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<document id="9E01F36C36D2CECCBDC7B0C332C9ACFF" ID="10.11646/zootaxa.4246.1.1SLASH10606" ID-CLB-Dataset="36212" ID-DOI="10.5281/zenodo.437611" ID-GBIF-Dataset="3ff74501-6241-43b2-a52f-0e7580bb2f45" ID-ISSN="1175-5326" ID-Zenodo-Dep="437611" ID-ZooBank="D8594E96-D561-4A37-9073-B138EC7E95A7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1490260213340" checkinUser="plazi" docAuthor="Zaragoza, Juan A." docDate="2017" docId="03889E18FFE4554DFF5C40D6FDF3F817" docLanguage="en" docName="zootaxa.4246.1.1SLASH10606.pdf" docOrigin="Zootaxa 4246 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Ephippiochthonius mariolae Carabajal Marquez, Garcia Carrillo &amp; Rodriguez Fernandez 2001, n. comb." docType="treatment" docVersion="9" lastPageNumber="92" masterDocId="FFB1E660FFBF5516FFCB4161FF86FFD2" masterDocTitle="Revision of the Ephippiochthonius complex in the Iberian Peninsula, Balearic Islands and Macaronesia, with proposed changes to the status of the Chthonius subgenera (Pseudoscorpiones, Chthoniidae)" masterLastPageNumber="221" masterPageNumber="1" pageNumber="92" updateTime="1698438337209" updateUser="ExternalLinkService">
<mods:mods id="5F40AC2BD099D245ABAA50CC2121FA8D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="084D9C63F4B8154BDFB6D49ED76A0FFE">
<mods:title id="3B589F3AB136CAB5820411783D4AE4BC">Revision of the Ephippiochthonius complex in the Iberian Peninsula, Balearic Islands and Macaronesia, with proposed changes to the status of the Chthonius subgenera (Pseudoscorpiones, Chthoniidae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="AC20ACD228D103D72B8D06417BBC2B84">Zaragoza, Juan A.</mods:namePart>
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<mods:part id="55E49B14C59952825D3374C4D914C669">
<mods:date id="D6C188B852C2F95A2912C057AFE66C13">2017</mods:date>
<mods:detail id="5E7AA96A7028EC97E7E92CF28E8AF54B" type="volume">
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<mods:classification id="32B21B38877A1771C559575706736A9B">journal article</mods:classification>
<mods:identifier id="D41FBFCFC97340FC3DB11F5933E67507" type="CLB-Dataset">36212</mods:identifier>
<mods:identifier id="091D53AD0D8E18623248D58D7E04A0EC" type="DOI">10.5281/zenodo.437611</mods:identifier>
<mods:identifier id="59861CF1C725F5982FC189BE1B8FC5AF" type="GBIF-Dataset">3ff74501-6241-43b2-a52f-0e7580bb2f45</mods:identifier>
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<mods:identifier id="9DC44D39CB61B37E1572DAC38AC19CEA" type="Zenodo-Dep">437611</mods:identifier>
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<treatment id="03889E18FFE4554DFF5C40D6FDF3F817" ID-DOI="http://doi.org/10.5281/zenodo.5687093" ID-GBIF-Taxon="127852053" ID-Zenodo-Dep="5687093" LSID="urn:lsid:plazi:treatment:03889E18FFE4554DFF5C40D6FDF3F817" httpUri="http://treatment.plazi.org/id/03889E18FFE4554DFF5C40D6FDF3F817" lastPageNumber="92" pageId="91" pageNumber="92">
<subSubSection id="C33B7C85FFE4554DFF5C40D6FF65FE26" pageId="91" pageNumber="92" type="nomenclature">
<paragraph id="8B9E2F0EFFE4554DFF5C40D6FF65FE26" blockId="91.[151,1402,439,500]" pageId="91" pageNumber="92">
<heading id="D0D69862FFE4554DFF5C40D6FF65FE26" bold="true" fontSize="11" level="1" pageId="91" pageNumber="92" reason="1">
<emphasis id="B955F31CFFE4554DFF5C40D6FF65FE26" bold="true" pageId="91" pageNumber="92">
<taxonomicName id="4C21548DFFE4554DFF5C40D6FAD3FE00" ID-CoL="6FPZ2" authority="Carabajal Marquez, Garcia Carrillo &amp; Rodriguez Fernandez, 2001" authorityName="Carabajal Marquez, Garcia Carrillo &amp; Rodriguez Fernandez" authorityYear="2001" box="[151,1365,439,466]" class="Arachnida" family="Chthoniidae" genus="Ephippiochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="mariolae" status="comb. nov.">
<emphasis id="B955F31CFFE4554DFF5C40D6FE77FE03" bold="true" box="[151,497,439,465]" italics="true" pageId="91" pageNumber="92">Ephippiochthonius mariolae</emphasis>
(Carabajal Márquez, García Carrillo &amp; Rodríguez Fernández, 2001)
</taxonomicName>
,
<taxonomicNameLabel id="A2664E67FFE4554DFAA940D9FF65FE26" pageId="91" pageNumber="92">n. comb.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FFE4554DFF5C4340FCA9FDA6" pageId="91" pageNumber="92" type="reference_group">
<paragraph id="8B9E2F0EFFE4554DFF5C4340FEC6FD84" blockId="91.[151,1436,545,628]" pageId="91" pageNumber="92">
<treatmentCitationGroup id="AB310820FFE4554DFF5C4340FEC6FD84" pageId="91" pageNumber="92">
<taxonomicName id="4C21548DFFE4554DFF5C4340FAB3FDE5" ID-CoL="V3DV" authority="Carabajal Marquez, Garcia Carrillo &amp; Rodriguez Fernandez 2001b: 170" authorityName="Carabajal Marquez, Garcia Carrillo &amp; Rodriguez Fernandez" authorityPageNumber="170" authorityYear="2001" box="[151,1333,545,567]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="mariolae" subGenus="Ephippiochthonius">
<emphasis id="B955F31CFFE4554DFF5C4340FF79FDE5" box="[151,255,545,567]" italics="true" pageId="91" pageNumber="92">Chthonius</emphasis>
(
<emphasis id="B955F31CFFE4554DFEDB4340FE49FDE5" box="[272,463,545,567]" italics="true" pageId="91" pageNumber="92">Ephippiochthonius</emphasis>
)
<emphasis id="B955F31CFFE4554DFE2A4340FDBBFDE5" box="[481,573,545,567]" italics="true" pageId="91" pageNumber="92">mariolae</emphasis>
<treatmentCitation id="0A80091FFFE4554DFD8E4340FAB3FDE5" author="Carabajal" box="[581,1333,545,567]" page="170" pageId="91" pageNumber="92" year="2001">
<bibRefCitation id="EFB052FFFFE4554DFD8E4340FA87FDE5" author="Carabajal" box="[581,1281,545,567]" pageId="91" pageNumber="92" refString="Carabajal Marquez, E., Garcia Carrillo, J. &amp; Rodriguez Fernandez, F. (2001 b) Descripcion de cuatro nuevos pseudoscorpiones cavernicolas de Andalucia, Espana (Arachnida, Pseudoscorpionida, Chthoniidae). Zoologica Baetica, 12, 169 - 184." type="journal article" year="2001" yearSuffix="b">Carabajal Márquez, García Carrillo &amp; Rodríguez Fernández 2001b</bibRefCitation>
: 170
</treatmentCitation>
</taxonomicName>
–173, figs 1‒5, table I.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9E2F0EFFE4554DFF5C433FFCA9FDA6" blockId="91.[151,1436,545,628]" box="[151,815,606,628]" pageId="91" pageNumber="92">
<treatmentCitationGroup id="AB310820FFE4554DFF5C433FFCA9FDA6" box="[151,815,606,628]" pageId="91" pageNumber="92">
<taxonomicName id="4C21548DFFE4554DFF5C433FFC84FDA6" ID-CoL="V3DV" authority="Zaragoza 2007: 17" authorityName="Zaragoza" authorityPageNumber="17" authorityYear="2007" box="[151,770,606,628]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="mariolae" subGenus="Ephippiochthonius">
<emphasis id="B955F31CFFE4554DFF5C433FFF79FDA6" box="[151,255,606,628]" italics="true" pageId="91" pageNumber="92">Chthonius</emphasis>
(
<emphasis id="B955F31CFFE4554DFEC6433FFE4AFDA6" box="[269,460,606,628]" italics="true" pageId="91" pageNumber="92">Ephippiochthonius</emphasis>
)
<emphasis id="B955F31CFFE4554DFE10433FFDB1FDA6" box="[475,567,606,628]" italics="true" pageId="91" pageNumber="92">mariolae</emphasis>
:
<treatmentCitation id="0A80091FFFE4554DFD8F433EFC84FDA6" author="Zaragoza" box="[580,770,606,628]" page="17" pageId="91" pageNumber="92" year="2007">
<bibRefCitation id="EFB052FFFFE4554DFD8F433EFD59FDA6" author="Zaragoza" box="[580,735,606,628]" pageId="91" pageNumber="92" refString="Zaragoza, J. A. (2007) Catalogo de los pseudoescorpiones de la Peninsula Iberica e Islas Baleares (Arachnida: Pseudoscorpiones). Revista Iberica Aracnologia, 13, 3 - 91." type="journal article" year="2007">Zaragoza 2007</bibRefCitation>
: 17
</treatmentCitation>
</taxonomicName>
–18.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FFE4554DFF5C43FDFE46FD0A" pageId="91" pageNumber="92" type="materials_examined">
<paragraph id="8B9E2F0EFFE4554DFF5C43FDFE46FD0A" blockId="91.[151,1437,668,1989]" pageId="91" pageNumber="92">
<materialsCitation id="3B492553FFE4554DFF5C43FDFD8FFD67" ID-GBIF-Occurrence="1449740845" box="[151,521,668,693]" country="Spain" location="Andalusia" pageId="91" pageNumber="92" specimenCount="1" stateProvince="Andalusia" typeStatus="holotype">
<emphasis id="B955F31CFFE4554DFF5C43FDFEBFFD67" bold="true" box="[151,313,668,693]" pageId="91" pageNumber="92">
<typeStatus id="549A91ACFFE4554DFF5C43FDFF54FD67" box="[151,210,668,693]" pageId="91" pageNumber="92">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3366F9EFFE4554DFE8843FDFE00FD67" box="[323,390,668,693]" name="Spain" pageId="91" pageNumber="92">Spain</collectingCountry>
,
<collectingRegion id="49E5E1ECFFE4554DFE5E43FDFD8FFD67" box="[405,521,668,693]" country="Spain" name="Andalucia" pageId="91" pageNumber="92">Andalusia</collectingRegion>
</materialsCitation>
,
<materialsCitation id="3B492553FFE4554DFDD343FDFE3DFD0A" ID-GBIF-Occurrence="1449740852" country="Grenada" county="Salobrena" elevation="140" latitude="36.78361" location="Cueva" longLatPrecision="19" longitude="-3.5491667" municipality="Lobres" pageId="91" pageNumber="92" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F3366F9EFFE4554DFDD343FDFDF1FD67" box="[536,631,668,693]" name="Grenada" pageId="91" pageNumber="92">Granada</collectingCountry>
province,
<collectingCounty id="62FF5782FFE4554DFD3E43FDFCEEFD67" box="[757,872,668,693]" pageId="91" pageNumber="92">Salobreña</collectingCounty>
,
<collectingMunicipality id="6BFAB574FFE4554DFCBC43FDFC4EFD67" box="[887,968,668,693]" pageId="91" pageNumber="92">Lobres</collectingMunicipality>
,
<location id="8EFE79D5FFE4554DFC1D43FDFB9BFD66" LSID="urn:lsid:plazi:treatment:03889E18FFE4554DFF5C40D6FDF3F817:8EFE79D5FFE4554DFC1D43FDFB9BFD66" box="[982,1053,668,692]" country="Grenada" county="Salobrena" latitude="36.78361" longLatPrecision="19" longitude="-3.5491667" municipality="Lobres" name="Cueva" pageId="91" pageNumber="92">Cueva</location>
del
<location id="8EFE79D5FFE4554DFB9E43FCFB79FD66" LSID="urn:lsid:plazi:treatment:03889E18FFE4554DFF5C40D6FDF3F817:8EFE79D5FFE4554DFB9E43FCFB79FD66" box="[1109,1279,669,692]" country="Grenada" county="Salobrena" latitude="36.78361" longLatPrecision="19" longitude="-3.5491667" municipality="Lobres" name="Barranco Inate" pageId="91" pageNumber="92">Barranco Iñate</location>
(
<geoCoordinate id="EE1549C9FFE4554DFADA43FDFA1EFD67" box="[1297,1432,668,693]" direction="north" orientation="latitude" pageId="91" pageNumber="92" precision="15" value="36.78361">36°47′01″N</geoCoordinate>
,
<geoCoordinate id="EE1549C9FFE4554DFF5C43DEFEA4FD0A" box="[151,290,703,728]" direction="west" orientation="longitude" pageId="91" pageNumber="92" precision="15" value="-3.5491667">03°32′57″W</geoCoordinate>
;
<elevation id="000CC83DFFE4554DFEE643A1FE37FD0A" box="[301,433,703,728]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="91" pageNumber="92" unit="m" value="140.0">
<quantity id="4CD982EBFFE4554DFEE643A1FEF2FD05" box="[301,372,704,728]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="91" pageNumber="92" unit="m" value="140.0">140 m</quantity>
a.s.l.
</elevation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FFE4554DFF0C4385FDA9FD2F" box="[199,559,740,765]" pageId="91" pageNumber="92" type="distribution">
<paragraph id="8B9E2F0EFFE4554DFF0C4385FDA9FD2F" blockId="91.[151,1437,668,1989]" box="[199,559,740,765]" pageId="91" pageNumber="92">
<emphasis id="B955F31CFFE4554DFF0C4385FEE6FD2F" bold="true" box="[199,352,740,765]" pageId="91" pageNumber="92">Distribution.</emphasis>
<collectingCountry id="F3366F9EFFE4554DFEAC4385FE2DFD2F" box="[359,427,740,765]" name="Spain" pageId="91" pageNumber="92">Spain</collectingCountry>
:
<collectingRegion id="49E5E1ECFFE4554DFE7E4385FDACFD2F" box="[437,554,740,765]" country="Spain" name="Andalucia" pageId="91" pageNumber="92">Andalusia</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FFE4554DFF0C4266FE44FBCF" pageId="91" pageNumber="92" type="diagnosis">
<paragraph id="8B9E2F0EFFE4554DFF0C4266FE44FBCF" blockId="91.[151,1437,668,1989]" pageId="91" pageNumber="92">
<emphasis id="B955F31CFFE4554DFF0C4266FED3FCF2" bold="true" box="[199,341,775,800]" pageId="91" pageNumber="92">Diagnosis (</emphasis>
<specimenCount id="9D27E487FFE4554DFE9E4269FEECFCCD" box="[341,362,776,799]" pageId="91" pageNumber="92" type="female">♀</specimenCount>
<emphasis id="B955F31CFFE4554DFEA14266FEFCFCF2" bold="true" box="[362,378,775,800]" pageId="91" pageNumber="92">).</emphasis>
A moderately large, hypogean
<taxonomicName id="4C21548DFFE4554DFCDB4266FC62FCF2" box="[784,996,775,800]" class="Arachnida" family="Chthoniidae" genus="Ephippiochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B955F31CFFE4554DFCDB4266FC62FCF2" box="[784,996,775,800]" italics="true" pageId="91" pageNumber="92">Ephippiochthonius</emphasis>
</taxonomicName>
species tentatively assigned to the
<taxonomicName id="4C21548DFFE4554DFF5C424DFEAFFC97" box="[151,297,812,837]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="tetrachelatus">
<emphasis id="B955F31CFFE4554DFF5C424DFEAFFC97" box="[151,297,812,837]" italics="true" pageId="91" pageNumber="92">tetrachelatus</emphasis>
</taxonomicName>
-group. Movable cheliceral finger without isolated subapical tooth (
<emphasis id="B955F31CFFE4554DFBE8424DFBBEFC97" box="[1059,1080,812,837]" italics="true" pageId="91" pageNumber="92">di</emphasis>
) and spinneret small in female, unknown for male; no eyes or eye-spots, 1 preocular microseta on each side, posterior margin of carapace with 2 macrosetae; coxa II with 7–9 coxal spines; chelal hand weakly depressed at level of
<emphasis id="B955F31CFFE4554DFB8B4215FBD0FC5F" box="[1088,1110,884,909]" italics="true" pageId="91" pageNumber="92">ib</emphasis>
/
<emphasis id="B955F31CFFE4554DFB954215FBF9FC5F" box="[1118,1151,884,909]" italics="true" pageId="91" pageNumber="92">isb</emphasis>
, with long and low hump distad of
<emphasis id="B955F31CFFE4554DFECC42F6FE9BFC62" box="[263,285,919,944]" italics="true" pageId="91" pageNumber="92">ib</emphasis>
/
<emphasis id="B955F31CFFE4554DFEEF42F6FEC3FC62" box="[292,325,919,944]" italics="true" pageId="91" pageNumber="92">isb</emphasis>
and an abrupt slope between trichobothria
<emphasis id="B955F31CFFE4554DFC8D42F6FCDAFC62" box="[838,860,919,944]" italics="true" pageId="91" pageNumber="92">ib</emphasis>
/
<emphasis id="B955F31CFFE4554DFCA842F6FC02FC62" box="[867,900,919,944]" italics="true" pageId="91" pageNumber="92">isb</emphasis>
and
<emphasis id="B955F31CFFE4554DFC0942F6FC5BFC62" box="[962,989,919,944]" italics="true" pageId="91" pageNumber="92">eb</emphasis>
; fixed chelal finger with 23–27 teeth; movable chelal finger with 10–11 teeth, without marginal lamina; pedipalpal femur (
<specimenCount id="9D27E487FFE4554DFB9C42DDFBEFFC06" box="[1111,1129,956,980]" pageId="91" pageNumber="92" type="female">♀</specimenCount>
) 7.0–7.3 times longer than broad, length
<quantity id="4CD982EBFFE4554DFEFA4281FE51FC25" box="[305,471,992,1016]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" metricValueMax="10.2" metricValueMin="9.399999999999999" pageId="91" pageNumber="92" unit="mm" value="0.98" valueMax="1.02" valueMin="0.94">0.94–1.02 mm</quantity>
; chela (
<specimenCount id="9D27E487FFE4554DFDE44281FDC7FC2A" box="[559,577,992,1016]" pageId="91" pageNumber="92" type="female">♀</specimenCount>
) 5.8–5.9 times longer than deep, length
<quantity id="4CD982EBFFE4554DFBCC42BEFB2BFC25" box="[1031,1197,991,1016]" metricMagnitude="-3" metricUnit="m" metricValue="1.455" metricValueMax="1.5" metricValueMin="1.41" pageId="91" pageNumber="92" unit="mm" value="1.455" valueMax="1.5" valueMin="1.41">1.41–1.50 mm</quantity>
; ratio movable chelal finger/chelal hand (
<specimenCount id="9D27E487FFE4554DFEB84565FE03FBCE" box="[371,389,1028,1052]" pageId="91" pageNumber="92" type="female">♀</specimenCount>
) 1.6.
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FFE4554DFF0C4546FD00FB92" box="[199,646,1063,1088]" pageId="91" pageNumber="92" type="materials_examined">
<paragraph id="8B9E2F0EFFE4554DFF0C4546FD00FB92" blockId="91.[151,1437,668,1989]" box="[199,646,1063,1088]" pageId="91" pageNumber="92">
<emphasis id="B955F31CFFE4554DFF0C4546FEB0FB92" bold="true" box="[199,310,1063,1088]" pageId="91" pageNumber="92">Material.</emphasis>
None available for this study.
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FFE4554DFF0C452DFEC8F91A" pageId="91" pageNumber="92" type="description">
<paragraph id="8B9E2F0EFFE4554DFF0C452DFA9FFBB7" blockId="91.[151,1437,668,1989]" box="[199,1305,1100,1125]" pageId="91" pageNumber="92">
<emphasis id="B955F31CFFE4554DFF0C452DFED4FBB7" bold="true" box="[199,338,1100,1125]" pageId="91" pageNumber="92">Description</emphasis>
(
<specimenCount id="9D27E487FFE4554DFEA8452DFEF0FBB6" box="[355,374,1100,1124]" pageId="91" pageNumber="92" type="female">♀</specimenCount>
;
<specimenCount id="9D27E487FFE4554DFE4C452CFE1AFBB6" box="[391,412,1101,1124]" pageId="91" pageNumber="92" type="male">♂</specimenCount>
<emphasis id="B955F31CFFE4554DFE68452DFDA2FBB7" bold="true" box="[419,548,1100,1125]" pageId="91" pageNumber="92">unknown).</emphasis>
Large, hypogean species. Integument depigmented, troglomorphic.
</paragraph>
<paragraph id="8B9E2F0EFFE4554DFF0C4511FEA9FB02" blockId="91.[151,1437,668,1989]" pageId="91" pageNumber="92">
Carapace slightly longer than broad and posteriorly constricted; medial part of anterior margin strongly dentate; without eyes or eye-spots. Chaetotaxy: 18 macrosetae and 1 preocular microseta on each side, formula m
<quantity id="4CD982EBFFE4554DFF7A45D6FF5BFB02" box="[177,221,1207,1232]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="91" pageNumber="92" unit="m" value="4.0">4m</quantity>
:6:4:2:2.
</paragraph>
<paragraph id="8B9E2F0EFFE4554DFF0C45BDFB21FACA" blockId="91.[151,1437,668,1989]" pageId="91" pageNumber="92">
Pedipalpal coxa with 3 setae (setae on manducatory process not included);
<date id="FF9F09CEFFE4554DFBD545BCFBC1FB26" box="[1054,1095,1244,1268]" pageId="91" pageNumber="92">II 4</date>
+ 7–9 bipinnate coxal spines,
<date id="FF9F09CEFFE4554DFF5C4461FF4EFACA" box="[151,200,1279,1304]" pageId="91" pageNumber="92">III 5</date>
+ 5–7 bipinnate coxal spines and
<date id="FF9F09CEFFE4554DFD8D4461FD1AFACA" box="[582,668,1279,1304]" pageId="91" pageNumber="92">IV 5–6.</date>
Chaetotaxy of tergites I–IX: 4:4:4:4:6:6:6:6:6.
</paragraph>
<paragraph id="8B9E2F0EFFE4554DFF0C4445FCE0FA57" blockId="91.[151,1437,668,1989]" pageId="91" pageNumber="92">
Cheliceral fixed finger with 7 teeth decreasing in size proximally, 1–2 distal teeth distinctly larger than others. Movable finger without an isolated subapical tooth (
<emphasis id="B955F31CFFE4554DFD2F4426FD7CFAB2" box="[740,762,1351,1376]" italics="true" pageId="91" pageNumber="92">di</emphasis>
), with 5 teeth decreasing in size proximally, the distal tooth larger than others; spinneret small in female, unknown for male.
</paragraph>
<paragraph id="8B9E2F0EFFE4554DFF0C44EEFC17F9C6" blockId="91.[151,1437,668,1989]" pageId="91" pageNumber="92">
Chelal hand weakly depressed at level of
<emphasis id="B955F31CFFE4554DFD6A44EEFD31FA7A" box="[673,695,1423,1448]" italics="true" pageId="91" pageNumber="92">ib</emphasis>
/
<emphasis id="B955F31CFFE4554DFD7544EEFD59FA7A" box="[702,735,1423,1448]" italics="true" pageId="91" pageNumber="92">isb</emphasis>
, with long and low hump distad of
<emphasis id="B955F31CFFE4554DFBBC44EEFB0BFA7A" box="[1143,1165,1423,1448]" italics="true" pageId="91" pageNumber="92">ib</emphasis>
/
<emphasis id="B955F31CFFE4554DFB5E44EEFB30FA7A" box="[1173,1206,1423,1448]" italics="true" pageId="91" pageNumber="92">isb</emphasis>
and an abrupt slope between trichobothria
<emphasis id="B955F31CFFE4554DFE5744D5FE34FA1F" box="[412,434,1460,1485]" italics="true" pageId="91" pageNumber="92">ib</emphasis>
/
<emphasis id="B955F31CFFE4554DFE7244D5FE5CFA1F" box="[441,474,1460,1485]" italics="true" pageId="91" pageNumber="92">isb</emphasis>
and
<emphasis id="B955F31CFFE4554DFDD144D5FDB2FA1F" box="[538,564,1460,1485]" italics="true" pageId="91" pageNumber="92">eb</emphasis>
. Fixed finger with 23–27 teeth Movable finger with 10–11 teeth, without marginal lamina, subdistally teeth are absent up to level of the 6th distal tooth of fixed finger, dental row reaching level of trichobothrium
<emphasis id="B955F31CFFE4554DFE69449DFE3DF9C7" box="[418,443,1532,1557]" italics="true" pageId="91" pageNumber="92">sb</emphasis>
. Trichobothrium
<emphasis id="B955F31CFFE4554DFD4B449CFD1DF9C6" box="[640,667,1533,1556]" italics="true" pageId="91" pageNumber="92">ist</emphasis>
slightly distad of
<emphasis id="B955F31CFFE4554DFCAF449DFC0FF9C7" box="[868,905,1532,1557]" italics="true" pageId="91" pageNumber="92">esb</emphasis>
.
</paragraph>
<paragraph id="8B9E2F0EFFE4554DFF0C4740FEC8F91A" blockId="91.[151,1437,668,1989]" pageId="91" pageNumber="92">
<emphasis id="B955F31CFFE4554DFF0C4740FE63F9EA" box="[199,485,1567,1592]" italics="true" pageId="91" pageNumber="92">Measurements and ratios</emphasis>
. Female
<typeStatus id="549A91ACFFE4554DFD84477EFD32F9EA" box="[591,692,1567,1592]" pageId="91" pageNumber="92" type="holotype">holotype</typeStatus>
, followed, when different, by female
<typeStatus id="549A91ACFFE4554DFB944741FB39F9EA" box="[1119,1215,1568,1592]" pageId="91" pageNumber="92" type="paratype">paratype</typeStatus>
in square brackets: Body 1.86 [2.20]. Carapace 0.57/0.55 (1.0). Pedipalp: femur 1.02/0.15 (7.0)[0.94/0.13 (7.3)], patella 0.41/0.16 (2.5) [0.39/0.17 (2.3)], chela 1.50/0.26 (5.8) [1.41/0.24 (5.9)], hand 0.57 (2.2) [0.54 (2.3)], movable finger 0.93 [0.88]; ratio movable finger/hand 1.6, femur/movable finger 1.1, femur/carapace 1.8 [1.6], chela/carapace 2.6 [2.5], chela/femur 1.5.
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FFE4554DFF0C47B5FDF3F817" pageId="91" pageNumber="92" type="discussion">
<paragraph id="8B9E2F0EFFE4554DFF0C47B5FDF3F817" blockId="91.[151,1437,668,1989]" pageId="91" pageNumber="92">
<emphasis id="B955F31CFFE4554DFF0C47B5FEBDF93F" bold="true" box="[199,315,1748,1773]" pageId="91" pageNumber="92">Remarks.</emphasis>
Within the
<taxonomicName id="4C21548DFFE4554DFE0E47B5FDD1F93F" box="[453,599,1748,1773]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="tetrachelatus">
<emphasis id="B955F31CFFE4554DFE0E47B5FDD1F93F" box="[453,599,1748,1773]" italics="true" pageId="91" pageNumber="92">tetrachelatus</emphasis>
</taxonomicName>
-group,
<taxonomicName id="4C21548DFFE4554DFD7947B4FCBEF93F" box="[690,824,1748,1773]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="mariolae">
<emphasis id="B955F31CFFE4554DFD7947B4FCBEF93F" box="[690,824,1748,1773]" italics="true" pageId="91" pageNumber="92">E. mariolae</emphasis>
</taxonomicName>
shares with
<taxonomicName id="4C21548DFFE4554DFC0047B4FBC2F93F" box="[971,1092,1748,1773]" class="Arachnida" family="Chthoniidae" genus="Ephippiochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis id="B955F31CFFE4554DFC0047B4FBC2F93F" box="[971,1092,1748,1773]" italics="true" pageId="91" pageNumber="92">E. bolivari</emphasis>
</taxonomicName>
the following combination of characteristics: anophthalmic, hypogean, with 1–2 preocular microsetae on each side, only 2 setae in posterior row of carapace, without isolated subapical tooth on movable cheliceral finger and reduced number of teeth (9–11) with dental canals on the movable chelal finger.
<taxonomicName id="4C21548DFFE4554DFDB24620FD7BF88A" box="[633,765,1855,1880]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="mariolae">
<emphasis id="B955F31CFFE4554DFDB24620FD7BF88A" box="[633,765,1855,1880]" italics="true" pageId="91" pageNumber="92">E. mariolae</emphasis>
</taxonomicName>
is easily distinguishable from
<taxonomicName id="4C21548DFFE4554DFB994620FB4FF88A" box="[1106,1225,1855,1880]" class="Arachnida" family="Chthoniidae" genus="Ephippiochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis id="B955F31CFFE4554DFB994620FB4FF88A" box="[1106,1225,1855,1880]" italics="true" pageId="91" pageNumber="92">E. bolivari</emphasis>
</taxonomicName>
by the presence of only 7–9 coxal spines on coxa II when compared with
<quantity id="4CD982EBFFE4554DFCD64605FC05F8AF" box="[797,899,1892,1917]" metricMagnitude="-1" metricUnit="m" metricValue="4.445" metricValueMax="4.826" metricValueMin="4.064" pageId="91" pageNumber="92" unit="in" value="17.5" valueMax="19.0" valueMin="16.0">16–19 in</quantity>
the latter. Although no material of any of the species has been available for this study, it is tentatively assigned to the
<taxonomicName id="4C21548DFFE4554DFC7246E6FBCDF872" box="[953,1099,1927,1952]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="tetrachelatus">
<emphasis id="B955F31CFFE4554DFC7246E6FBCDF872" box="[953,1099,1927,1952]" italics="true" pageId="91" pageNumber="92">tetrachelatus</emphasis>
</taxonomicName>
-group. The above description is based in that of
<bibRefCitation id="EFB052FFFFE4554DFEA846CDFDF6F817" box="[355,624,1964,1989]" pageId="91" pageNumber="92" refString="Carabajal Marquez, E., Garcia Carrillo, J. &amp; Rodriguez Fernandez, F. (2001 b) Descripcion de cuatro nuevos pseudoscorpiones cavernicolas de Andalucia, Espana (Arachnida, Pseudoscorpionida, Chthoniidae). Zoologica Baetica, 12, 169 - 184." type="journal article">
Carabajal
<emphasis id="B955F31CFFE4554DFE1E46CCFD88F817" box="[469,526,1964,1989]" italics="true" pageId="91" pageNumber="92">et al.</emphasis>
(2001b)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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