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<document id="D70CB7D1E0A5E0A85F4F24466034B6DF" ID-DOI="10.5852/ejt.2024.958.2675" ID-ISSN="2118-9773" ID-Zenodo-Dep="13914492" ID-ZooBank="18DC2A7C-175B-4375-A82E-E1AFF4029A94" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1728579298640" checkinUser="plazi" docAuthor="Meyer, Marc De, Goergen, Georg, Midgley, John &amp; Jordaens, Kurt" docDate="2024" docId="CE3D0D146542A4199C26A87241BEB407" docLanguage="en" docName="EJT.2024.958.242-290.pdf" docOrigin="European Journal of Taxonomy 958 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2675/12391" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Mallota meromacrimima Hull 1941" docType="treatment" docVersion="1" lastPageNumber="253" masterDocId="3204756C6548A4129E6CA9074335B25D" masterDocTitle="On the identity of the Afrotropical species of Mallota Meigen (Diptera: Syrphidae)" masterLastPageNumber="290" masterPageNumber="242" pageNumber="252" updateTime="1729092679658" updateUser="guilherme" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="1B5B3A961E457DF4C08D7F571A160916">On the identity of the Afrotropical species of Mallota Meigen (Diptera: Syrphidae)</mods:title>
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<mods:namePart id="6446850AC96BA296419F6A9722F34EA4">Meyer, Marc De</mods:namePart>
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<mods:affiliation id="D79C7EBF2C195ED23DE88AA1043C5DD5">Royal Museum for Central Africa, Invertebrates Section, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium.</mods:affiliation>
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<mods:namePart id="1402BB798281E0F74F6DBF0259DBBFE9">Goergen, Georg</mods:namePart>
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<mods:affiliation id="35AFEFA5C24FB02A66E776E4C469A266">International Institute for Tropical Agriculture (IITA), Biodiversity Centre, 08 BP 0932 Tri Postal, Cotonou, Benin.</mods:affiliation>
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<mods:affiliation id="F11EE312F876BD584E6DCF2D96648DE6">KwaZulu-Natal Museum, Department Natural Sciences, 237 Jabu Ndlovu Street, Pietermaritzburg, KwaZulu-Natal, South Africa, 3201. &amp; Department of Zoology and Entomology, Rhodes University, Makhanda, South Africa, 6139.</mods:affiliation>
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<mods:affiliation id="6070913E3C0C9C4526FE04058EFC473F">Royal Museum for Central Africa, Invertebrates Section, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium.</mods:affiliation>
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<mods:date id="EC217310AFD80065320514F8C3A4B58B">2024</mods:date>
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<taxonomicName id="8194C7816542A4189C26A87240DCB3CD" ID-CoL="3XPS2" authority="Hull, 1941" authorityName="Hull" authorityYear="1941" box="[586,1001,373,400]" class="Insecta" family="Syrphidae" genus="Mallota" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="252" phylum="Arthropoda" rank="species" species="meromacrimima">
<emphasis id="74E060106542A4189C26A872405FB3CD" bold="true" box="[586,874,373,400]" italics="true" pageId="10" pageNumber="252">Mallota meromacrimima</emphasis>
<bibRefCitation id="2205C1F36542A4189D1EA87140DCB3CD" author="Hull F. M." box="[882,1001,374,400]" pageId="10" pageNumber="252" pagination="309 - 334" refId="ref20814" refString="Hull F. M. 1941. A study of syrphid flies from Madagascar (Diptera). Proceedings of the Academy of Natural Sciences of Philadelphia 92: 309 - 334." type="journal article" year="1941">Hull, 1941</bibRefCitation>
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<subSubSection id="0E8EEF896542A4189CCFA89040A5B3EF" box="[675,912,407,434]" pageId="10" pageNumber="252" type="description">
<paragraph id="462BBC026542A4189CCFA89040A5B3EF" blockId="10.[586,1001,373,434]" box="[675,912,407,434]" pageId="10" pageNumber="252">
<figureCitation id="DEAFA0876542A4189CCFA89041DFB3EF" box="[675,746,407,434]" captionStart="Figs 1–2" captionStartId="8.[189,242,1906,1932]" captionTargetBox="[189,1349,270,1858]" captionTargetId="figure-12@8.[189,1398,270,1858]" captionTargetPageId="8" captionText="Figs 1–2. Species of Mallota Bezzi, 1822. Habitus, lateral view. 1. M. hirsuta Hull, 1941, holotype, ♀ (ANSP ENT 142915 (6591)). 2. M. meromacrimima Hull, 1941, holotype, ♂ (ANSP ENT 142916 (6592))." figureDoi="http://doi.org/10.5281/zenodo.13914494" httpUri="https://zenodo.org/record/13914494/files/figure.png" pageId="10" pageNumber="252">Figs 2</figureCitation>
,
<figureCitation id="DEAFA0876542A4189C9AA89F4021B3EF" box="[758,788,408,434]" captionStart="Figs 11–16" captionStartId="31.[189,242,1857,1883]" captionTargetBox="[216,1384,265,1815]" captionTargetId="figure-13@31.[203,1384,265,1815]" captionTargetPageId="31" captionText="Figs 11–16. Species of Mallota Bezzi, 1822. Head, lateral view. 11. M. hirsuta Hull, 1941, holotype, ♀ (ANSP ENT 142915 (6591)). 12. M. meromacrimima Hull, 1941, holotype, ♂ (ANSP ENT 142916 (6592)). 13. M. aperta (Bezzi, 1912), ♀ (RBINS, Eala, Aug. 1935, J. Ghesquière). 14. M. dasyops (Wiedemann, 1819), ♂ (NMSA DIP 211695). 15. M. glabra sp. nov., paratype, ♂ (NMSA DIP 65015). 16. M. hircus sp. nov., paratype, ♀ (RMCA ENT 000037957)." figureDoi="http://doi.org/10.5281/zenodo.13914504" httpUri="https://zenodo.org/record/13914504/files/figure.png" pageId="10" pageNumber="252">12</figureCitation>
,
<figureCitation id="DEAFA0876542A4189D4CA890400BB3EC" box="[800,830,407,433]" captionStart="Figs 25–30" captionStartId="34.[189,242,1857,1883]" captionTargetBox="[202,1386,265,1819]" captionTargetId="figure-11@34.[202,1386,265,1819]" captionTargetPageId="34" captionText="Figs 25–30. Species of Mallota Bezzi, 1822. Metaleg, anterior view.25. M. hirsuta Hull, 1941, holotype, ♀ (ANSP ENT 142915 (6591)). 26. M. meromacrimima Hull, 1941, holotype, ♂ (ANSP ENT 142916 (6592)). 27. M. aperta (Bezzi, 1912), ♀ (RBINS, Eala, 5 Oct. 1936, J. Ghesquière). 28. M. dasyops (Wiedemann, 1819), ♀ (NMSA DIP 175981). 29. M. glabra sp. nov., paratype, ♂ (NMSA DIP 65015). 30. M. hircus sp. nov., paratype, ♂ (RMCA ENT 000037959)." figureDoi="http://doi.org/10.5281/zenodo.13914512" httpUri="https://zenodo.org/record/13914512/files/figure.png" pageId="10" pageNumber="252">26</figureCitation>
,
<figureCitation id="DEAFA0876542A4189D26A89F405DB3EF" box="[842,872,408,434]" captionStart="Figs 37–39" captionStartId="36.[189,242,1901,1927]" captionTargetBox="[189,1377,265,1853]" captionTargetId="figure-12@36.[189,1399,265,1853]" captionTargetPageId="36" captionText="Figs 37–39. Species of Mallota Bezzi, 1822. Right wing. 37. M. hirsuta Hull, 1941, holotype, ♀ (ANSP ENT 142915 (6591)). 38. M. meromacrimima Hull, 1941, holotype, ♂ (ANSP ENT 142916 (6592)). 39. M. aperta (Bezzi, 1912), holotype, ♂ (MCSNG)." figureDoi="http://doi.org/10.5281/zenodo.13914516" httpUri="https://zenodo.org/record/13914516/files/figure.png" pageId="10" pageNumber="252">38</figureCitation>
,
<figureCitation id="DEAFA0876542A4189D18A89F40A5B3EF" box="[884,912,408,434]" captionStart="Figs 47–50" captionStartId="40.[189,242,1892,1918]" captionTargetBox="[190,1398,266,1839]" captionTargetId="figure-11@40.[190,1398,266,1849]" captionTargetPageId="40" captionText="Figs 47–50. Species of Mallota Bezzi, 1822. Abdomen, dorsal view.47. M. hirsuta Hull, 1941, holotype, ♀ (ANSP ENT 142915 (6591)). 48. M. meromacrimima Hull, 1941, holotype, ♂ (ANSP ENT 142916 (6592)). 49. M. aperta (Bezzi, 1912), holotype, ♂ (MCSNG). 50. M. dasyops (Wiedemann, 1819), ♂ (NMSA DIP 211695)." figureDoi="http://doi.org/10.5281/zenodo.13914524" httpUri="https://zenodo.org/record/13914524/files/figure.png" pageId="10" pageNumber="252">48</figureCitation>
</paragraph>
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<treatmentCitation id="C7359A136542A4189ED1A8DE4193B3AE" author="Hull F. M." box="[189,678,473,499]" page="328" pageId="10" pageNumber="252" year="1941">
<taxonomicName id="8194C7816542A4189ED1A8DE4193B3AE" ID-CoL="3XPS2" authority="Hull, 1941: 328" authorityName="Hull" authorityPageNumber="328" authorityYear="1941" box="[189,678,473,499]" class="Insecta" family="Syrphidae" genus="Mallota" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="252" phylum="Arthropoda" rank="species" species="meromacrimima">
<emphasis id="74E060106542A4189ED1A8DE42D7B3AE" box="[189,482,473,499]" italics="true" pageId="10" pageNumber="252">Mallota meromacrimima</emphasis>
<bibRefCitation id="2205C1F36542A4189F85A8DE4193B3AE" author="Hull F. M." box="[489,678,473,499]" pageId="10" pageNumber="252" pagination="309 - 334" refId="ref20814" refString="Hull F. M. 1941. A study of syrphid flies from Madagascar (Diptera). Proceedings of the Academy of Natural Sciences of Philadelphia 92: 309 - 334." type="journal article" year="1941">Hull, 1941: 328</bibRefCitation>
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.
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<subSubSection id="0E8EEF896542A4189ED1AB1E40C3B168" pageId="10" pageNumber="252" type="diagnosis">
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<heading id="1D630B6E6542A4189ED1AB1E4289B06E" bold="true" box="[189,444,537,563]" fontSize="11" level="3" pageId="10" pageNumber="252" reason="3">
<emphasis id="74E060106542A4189ED1AB1E4289B06E" bold="true" box="[189,444,537,563]" pageId="10" pageNumber="252">Differential diagnosis</emphasis>
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<paragraph id="462BBC026542A4189ED1AB4040C3B168" blockId="10.[189,1399,583,821]" pageId="10" pageNumber="252">
One of the two Madagascan species. It can be differentiated from the other Madagascan species (
<taxonomicName id="8194C7816542A4189EABAB6C4272B0D9" authorityName="Hull" authorityYear="1941" box="[199,327,618,645]" class="Insecta" family="Syrphidae" genus="Mallota" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="252" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="74E060106542A4189EABAB6C43EAB0D9" box="[199,223,619,644]" italics="true" pageId="10" pageNumber="252">M</emphasis>
.
<emphasis id="74E060106542A4189E98AB6D4272B0D9" box="[244,327,618,644]" italics="true" pageId="10" pageNumber="252">hirsuta</emphasis>
</taxonomicName>
) by the yellow face (darker orange in
<taxonomicName id="8194C7816542A4189D52AB6D40A4B0D9" authorityName="Hull" authorityYear="1941" box="[830,913,618,644]" class="Insecta" family="Syrphidae" genus="Mallota" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="252" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="74E060106542A4189D52AB6D40A4B0D9" box="[830,913,618,644]" italics="true" pageId="10" pageNumber="252">hirsuta</emphasis>
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), the posterior anepisternum with pale yellow pile (yellow-orange in
<taxonomicName id="8194C7816542A4189C44AB89414FB0F5" authorityName="Hull" authorityYear="1941" box="[552,634,654,680]" class="Insecta" family="Syrphidae" genus="Mallota" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="252" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="74E060106542A4189C44AB89414FB0F5" box="[552,634,654,680]" italics="true" pageId="10" pageNumber="252">hirsuta</emphasis>
</taxonomicName>
), the more reddish abdominal terga, and the presence of narrow interrupted fasciae of white pollinosity along the anterior margin of terga 3 and 4 (absent in
<taxonomicName id="8194C7816542A4189A81ABB64675B096" authorityName="Hull" authorityYear="1941" box="[1261,1344,689,715]" class="Insecta" family="Syrphidae" genus="Mallota" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="252" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="74E060106542A4189A81ABB64675B096" box="[1261,1344,689,715]" italics="true" pageId="10" pageNumber="252">hirsuta</emphasis>
</taxonomicName>
) but see comments below. Both can be differentiated from the only other Afrotropical species with bare eyes (
<taxonomicName id="8194C7816542A4189EABABFF420EB14C" baseAuthorityName="Bezzi" baseAuthorityYear="1912" box="[199,315,760,785]" class="Insecta" family="Syrphidae" genus="Protylocera" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="252" phylum="Arthropoda" rank="species" species="aperta">
<emphasis id="74E060106542A4189EABABFF420EB14C" box="[199,315,760,785]" italics="true" pageId="10" pageNumber="252">M. aperta</emphasis>
</taxonomicName>
) by the absence of a distinct medial dark brown macula in the wing (present in
<taxonomicName id="8194C7816542A4189A8BABFF4606B14C" baseAuthorityName="Bezzi" baseAuthorityYear="1912" box="[1255,1331,760,785]" class="Insecta" family="Syrphidae" genus="Protylocera" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="252" phylum="Arthropoda" rank="species" species="aperta">
<emphasis id="74E060106542A4189A8BABFF4606B14C" box="[1255,1331,760,785]" italics="true" pageId="10" pageNumber="252">aperta</emphasis>
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), and the unmarginated rounded scutellum (distinctly marginated in
<taxonomicName id="8194C7816542A4189DF4AA1C40D0B169" baseAuthorityName="Bezzi" baseAuthorityYear="1912" box="[920,997,795,820]" class="Insecta" family="Syrphidae" genus="Protylocera" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="252" phylum="Arthropoda" rank="species" species="aperta">
<emphasis id="74E060106542A4189DF4AA1C40D0B169" box="[920,997,795,820]" italics="true" pageId="10" pageNumber="252">aperta</emphasis>
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).
</paragraph>
</subSubSection>
<subSubSection id="0E8EEF896542A4189ED1AA5B477EB60D" pageId="10" pageNumber="252" type="materials_examined">
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<emphasis id="74E060106542A4189ED1AA5B4259B12B" bold="true" box="[189,364,860,886]" pageId="10" pageNumber="252">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="462BBC026542A4189E86AA8C4269B1F8" blockId="10.[189,1399,907,1004]" box="[234,348,907,933]" pageId="10" pageNumber="252">
<emphasis id="74E060106542A4189E86AA8C4269B1F8" bold="true" box="[234,348,907,933]" pageId="10" pageNumber="252">
<typeStatus id="992F02A06542A4189E86AA8C4269B1F8" box="[234,348,907,933]" pageId="10" pageNumber="252" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="462BBC026542A4189ED1AAA941A4B1B1" blockId="10.[189,1399,907,1004]" pageId="10" pageNumber="252">
<materialsCitation id="F6FCB65F6542A4189ED1AAA941A4B1B1" collectingDate="1937-01" collectingDateMax="1937-05-31" collectingDateMin="1937-01" collectionCode="ANSP" collectorName="C. Lamberton" country="Madagascar" location="Oriental Forest" municipality="Fanovana" pageId="10" pageNumber="252" specimenCode="ANSP ENT 142916" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa Dist." typeStatus="holotype">
<collectingCountry id="3E83FC926542A4189ED1AAA942BBB195" box="[189,398,942,968]" name="Madagascar" pageId="10" pageNumber="252">MADAGASCAR</collectingCountry>
•
<specimenCount id="5092778B6542A4189FC0AAA942F0B195" box="[428,453,942,968]" count="1" pageId="10" pageNumber="252" type="male">♂</specimenCount>
;
<collectingRegion id="845072E06542A4189FB8AAA9419AB195" box="[468,687,942,968]" country="Madagascar" name="Fianarantsoa" pageId="10" pageNumber="252">Fianarantsoa Dist.</collectingRegion>
,
<collectingMunicipality id="A64F26786542A4189CD2AAA94006B195" box="[702,819,942,968]" pageId="10" pageNumber="252">Fanovana</collectingMunicipality>
,
<location id="434BEAD96542A4189D2DAAA940CDB195" LSID="urn:lsid:plazi:treatment:CE3D0D146542A4199C26A87241BEB407:434BEAD96542A4189D2DAAA940CDB195" box="[833,1016,942,968]" country="Madagascar" municipality="Fanovana" name="Oriental Forest" pageId="10" pageNumber="252" stateProvince="Fianarantsoa Dist.">Oriental Forest</location>
;
<date id="322A9AC26542A4189A6BAAA947F7B195" box="[1031,1218,942,968]" pageId="10" pageNumber="252" value="1937-01" valueMax="1937-05-31" valueMin="1937-01-00">
<collectingDate id="226E632A6542A4189A6BAAA947F7B195" box="[1031,1218,942,968]" pageId="10" pageNumber="252" value="1937-01" valueMax="1937-05-31" valueMin="1937-01-00">Jan.–May 1937</collectingDate>
</date>
;
<collectorName id="EB61D9D46542A4189ABDAAA94642B195" box="[1233,1399,942,968]" pageId="10" pageNumber="252">C. Lamberton</collectorName>
leg.;
<collectionCode id="208524C76542A4189E9AAAD64274B1B6" box="[246,321,977,1003]" collectionName="USA, Pennsylvania, Philadelphia, Academy of Natural Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="10" pageNumber="252">ANSP</collectionCode>
,
<specimenCode id="163214796542A4189F27AAD64101B1B6" box="[331,564,977,1003]" pageId="10" pageNumber="252">ANSP ENT 142916</specimenCode>
(6592).
</materialsCitation>
</paragraph>
<paragraph id="462BBC026542A4189E86AD154261B671" blockId="10.[189,1099,1042,1104]" box="[234,340,1042,1068]" pageId="10" pageNumber="252">
<emphasis id="74E060106542A4189E86AD154261B671" bold="true" box="[234,340,1042,1068]" pageId="10" pageNumber="252">
<typeStatus id="992F02A06542A4189E86AD154261B671" box="[234,340,1042,1068]" pageId="10" pageNumber="252" type="allotype">Allotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="462BBC026542A4189ED1AD31477EB60D" blockId="10.[189,1099,1042,1104]" box="[189,1099,1077,1104]" pageId="10" pageNumber="252">
<materialsCitation id="F6FCB65F6542A4189ED1AD31477EB60D" box="[189,1099,1077,1104]" collectingDate="1937-01" collectingDateMax="1937-05-31" collectingDateMin="1937-01" collectionCode="ANSP" collectorName="C. Lamberton" country="Madagascar" location="MADAGASCAR" municipality="Fanovana" pageId="10" pageNumber="252" specimenCode="ENT 142917" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa" typeStatus="allotype">
<collectingCountry id="3E83FC926542A4189ED1AD3142BBB60D" box="[189,398,1078,1104]" name="Madagascar" pageId="10" pageNumber="252">MADAGASCAR</collectingCountry>
•
<specimenCount id="5092778B6542A4189FCAAD314288B60D" box="[422,445,1078,1104]" count="1" pageId="10" pageNumber="252" type="female">♀</specimenCount>
; same data as for holotype;
<collectionCode id="208524C76542A4189D6AAD314064B60D" box="[774,849,1078,1104]" collectionName="USA, Pennsylvania, Philadelphia, Academy of Natural Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="10" pageNumber="252">ANSP</collectionCode>
,
<collectionCode id="208524C76542A4189D36AD314091B60D" box="[858,932,1078,1104]" collectionName="USA, Pennsylvania, Philadelphia, Academy of Natural Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="10" pageNumber="252">ANSP</collectionCode>
<specimenCode id="163214796542A4189DC0AD314772B60D" box="[940,1095,1078,1104]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="10" pageNumber="252">ENT 142917</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="0E8EEF896542A4199ED1AD72478DB1D3" lastPageId="11" lastPageNumber="253" pageId="10" pageNumber="252" type="description">
<paragraph id="462BBC026542A4189ED1AD724272B6D2" blockId="10.[189,327,1141,1167]" box="[189,327,1141,1167]" pageId="10" pageNumber="252">
<heading id="1D630B6E6542A4189ED1AD724272B6D2" bold="true" box="[189,327,1141,1167]" fontSize="11" level="3" pageId="10" pageNumber="252" reason="3">
<emphasis id="74E060106542A4189ED1AD724272B6D2" bold="true" box="[189,327,1141,1167]" pageId="10" pageNumber="252">Description</emphasis>
</heading>
</paragraph>
<paragraph id="462BBC026542A4189ED1ADA34052B6E3" blockId="10.[189,871,1188,1215]" box="[189,871,1188,1215]" pageId="10" pageNumber="252">
Body length: 10.5–17.0 mm. Wing length:
<quantity id="816C11E76542A4189CD9ADA34056B6E3" box="[693,867,1188,1215]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.25" metricValueMin="1.05" pageId="10" pageNumber="252" unit="mm" value="11.5" valueMax="12.5" valueMin="10.5">10.5–12.5 mm</quantity>
.
</paragraph>
<paragraph id="462BBC026542A4189E86ADE34212B6A3" blockId="10.[187,1401,1252,1525]" box="[234,295,1252,1278]" pageId="10" pageNumber="252">
<emphasis id="74E060106542A4189E86ADE34212B6A3" bold="true" box="[234,295,1252,1278]" pageId="10" pageNumber="252">Male</emphasis>
</paragraph>
<paragraph id="462BBC026542A4189ED1AC0F4734B7A9" blockId="10.[187,1401,1252,1525]" pageId="10" pageNumber="252">
<smallCapsWord id="40CD2ADE6542A4189ED1AC0F43C9B77D" baselines="1307,1307" box="[189,252,1288,1314]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="10" pageNumber="252">HEAD</smallCapsWord>
(
<figureCitation id="DEAFA0876542A4189F61AC004256B77C" box="[269,355,1287,1314]" captionStart="Figs 11–16" captionStartId="31.[189,242,1857,1883]" captionTargetBox="[216,1384,265,1815]" captionTargetId="figure-13@31.[203,1384,265,1815]" captionTargetPageId="31" captionText="Figs 11–16. Species of Mallota Bezzi, 1822. Head, lateral view. 11. M. hirsuta Hull, 1941, holotype, ♀ (ANSP ENT 142915 (6591)). 12. M. meromacrimima Hull, 1941, holotype, ♂ (ANSP ENT 142916 (6592)). 13. M. aperta (Bezzi, 1912), ♀ (RBINS, Eala, Aug. 1935, J. Ghesquière). 14. M. dasyops (Wiedemann, 1819), ♂ (NMSA DIP 211695). 15. M. glabra sp. nov., paratype, ♂ (NMSA DIP 65015). 16. M. hircus sp. nov., paratype, ♀ (RMCA ENT 000037957)." figureDoi="http://doi.org/10.5281/zenodo.13914504" httpUri="https://zenodo.org/record/13914504/files/figure.png" pageId="10" pageNumber="252">Fig. 12</figureCitation>
). Eye bare; holoptic, eye contiguity for distance equal to ocellar triangle, ommatidia equal in size. Frons strongly protruding, in lateral view beyond facial tubercle; ground colour yellow; with medium long to long pale yellow pile; anterior of ocellar triangle black medium long pile. Ocellar triangle black; medium long black pilose. Face ground colour yellow, genae black; with whitish pollinosity, facial tubercle and medial part ventral of tubercle non-pollinose; with dispersed long yellow pile along dorsolateral margins, otherwise bare; facial tubercle distinctly pronounced. Antennal segments orange; arista bare, orange at base, darker distally; postpedicel wider than long.
</paragraph>
<paragraph id="462BBC026542A4189ED1AF1B43C2B571" blockId="10.[188,1400,1564,1836]" pageId="10" pageNumber="252">
<smallCapsWord id="40CD2ADE6542A4189ED1AF1B422CB469" baselines="1583,1583" box="[189,281,1564,1590]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="10" pageNumber="252">THORAX</smallCapsWord>
(
<figureCitation id="DEAFA0876542A4189F46AF1B4244B46B" box="[298,369,1564,1590]" captionStart="Figs 1–2" captionStartId="8.[189,242,1906,1932]" captionTargetBox="[189,1349,270,1858]" captionTargetId="figure-12@8.[189,1398,270,1858]" captionTargetPageId="8" captionText="Figs 1–2. Species of Mallota Bezzi, 1822. Habitus, lateral view. 1. M. hirsuta Hull, 1941, holotype, ♀ (ANSP ENT 142915 (6591)). 2. M. meromacrimima Hull, 1941, holotype, ♂ (ANSP ENT 142916 (6592))." figureDoi="http://doi.org/10.5281/zenodo.13914494" httpUri="https://zenodo.org/record/13914494/files/figure.png" pageId="10" pageNumber="252">Fig. 2</figureCitation>
). Scutum subshining black; anteriorly, posteriorly and along transverse suture with grey pollinosity; anterior third with medium long to short yellow pile intermixed with shorter black pile, black pile predominant in central part; posterior two-thirds with short black pile, along posterior margin with few pale yellow hairs, postalar callus with longer yellow pile. Scutellum yellow to dark yellow, anterior margin narrowly darker; with long pale yellow pile; unmarginated. Pleura ground colour black; posterior anepisternum with long and dense pale yellow pile, katepisternum with dispersed long pale yellow pile, anterior and dorsomedial parts anepimeron with dispersed long black pile; otherwise pleura bare.
</paragraph>
<paragraph id="462BBC026542A4199ED1AE5347A5B31B" blockId="10.[189,1400,1875,1972]" lastBlockId="11.[189,1399,265,327]" lastPageId="11" lastPageNumber="253" pageId="10" pageNumber="252">
<smallCapsWord id="40CD2ADE6542A4189ED1AE5343C3B531" baselines="1895,1895" box="[189,246,1876,1902]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Legs" pageId="10" pageNumber="252">LEGS</smallCapsWord>
. Femora all thickened, black; with short dense black pile except ventrally where long black pile, sometimes intermixed with few pale yellow hairs; profemur posteriorly at base with longer pale yellow pile; mesofemur posteriorly along basal two-thirds with longer pale yellow pile continued distally by longer black pile; metafemur (
<figureCitation id="DEAFA0876543A4199C46A80E41B6B37E" box="[554,643,265,291]" captionStart="Figs 25–30" captionStartId="34.[189,242,1857,1883]" captionTargetBox="[202,1386,265,1819]" captionTargetId="figure-11@34.[202,1386,265,1819]" captionTargetPageId="34" captionText="Figs 25–30. Species of Mallota Bezzi, 1822. Metaleg, anterior view.25. M. hirsuta Hull, 1941, holotype, ♀ (ANSP ENT 142915 (6591)). 26. M. meromacrimima Hull, 1941, holotype, ♂ (ANSP ENT 142916 (6592)). 27. M. aperta (Bezzi, 1912), ♀ (RBINS, Eala, 5 Oct. 1936, J. Ghesquière). 28. M. dasyops (Wiedemann, 1819), ♀ (NMSA DIP 175981). 29. M. glabra sp. nov., paratype, ♂ (NMSA DIP 65015). 30. M. hircus sp. nov., paratype, ♂ (RMCA ENT 000037959)." figureDoi="http://doi.org/10.5281/zenodo.13914512" httpUri="https://zenodo.org/record/13914512/files/figure.png" pageId="11" pageNumber="253">Fig. 26</figureCitation>
) anterodorsally along basal two-thirds with row of longer pale yellow pile. Tibia and tarsal segments orange to red-brown; with short whitish pile.
</paragraph>
<paragraph id="462BBC026543A4199ED1A86942C6B393" blockId="11.[189,1398,366,462]" pageId="11" pageNumber="253">
<smallCapsWord id="40CD2ADE6543A4199ED1A86943CBB3DB" baselines="385,385" box="[189,254,366,392]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Wing" pageId="11" pageNumber="253">WING</smallCapsWord>
(
<figureCitation id="DEAFA0876543A4199F7BA8694242B3D5" box="[279,375,366,392]" captionStart="Figs 37–39" captionStartId="36.[189,242,1901,1927]" captionTargetBox="[189,1377,265,1853]" captionTargetId="figure-12@36.[189,1399,265,1853]" captionTargetPageId="36" captionText="Figs 37–39. Species of Mallota Bezzi, 1822. Right wing. 37. M. hirsuta Hull, 1941, holotype, ♀ (ANSP ENT 142915 (6591)). 38. M. meromacrimima Hull, 1941, holotype, ♂ (ANSP ENT 142916 (6592)). 39. M. aperta (Bezzi, 1912), holotype, ♂ (MCSNG)." figureDoi="http://doi.org/10.5281/zenodo.13914516" httpUri="https://zenodo.org/record/13914516/files/figure.png" pageId="11" pageNumber="253">Fig. 38</figureCitation>
). Slightly fumose, more so along anterior margin, widening distally; most areas microtrichose. Stigmal cross-vein present between distal end of vein Sc and middle of vein R 1. Vein R 4+5 sinuate, without appendix.
</paragraph>
<paragraph id="462BBC026543A4199ED1A8F147FFB0E2" blockId="11.[188,1401,501,704]" pageId="11" pageNumber="253">
<smallCapsWord id="40CD2ADE6543A4199ED1A8F14219B053" baselines="521,521" box="[189,300,502,528]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Abdomen" pageId="11" pageNumber="253">ABDOMEN</smallCapsWord>
(
<figureCitation id="DEAFA0876543A4199F2CA8F242AFB04D" box="[320,410,501,528]" captionStart="Figs 47–50" captionStartId="40.[189,242,1892,1918]" captionTargetBox="[190,1398,266,1839]" captionTargetId="figure-11@40.[190,1398,266,1849]" captionTargetPageId="40" captionText="Figs 47–50. Species of Mallota Bezzi, 1822. Abdomen, dorsal view.47. M. hirsuta Hull, 1941, holotype, ♀ (ANSP ENT 142915 (6591)). 48. M. meromacrimima Hull, 1941, holotype, ♂ (ANSP ENT 142916 (6592)). 49. M. aperta (Bezzi, 1912), holotype, ♂ (MCSNG). 50. M. dasyops (Wiedemann, 1819), ♂ (NMSA DIP 211695)." figureDoi="http://doi.org/10.5281/zenodo.13914524" httpUri="https://zenodo.org/record/13914524/files/figure.png" pageId="11" pageNumber="253">Fig. 48</figureCitation>
). Mainly dark reddish brown, along anterior margin of tergum 2 black; with short black pile except tergum 1 with longer yellow-orange pilosity and white pollinosity; tergum 2 along anterolateral margins with longer yellow-orange pile; anterior margin of terga 3 and 4 with narrow fascia of white pollinosity, interrupted medially, and short whitish pile, anterolateral slightly widening; other terga laterally with longer whitish pile. Sterna black to red-brown; sterna 1–3 with very long, dispersed whitish pile intermixed with few shorter black pile, sterna 4–5 with dispersed black pile.
</paragraph>
<paragraph id="462BBC026543A4199E86ABE04270B15C" blockId="11.[187,1399,743,910]" box="[234,325,743,769]" pageId="11" pageNumber="253">
<emphasis id="74E060106543A4199E86ABE04270B15C" bold="true" box="[234,325,743,769]" pageId="11" pageNumber="253">Female</emphasis>
</paragraph>
<paragraph id="462BBC026543A4199ED1AA0D478DB1D3" blockId="11.[187,1399,743,910]" pageId="11" pageNumber="253">As male except eyes dichoptic, frons yellow, in dorsal half medially slightly darker; with medium long to long pale yellow pile. Thorax, along posterior margin of scutum with pale yellow pile. Femora ventrally without long dense pilosity; only metafemur with dispersed black pile. Metafemur only thickened. Wing, fumose area along anterior margin less pronounced. Abdominal terga more orange-red.</paragraph>
</subSubSection>
<subSubSection id="0E8EEF896543A4199ED1AAB24264B1A3" box="[189,342,949,1022]" pageId="11" pageNumber="253" type="distribution">
<paragraph id="462BBC026543A4199ED1AAB24263B192" blockId="11.[189,342,949,975]" box="[189,342,949,975]" pageId="11" pageNumber="253">
<heading id="1D630B6E6543A4199ED1AAB24263B192" bold="true" box="[189,342,949,975]" fontSize="11" level="3" pageId="11" pageNumber="253" reason="3">
<emphasis id="74E060106543A4199ED1AAB24263B192" bold="true" box="[189,342,949,975]" pageId="11" pageNumber="253">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="462BBC026543A4199ED1AAE34264B1A3" blockId="11.[189,337,996,1022]" box="[189,337,996,1022]" pageId="11" pageNumber="253">
<collectingCountry id="3E83FC926543A4199ED1AAE34278B1A3" box="[189,333,996,1022]" name="Madagascar" pageId="11" pageNumber="253">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="0E8EEF896543A4199ED1AD2241BEB407" pageId="11" pageNumber="253" type="discussion">
<paragraph id="462BBC026543A4199ED1AD224271B662" blockId="11.[189,324,1061,1087]" box="[189,324,1061,1087]" pageId="11" pageNumber="253">
<heading id="1D630B6E6543A4199ED1AD224271B662" bold="true" box="[189,324,1061,1087]" fontSize="11" level="3" pageId="11" pageNumber="253" reason="3">
<emphasis id="74E060106543A4199ED1AD224271B662" bold="true" box="[189,324,1061,1087]" pageId="11" pageNumber="253">Comments</emphasis>
</heading>
</paragraph>
<paragraph id="462BBC026543A4199ED1AD5341BEB407" blockId="11.[188,1399,1108,1627]" pageId="11" pageNumber="253">
Both species (
<taxonomicName id="8194C7816543A4199F1EAD5342C1B633" authorityName="Hull" authorityYear="1941" box="[370,500,1108,1134]" class="Insecta" family="Syrphidae" genus="Mallota" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="253" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="74E060106543A4199F1EAD5342C1B633" box="[370,500,1108,1134]" italics="true" pageId="11" pageNumber="253">M. hirsuta</emphasis>
</taxonomicName>
and
<taxonomicName id="8194C7816543A4199C50AD534019B630" authorityName="Hull" authorityYear="1941" box="[572,812,1108,1133]" class="Insecta" family="Syrphidae" genus="Mallota" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="253" phylum="Arthropoda" rank="species" species="meromacrimima">
<emphasis id="74E060106543A4199C50AD534019B630" box="[572,812,1108,1133]" italics="true" pageId="11" pageNumber="253">M. meromacrimima</emphasis>
</taxonomicName>
) are similar in most respects especially when comparing the female
<typeStatus id="992F02A06543A4199FBDAD70410DB6CC" box="[465,568,1143,1169]" pageId="11" pageNumber="253" type="holotype">holotype</typeStatus>
of
<taxonomicName id="8194C7816543A4199C0BAD7F41D3B6CC" authorityName="Hull" authorityYear="1941" box="[615,742,1143,1169]" class="Insecta" family="Syrphidae" genus="Mallota" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="253" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="74E060106543A4199C0BAD7F41D3B6CC" box="[615,742,1143,1169]" italics="true" pageId="11" pageNumber="253">M. hirsuta</emphasis>
</taxonomicName>
with the female
<typeStatus id="992F02A06543A4199DD0AD70472EB6CC" box="[956,1051,1143,1169]" pageId="11" pageNumber="253" type="allotype">allotype</typeStatus>
of
<taxonomicName id="8194C7816543A4199A26AD7F4602B6CC" authorityName="Hull" authorityYear="1941" box="[1098,1335,1143,1169]" class="Insecta" family="Syrphidae" genus="Mallota" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="253" phylum="Arthropoda" rank="species" species="meromacrimima">
<emphasis id="74E060106543A4199A26AD7F4757B6CC" box="[1098,1122,1144,1169]" italics="true" pageId="11" pageNumber="253">M</emphasis>
.
<emphasis id="74E060106543A4199A1AAD7F4602B6CC" box="[1142,1335,1144,1169]" italics="true" pageId="11" pageNumber="253">meromacrimima</emphasis>
</taxonomicName>
. The main difference is the colouration of abdominal terga, the presence of distinct narrow fasciae of white pollinosity along the anterior margin of terga 3 and
<quantity id="816C11E76543A4199D48ADBA4067B68A" box="[804,850,1213,1239]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="11" pageNumber="253" unit="in" value="4.0">4 in</quantity>
<taxonomicName id="8194C7816543A4199D36ADB9472EB68A" authorityName="Hull" authorityYear="1941" box="[858,1051,1214,1239]" class="Insecta" family="Syrphidae" genus="Mallota" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="253" phylum="Arthropoda" rank="species" species="meromacrimima">
<emphasis id="74E060106543A4199D36ADB9472EB68A" box="[858,1051,1214,1239]" italics="true" pageId="11" pageNumber="253">meromacrimima</emphasis>
</taxonomicName>
(see
<figureCitation id="DEAFA0876543A4199A36ADBA479AB68A" box="[1114,1199,1213,1239]" captionStart="Figs 47–50" captionStartId="40.[189,242,1892,1918]" captionTargetBox="[190,1398,266,1839]" captionTargetId="figure-11@40.[190,1398,266,1849]" captionTargetPageId="40" captionText="Figs 47–50. Species of Mallota Bezzi, 1822. Abdomen, dorsal view.47. M. hirsuta Hull, 1941, holotype, ♀ (ANSP ENT 142915 (6591)). 48. M. meromacrimima Hull, 1941, holotype, ♂ (ANSP ENT 142916 (6592)). 49. M. aperta (Bezzi, 1912), holotype, ♂ (MCSNG). 50. M. dasyops (Wiedemann, 1819), ♂ (NMSA DIP 211695)." figureDoi="http://doi.org/10.5281/zenodo.13914524" httpUri="https://zenodo.org/record/13914524/files/figure.png" pageId="11" pageNumber="253">Fig. 48</figureCitation>
) and the pilosity on the scutum. However, abdominal tergal colouration has been shown to be very variable in some eristaline genera; for example, see revisions of
<taxonomicName id="8194C7816543A4199D69AC034790B743" authority="(De Meyer et al. 2020 b)" baseAuthorityName="De Meyer" baseAuthorityYear="2020" box="[773,1189,1284,1310]" class="Insecta" family="Syrphidae" genus="Senaspis" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="74E060106543A4199D69AC034059B743" box="[773,876,1284,1310]" italics="true" pageId="11" pageNumber="253">Senaspis</emphasis>
(
<bibRefCitation id="2205C1F36543A4199DECAC0347A8B743" author="De Meyer M. &amp; Goergen G. &amp; Jordaens K." box="[896,1181,1284,1310]" pageId="11" pageNumber="253" pagination="83 - 160" refId="ref20694" refString="De Meyer M., Goergen G. &amp; Jordaens K. 2020 b. Taxonomic revision of the Afrotropical Senaspis Macquart (Diptera: Syrphidae). ZooKeys 1003: 83 - 160. https: // doi. org / 10.3897 / zookeys. 1003.56557" type="journal article" year="2020">
De Meyer
<emphasis id="74E060106543A4199A64AC034772B743" box="[1032,1095,1284,1310]" italics="true" pageId="11" pageNumber="253">et al.</emphasis>
2020b
</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="8194C7816543A4199A8BAC034258B71F" authority="Rondani, 1857" authorityName="Rondani" authorityYear="1857" class="Insecta" family="Syrphidae" genus="Mesembrius" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="74E060106543A4199A8BAC034643B743" box="[1255,1398,1284,1310]" italics="true" pageId="11" pageNumber="253">Mesembrius</emphasis>
Rondani, 1857
</taxonomicName>
(
<bibRefCitation id="2205C1F36543A4199F13AC20414DB71C" author="Jordaens K. &amp; Goergen G. &amp; Skevington J. H. &amp; Kelso S. &amp; De Meyer M." box="[383,632,1319,1345]" pageId="11" pageNumber="253" pagination="1 - 141" refId="ref21027" refString="Jordaens K., Goergen G., Skevington J. H., Kelso S. &amp; De Meyer M. 2021. Morphological and molecular revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera: Syrphidae). ZooKeys 1046: 1 - 141. https: // doi. org / 10.3897 / zookeys. 1046.57052" type="journal article" year="2021">
Jordaens
<emphasis id="74E060106543A4199F9DAC2F411AB71C" box="[497,559,1319,1345]" italics="true" pageId="11" pageNumber="253">et al.</emphasis>
2021
</bibRefCitation>
). The poor condition of the sole specimen of
<taxonomicName id="8194C7816543A4199AC8AC2F4614B71C" authorityName="Hull" authorityYear="1941" box="[1188,1313,1319,1345]" class="Insecta" family="Syrphidae" genus="Mallota" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="253" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="74E060106543A4199AC8AC2F4614B71C" box="[1188,1313,1319,1345]" italics="true" pageId="11" pageNumber="253">M. hirsuta</emphasis>
</taxonomicName>
makes comparison between both with regard to scutal and abdominal character states difficult.
<specimenCount id="5092778B6543A4199AB4AC4D4643B738" box="[1240,1398,1354,1381]" count="1" pageId="11" pageNumber="253" type="generic" typeStatus="paratype">One paratype</specimenCount>
(not studied) of
<taxonomicName id="8194C7816543A4199F1BAC69416BB7DA" authorityName="Hull" authorityYear="1941" box="[375,606,1390,1415]" class="Insecta" family="Syrphidae" genus="Mallota" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="253" phylum="Arthropoda" rank="species" species="meromacrimima">
<emphasis id="74E060106543A4199F1BAC69416BB7DA" box="[375,606,1390,1415]" italics="true" pageId="11" pageNumber="253">M. meromacrimima</emphasis>
</taxonomicName>
is collected at the same spot as the
<typeStatus id="992F02A06543A4199D90AC6A4756B7DA" box="[1020,1123,1389,1415]" pageId="11" pageNumber="253" type="holotype">holotype</typeStatus>
of
<taxonomicName id="8194C7816543A4199AE4AC694634B7DA" authorityName="Hull" authorityYear="1941" box="[1160,1281,1389,1415]" class="Insecta" family="Syrphidae" genus="Mallota" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="253" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="74E060106543A4199AE4AC694634B7DA" box="[1160,1281,1389,1415]" italics="true" pageId="11" pageNumber="253">M. hirsuta</emphasis>
</taxonomicName>
. The type locality for
<taxonomicName id="8194C7816543A4199F25AC964106B7F7" authorityName="Hull" authorityYear="1941" box="[329,563,1425,1450]" class="Insecta" family="Syrphidae" genus="Mallota" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="253" phylum="Arthropoda" rank="species" species="meromacrimima">
<emphasis id="74E060106543A4199F25AC964106B7F7" box="[329,563,1425,1450]" italics="true" pageId="11" pageNumber="253">M. meromacrimima</emphasis>
</taxonomicName>
is about
<quantity id="816C11E76543A4199CCCAC9641DFB7F7" box="[672,746,1424,1451]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="253" unit="km" value="30.0">30 km</quantity>
east of the one for
<taxonomicName id="8194C7816543A4199DBAAC964765B7F7" authorityName="Hull" authorityYear="1941" box="[982,1104,1424,1450]" class="Insecta" family="Syrphidae" genus="Mallota" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="253" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="74E060106543A4199DBAAC964765B7F7" box="[982,1104,1424,1450]" italics="true" pageId="11" pageNumber="253">M. hirsuta</emphasis>
</taxonomicName>
. It is likely that both are actually varied representatives of the same species and the late F.C. Thompson already indicated this as such in the
<emphasis id="74E060106543A4199F2FACD04106B7AC" box="[323,563,1495,1521]" italics="true" pageId="11" pageNumber="253">Systema Dipterorum</emphasis>
website (http://www.diptera.org/), but the proposed synonymy was not officially published. However, because of the limited material available which does not allow a detailed investigation of the intra-specific variability and lacking molecular data, we do not propose a synonymy here. No DNA barcodes were obtained.
</paragraph>
</subSubSection>
</treatment>
</document>
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