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<document id="0425123085B9E0E771B83996A15E0095" ID-DOI="10.5281/zenodo.210864" ID-GBIF-Dataset="bc8606df-9e8b-44af-a522-0fd0ae7b1349" ID-ISSN="1175-5326" ID-Zenodo-Dep="210864" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460475468456" checkinUser="plazi" docAuthor="Borkent, Art &amp; Grafe, Ulmar" docDate="2012" docId="9720782FAA08FFADFF78FC9446DB0EF9" docLanguage="en" docName="zt03279p045.pdf" docOrigin="Zootaxa 3279" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Corethrella (Corethrella) lutea Borkent &amp; Grafe, new species" docType="treatment" docVersion="8" lastPageNumber="9" masterDocId="6B190057AA0FFFA5FFEFFFE147380A5C" masterDocTitle="The Frog-Biting Midges of Borneo — From Two to Eleven Species (Corethrellidae: Diptera)" masterLastPageNumber="45" masterPageNumber="1" pageNumber="8" updateTime="1698306645417" updateUser="plazi">
<mods:mods id="3AEC8389780EDA8D1FB903DD66803CA9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="38638811E266A39B3D2DEA8B7428AB7C">The Frog-Biting Midges of Borneo — From Two to Eleven Species (Corethrellidae: Diptera)</mods:title>
</mods:titleInfo>
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<mods:namePart id="82C2D02929C33C65668858187A699922">Borkent, Art</mods:namePart>
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<mods:namePart id="CED88C03D4D90C8B90EE8F8CD1B92C12">Grafe, Ulmar</mods:namePart>
</mods:name>
<mods:typeOfResource id="4911628D56A96AD198F698329ED0012A">text</mods:typeOfResource>
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<mods:date id="2A1ECF9DE30B2A6EBC34536F577C4FE6">2012</mods:date>
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<mods:number id="A206438FCCC2951B632CBA717592472D">3279</mods:number>
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<mods:identifier id="314F517D6B5123DA430EC0E7C638B55A" type="DOI">10.5281/zenodo.210864</mods:identifier>
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<mods:identifier id="60E469619BF2DEC53A334E6403A98650" type="ISSN">1175-5326</mods:identifier>
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<treatment id="9720782FAA08FFADFF78FC9446DB0EF9" ID-DOI="http://doi.org/10.5281/zenodo.5619820" ID-GBIF-Taxon="119516206" ID-Zenodo-Dep="5619820" LSID="urn:lsid:plazi:treatment:9720782FAA08FFADFF78FC9446DB0EF9" httpUri="http://treatment.plazi.org/id/9720782FAA08FFADFF78FC9446DB0EF9" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<subSubSection id="57939AB2AA08FFA2FF78FC94454209EC" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="1F36C939AA08FFA2FF78FC9444BF09D3" blockId="7.[151,903,885,944]" box="[151,903,885,911]" pageId="7" pageNumber="8">
<heading id="447E7E55AA08FFA2FF78FC9444BF09D3" bold="true" box="[151,903,885,911]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<emphasis id="2DFD152BAA08FFA2FF78FC9444BF09D3" bold="true" box="[151,903,885,911]" pageId="7" pageNumber="8">
<taxonomicName id="D889B2BAAA08FFA2FF78FC9445D209D3" ID-CoL="BCR9H" authority="Borkent &amp; Grafe" authorityName="Borkent &amp; Grafe" box="[151,746,885,911]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lutea" status="sp. nov." subGenus="Corethrella">
<emphasis id="2DFD152BAA08FFA2FF78FC94461A09D3" bold="true" box="[151,290,885,911]" italics="true" pageId="7" pageNumber="8">Corethrella</emphasis>
(
<emphasis id="2DFD152BAA08FFA2FEDDFC94468409D3" bold="true" box="[306,444,885,911]" italics="true" pageId="7" pageNumber="8">Corethrella</emphasis>
)
<emphasis id="2DFD152BAA08FFA2FE21FC94453209D3" bold="true" box="[462,522,885,911]" italics="true" pageId="7" pageNumber="8">lutea</emphasis>
Borkent &amp; Grafe
</taxonomicName>
,
<taxonomicNameLabel id="36CEA850AA08FFA2FD18FC9444BF09D3" box="[759,903,885,911]" pageId="7" pageNumber="8" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="1F36C939AA08FFA2FF78FC79454209EC" blockId="7.[151,903,885,944]" box="[151,634,920,944]" pageId="7" pageNumber="8">
(
<figureCitation id="87B2D5BCAA08FFA2FF70FC7947D509EC" box="[159,237,920,944]" captionStart="FIGURE 2" captionStartId="24.[151,248,1916,1939]" captionTargetBox="[172,1414,180,1895]" captionTargetId="figure@24.[172,1416,180,1895]" captionTargetPageId="24" captionText="FIGURE 2. Female adult Corethrella feeding on calling male frogs in Borneo. A. C. tigrina on Metaphrynella sundana (Peters) (Ashton Trail in the Ulu Temburong National Park, Brunei Darussalam). B. C. lutea resting on somewhat drooping leaf after feeding on Rhacophorus appendiculatus (Günther) (Sungai Ingei Conservation Forest, 8.2 km S. Melilas, Brunei Darussalam). C. C. bipigmenta on Megophrys nasuta (Ulu Temburong National Park, Brunei). All photographs by second author." httpUri="https://zenodo.org/record/210866/files/figure.png" pageId="7" pageNumber="8">Figs. 2</figureCitation>
B, 4B, 6B, 8B, 9B, 12B, 13B, 15A)
</paragraph>
</subSubSection>
<subSubSection id="57939AB2AA08FFA2FF78FC3F46630EF9" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="1F36C939AA08FFA2FF78FC3F46630EF9" blockId="7.[151,1436,990,2030]" pageId="7" pageNumber="8">
DIAGNOSIS:
<emphasis id="2DFD152BAA08FFA2FEA0FC3F46EF09AA" box="[335,471,990,1014]" italics="true" pageId="7" pageNumber="8">
<collectingCountry id="679E89A9AA08FFA2FEA0FC3F46B109AA" box="[335,393,990,1014]" name="Malta" pageId="7" pageNumber="8">Male</collectingCountry>
adult.
</emphasis>
Unknown.
<emphasis id="2DFD152BAA08FFA2FD9CFC3F442E09AA" box="[627,790,990,1014]" italics="true" pageId="7" pageNumber="8">Female adult.</emphasis>
Only extant species of
<taxonomicName id="D889B2BAAA08FFA2FBD5FC3F438309AA" box="[1082,1211,990,1014]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="2DFD152BAA08FFA2FBD5FC3F438309AA" box="[1082,1211,990,1014]" italics="true" pageId="7" pageNumber="8">Corethrella</emphasis>
</taxonomicName>
in the Palaearctic, Oriental, Oceanian and northern Australasian Regions with the combination of a plain wing (
<figureCitation id="87B2D5BCAA08FFA2FAF0FBE042B60E45" box="[1311,1422,1025,1049]" captionStart="FIGURE 12. A – K" captionStartId="34.[151,250,1944,1967]" captionTargetBox="[151,1435,193,1921]" captionTargetId="figure@34.[151,1436,193,1922]" captionTargetPageId="34" captionText="FIGURE 12. A – K. Wings of female Corethrella." httpUri="https://zenodo.org/record/210876/files/figure.png" pageId="7" pageNumber="8">Fig. 12B</figureCitation>
), flagellomeres 1–4 relatively short (
<figureCitation id="87B2D5BCAA08FFA2FDCDFBC5455C0E60" box="[546,612,1060,1084]" captionStart="FIGURE 8" captionStartId="30.[151,250,1973,1996]" captionTargetBox="[185,1417,209,1928]" captionTargetId="figure@30.[178,1422,193,1959]" captionTargetPageId="30" captionText="FIGURE 8. Antennae and male genitalia of Corethrella. A – K. Antennae of female Corethrella. L. Antenna of male C. tigrina. M. Male genitalia of C. tigrina, dorsal view; parts drawn in proportion to each other." httpUri="https://zenodo.org/record/210872/files/figure.png" pageId="7" pageNumber="8">Fig. 8</figureCitation>
B), scutum a lighter brown, contrasting with darker, mottled, pleura (
<figureCitation id="87B2D5BCAA08FFA2FA80FBC547800E03" captionStart="FIGURE 9. A – D" captionStartId="31.[151,250,1975,1998]" captionTargetBox="[157,1431,200,1947]" captionTargetId="figure@31.[151,1436,193,1952]" captionTargetPageId="31" captionText="FIGURE 9. A – D. Thoraces and legs of female Corethrella, lateral view; slide-mounted." httpUri="https://zenodo.org/record/210873/files/figure.png" pageId="7" pageNumber="8">Fig. 9B</figureCitation>
) and midfemur relatively thick (
<figureCitation id="87B2D5BCAA08FFA2FDD6FBA645AB0E03" box="[569,659,1095,1119]" captionStart="FIGURE 9. A – D" captionStartId="31.[151,250,1975,1998]" captionTargetBox="[157,1431,200,1947]" captionTargetId="figure@31.[151,1436,193,1952]" captionTargetPageId="31" captionText="FIGURE 9. A – D. Thoraces and legs of female Corethrella, lateral view; slide-mounted." httpUri="https://zenodo.org/record/210873/files/figure.png" pageId="7" pageNumber="8">Fig. 9B</figureCitation>
) (compared to that of
<taxonomicName id="D889B2BAAA08FFA2FC4DFBA9432D0E03" box="[930,1045,1096,1119]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nippon">
<emphasis id="2DFD152BAA08FFA2FC4DFBA9432D0E03" box="[930,1045,1096,1119]" italics="true" pageId="7" pageNumber="8">C. nippon</emphasis>
</taxonomicName>
Miyagi). Only extant species in Borneo with the combination of a plain wing (
<figureCitation id="87B2D5BCAA08FFA2FD48FB8B44340EDE" box="[679,780,1130,1154]" captionStart="FIGURE 12. A – K" captionStartId="34.[151,250,1944,1967]" captionTargetBox="[151,1435,193,1921]" captionTargetId="figure@34.[151,1436,193,1922]" captionTargetPageId="34" captionText="FIGURE 12. A – K. Wings of female Corethrella." httpUri="https://zenodo.org/record/210876/files/figure.png" pageId="7" pageNumber="8">Fig. 12B</figureCitation>
) and a thorax with patterned pigmentation (especially the pleura) (
<figureCitation id="87B2D5BCAA08FFA2FF1AFB6C46760EF9" box="[245,334,1165,1189]" captionStart="FIGURE 9. A – D" captionStartId="31.[151,250,1975,1998]" captionTargetBox="[157,1431,200,1947]" captionTargetId="figure@31.[151,1436,193,1952]" captionTargetPageId="31" captionText="FIGURE 9. A – D. Thoraces and legs of female Corethrella, lateral view; slide-mounted." httpUri="https://zenodo.org/record/210873/files/figure.png" pageId="7" pageNumber="8">Fig. 9B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="57939AB2AA08FFA2FF28FB4E45B60DD9" pageId="7" pageNumber="8" type="description">
<paragraph id="1F36C939AA08FFA2FF28FB4E44DB0D3E" blockId="7.[151,1436,990,2030]" pageId="7" pageNumber="8">
DESCRIPTION:
<emphasis id="2DFD152BAA08FFA2FE7CFB5145350E94" box="[403,525,1200,1224]" italics="true" pageId="7" pageNumber="8">
<collectingCountry id="679E89A9AA08FFA2FE7CFB5146F50E94" box="[403,461,1200,1224]" name="Malta" pageId="7" pageNumber="8">Male</collectingCountry>
adult
</emphasis>
. Unknown.
<emphasis id="2DFD152BAA08FFA2FD70FB5144040E94" box="[671,828,1200,1224]" italics="true" pageId="7" pageNumber="8">Female adult.</emphasis>
Descriptive statistics: See
<tableCitation id="520BFC82AA08FFA2FB85FB4E43FD0E9B" box="[1130,1221,1199,1223]" captionStart="TABLE 1" captionStartId="42.[151,235,154,175]" captionTargetBox="[151,1259,203,680]" captionTargetPageId="42" captionText="TABLE 1. Length (in mm) of wing of females of species of Corethrella." httpUri="http://table.plazi.org/id/4BF699B1AA25FF8FFF78FF7B44440AF3" pageId="7" pageNumber="8" tableUuid="4BF699B1AA25FF8FFF78FF7B44440AF3">Tables 1</tableCitation>
–5.
<emphasis id="2DFD152BAA08FFA2FB16FB5142010E94" bold="true" box="[1273,1337,1200,1224]" pageId="7" pageNumber="8">Head</emphasis>
: Outline in anterior view somewhat circular (
<figureCitation id="87B2D5BCAA08FFA2FDDDFB32454E0EB7" box="[562,630,1235,1259]" captionStart="FIGURE 6" captionStartId="28.[151,250,1929,1952]" captionTargetBox="[151,1435,193,1908]" captionTargetId="figure@28.[151,1436,193,1908]" captionTargetPageId="28" captionText="FIGURE 6. Head capsules of female Corethrella, anterior view." httpUri="https://zenodo.org/record/210870/files/figure.png" pageId="7" pageNumber="8">Fig. 6</figureCitation>
B). Coronal suture elongate, extending ventrally past ommatidia. Two large setae on frons between ventromedial area of ommatidia. Antenna uniformly medium brown; pedicel without distinctive elongate, stout setae. Antennal flagellomeres as in
<figureCitation id="87B2D5BCAA08FFA2FCB6FAF944A30F6C" box="[857,923,1304,1328]" captionStart="FIGURE 8" captionStartId="30.[151,250,1973,1996]" captionTargetBox="[185,1417,209,1928]" captionTargetId="figure@30.[178,1422,193,1959]" captionTargetPageId="30" captionText="FIGURE 8. Antennae and male genitalia of Corethrella. A – K. Antennae of female Corethrella. L. Antenna of male C. tigrina. M. Male genitalia of C. tigrina, dorsal view; parts drawn in proportion to each other." httpUri="https://zenodo.org/record/210872/files/figure.png" pageId="7" pageNumber="8">Fig. 8</figureCitation>
B; sensilla coeloconica on flagellomeres 1–2, 9–13, flagellomeres 1 with several, (10), 11–12 with 2 sensilla; flagellomere 13 with well-developed apical bifurcation. Clypeus (
<figureCitation id="87B2D5BCAA08FFA2FE7FFABF46EA0F2A" box="[400,466,1374,1398]" captionStart="FIGURE 6" captionStartId="28.[151,250,1929,1952]" captionTargetBox="[151,1435,193,1908]" captionTargetId="figure@28.[151,1436,193,1908]" captionTargetPageId="28" captionText="FIGURE 6. Head capsules of female Corethrella, anterior view." httpUri="https://zenodo.org/record/210870/files/figure.png" pageId="7" pageNumber="8">Fig. 6</figureCitation>
B) somewhat square. Mandible with moderately sized, pointed teeth. Palpus (
<figureCitation id="87B2D5BCAA08FFA2FAD1FABF42B80F2A" box="[1342,1408,1374,1398]" captionStart="FIGURE 6" captionStartId="28.[151,250,1929,1952]" captionTargetBox="[151,1435,193,1908]" captionTargetId="figure@28.[151,1436,193,1908]" captionTargetPageId="28" captionText="FIGURE 6. Head capsules of female Corethrella, anterior view." httpUri="https://zenodo.org/record/210870/files/figure.png" pageId="7" pageNumber="8">Fig. 6</figureCitation>
B) brown; segment 3 swollen apically.
<emphasis id="2DFD152BAA08FFA2FDD2FA6345AE0FC6" bold="true" box="[573,662,1410,1434]" pageId="7" pageNumber="8">Thorax</emphasis>
(
<figureCitation id="87B2D5BCAA08FFA2FD48FA60443A0FC5" box="[679,770,1409,1433]" captionStart="FIGURE 9. A – D" captionStartId="31.[151,250,1975,1998]" captionTargetBox="[157,1431,200,1947]" captionTargetId="figure@31.[151,1436,193,1952]" captionTargetPageId="31" captionText="FIGURE 9. A – D. Thoraces and legs of female Corethrella, lateral view; slide-mounted." httpUri="https://zenodo.org/record/210873/files/figure.png" pageId="7" pageNumber="8">Fig. 9B</figureCitation>
): Scutum mostly light to medium brown, contrasting with darker brown pleura; scutellum, mediotergite, most of pleura dark brown, katepisternum with pale area at midheight, posterior portion of anepimeron paler, pale sclerites around base of wing. Posterior portion of dorsocentral row with 2 elongate setae situated somewhat lateral to one another. Prescutal suture elongate, thick, uninterrupted, extending to near dorsocentral row of setae. Anterior anepisternum divided diagonally by sinuous suture, dorsal portion about equal to ventral portion. Ventral portion of posterior anepisternum triangular, uniformly brown, with anterodorsal margin not thick.
<emphasis id="2DFD152BAA08FFA2FE1CF9B3450B0C37" bold="true" box="[499,563,1618,1643]" pageId="7" pageNumber="8">Wing</emphasis>
(
<figureCitation id="87B2D5BCAA08FFA2FDACF9B245910C37" box="[579,681,1619,1643]" captionStart="FIGURE 12. A – K" captionStartId="34.[151,250,1944,1967]" captionTargetBox="[151,1435,193,1921]" captionTargetId="figure@34.[151,1436,193,1922]" captionTargetPageId="34" captionText="FIGURE 12. A – K. Wings of female Corethrella." httpUri="https://zenodo.org/record/210876/files/figure.png" pageId="7" pageNumber="8">Fig. 12B</figureCitation>
): Apex of R2 slightly distal to apex of M1. Plain, without pattern of pigmented veins and/or scales; veins (other than costa and wing margin) with slender scales. Halter light to medium brown, equal to or somewhat lighter than scutellum.
<emphasis id="2DFD152BAA08FFA2FCB3F97D44AC0CE8" bold="true" box="[860,916,1692,1716]" pageId="7" pageNumber="8">Legs</emphasis>
(
<figureCitation id="87B2D5BCAA08FFA2FC4AF97D44C70CE8" box="[933,1023,1692,1716]" captionStart="FIGURE 9. A – D" captionStartId="31.[151,250,1975,1998]" captionTargetBox="[157,1431,200,1947]" captionTargetId="figure@31.[151,1436,193,1952]" captionTargetPageId="31" captionText="FIGURE 9. A – D. Thoraces and legs of female Corethrella, lateral view; slide-mounted." httpUri="https://zenodo.org/record/210873/files/figure.png" pageId="7" pageNumber="8">Fig. 9B</figureCitation>
): Light brown, forecoxa lighter than mid-, hind coxae, with about basal 0.4 of hind femur dark brown. With only slender setae, lacking scales. Midleg with thick, subapical setae on each of at least tarsomeres 1–3. Claws of each leg equal to those of others; equal on each leg, simple (without inner teeth). Empodia thick.
<emphasis id="2DFD152BAA08FFA2FCF0F8E544AC0D41" bold="true" box="[799,916,1796,1821]" pageId="7" pageNumber="8">Abdomen</emphasis>
(
<figureCitation id="87B2D5BCAA08FFA2FC4AF8E543360D40" box="[933,1038,1796,1820]" captionStart="FIGURE 13. A – J" captionStartId="35.[151,250,2013,2036]" captionTargetBox="[151,1435,193,1989]" captionTargetId="figure@35.[151,1436,193,1991]" captionTargetPageId="35" captionText="FIGURE 13. A – J. Abdomens of female Corethrella, ventral view." httpUri="https://zenodo.org/record/210877/files/figure.png" pageId="7" pageNumber="8">Fig. 13B</figureCitation>
): Pale or light yellow, tergites 2–6 with anterolateral light brown pigmentation, sternites medium brown, 3–5 somewhat darker anteriorly, segment 7 light to dark brown, segments 8–9 dark brown. Cercus light brown or pale.
</paragraph>
<paragraph id="1F36C939AA08FFA2FF78F88E45B60DD9" blockId="7.[151,1436,990,2030]" box="[151,654,1901,1926]" pageId="7" pageNumber="8">
<emphasis id="2DFD152BAA08FFA2FF78F88E46680DDA" box="[151,336,1903,1926]" italics="true" pageId="7" pageNumber="8">Immature stages</emphasis>
. Unknown (but see below).
</paragraph>
</subSubSection>
<subSubSection id="57939AB2AA08FFADFF28F87146DB0EF9" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="distribution">
<paragraph id="1F36C939AA08FFADFF28F87146C10A8E" blockId="7.[151,1436,990,2030]" lastBlockId="8.[151,1437,151,1189]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
DISTRIBUTION AND BIONOMICS:
<taxonomicName id="D889B2BAAA08FFA2FD63F87144750DF4" box="[652,845,1936,1960]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lutea">
<emphasis id="2DFD152BAA08FFA2FD63F87144750DF4" box="[652,845,1936,1960]" italics="true" pageId="7" pageNumber="8">Corethrella lutea</emphasis>
</taxonomicName>
is known from
<collectingCountry id="679E89A9AA08FFA2FBECF87143770DF4" box="[1027,1103,1936,1960]" name="Brunei" pageId="7" pageNumber="8">Brunei</collectingCountry>
and
<collectingCountry id="679E89A9AA08FFA2FB68F87143D20DF4" box="[1159,1258,1936,1960]" name="Malaysia" pageId="7" pageNumber="8">Sarawak</collectingCountry>
,
<collectingCountry id="679E89A9AA08FFA2FB18F87142660DF4" box="[1271,1374,1936,1960]" name="Malaysia" pageId="7" pageNumber="8">Malaysia</collectingCountry>
from
<quantity id="D87164DCAA08FFA2FF78F85246320D97" box="[151,266,1971,1995]" metricMagnitude="2" metricUnit="m" metricValue="1.15" metricValueMax="2.0" metricValueMin="0.3" pageId="7" pageNumber="8" unit="m" value="115.0" valueMax="200.0" valueMin="30.0">30–200 m</quantity>
(
<figureCitation id="87B2D5BCAA08FFA2FEF3F85246B00D97" box="[284,392,1971,1995]" captionStart="FIGURE 15. A – B" captionStartId="37.[151,250,1947,1970]" captionTargetBox="[202,1408,229,1887]" captionTargetId="figure@37.[151,1436,193,1924]" captionTargetPageId="37" captionText="FIGURE 15. A – B. Distributions of species of Corethrella from Borneo." httpUri="https://zenodo.org/record/210879/files/figure.png" pageId="7" pageNumber="8">Fig. 15A</figureCitation>
). Habitats include lowland peat swamps (including standing water and very slow moving streams) and steep terrain in mature mixed dipterocarp forest (
<figureCitation id="87B2D5BCAA08FFA2FC86F837448D0DB2" box="[873,949,2006,2030]" captionStart="FIGURE 1" captionStartId="23.[151,250,1857,1880]" captionTargetBox="[211,1379,193,1836]" captionTargetId="figure@23.[211,1379,193,1837]" captionTargetPageId="23" captionText="FIGURE 1. Habitats from which Corethrella adults were collected with a frog-call trap in Brunei (species collected given in parentheses). A. 12 km S. Liang, Brunei (C. tigrina, C. lutea, C. calathicola, C. pauciseta, C. gilva) (authors, for scale). B. Nepenthes ampullaria, 12 km S. Liang, Brunei (C. calathicola). C. 12.5 km S. Liang, Brunei (same species as for A) (first author, for scale). D. 20 km N. Labi, Luagan Lalak Forestry Rec. Site, Brunei (C. lutea). E. Belalong River, Kuala Belalong Field Studies Centre, Brunei (C. lutea, C. bipigmenta, C. mitra). F. Sungai Esu, Ulu Temburong NP, Brunei (C. tigrina, C. bipigmenta, C. mitra) (first author's wife Annette, for scale). G. Pa Umor, 5 km E. Bario (C. bicincta, C. unizona)." httpUri="https://zenodo.org/record/210865/files/figure.png" pageId="7" pageNumber="8">Figs. 1</figureCitation>
A, D, E). They are relatively common in frog-call traps in
<collectingCountry id="679E89A9AA07FFADFEB5FF76469E0AF3" box="[346,422,151,175]" name="Brunei" pageId="8" pageNumber="9">Brunei</collectingCountry>
(
<bibRefCitation id="7B18B4C8AA07FFADFE59FF7645420AF3" author="Grafe" box="[438,634,151,176]" pageId="8" pageNumber="9" refString="Grafe, T. U., Saat, H. B. M., Hagen, N., Kaluza, B., Berudin, Z. B. H. &amp; Wahab, M. A. B. A. (2008) Acoustic localisation of frog hosts by blood-sucking flies Corethrella Coquillet (Diptera: Corethrellidae) in Borneo. Australian Journal of Entomology 47,350 - 354." type="book" year="2008">
Grafe
<emphasis id="2DFD152BAA07FFADFE10FF7845080AEC" box="[511,560,152,176]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. 2008
</bibRefCitation>
, as morphotype 2) and have been collected feeding on calling males of three species of frogs (Table 7).
</paragraph>
<paragraph id="1F36C939AA07FFADFF28FF3F42B40B44" blockId="8.[151,1437,151,1189]" pageId="8" pageNumber="9">
TAXONOMIC DISCUSSION: The seven females noted by
<bibRefCitation id="7B18B4C8AA07FFADFC9DFF3F43250AAA" author="Borkent" box="[882,1053,222,246]" pageId="8" pageNumber="9" refString="Borkent, A. (2008) The Frog-Biting Midges of the World (Corethrellidae: Diptera). Zootaxa, 1804,1 - 456." type="book" year="2008">Borkent (2008)</bibRefCitation>
in the taxonomic discussion of
<taxonomicName id="D889B2BAAA07FFADFA6DFF3E47DD0B45" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nippon">
<emphasis id="2DFD152BAA07FFADFA6DFF3E47DD0B45" italics="true" pageId="8" pageNumber="9">C. nippon</emphasis>
</taxonomicName>
, collected from a tributary of Sungai Belalong, Temburong district,
<collectingCountry id="679E89A9AA07FFADFC0FFEE143170B44" box="[992,1071,256,280]" name="Brunei" pageId="8" pageNumber="9">Brunei</collectingCountry>
, are
<typeStatus id="C032779BAA07FFADFB8BFEE043F70B45" box="[1124,1231,257,281]" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="D889B2BAAA07FFADFB1BFEE042750B45" box="[1268,1357,257,281]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lutea">
<emphasis id="2DFD152BAA07FFADFB1BFEE042750B45" box="[1268,1357,257,281]" italics="true" pageId="8" pageNumber="9">C. lutea</emphasis>
</taxonomicName>
here.
</paragraph>
<paragraph id="1F36C939AA07FFADFF28FEC244280BF8" blockId="8.[151,1437,151,1189]" pageId="8" pageNumber="9">
Six females collected from the headquarters of Gunung Mulu National Park,
<collectingCountry id="679E89A9AA07FFADFBCCFEC243BE0B67" box="[1059,1158,291,315]" name="Malaysia" pageId="8" pageNumber="9">Sarawak</collectingCountry>
,
<collectingCountry id="679E89A9AA07FFADFB7EFEC243CF0B67" box="[1169,1271,291,315]" name="Malaysia" pageId="8" pageNumber="9">Malaysia</collectingCountry>
(
<geoCoordinate id="7ABDAFFEAA07FFADFAE8FEC2424F0B67" box="[1287,1399,291,315]" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="113.916664">113° 55'E</geoCoordinate>
,
<geoCoordinate id="7ABDAFFEAA07FFADFA6CFEC247F50B02" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="4.383333">4° 23'N</geoCoordinate>
) on
<date id="6B37EFF9AA07FFADFEFEFEA746AD0B02" box="[273,405,326,350]" pageId="8" pageNumber="9" value="2009-11-14">14-XI-2009</date>
with a frog-call trap differed from other
<taxonomicName id="D889B2BAAA07FFADFC4AFEA6433E0B02" box="[933,1030,326,350]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lutea">
<emphasis id="2DFD152BAA07FFADFC4AFEA6433E0B02" box="[933,1030,326,350]" italics="true" pageId="8" pageNumber="9">C. lutea</emphasis>
</taxonomicName>
in having only a single sensilla coeloconicum on each of flagellomeres 10–12 and relatively shorter flagellomeres 2–3 (only one slide-mounted). For the present we consider these as variants of
<taxonomicName id="D889B2BAAA07FFADFD5FFE6C44310BF8" box="[688,777,396,420]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lutea">
<emphasis id="2DFD152BAA07FFADFD5FFE6C44310BF8" box="[688,777,396,420]" italics="true" pageId="8" pageNumber="9">C. lutea</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="1F36C939AA07FFADFF28FE4E45BB080F" blockId="8.[151,1437,151,1189]" pageId="8" pageNumber="9">
The females of
<taxonomicName id="D889B2BAAA07FFADFE99FE5146F70B9B" box="[374,463,431,455]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lutea">
<emphasis id="2DFD152BAA07FFADFE99FE5146F70B9B" box="[374,463,431,455]" italics="true" pageId="8" pageNumber="9">C. lutea</emphasis>
</taxonomicName>
are very similar to those of
<taxonomicName id="D889B2BAAA07FFADFCE3FE5144430B9B" box="[780,891,432,455]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nippon">
<emphasis id="2DFD152BAA07FFADFCE3FE5144430B9B" box="[780,891,432,455]" italics="true" pageId="8" pageNumber="9">C. nippon</emphasis>
</taxonomicName>
but the latter have relatively thicker midfemora. Although the male of
<taxonomicName id="D889B2BAAA07FFADFE47FE32453F0BB6" box="[424,519,466,490]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lutea">
<emphasis id="2DFD152BAA07FFADFE47FE32453F0BB6" box="[424,519,466,490]" italics="true" pageId="8" pageNumber="9">C. lutea</emphasis>
</taxonomicName>
is unknown, and therefore we do not know if this species shares the only synapomorphy for the group (unequal male midclaws), we consider it a member of the
<emphasis id="2DFD152BAA07FFADFB90FE14427A0851" box="[1151,1346,501,525]" italics="true" pageId="8" pageNumber="9">drakensbergensis</emphasis>
species group (
<figureCitation id="87B2D5BCAA07FFADFF07FDF94604086C" box="[232,316,536,560]" captionStart="FIGURE 19" captionStartId="41.[151,250,1568,1591]" captionTargetBox="[189,1400,240,1507]" captionTargetId="figure@41.[165,1422,193,1547]" captionTargetPageId="41" captionText="FIGURE 19. Phylogenetic relationships of Corethrella from Borneo to other species and showing their distributions. Numbers refer to character states given in Borkent (2008). Evidence for cladistic relationships given in Borkent (2008)." httpUri="https://zenodo.org/record/210883/files/figure.png" pageId="8" pageNumber="9">Fig. 19</figureCitation>
). It has synapomorphy 25 but lacks 26 and 27 (hence, an unresolved lineage along with
<taxonomicName id="D889B2BAAA07FFADFAC3FDF842A4086C" box="[1324,1436,537,560]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nippon">
<emphasis id="2DFD152BAA07FFADFAC3FDF842A4086C" box="[1324,1436,537,560]" italics="true" pageId="8" pageNumber="9">C. nippon</emphasis>
</taxonomicName>
and
<taxonomicName id="D889B2BAAA07FFADFF29FDDD4546080F" authority="Freeman (Borkent, 2008)" authorityName="Freeman (Borkent" authorityYear="2008" box="[198,638,571,595]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="harrisoni">
<emphasis id="2DFD152BAA07FFADFF29FDDD4677080F" box="[198,335,571,595]" italics="true" pageId="8" pageNumber="9">C. harrisoni</emphasis>
Freeman
<bibRefCitation id="7B18B4C8AA07FFADFE50FDDA4546080F" author="Borkent" box="[447,638,571,595]" pageId="8" pageNumber="9" refString="Borkent, A. (2008) The Frog-Biting Midges of the World (Corethrellidae: Diptera). Zootaxa, 1804,1 - 456." type="book" year="2008">(Borkent, 2008))</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="1F36C939AA07FFADFF28FDBF421A0E03" blockId="8.[151,1437,151,1189]" pageId="8" pageNumber="9">
<typeStatus id="C032779BAA07FFADFF28FDBF4619082A" box="[199,289,606,630]" pageId="8" pageNumber="9">TYPES</typeStatus>
:
<typeStatus id="C032779BAA07FFADFEC0FDBF46A2082A" box="[303,410,606,630]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
, female adult on microscope slide, labeled &quot;
<typeStatus id="C032779BAA07FFADFC70FDBF430E082A" box="[927,1078,606,630]" pageId="8" pageNumber="9" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="D889B2BAAA07FFADFBAEFDBF47E408C5" authority="Borkent and Grafe" authorityName="Borkent and Grafe" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lutea">
<emphasis id="2DFD152BAA07FFADFBAEFDBF423B082A" box="[1089,1283,606,630]" italics="true" pageId="8" pageNumber="9">Corethrella lutea</emphasis>
Borkent and Grafe
</taxonomicName>
&quot;, &quot;
<quantity id="D87164DCAA07FFADFF14FD60467C08C5" box="[251,324,641,665]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="9" unit="km" value="12.0">12 km</quantity>
S. Liang,
<collectingCountry id="679E89A9AA07FFADFE54FD60453208C5" box="[443,522,641,665]" name="Brunei" pageId="8" pageNumber="9">Brunei</collectingCountry>
,
<quantity id="D87164DCAA07FFADFDF9FD60456A08C5" box="[534,594,641,665]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="9" unit="m" value="30.0">30 m</quantity>
,
<date id="6B37EFF9AA07FFADFDB0FD6045E308C5" box="[607,731,641,665]" pageId="8" pageNumber="9" value="2007-05-10">10-V-2007</date>
; on
<taxonomicName id="D889B2BAAA07FFADFCE4FD6044C308C5" box="[779,1019,641,665]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="glandulosa">
<emphasis id="2DFD152BAA07FFADFCE4FD6044C308C5" box="[779,1019,641,665]" italics="true" pageId="8" pageNumber="9">Hylarana glandulosa</emphasis>
</taxonomicName>
; U. Grafe; 18-2/3&quot;, &quot;18-2/3
<date id="6B37EFF9AA07FFADFAD1FD6042AC08C5" box="[1342,1428,641,665]" pageId="8" pageNumber="9" value="07-11-26">261107</date>
&quot; (CNCI).
<typeStatus id="C032779BAA07FFADFEEBFD44464908E1" box="[260,369,677,701]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
on microscope slides: 1 Ƥ labeled as for
<typeStatus id="C032779BAA07FFADFCA5FD45449408E0" box="[842,940,676,700]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
but 18-2/2,
<date id="6B37EFF9AA07FFADFBD7FD4543B408E0" box="[1080,1164,676,700]" pageId="8" pageNumber="9" value="07-10-23">231007</date>
(CNCI); 3 Ƥ from
<typeStatus id="C032779BAA07FFADFA83FD4442A408E1" box="[1388,1436,677,701]" pageId="8" pageNumber="9">type</typeStatus>
locality but
<date id="6B37EFF9AA07FFADFEF2FD26469A0883" box="[285,418,711,735]" pageId="8" pageNumber="9" value="2007-04-21">21-IV-2007</date>
,
<date id="6B37EFF9AA07FFADFE42FD26450A0883" box="[429,562,711,735]" pageId="8" pageNumber="9" value="2007-04-30">30-IV-2007</date>
,
<date id="6B37EFF9AA07FFADFDD2FD26458D0883" box="[573,693,711,735]" pageId="8" pageNumber="9" value="2007-05-15">15-V-2007</date>
(CNCI); 1 Ƥ,
<quantity id="D87164DCAA07FFADFCB5FD2644990883" box="[858,929,711,735]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="9" unit="km" value="15.0">15 km</quantity>
S. Liang,
<collectingCountry id="679E89A9AA07FFADFBF9FD26435D0883" box="[1046,1125,711,735]" name="Brunei" pageId="8" pageNumber="9">Brunei</collectingCountry>
,
<quantity id="D87164DCAA07FFADFB9FFD2643940883" box="[1136,1196,711,735]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="9" unit="m" value="40.0">40 m</quantity>
,
<date id="6B37EFF9AA07FFADFB57FD26421A0883" box="[1208,1314,711,735]" pageId="8" pageNumber="9" value="2007-05-09">9-V-2007</date>
(CNCI); 1 Ƥ,
<quantity id="D87164DCAA07FFADFF52FD084630095D" box="[189,264,745,769]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="9" unit="km" value="17.0">17 km</quantity>
N. Labi,
<collectingCountry id="679E89A9AA07FFADFE91FD0846F5095D" box="[382,461,745,769]" name="Brunei" pageId="8" pageNumber="9">Brunei</collectingCountry>
,
<quantity id="D87164DCAA07FFADFE33FD084522095D" box="[476,538,745,769]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="9" unit="m" value="40.0">40 m</quantity>
,
<date id="6B37EFF9AA07FFADFDC5FD0845A5095D" box="[554,669,745,769]" pageId="8" pageNumber="9" value="2009-04-06">6-IV-2009</date>
(UBD); 1 Ƥ, from previous locality but
<date id="6B37EFF9AA07FFADFB6EFD0843CD095D" box="[1153,1269,745,769]" pageId="8" pageNumber="9" value="2009-04-08">8-IV-2009</date>
(CNCI); 7 Ƥ, tributary of Sungai Belalong, Temburong district,
<collectingCountry id="679E89A9AA07FFADFD25FCED44210978" box="[714,793,780,804]" name="Brunei" pageId="8" pageNumber="9">Brunei</collectingCountry>
,
<quantity id="D87164DCAA07FFADFCCAFCED44560978" box="[805,878,780,804]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="9" unit="m" value="110.0">110 m</quantity>
,
<geoCoordinate id="7ABDAFFEAA07FFADFC95FCED44D20978" box="[890,1002,780,804]" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="115.15">115° 09'E</geoCoordinate>
,
<geoCoordinate id="7ABDAFFEAA07FFADFC19FCED43760978" box="[1014,1102,780,804]" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="4.55">4° 33'N</geoCoordinate>
,
<quantity id="D87164DCAA07FFADFBB5FCED43AE0978" box="[1114,1174,780,804]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="9" unit="m" value="50.0">50 m</quantity>
,
<date id="6B37EFF9AA07FFADFB4DFCED42130978" box="[1186,1323,780,804]" pageId="8" pageNumber="9" value="2006-08-07">7-VIII-2006</date>
(6 CNCI; 1 UBD); 1 Ƥ, Gunung Mulu NP,
<collectingCountry id="679E89A9AA07FFADFDE4FCCE4556091B" box="[523,622,815,839]" name="Malaysia" pageId="8" pageNumber="9">Sarawak</collectingCountry>
,
<collectingCountry id="679E89A9AA07FFADFD96FCCE45DA091B" box="[633,738,815,839]" name="Malaysia" pageId="8" pageNumber="9">Malaysia</collectingCountry>
,
<quantity id="D87164DCAA07FFADFD01FCCE4411091B" box="[750,809,815,839]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="9" unit="m" value="30.0">30 m</quantity>
,
<date id="6B37EFF9AA07FFADFCDAFCCE4481091B" box="[821,953,815,839]" pageId="8" pageNumber="9" value="2008-11-14">14-XI-2008</date>
(CNCI); 5 Ƥ, Matang,
<collectingCountry id="679E89A9AA07FFADFB2FFCCE421B091B" box="[1216,1315,815,839]" name="Malaysia" pageId="8" pageNumber="9">Sarawak</collectingCountry>
,
<collectingCountry id="679E89A9AA07FFADFAC1FCCE42AF091B" box="[1326,1431,815,839]" name="Malaysia" pageId="8" pageNumber="9">Malaysia</collectingCountry>
,
<quantity id="D87164DCAA07FFADFF78FCB347E70936" box="[151,223,850,874]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="9" unit="m" value="200.0">200 m</quantity>
,
<date id="6B37EFF9AA07FFADFF05FCB346590936" box="[234,353,850,874]" pageId="8" pageNumber="9" value="2008-09-04">4-IX-2008</date>
(CNCI).
<typeStatus id="C032779BAA07FFADFE25FCB2450F0937" box="[458,567,851,875]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
on pins: 1 Ƥ, Kuala Belalong Field Studies Centre,
<collectingCountry id="679E89A9AA07FFADFB97FCB343FF0936" box="[1144,1223,850,874]" name="Brunei" pageId="8" pageNumber="9">Brunei</collectingCountry>
,
<quantity id="D87164DCAA07FFADFB3DFCB342340936" box="[1234,1292,850,874]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="9" unit="m" value="50.0">50 m</quantity>
,
<date id="6B37EFF9AA07FFADFAF7FCB342A40936" box="[1304,1436,850,874]" pageId="8" pageNumber="9" value="2010-11-17">17-XI-2010</date>
(CNCI); 1 Ƥ, Belalong Münd (estuary), Ulu Temburong NP,
<collectingCountry id="679E89A9AA07FFADFCAFFC9444B709D1" box="[832,911,885,909]" name="Brunei" pageId="8" pageNumber="9">Brunei</collectingCountry>
,
<date id="6B37EFF9AA07FFADFC74FC94432209D1" box="[923,1050,885,909]" pageId="8" pageNumber="9" value="2008-07-04">4-VII-2008</date>
(CNCI); 1 Ƥ, lower Sungai Apan, Ulu Temburong NP,
<collectingCountry id="679E89A9AA07FFADFE92FC7946F409EC" box="[381,460,920,944]" name="Brunei" pageId="8" pageNumber="9">Brunei</collectingCountry>
,
<quantity id="D87164DCAA07FFADFE38FC79452609EC" box="[471,542,920,944]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="9" unit="m" value="110.0">110 m</quantity>
,
<date id="6B37EFF9AA07FFADFDC6FC79459009EC" box="[553,680,920,944]" pageId="8" pageNumber="9" value="2008-07-08">8-VII-2008</date>
(CNCI); 1 Ƥ, from previous locality but
<date id="6B37EFF9AA07FFADFB9CFC79423909EC" box="[1139,1281,920,944]" pageId="8" pageNumber="9" value="2008-07-13">13-VII-2008</date>
(CNCI); 8 Ƥ,
<quantity id="D87164DCAA07FFADFF78FC5A47CA098F" box="[151,242,955,979]" metricMagnitude="4" metricUnit="m" metricValue="1.25" pageId="8" pageNumber="9" unit="km" value="12.5">12.5 km</quantity>
S. Liang,
<collectingCountry id="679E89A9AA07FFADFE8BFC5A468B098F" box="[356,435,955,979]" name="Brunei" pageId="8" pageNumber="9">Brunei</collectingCountry>
,
<quantity id="D87164DCAA07FFADFE51FC5A46C1098F" box="[446,505,955,979]" metricMagnitude="1" metricUnit="m" metricValue="3.3" pageId="8" pageNumber="9" unit="m" value="33.0">33 m</quantity>
,
<date id="6B37EFF9AA07FFADFDEBFC5A454B098F" box="[516,627,955,979]" pageId="8" pageNumber="9" value="2011-01-28">28-I-2011</date>
(6, CNCI; 2 UBD); 12 Ƥ, from previous locality but
<date id="6B37EFF9AA07FFADFB2BFC5A4200098F" box="[1220,1336,955,979]" pageId="8" pageNumber="9" value="2009-04-02">2-IV-2009</date>
(CNCI); 6 Ƥ,
<quantity id="D87164DCAA07FFADFF3CFC3F462409AA" box="[211,284,990,1014]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="9" unit="km" value="20.0">20 km</quantity>
N. Labi, Luagan Lalak Forestry Rec. Site,
<collectingCountry id="679E89A9AA07FFADFCE0FC3F446609AA" box="[783,862,990,1014]" name="Brunei" pageId="8" pageNumber="9">Brunei</collectingCountry>
,
<date id="6B37EFF9AA07FFADFC84FC3F44D909AA" box="[875,993,990,1014]" pageId="8" pageNumber="9" value="2010-11-06">6-XI-2010</date>
(CNCI); 2 Ƥ, Sungai Ingei,
<quantity id="D87164DCAA07FFADFAC3FC3F424509AA" box="[1324,1405,990,1014]" metricMagnitude="3" metricUnit="m" metricValue="8.2" pageId="8" pageNumber="9" unit="km" value="8.2">8.2 km</quantity>
S. Melilas,
<collectingCountry id="679E89A9AA07FFADFF12FBE046740E45" box="[253,332,1025,1049]" name="Brunei" pageId="8" pageNumber="9">Brunei</collectingCountry>
,
<quantity id="D87164DCAA07FFADFEB2FBE046A40E45" box="[349,412,1025,1049]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="9" unit="m" value="50.0">50 m</quantity>
,
<date id="6B37EFF9AA07FFADFE42FBE045140E45" box="[429,556,1025,1049]" pageId="8" pageNumber="9" value="2010-07-09">9-VII-2010</date>
(CNCI); 11 Ƥ, from previous locality but
<date id="6B37EFF9AA07FFADFBCBFBE043970E45" box="[1060,1199,1025,1049]" pageId="8" pageNumber="9" value="2010-07-12">12-VII-2010</date>
(CNCI); 4 Ƥ, from previous locality but
<date id="6B37EFF9AA07FFADFE67FBC546C80E60" box="[392,496,1060,1084]" pageId="8" pageNumber="9" value="2010-07">VII-2010</date>
(CNCI); 2 Ƥ, from previous locality but
<date id="6B37EFF9AA07FFADFC28FBC5436D0E60" box="[967,1109,1060,1084]" pageId="8" pageNumber="9" value="2010-07-10">10-VII-2010</date>
(CNCI); 5 Ƥ, Gunung Mulu NP,
<collectingCountry id="679E89A9AA07FFADFF2AFBA646100E03" box="[197,296,1095,1119]" name="Malaysia" pageId="8" pageNumber="9">Sarawak</collectingCountry>
,
<collectingCountry id="679E89A9AA07FFADFEDBFBA646A60E03" box="[308,414,1095,1119]" name="Malaysia" pageId="8" pageNumber="9">Malaysia</collectingCountry>
,
<quantity id="D87164DCAA07FFADFE47FBA646DA0E03" box="[424,482,1095,1119]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="9" unit="m" value="30.0">30 m</quantity>
,
<date id="6B37EFF9AA07FFADFE08FBA645540E03" box="[487,620,1095,1119]" pageId="8" pageNumber="9" value="2008-11-14">14-XI-2008</date>
(CNCI); 3 Ƥ, from previous locality but
<date id="6B37EFF9AA07FFADFBD5FBA643870E03" box="[1082,1215,1095,1119]" pageId="8" pageNumber="9" value="2009-11-26">26-XI-2009</date>
(CNCI).
</paragraph>
<paragraph id="1F36C939AA07FFADFF28FB8B46DB0EF9" blockId="8.[151,1437,151,1189]" pageId="8" pageNumber="9">
DERIVATION OF SPECIFIC EPITHET: The name
<taxonomicName id="D889B2BAAA07FFADFCF1FB8B446E0EDE" box="[798,854,1130,1154]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lutea">
<emphasis id="2DFD152BAA07FFADFCF1FB8B446E0EDE" box="[798,854,1130,1154]" italics="true" pageId="8" pageNumber="9">lutea</emphasis>
</taxonomicName>
(yellow) refers to the overall yellowish appearance of the females of this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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