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<document id="37340EB490F21BFE748B525519BA61C0" ID="10.11646/zootaxa.4169.3.7" ID-DOI="10.11646/zootaxa.4169.3.7" ID-GBIF-Dataset="faeafa7a-a105-4443-bbc3-2bea598d8cd5" ID-ISSN="1175-5326" ID-Zenodo-Dep="258027" ID-ZooBank="8A1FB454-AC73-4D54-A5CB-D12D6CD82AC6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1474440730591" checkinUser="plazi" docAuthor="Tian, Mingyi &amp; Deuve, Thierry" docDate="2016" docId="8706BE21FFB9FF9B1DC8FADEFDBCA2B7" docLanguage="en" docName="zootaxa.4169.3.7.pdf" docOrigin="Zootaxa 4169 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hexachaetus lateralis subsp. maindroni Tian et Deuve 2006, n. stat." docType="treatment" docVersion="7" lastPageNumber="543" masterDocId="7B3FC659FFBAFF981D5FFF88FFF0A546" masterDocTitle="Re-definition and review of the Oriental genus Hexachaetus Chaudoir, 1871 (Coleoptera: Carabidae: Orthogoniini)" masterLastPageNumber="554" masterPageNumber="540" pageNumber="543" updateTime="1698386596725" updateUser="plazi">
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<mods:title id="5D4F4E81EDC16185A53C0321DD17AD38">Re-definition and review of the Oriental genus Hexachaetus Chaudoir, 1871 (Coleoptera: Carabidae: Orthogoniini)</mods:title>
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<mods:namePart id="06395C60E15090C64B9BFD5DDA8AEE17">Tian, Mingyi</mods:namePart>
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<mods:roleTerm id="F170B17895BCF04D3A73393A8306E2D3">Author</mods:roleTerm>
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<mods:namePart id="0977E29C8308596725EA79266B90E63D">Deuve, Thierry</mods:namePart>
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<mods:title id="9CD1E3408961AC9D6E7D2FFB0AB3051E">Zootaxa</mods:title>
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<mods:date id="36616C72B97C8576A8370DA7A154D795">2016</mods:date>
<mods:detail id="26CEF9F593ECCCB3D1051829395452C6" type="volume">
<mods:number id="0EBC5854EFFA9C5BA8252EF040F35EDE">4169</mods:number>
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<mods:identifier id="07F8AE4FCADBACFB059EF4ABD0AD4CA2" type="DOI">10.11646/zootaxa.4169.3.7</mods:identifier>
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<treatment id="8706BE21FFB9FF9B1DC8FADEFDBCA2B7" ID-DOI="http://doi.org/10.5281/zenodo.6075819" ID-GBIF-Taxon="124552515" ID-Zenodo-Dep="6075819" LSID="urn:lsid:plazi:treatment:8706BE21FFB9FF9B1DC8FADEFDBCA2B7" httpUri="http://treatment.plazi.org/id/8706BE21FFB9FF9B1DC8FADEFDBCA2B7" lastPageNumber="543" pageId="3" pageNumber="543">
<subSubSection id="47B55CBCFFB9FF9B1DC8FADEFEDFA0D4" pageId="3" pageNumber="543" type="nomenclature">
<paragraph id="0F100F37FFB9FF9B1DC8FADEFC74A036" blockId="3.[151,900,1366,1426]" box="[151,900,1366,1392]" pageId="3" pageNumber="543">
<heading id="5458B85BFFB9FF9B1DC8FADEFC74A036" bold="true" box="[151,900,1366,1392]" fontSize="11" level="1" pageId="3" pageNumber="543" reason="1">
<emphasis id="3DDBD325FFB9FF9B1DC8FADEFC74A036" bold="true" box="[151,900,1366,1392]" pageId="3" pageNumber="543">
<taxonomicName id="C8AF74B4FFB9FF9B1DC8FADEFCD3A036" ID-CoL="7GWRY" authority="Tian et Deuve, 2006" authorityName="Tian et Deuve" authorityYear="2006" box="[151,803,1366,1392]" class="Insecta" family="Carabidae" genus="Hexachaetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="543" phylum="Arthropoda" rank="subSpecies" species="lateralis" status="stat. nov." subSpecies="maindroni">
<emphasis id="3DDBD325FFB9FF9B1DC8FADEFDD4A036" bold="true" box="[151,548,1366,1392]" italics="true" pageId="3" pageNumber="543">Hexachaetus lateralis maindroni</emphasis>
<bibRefCitation id="6B3E72C6FFB9FF9B1F74FADEFCD3A036" author="Tian" box="[555,803,1366,1392]" pageId="3" pageNumber="543" refString="Tian, M. Y. &amp; Deuve, T. (2006) Contribution to the knowledge of the tribe Orthogoniini of the Oriental Region. II. (Coleoptera: Caraboidea). Coleopteres, 12 (9), 111 - 154." type="journal article" year="2006">Tian et Deuve, 2006</bibRefCitation>
</taxonomicName>
,
<taxonomicNameLabel id="26E86E5EFFB9FF9B1E6EFADEFC74A036" box="[817,900,1366,1392]" pageId="3" pageNumber="543">n. stat.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="0F100F37FFB9FF9B1DC8FAF1FEDFA0D4" blockId="3.[151,900,1366,1426]" box="[151,303,1401,1426]" pageId="3" pageNumber="543">
(
<figureCitation id="979413B2FFB9FF9B1DC1FAF1FED6A0D4" box="[158,294,1401,1426]" captionStart="FIGURES 1 – 4" captionStartId="4.[151,264,1491,1514]" captionTargetBox="[395,1186,207,1455]" captionTargetId="figure@4.[386,1201,195,1468]" captionTargetPageId="4" captionText="FIGURES 1 – 4. Hexachaetus lateralis Guérin (1. H. lateralis lateralis Guérin, habitus, male (photo: E. Kirschhofer); 2. H. lateralis maindroni Tian et Deuve, habitus, male, holotype; 3. Median lobe of aedeagus of H. lateralis maindroni, lateral view; 4. ibid. apical lamella, dorsal view; Scale bar: 5.0 mm)" httpUri="https://zenodo.org/record/258028/files/figure.png" pageId="3" pageNumber="543">Figures 2–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="47B55CBCFFB9FF9B1DC8FA49FD0BA35A" pageId="3" pageNumber="543" type="reference_group">
<paragraph id="0F100F37FFB9FF9B1DC8FA49FD61A091" blockId="3.[151,657,1473,1495]" box="[151,657,1473,1495]" pageId="3" pageNumber="543">
<treatmentCitationGroup id="2FBF2819FFB9FF9B1DC8FA49FD61A091" box="[151,657,1473,1495]" pageId="3" pageNumber="543">
<taxonomicName id="C8AF74B4FFB9FF9B1DC8FA49FD61A091" authority="Tian &amp; Deuve, 2006: 149" authorityName="Tian &amp; Deuve" authorityPageNumber="149" authorityYear="2006" box="[151,657,1473,1495]" class="Insecta" family="Carabidae" genus="Hexachaetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="543" phylum="Arthropoda" rank="species" species="maindroni">
<emphasis id="3DDBD325FFB9FF9B1DC8FA49FE78A091" box="[151,392,1473,1495]" italics="true" pageId="3" pageNumber="543">Hexachaetus maindroni</emphasis>
<treatmentCitation id="8E0E2926FFB9FF9B1CD0FA49FD61A091" author="Tian" box="[399,657,1473,1495]" page="149" pageId="3" pageNumber="543" year="2006">
<bibRefCitation id="6B3E72C6FFB9FF9B1CD0FA49FD92A091" author="Tian" box="[399,610,1473,1495]" pageId="3" pageNumber="543" refString="Tian, M. Y. &amp; Deuve, T. (2006) Contribution to the knowledge of the tribe Orthogoniini of the Oriental Region. II. (Coleoptera: Caraboidea). Coleopteres, 12 (9), 111 - 154." type="journal article" year="2006">Tian &amp; Deuve, 2006</bibRefCitation>
: 149
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="0F100F37FFB9FF9B1DC8F98CFD0BA35A" blockId="3.[151,1437,1540,2033]" box="[151,763,1540,1565]" pageId="3" pageNumber="543">
Length: 19.0 mm; width: 8.0 mm. Habitus as in
<figureCitation id="979413B2FFB9FF9B1FECF98CFD07A35A" box="[691,759,1540,1565]" captionStart="FIGURES 1 – 4" captionStartId="4.[151,264,1491,1514]" captionTargetBox="[395,1186,207,1455]" captionTargetId="figure@4.[386,1201,195,1468]" captionTargetPageId="4" captionText="FIGURES 1 – 4. Hexachaetus lateralis Guérin (1. H. lateralis lateralis Guérin, habitus, male (photo: E. Kirschhofer); 2. H. lateralis maindroni Tian et Deuve, habitus, male, holotype; 3. Median lobe of aedeagus of H. lateralis maindroni, lateral view; 4. ibid. apical lamella, dorsal view; Scale bar: 5.0 mm)" httpUri="https://zenodo.org/record/258028/files/figure.png" pageId="3" pageNumber="543">Fig. 2</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="47B55CBCFFB9FF9B1D98F9AFFD31A3B2" pageId="3" pageNumber="543" type="diagnosis">
<paragraph id="0F100F37FFB9FF9B1D98F9AFFD31A3B2" blockId="3.[151,1437,1540,2033]" pageId="3" pageNumber="543">
<emphasis id="3DDBD325FFB9FF9B1D98F9AFFEB1A306" bold="true" box="[199,321,1575,1600]" pageId="3" pageNumber="543">Diagnosis.</emphasis>
Large sized, polish, glabrous and shiny species; head, pronotum, side margin of elytra, whole apical margin of abdominal ventrite VII, tibiae and tarsi of legs dark brown, underside, femora and coxae yellowish brown; elytral marking narrow, not anchoreaform; labial palpomere 3 glabrous, slightly shorter than palpomere 2; labial palpomere 2 bisetose on inner margin, and with two more shorter setae near apex; maxillary palpomere 3 longer than 4; the median lobe of aedeagus stout, dramatically constricted at apex in profile; lamella broadly and extraordinary dilated in lateral aspect (
<figureCitation id="979413B2FFB9FF9B1F18F954FD44A3B2" box="[583,692,1756,1781]" captionStart="FIGURES 1 – 4" captionStartId="4.[151,264,1491,1514]" captionTargetBox="[395,1186,207,1455]" captionTargetId="figure@4.[386,1201,195,1468]" captionTargetPageId="4" captionText="FIGURES 1 – 4. Hexachaetus lateralis Guérin (1. H. lateralis lateralis Guérin, habitus, male (photo: E. Kirschhofer); 2. H. lateralis maindroni Tian et Deuve, habitus, male, holotype; 3. Median lobe of aedeagus of H. lateralis maindroni, lateral view; 4. ibid. apical lamella, dorsal view; Scale bar: 5.0 mm)" httpUri="https://zenodo.org/record/258028/files/figure.png" pageId="3" pageNumber="543">Figs. 3–4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="47B55CBCFFB9FF9B1D98F977FBD5A219" pageId="3" pageNumber="543" type="discussion">
<paragraph id="0F100F37FFB9FF9B1D98F977FBD5A219" blockId="3.[151,1437,1540,2033]" pageId="3" pageNumber="543">
<emphasis id="3DDBD325FFB9FF9B1D98F977FECBA25E" bold="true" box="[199,315,1791,1816]" pageId="3" pageNumber="543">Remarks.</emphasis>
Similar to
<taxonomicName id="C8AF74B4FFB9FF9B1CE6F889FD6DA25E" box="[441,669,1792,1816]" class="Insecta" family="Carabidae" genus="Hexachaetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="543" phylum="Arthropoda" rank="subSpecies" species="lateralis" subSpecies="lateralis">
<emphasis id="3DDBD325FFB9FF9B1CE6F889FD6DA25E" box="[441,669,1792,1816]" italics="true" pageId="3" pageNumber="543">H. lateralis lateralis</emphasis>
</taxonomicName>
not only on appearance, but also on genital structure. However, it is slightly larger than
<taxonomicName id="C8AF74B4FFB9FF9B1C2BF8ADFE03A27A" box="[372,499,1828,1852]" class="Insecta" family="Carabidae" genus="Hexachaetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="543" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="3DDBD325FFB9FF9B1C2BF8ADFE03A27A" box="[372,499,1828,1852]" italics="true" pageId="3" pageNumber="543">H. lateralis</emphasis>
</taxonomicName>
, elytra marking narrower at middle, extending only to intervals 2 (to intervals
<quantity id="C857A2D2FFB9FF9B182EF8ACFA6CA27B" box="[1393,1436,1828,1853]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="3" pageNumber="543" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="C8AF74B4FFB9FF9B1DC8F8C1FEE6A226" box="[151,278,1864,1888]" class="Insecta" family="Carabidae" genus="Hexachaetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="543" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="3DDBD325FFB9FF9B1DC8F8C1FEE6A226" box="[151,278,1864,1888]" italics="true" pageId="3" pageNumber="543">H. lateralis</emphasis>
</taxonomicName>
), and inner apical angle of elytra blunter than in
<taxonomicName id="C8AF74B4FFB9FF9B1E64F8C1FBEEA226" box="[827,1054,1864,1888]" class="Insecta" family="Carabidae" genus="Hexachaetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="543" phylum="Arthropoda" rank="subSpecies" species="lateralis" subSpecies="lateralis">
<emphasis id="3DDBD325FFB9FF9B1E64F8C1FBEEA226" box="[827,1054,1864,1888]" italics="true" pageId="3" pageNumber="543">H. lateralis lateralis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="47B55CBCFFB9FF9B1D98F8E4FE6AA28A" pageId="3" pageNumber="543" type="materials_examined">
<paragraph id="0F100F37FFB9FF9B1D98F8E4FE6AA28A" blockId="3.[151,1437,1540,2033]" pageId="3" pageNumber="543">
<materialsCitation id="BFC7056AFFB9FF9B1D98F8E4FE6AA28A" ID-GBIF-Occurrence="1313880548" collectingDate="1894-10-16" collectionCode="MNHN" collectorName="Museum Paris &amp; Ex Coll. M." country="Malaysia" determinerName="Tian &amp; Deuve &amp; Maindron M." location="Borneo" municipality="Kuching" pageId="3" pageNumber="543" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="3DDBD325FFB9FF9B1D98F8E4FEC6A2C3" bold="true" box="[199,310,1900,1925]" pageId="3" pageNumber="543">Material.</emphasis>
<specimenCount id="19A9C4BEFFB9FF9B1C62F8E4FE7BA2C3" box="[317,395,1900,1925]" pageId="3" pageNumber="543" type="male">1 male</specimenCount>
, the
<typeStatus id="D014B195FFB9FF9B1CE0F8E4FDD4A2C3" box="[447,548,1900,1925]" pageId="3" pageNumber="543" type="holotype">holotype</typeStatus>
, “
<location id="0A7059ECFFB9FF9B1F65F8E4FD61A2C2" LSID="urn:lsid:plazi:treatment:8706BE21FFB9FF9B1DC8FADEFDBCA2B7:0A7059ECFFB9FF9B1F65F8E4FD61A2C2" box="[570,657,1900,1924]" country="Malaysia" municipality="Kuching" name="Borneo" pageId="3" pageNumber="543">Borneo</location>
,
<collectingCountry id="77B84FA7FFB9FF9B1FC3F8E4FD0FA2C3" box="[668,767,1900,1925]" name="Malaysia" pageId="3" pageNumber="543">Sarawak</collectingCountry>
,
<collectingMunicipality id="EF74954DFFB9FF9B1E54F8E4FC80A2C3" box="[779,880,1900,1925]" pageId="3" pageNumber="543">Kuching</collectingMunicipality>
”, “
<collectingDate id="6B55D01FFFB9FF9B1ECEF8E4FBD8A2C2" box="[913,1064,1900,1925]" pageId="3" pageNumber="543" value="1894-10-16">
<date id="7B1129F7FFB9FF9B1ECEF8E4FBD8A2C2" box="[913,1064,1900,1925]" pageId="3" pageNumber="543" value="1894-10-16">16 Oct. 1894</date>
</collectingDate>
”, “
<taxonomicName id="C8AF74B4FFB9FF9B1914F8E4FAA6A2C2" box="[1099,1366,1900,1924]" class="Insecta" family="Carabidae" genus="Hexachaetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="543" phylum="Arthropoda" rank="species" species="maindroni" status="sp. nov.">
<emphasis id="3DDBD325FFB9FF9B1914F8E4FAA6A2C2" box="[1099,1366,1900,1924]" italics="true" pageId="3" pageNumber="543">Hexachaetus maindroni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="26E86E5EFFB9FF9B1802F8E4FA6CA2C3" box="[1373,1436,1900,1925]" pageId="3" pageNumber="543" rank="species">
<emphasis id="3DDBD325FFB9FF9B1802F8E4FA6CA2C3" bold="true" box="[1373,1436,1900,1925]" pageId="3" pageNumber="543">n. sp.</emphasis>
</taxonomicNameLabel>
det.
<bibRefCitation id="6B3E72C6FFB9FF9B1D95F807FE4CA2EE" author="Tian" box="[202,444,1935,1960]" pageId="3" pageNumber="543" refString="Tian, M. Y. &amp; Deuve, T. (2004) Descriptions of three new species of Orthogoniini (Coleoptera: Caraboidea) from Sumatra Island, Indonesia. Entomotaxonomia, 26 (1), 23 - 28." type="journal article" year="2004">
<determinerName id="FACCE3F3FFB9FF9B1D95F807FF0CA2EE" box="[202,252,1935,1960]" pageId="3" pageNumber="543">Tian</determinerName>
&amp;
<determinerName id="FACCE3F3FFB9FF9B1C7AF818FE81A2E1" box="[293,369,1936,1959]" pageId="3" pageNumber="543">Deuve</determinerName>
, 2004
</bibRefCitation>
”, “
<taxonomicName id="C8AF74B4FFB9FF9B1CBFF818FD13A2EE" box="[480,739,1936,1960]" class="Insecta" family="Carabidae" genus="Hexachaetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="543" phylum="Arthropoda" rank="species" species="laeviceps">
<emphasis id="3DDBD325FFB9FF9B1CBFF818FD13A2EE" box="[480,739,1936,1960]" italics="true" pageId="3" pageNumber="543">Hexachaetus laeviceps</emphasis>
</taxonomicName>
M.”(det.
<determinerName id="FACCE3F3FFB9FF9B1E0AF807FC1FA2EE" box="[853,1007,1935,1960]" pageId="3" pageNumber="543">Maindron M.</determinerName>
), “
<collectorName id="A25A6AE1FFB9FF9B194EF818FB4CA2EE" box="[1041,1212,1935,1960]" pageId="3" pageNumber="543">Museum Paris</collectorName>
”, and “
<collectorName id="A25A6AE1FFB9FF9B184EF818FA6CA2E1" box="[1297,1436,1935,1960]" pageId="3" pageNumber="543">Ex Coll. M.</collectorName>
Maindron”, in
<collectionCode id="69BE97F2FFB9FF9B1C62F83DFE66A28A" box="[317,406,1973,1996]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="543">MNHN</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="47B55CBCFFB9FF9B1D98F850FDBCA2B7" box="[199,588,2008,2033]" pageId="3" pageNumber="543" type="distribution">
<paragraph id="0F100F37FFB9FF9B1D98F850FDBCA2B7" blockId="3.[151,1437,1540,2033]" box="[199,588,2008,2033]" pageId="3" pageNumber="543">
<emphasis id="3DDBD325FFB9FF9B1D98F850FE90A2B7" bold="true" box="[199,352,2008,2033]" pageId="3" pageNumber="543">Distribution.</emphasis>
<collectingCountry id="77B84FA7FFB9FF9B1C38F850FE3EA2B7" box="[359,462,2008,2033]" name="Malaysia" pageId="3" pageNumber="543">Malaysia</collectingCountry>
(
<figureCitation id="979413B2FFB9FF9B1C82F850FDC0A2B7" box="[477,560,2008,2033]" captionStart="FIGURE 20" captionStartId="12.[151,250,1789,1811]" captionTargetBox="[226,1360,198,1765]" captionTargetId="figure@12.[223,1363,195,1768]" captionTargetPageId="12" captionText="FIGURE 20. Distribution of Hexachaetus species (square: angulatus species group; round: lateralis species group; star: laevissimus species group; heart: vietnamensis species group; triangle: mulan species group; a. H. lateralis lateralis; b. H. lateralis maindroni; c. H. maculatus; d. H. kirschenhoferi n. sp.; e. H. brunki n. sp.; f. H. laevissimus; g. H. vietnamensis n. sp.; h. H. prodigus; i. H. mulan n. sp.)" httpUri="https://zenodo.org/record/258032/files/figure.png" pageId="3" pageNumber="543">Fig. 20</figureCitation>
b).
</paragraph>
</subSubSection>
</treatment>
</document>
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