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<document ID-DOI="http://dx.doi.org/10.3897/zitteliana.96.87253" ID-Pensoft-Pub="2747-8106-96-153" ID-Pensoft-UUID="AFA9059B36235BFF9BA3E9B7BE816DC1" ID-ZooBank="191199E07F3E4E09A3774ADFBF93A248" ModsDocID="2747-8106-96-153" checkinTime="1660292272395" checkinUser="pensoft" docAuthor="Fuersich, Franz T., Alberti, Matthias, Pandey, Dhirendra K. &amp; Ayoub-Hannaa, Wagih S." docDate="2022" docId="5C6B218E4EA859238D2E8F47BB5F4270" docLanguage="en" docName="Zitteliana 96: 153-178" docOrigin="Zitteliana 96" docPubDate="2022-08-11" docSource="http://dx.doi.org/10.3897/zitteliana.96.87253" docTitle="Megacucullaea kraussii" docType="treatment" docVersion="1" id="AFA9059B36235BFF9BA3E9B7BE816DC1" lastPageNumber="153" masterDocId="AFA9059B36235BFF9BA3E9B7BE816DC1" masterDocTitle="Jurassic bivalves from the Spiti area of the Himalayas, northern India" masterLastPageNumber="178" masterPageNumber="153" pageNumber="153" updateTime="1660292272395" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Jurassic bivalves from the Spiti area of the Himalayas, northern India</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fuersich, Franz T.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0844-9297</mods:nameIdentifier>
<mods:affiliation>Universitaet Erlangen-Nuernberg, GeoZentrum Nordbayern, FG Palaeoumwelt, Lowenichstrasse 28, 91054 Erlangen, Germany</mods:affiliation>
<mods:nameIdentifier type="email">franz.fuersich@fau.de</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Alberti, Matthias</mods:namePart>
<mods:affiliation>State Key Laboratory for Mineral Deposits Research, School of Earth Sciences and Engineering, Centre for Research and Education on Biological Evolution and Environment and Frontiers Science Center for Critical Earth Material Cycling, Nanjing University, Nanjing 210023, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pandey, Dhirendra K.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7721-9604</mods:nameIdentifier>
<mods:affiliation>Department of Earth and Environmental Science, KSKV Kachchh University, Bhuj, India</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ayoub-Hannaa, Wagih S.</mods:namePart>
<mods:affiliation>Geology Department, Faculty of Science, Minufiya University, El-Minufiya, Shibin El Kom, Egypt</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zitteliana</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-08-11</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>96</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>153</mods:start>
<mods:end>178</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zitteliana.96.87253</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zitteliana.96.87253</mods:identifier>
<mods:identifier type="Pensoft-Pub">2747-8106-96-153</mods:identifier>
<mods:identifier type="ZooBank">191199E07F3E4E09A3774ADFBF93A248</mods:identifier>
<mods:identifier type="Pensoft-UUID">AFA9059B36235BFF9BA3E9B7BE816DC1</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:5C6B218E4EA859238D2E8F47BB5F4270" httpUri="http://treatment.plazi.org/id/5C6B218E4EA859238D2E8F47BB5F4270" lastPageNumber="153" pageId="0" pageNumber="153">
<subSubSection pageId="0" pageNumber="153" type="nomenclature">
<paragraph pageId="0" pageNumber="153">
<taxonomicName LSID="5C6B218E-4EA8-5923-8D2E-8F47BB5F4270" authority="(Tate, 1867)" baseAuthorityName="Tate" baseAuthorityYear="1867" class="Bivalvia" family="Cucullaeidae" genus="Megacucullaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Megacucullaea kraussii" order="Arcida" pageId="0" pageNumber="153" phylum="Mollusca" rank="species" species="kraussii">Megacucullaea cf. kraussii (Tate, 1867)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="153" type="description">
<paragraph pageId="0" pageNumber="153">
<figureCitation captionStart="Plate 1" captionStartId="F3" captionText="Plate 1. (1, 2) Palaeonucula cuneiformis (J. de C. Sowerby, 1840). 1. Left valve from the lower member at Langza (SNSB-BSPG 2020 XCIX 26 a). 2. Left valve from the lower member at Langza (SNSB-BSPG 2020 XCIX 26 b). (3) Palaeonucula stoliczkai Cox, 1940. Laterally crushed articulated specimen, lower member at Langza (SNSB-BSPG 2020 XCIX 34); a, left valve view; b, right valve view. (4) Megacucullaea cf. kraussii (Tate, 1867). Internal mould of articulated specimen with remains of shell from the lower member near Chichim (SNSB-BSPG 2020 XCIX 1); a, right valve view; b, dorsal view. (5 - 7) Grammatodon (Indogrammatodon) egertonianus (Stoliczka, 1866). 5. Articulated specimen from the lower member near the pass to Tashigeng (SNSB-BSPG 2020 XCIX 4). a, left valve view; b, dorsal view. 6. Laterally crushed articulated specimen from the lower member along the Kaza-Hikkim road (SNSB-BSPG 2020 XCIX 3 b); right valve view. 7. Dorso-ventrally compacted articulated specimen from the lower member at Langza (SNSB-BSPG 2020 XCIX 6); right valve view. (8, 9) Grammatodon (Cosmetodon) sp. from the lower member along the Kaza-Hikkim road. 8. Right valve (SNSB-BSPG 2020 XCIX 2 a). 9. Right valve (SNSB-BSPG 2020 XCIX 2 b). a, lateral view; b, dorsal view. (10) Anopaea? Fragmented right valve from the upper? member at Langza (SNSB-BSPG 2020 XCIX 67)." figureDoi="10.3897/zitteliana.96.87253.plate1" httpUri="https://binary.pensoft.net/fig/726364" pageId="0" pageNumber="153">Plate 1, fig. 4a, b</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="153" type="reference_group">
<paragraph pageId="0" pageNumber="153">
<taxonomicName class="Bivalvia" family="Cucullaeidae" genus="Cucullaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cucullaea cancellata" order="Arcida" pageId="0" pageNumber="153" phylum="Mollusca" rank="species" species="cancellata">cf. 1850 Cucullaea cancellata</taxonomicName>
- Krauss: 452, pl. 48, fig. 2a, b (non Phillips, 1829)
</paragraph>
<paragraph pageId="0" pageNumber="153">
<taxonomicName class="Bivalvia" family="Cucullaeidae" genus="Cucullaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cucullaea kraussii" order="Arcida" pageId="0" pageNumber="153" phylum="Mollusca" rank="species" species="kraussii">*cf. 1867 Cucullaea kraussii</taxonomicName>
- Tate: 161.
</paragraph>
<paragraph pageId="0" pageNumber="153">
<taxonomicName authorityName="Lamarck" authorityYear="1801" class="Bivalvia" family="Cucullaeidae" genus="Cucullaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cucullaea" order="Arcida" pageId="0" pageNumber="153" phylum="Mollusca" rank="genus">cf. 1882 Cucullaea</taxonomicName>
Kraussi Tate - Holub and Neumayr: 375, pl. 2, fig. 2a-c.
</paragraph>
<paragraph pageId="0" pageNumber="153">
<taxonomicName class="Bivalvia" family="Cucullaeidae" genus="Cucullaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cucullaea (Megacucullaea) kraussii" order="Arcida" pageId="0" pageNumber="153" phylum="Mollusca" rank="species" species="kraussii" subGenus="Megacucullaea">cf. 1940 Cucullaea (Megacucullaea) kraussii</taxonomicName>
Tate - Cox: 57, pl. 4, figs 3-4.
</paragraph>
<paragraph pageId="0" pageNumber="153">
<taxonomicName class="Bivalvia" family="Cucullaeidae" genus="Megacucullaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Megacucullaea kraussi" order="Arcida" pageId="0" pageNumber="153" phylum="Mollusca" rank="species" species="kraussi">cf. 1998 Megacucullaea kraussi</taxonomicName>
(Tate, 1867) - Kanjilal and Pathak: 34, pl. 1, fig. 6.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="153" type="materials_examined">
<paragraph pageId="0" pageNumber="153">Material.</paragraph>
<paragraph pageId="0" pageNumber="153">A single poorly preserved articulated internal mould with remains of shell from the lower member of the Spiti Shale Formation (Oxfordian) near Chichim (SNSB- BSPG 2020 XCIX 1).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="153" type="description">
<paragraph pageId="0" pageNumber="153">Description.</paragraph>
<paragraph pageId="0" pageNumber="153">Specimen large, longer than high (L ~79 mm, H ~60 mm, I ~70), thick-shelled, strongly inflated. Maximum inflation at around one-third of shell height from umbo. Ventral margin faintly arched, anterior margin convex, forming an angle of ~90° with hinge margin. Posterior margin straight, oblique, posteroventral corner rounded. Umbones only partly preserved, prominent, straight, incurved, situated anterior of mid-line of shell. Posterodorsal carina distinct but rounded, posterodorsal area flat to slightly convex. Cardinal area large, largest width anterior of umbo, slightly concave. Ornamentation poorly preserved; there are at least four widely spaced, strong, rounded radial ribs with concave interstices. Hinge line straight, dentition and other internal features not seen.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="153" type="remarks">
<paragraph pageId="0" pageNumber="153">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="153">The two valves are sligthly laterally sheared. Moreover, the specimen may be somewhat compressed dorso-ventrally. In this case, the inflation would be lower and the height greater than decribed above, and the point of maximum inflation would also be slightly off. The umbones are only partly preserved.</paragraph>
<paragraph pageId="0" pageNumber="153">
The various Jurassic species of
<taxonomicName authorityName="Rennie" authorityYear="1936" class="Bivalvia" family="Cucullaeidae" genus="Megacucullaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Megacucullaea" order="Arcida" pageId="0" pageNumber="153" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="153">Megacucullaea</emphasis>
</taxonomicName>
, i.e.
<taxonomicName genus="M." lsidName="M. eminens" pageId="0" pageNumber="153" rank="species" species="eminens">
<emphasis italics="true" pageId="0" pageNumber="153">M. eminens</emphasis>
</taxonomicName>
<bibRefCitation author="Cox, LR" journalOrPublisher="Proceedings of the Malacological Society" pageId="0" pageNumber="153" refId="B16" refString="Cox, LR, 1940. The Jurassic lamellibranch fauna of Kachh (Cutch). Memoirs of the Geological Survey of India, Palaeontologica Indica, New Series (9), 3(3): 1-157." title="The Jurassic lamellibranch fauna of Kachh (Cutch). Memoirs of the Geological Survey of India, Palaeontologica Indica, New Series (9), 3 (3): 1 - 157." year="1940">Cox (1940</bibRefCitation>
, p. 59, pl. 4, figs 5-7),
<taxonomicName genus="M." lsidName="M. kraussii" pageId="0" pageNumber="153" rank="species" species="kraussii">
<emphasis italics="true" pageId="0" pageNumber="153">M. kraussii</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.1144/GSL.JGS.1867.023.01-02.24" author="Tate, R" journalOrPublisher="Quarterly Journal of the Geological Society" pageId="0" pageNumber="153" pagination="139 - 175" refId="B86" refString="Tate, R, 1867. On some Secondary fossils from South Africa. Quarterly Journal of the Geological Society 23: 139 - 175, DOI: https://doi.org/10.1144/GSL.JGS.1867.023.01-02.24" title="On some Secondary fossils from South Africa." url="https://doi.org/10.1144/GSL.JGS.1867.023.01-02.24" volume="23" year="1867">Tate 1867</bibRefCitation>
, p. 161), and &quot;
<taxonomicName genus="M." lsidName="M." pageId="0" pageNumber="153" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="153">M.</emphasis>
</taxonomicName>
sp. nov. aff.
<taxonomicName genus="C." lsidName="C. (M.) eminens" pageId="0" pageNumber="153" rank="species" species="eminens" subGenus="M.">C. (M.) eminens</taxonomicName>
Cox&quot; of Agrawal (1956 p. 14, pl. 1, fig. 1) differ in outline and the number of radial ribs (
<taxonomicName genus="M." lsidName="M. kraussii" pageId="0" pageNumber="153" rank="species" species="kraussii">
<emphasis italics="true" pageId="0" pageNumber="153">M. kraussii</emphasis>
</taxonomicName>
: 6-8;
<taxonomicName genus="M." lsidName="M. eminens" pageId="0" pageNumber="153" rank="species" species="eminens">M. eminens</taxonomicName>
: 16-19;
<normalizedToken originalValue="Agrawal’s">Agrawal's</normalizedToken>
specimen: approximately 10). The present specimen is closest to
<taxonomicName genus="M." lsidName="M. kraussii" pageId="0" pageNumber="153" rank="species" species="kraussii">
<emphasis italics="true" pageId="0" pageNumber="153">M. kraussii</emphasis>
</taxonomicName>
, having a similar number of ribs, whereas in the other two species the number of ribs is distinctly higher. Due to the poor preservation of the specimen, a detailed comparison with
<taxonomicName genus="M." lsidName="M. kraussii" pageId="0" pageNumber="153" rank="species" species="kraussii">
<emphasis italics="true" pageId="0" pageNumber="153">M. kraussii</emphasis>
</taxonomicName>
is not possible. The material of
<bibRefCitation author="Cox, LR" journalOrPublisher="Proceedings of the Malacological Society" pageId="0" pageNumber="153" refId="B16" refString="Cox, LR, 1940. The Jurassic lamellibranch fauna of Kachh (Cutch). Memoirs of the Geological Survey of India, Palaeontologica Indica, New Series (9), 3(3): 1-157." title="The Jurassic lamellibranch fauna of Kachh (Cutch). Memoirs of the Geological Survey of India, Palaeontologica Indica, New Series (9), 3 (3): 1 - 157." year="1940">Cox (1940</bibRefCitation>
, p. 57, pl. 4, figs 3-4) exhibits eight radial ribs, of which three are only faintly developed on one out of two specimens. Interestingly,
<taxonomicName genus="M." lsidName="M. eminens" pageId="0" pageNumber="153" rank="species" species="eminens">
<emphasis italics="true" pageId="0" pageNumber="153">M. eminens</emphasis>
</taxonomicName>
and
<taxonomicName genus="M." lsidName="M. kraussii" pageId="0" pageNumber="153" rank="species" species="kraussii">
<emphasis italics="true" pageId="0" pageNumber="153">M. kraussii</emphasis>
</taxonomicName>
co-occur in Upper Tithonian rocks of the Kachchh Basin.
<taxonomicName genus="M." lsidName="M. eminens" pageId="0" pageNumber="153" rank="species" species="eminens">
<emphasis italics="true" pageId="0" pageNumber="153">M. eminens</emphasis>
</taxonomicName>
<bibRefCitation author="Cox, LR" journalOrPublisher="Proceedings of the Malacological Society" pageId="0" pageNumber="153" refId="B16" refString="Cox, LR, 1940. The Jurassic lamellibranch fauna of Kachh (Cutch). Memoirs of the Geological Survey of India, Palaeontologica Indica, New Series (9), 3(3): 1-157." title="The Jurassic lamellibranch fauna of Kachh (Cutch). Memoirs of the Geological Survey of India, Palaeontologica Indica, New Series (9), 3 (3): 1 - 157." year="1940">Cox (1940</bibRefCitation>
, p. 59, pl. 4, figs 5-7) is shorter, carries about 16 radial ribs differing in strength (weak close to the umbonal carina and strong on the remainder of the shell), and the area posterior of the umbonal ridge is distinctly concave. &quot;
<emphasis italics="true" pageId="0" pageNumber="153">
<taxonomicName authorityName="Lamarck" authorityYear="1801" class="Bivalvia" family="Parallelodontidae" genus="Cucullaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cucullaea" order="Arcida" pageId="0" pageNumber="153" phylum="Mollusca" rank="genus">Cucullaea</taxonomicName>
Kraussi
</emphasis>
Tate&quot; of
<bibRefCitation author="Barrabe, L" journalOrPublisher="Bulletin de la Societe Imperiale des Naturalistes de Moscou" pageId="0" pageNumber="153" refId="B6" refString="Barrabe, L, 1929. Contribution a l'etude stratigraphique et petrographique de la partie mediane du pays Sahalave (Madagascar). Memoires de la Societe Geologique de France, Nouvelle Serie, no. 12, 5(3-4): 1-270." title="Contribution a l'etude stratigraphique et petrographique de la partie mediane du pays Sahalave (Madagascar). Memoires de la Societe Geologique de France, Nouvelle Serie, no. 12, 5 (3 - 4): 1 - 270." year="1929">
<normalizedToken originalValue="Barrabé">Barrabe</normalizedToken>
(1929
</bibRefCitation>
, p. 147, pl. 8, fig. 13) has more radial ribs and resembles
<taxonomicName genus="M." lsidName="M. eminens" pageId="0" pageNumber="153" rank="species" species="eminens">
<emphasis italics="true" pageId="0" pageNumber="153">M. eminens</emphasis>
</taxonomicName>
. The early Tithonian specimen figured as
<taxonomicName baseAuthorityName="Tate" baseAuthorityYear="1867" class="Bivalvia" family="Cucullaeidae" genus="Megacucullaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Megacucullaea kraussii" order="Arcida" pageId="0" pageNumber="153" phylum="Mollusca" rank="species" species="kraussii">
<emphasis italics="true" pageId="0" pageNumber="153">Megacucullaea kraussii</emphasis>
</taxonomicName>
by
<bibRefCitation author="Kanjilal, S" journalOrPublisher="Himalayan Geology" pageId="0" pageNumber="153" pagination="29 - 37" refId="B50" refString="Kanjilal, S, Pathak, DB, 1998. Spiti Shale bivalves from Chichim and Gate, Spiti Valley, Himachal Pradesh, India. Himalayan Geology 19: 29 - 37" title="Spiti Shale bivalves from Chichim and Gate, Spiti Valley, Himachal Pradesh, India." volume="19" year="1998">Kanjilal and Pathak (1998)</bibRefCitation>
has seven ribs.
</paragraph>
</subSubSection>
</treatment>
</document>
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