FAFCFD6BFAC4" box="[633,664,1374,1401]" captionStart="FIGURES 5154" captionStartId="16.[151,264,1903,1927]" captionTargetBox="[155,1422,857,1865]" captionTargetId="figure-290@16.[151,1436,813,1880]" captionTargetPageId="16" captionText="FIGURES 5154. Sarju nigricollis (Westwood). 51, terminalia (dorsal); 52, spermatheca; 53, terminalia (ventral); 54, valvifers VIII. 5152: Scale: 0.5 mm; 5354: Scale: 0.25 mm. Lettering: lt IX—laterotergite IX; lt VIII—laterotergite VIII; rs—ring sclerite; sso—sclerite guarding spermathecal opening; vlf VIII—valvifers VIII." figureDoi="http://doi.org/10.5281/zenodo.4664164" httpUri="https://zenodo.org/record/4664164/files/figure.png" pageId="2" pageNumber="285">51</figureCitation>
). Shape of Valvifers VIII variable, sometimes posteriorly produced in to finger-like process (
<figureCitation id="1367178EFFC1FFA4BF98FA21FDE9FA20" box="[406,538,1410,1437]" captionStart="FIGURES 5154" captionStartId="16.[151,264,1903,1927]" captionTargetBox="[155,1422,857,1865]" captionTargetId="figure-290@16.[151,1436,813,1880]" captionTargetPageId="16" captionText="FIGURES 5154. Sarju nigricollis (Westwood). 51, terminalia (dorsal); 52, spermatheca; 53, terminalia (ventral); 54, valvifers VIII. 5152: Scale: 0.5 mm; 5354: Scale: 0.25 mm. Lettering: lt IX—laterotergite IX; lt VIII—laterotergite VIII; rs—ring sclerite; sso—sclerite guarding spermathecal opening; vlf VIII—valvifers VIII." figureDoi="http://doi.org/10.5281/zenodo.4664164" httpUri="https://zenodo.org/record/4664164/files/figure.png" pageId="2" pageNumber="285">Figs 51, 54</figureCitation>
), triangulin sometimes exposed externally. Spermatheca (
<figureCitation id="1367178EFFC1FFA4BAABFA21FB0EFA20" box="[1189,1277,1410,1437]" captionStart="FIGURES 3438" captionStartId="12.[151,264,1821,1845]" captionTargetBox="[182,1429,799,1793]" captionTargetId="figure-250@12.[151,1436,777,1797]" captionTargetPageId="12" captionText="FIGURES 3438. Sarju farida Ghauri. 34, terminalia (dorsal); 35, spermatheca; 36, apical receptacle; 37, terminalia (ventral); 38, valvifers VIII. 3435: Scale: 0.5 mm; 3638: Scale: 0.25 mm. Lettering: lt IX—laterotergite IX; lt VIII—laterotergite VIII; rs—ring sclerite; sso—sclerite guarding spermathecal opening; vlf VIII—valvifers VIII." figureDoi="http://doi.org/10.5281/zenodo.4664158" httpUri="https://zenodo.org/record/4664158/files/figure.png" pageId="2" pageNumber="285">Figs 35</figureCitation>
,
<figureCitation id="1367178EFFC1FFA4BB04FA20FAD4FA20" box="[1290,1319,1410,1437]" captionStart="FIGURES 5154" captionStartId="16.[151,264,1903,1927]" captionTargetBox="[155,1422,857,1865]" captionTargetId="figure-290@16.[151,1436,813,1880]" captionTargetPageId="16" captionText="FIGURES 5154. Sarju nigricollis (Westwood). 51, terminalia (dorsal); 52, spermatheca; 53, terminalia (ventral); 54, valvifers VIII. 5152: Scale: 0.5 mm; 5354: Scale: 0.25 mm. Lettering: lt IX—laterotergite IX; lt VIII—laterotergite VIII; rs—ring sclerite; sso—sclerite guarding spermathecal opening; vlf VIII—valvifers VIII." figureDoi="http://doi.org/10.5281/zenodo.4664164" httpUri="https://zenodo.org/record/4664164/files/figure.png" pageId="2" pageNumber="285">52</figureCitation>
) with median dilation elongate; apical receptacle with three narrow finger-like processes (except in
<taxonomicName id="4C5C7088FFC1FFA4BA85FA05FAD4FA7C" authority="Ghauri" authorityName="Ghauri" box="[1163,1319,1447,1473]" class="Insecta" family="Pentatomidae" genus="Sarju" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="285" phylum="Arthropoda" rank="species" species="lata">
<emphasis id="B928D719FFC1FFA4BA85FA05FB20FA7C" box="[1163,1235,1447,1473]" italics="true" pageId="2" pageNumber="285">S. lata</emphasis>
Ghauri
</taxonomicName>
according to
<bibRefCitation id="EFCD76FAFFC1FFA4BEBAFA69FEB6FA58" author="Ghauri, M. S. K." box="[180,325,1483,1509]" pageId="2" pageNumber="285" pagination="9 - 27" refId="ref12389" refString="Ghauri, M. S. K. (1977) Sarju - a new genus of Halyini (Heteroptera, Pentatomidae, Pentatominae) with new species. Tu ¨ rkiye Bitki Koruma Dergisi, 1 (1), 9 - 27." type="journal article" year="1977">Ghauri 1977</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3465880FFC1FFA4BEC9FA4CFB09F8CC" pageId="2" pageNumber="285" type="diagnosis">
<paragraph id="8BE30B0BFFC1FFA4BEC9FA4CFB09F8CC" blockId="2.[151,1437,151,2014]" pageId="2" pageNumber="285">
<emphasis id="B928D719FFC1FFA4BEC9FA4CFE3AF9B5" bold="true" box="[199,457,1518,1544]" pageId="2" pageNumber="285">Differential diagnosis</emphasis>
. The genus
<taxonomicName id="4C5C7088FFC1FFA4BC55FA4DFD6BF9B4" authorityName="Rider" authorityYear="2006" box="[603,664,1519,1545]" class="Insecta" family="Pentatomidae" genus="Sarju" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="B928D719FFC1FFA4BC55FA4DFD6BF9B4" box="[603,664,1519,1545]" italics="true" pageId="2" pageNumber="285">Sarju</emphasis>
</taxonomicName>
is quite similar to
<taxonomicName id="4C5C7088FFC1FFA4BD78FA4DFBD8F9B4" authority="Ghauri." authorityName="Ghauri." box="[886,1067,1519,1545]" class="Insecta" family="Pentatomidae" genus="Cahara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="B928D719FFC1FFA4BD78FA4DFC3FF9B4" box="[886,972,1519,1545]" italics="true" pageId="2" pageNumber="285">Cahara</emphasis>
Ghauri.
</taxonomicName>
The distinctive character which distinguishes this genus is the absence of median lobes in the concave ventral margin of genital capsule whereas in members of
<emphasis id="B928D719FFC1FFA4BF2AF995FE73F9EC" box="[292,384,1591,1617]" italics="true" pageId="2" pageNumber="285">
<taxonomicName id="4C5C7088FFC1FFA4BF2AF995FE8FF9EC" box="[292,380,1591,1617]" class="Insecta" family="Pentatomidae" genus="Cahara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">Cahara</taxonomicName>
,
</emphasis>
a pair of median lobes is present in concave ventral margin of genital capsule (
<bibRefCitation id="EFCD76FAFFC1FFA4BAF3F995FA63F9EC" author="Ghauri, M. S. K." box="[1277,1424,1591,1617]" pageId="2" pageNumber="285" pagination="9 - 27" refId="ref12389" refString="Ghauri, M. S. K. (1977) Sarju - a new genus of Halyini (Heteroptera, Pentatomidae, Pentatominae) with new species. Tu ¨ rkiye Bitki Koruma Dergisi, 1 (1), 9 - 27." type="journal article" year="1977">Ghauri 1977</bibRefCitation>
). The overall shape of the paramere (
<figureCitation id="1367178EFFC1FFA4BC2FF9F9FD85F9C8" box="[545,630,1627,1653]" captionStart="FIGURES 914" captionStartId="6.[151,264,1536,1560]" captionTargetBox="[184,1422,633,1489]" captionTargetId="figure-216@6.[151,1436,633,1513]" captionTargetPageId="6" captionText="FIGURES 914. Sarju brevirostrata sp. nov. 9, Genital capsule (ventral); 10, Genital capsule (caudal); 1113, paramere (different planes); 14, articulatory apparatus. Scale: 0.25 mm. Lettering: cp—capitate process; mb—mushroom shaped bodies." figureDoi="http://doi.org/10.5281/zenodo.4664144" httpUri="https://zenodo.org/record/4664144/files/figure.png" pageId="2" pageNumber="285">Figs 11</figureCitation>
,
<figureCitation id="1367178EFFC1FFA4BC8CF9F9FD53F9C8" box="[642,672,1627,1653]" captionStart="FIGURES 2833" captionStartId="11.[151,264,1472,1496]" captionTargetBox="[168,1405,436,1436]" captionTargetId="figure-90@11.[151,1436,417,1449]" captionTargetPageId="11" captionText="FIGURES 2833. Sarju farida Ghauri. 2829, paramere; 30, articulatory apparatus; 31, phallus (ventral); 32, phallus (dorsal); 33, phallus (lateral). Scale: 0.25 mm. Lettering: cp—capitate process; dcp—dorsal conjunctival processes; mcp—median conjunctival processes; pa—processes of aedeagus; sp—setose process by Ghauri, 1977; vcp—ventral conjunctival processes; vt—ventral tubercle." figureDoi="http://doi.org/10.5281/zenodo.4664156" httpUri="https://zenodo.org/record/4664156/files/figure.png" pageId="2" pageNumber="285">29</figureCitation>
,
<figureCitation id="1367178EFFC1FFA4BCA5F9F9FD3BF9CB" box="[683,712,1627,1654]" captionStart="FIGURES 4550" captionStartId="15.[151,264,1840,1864]" captionTargetBox="[170,1436,777,1786]" captionTargetId="figure-274@15.[151,1436,777,1817]" captionTargetPageId="15" captionText="FIGURES 4550. Sarju nigricollis (Westwood). 4546, paramere (different planes); 47, articulatory apparatus; 48, phallus (ventral); 49, phallus (dorsal); 50, phallus (lateral). Scale: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4664162" httpUri="https://zenodo.org/record/4664162/files/figure.png" pageId="2" pageNumber="285">46</figureCitation>
) and the presence of a short projection towards base of paramere (setose process by
<bibRefCitation id="EFCD76FAFFC1FFA4BF79F9DDFDF7F924" author="Ghauri, M. S. K." box="[375,516,1663,1689]" pageId="2" pageNumber="285" pagination="9 - 27" refId="ref12389" refString="Ghauri, M. S. K. (1977) Sarju - a new genus of Halyini (Heteroptera, Pentatomidae, Pentatominae) with new species. Tu ¨ rkiye Bitki Koruma Dergisi, 1 (1), 9 - 27." type="journal article" year="1977">Ghauri 1977</bibRefCitation>
) are other diagnostics for this genus, though these characters are shared with members of
<emphasis id="B928D719FFC1FFA4BEE0F901FEB8F900" box="[238,331,1699,1725]" italics="true" pageId="2" pageNumber="285">
<taxonomicName id="4C5C7088FFC1FFA4BEE0F901FEB4F900" box="[238,327,1699,1725]" class="Insecta" family="Pentatomidae" genus="Cahara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">Cahara</taxonomicName>
.
</emphasis>
All the members of
<emphasis id="B928D719FFC1FFA4BC39F901FD88F900" box="[567,635,1699,1725]" italics="true" pageId="2" pageNumber="285">
<taxonomicName id="4C5C7088FFC1FFA4BC39F901FD85F900" authorityName="Rider" authorityYear="2006" box="[567,630,1699,1725]" class="Insecta" family="Pentatomidae" genus="Sarju" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">Sarju</taxonomicName>
,
</emphasis>
so far reported, have the mandibular plates either as long as or longer than clypeus but not shorter than clypeus, though this character is shared with quite a few species in various related genera. The geniculate (bowed) second antennal segment with swollen apex (the degree of bending and swelling slightly variable between species) is an important distinguishing character of members of
<taxonomicName id="4C5C7088FFC1FFA4BA70F8ADFB48F894" authorityName="Rider" authorityYear="2006" box="[1150,1211,1807,1833]" class="Insecta" family="Pentatomidae" genus="Sarju" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="B928D719FFC1FFA4BA70F8ADFB48F894" box="[1150,1211,1807,1833]" italics="true" pageId="2" pageNumber="285">Sarju</emphasis>
</taxonomicName>
though it comprises species with normal straight second antennal segment like in other groups. Hence this character alone cannot vouch for the identity of this genus rather a set of characters mentioned above can distinguish its members.
</paragraph>
</subSubSection>
<subSubSection id="C3465880FFC1FFA5BEC9F8D9FC39FF44" lastPageId="3" lastPageNumber="286" pageId="2" pageNumber="285" type="discussion">
<paragraph id="8BE30B0BFFC1FFA5BEC9F8D9FC39FF44" blockId="2.[151,1437,151,2014]" lastBlockId="3.[151,1436,151,249]" lastPageId="3" lastPageNumber="286" pageId="2" pageNumber="285">
<emphasis id="B928D719FFC1FFA4BEC9F8D9FEC8F828" bold="true" box="[199,315,1915,1941]" pageId="2" pageNumber="285">Remarks.</emphasis>
Members of this genus are of typical Halyine
<typeStatus id="54E7B5A9FFC1FFA4BD36F8D9FC9BF828" box="[824,872,1915,1941]" pageId="2" pageNumber="285">type</typeStatus>
with respect to their external colouration, the presence of gradation of variation found in structural characters (morphological plasticity) and habitat (most of them are bark dwelling). Hence the examination of male genitalia is often necessary (especially if the second antennal segment is not geniculate) in order to separate
<taxonomicName id="4C5C7088FFC0FFA5BC7DFF35FD43FF0C" authorityName="Rider" authorityYear="2006" box="[627,688,151,177]" class="Insecta" family="Pentatomidae" genus="Sarju" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="286" phylum="Arthropoda" rank="genus">
<emphasis id="B928D719FFC0FFA5BC7DFF35FD43FF0C" box="[627,688,151,177]" italics="true" pageId="3" pageNumber="286">Sarju</emphasis>
</taxonomicName>
from similar genera in the
<taxonomicName id="4C5C7088FFC0FFA5BDD4FF35FBC1FF0C" authorityName="Amyot &amp; Serville" authorityYear="1843" box="[986,1074,151,177]" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="286" phylum="Arthropoda" rank="tribe" tribe="Halyini">Halyini</taxonomicName>
. The above mentioned similarity in external colouration and morphological plasticity are applicable to various species within
<taxonomicName id="4C5C7088FFC0FFA5BABEFF19FB1EFF68" authorityName="Rider" authorityYear="2006" box="[1200,1261,187,213]" class="Insecta" family="Pentatomidae" genus="Sarju" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="286" phylum="Arthropoda" rank="genus">
<emphasis id="B928D719FFC0FFA5BABEFF19FB1EFF68" box="[1200,1261,187,213]" italics="true" pageId="3" pageNumber="286">Sarju</emphasis>
</taxonomicName>
too. Therefore, the male genitalia are critical for accurate species determination in
<emphasis id="B928D719FFC0FFA5BD88FF7DFC39FF44" box="[902,970,223,249]" italics="true" pageId="3" pageNumber="286">
<taxonomicName id="4C5C7088FFC0FFA5BD88FF7DFC35FF44" authorityName="Rider" authorityYear="2006" box="[902,966,223,249]" class="Insecta" family="Pentatomidae" genus="Sarju" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="286" phylum="Arthropoda" rank="genus">Sarju</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>