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<mods:title id="DB52D5E149B145EA0A6448920D801ACE">“ Larger ” Benthic Foraminifera Of The Cenomanian. A Review Of The Identity And The Stratigraphic And Palaeogeographic Distribution Of Non-Fusiform Planispiral (Or Near-Planispiral) Forms</mods:title>
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<mods:namePart id="2E7FB6CF75B9442D83623B97F567EB24">SIMMONS, MICHAEL</mods:namePart>
<mods:affiliation id="6B99D7E125E5EB1A7B191D480A834709">Halliburton, 97 Milton Park, Abingdon, OX 14 4 RW, UK &amp; The Natural History Museum, Cromwell Road, London, SW 7 5 BD, UK</mods:affiliation>
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<paragraph id="8BF336A0FF9EA254FCB6FC6EA1E1C49B" blockId="71.[816,1436,886,976]" box="[816,1196,952,976]" pageId="71" pageNumber="110">
<taxonomicName id="4C4C4D23FF9EA254FCB6FC6EA1E1C49B" authority="De Castro 1965" authorityName="De Castro" authorityYear="1965" box="[816,1196,952,976]" class="Globothalamea" family="Charentiidae" genus="Peneroplis" kingdom="Chromista" order="Loftusiida" pageId="71" pageNumber="110" phylum="Foraminifera" rank="species" species="parvus">
<emphasis id="B938EAB2FF9EA254FCB6FC6EA1E1C49B" bold="true" box="[816,1196,952,976]" pageId="71" pageNumber="110">
<emphasis id="B938EAB2FF9EA254FCB6FC6EA6BBC49B" bold="true" box="[816,1014,952,976]" italics="true" pageId="71" pageNumber="110">Peneroplis parvus</emphasis>
<bibRefCitation id="EFDD4B51FF9EA254FC7BFC6EA1E1C49B" author="De Castro, P." box="[1021,1196,952,976]" pageId="71" pageNumber="110" pagination="317 - 372" refId="ref78519" refString="De Castro, P., 1965. Su alcune Soritidae (Foraminiferida) del Cretacico della Campania. Societa dei Naturalisti in Napoli, 74: 317 - 372." type="journal article" year="1965">De Castro 1965</bibRefCitation>
</emphasis>
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<subSubSection id="C356652BFF9EA254FCB6FC2CA61FC33E" pageId="71" pageNumber="110" type="reference_group">
<paragraph id="8BF336A0FF9EA254FCB6FC2CA1FBC359" blockId="71.[816,1206,1018,1042]" box="[816,1206,1018,1042]" pageId="71" pageNumber="110">Reference Illustration &amp; Description</paragraph>
<paragraph id="8BF336A0FF9EA254FCB6FBEBA61FC33E" blockId="71.[816,1437,1084,2068]" pageId="71" pageNumber="110">
De Castro in
<bibRefCitation id="EFDD4B51FF9EA254FC39FBEAA04BC31F" author="Schroeder, R. &amp; Neumann, M." box="[959,1286,1084,1108]" pageId="71" pageNumber="110" pagination="1 - 161" refId="ref98244" refString="Schroeder, R. &amp; Neumann, M., 1985. Les Grands foraminiferes du Cretace moyen de la region mediterraneenne. Geobios Memoire special, 7: 1 - 161." type="journal article" year="1985">Schroeder &amp; Neumann (1985)</bibRefCitation>
, Pl. 39, p. 86- 88.
</paragraph>
</subSubSection>
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<paragraph id="8BF336A0FF9EA254FCB6FBA9A198C16F" blockId="71.[816,1437,1084,2068]" pageId="71" pageNumber="110">
Originally described as
<taxonomicName id="4C4C4D23FF9EA254FBC6FBA9A620C3F3" authority="(Fichtel &amp; Moll)" authorityName="SIMMONS &amp; BIDGOOD" authorityYear="2023" baseAuthorityName="Fichtel &amp; Moll" baseAuthorityYear="1798" class="Globothalamea" family="Charentiidae" genus="Peneroplis" kingdom="Chromista" order="Loftusiida" pageId="71" pageNumber="110" phylum="Foraminifera" rank="species" species="planatus" status="subsp. nov.">
<emphasis id="B938EAB2FF9EA254FBC6FBA9A051C3DC" box="[1088,1308,1151,1175]" italics="true" pageId="71" pageNumber="110">Peneroplis planatus</emphasis>
(Fichtel &amp; Moll)
</taxonomicName>
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n.
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</taxonomicNameLabel>
<emphasis id="B938EAB2FF9EA254FC39FB77A14AC3F3" box="[959,1031,1185,1208]" italics="true" pageId="71" pageNumber="110">parvus</emphasis>
, the
<typeStatus id="54F78802FF9EA254FBBAFB76A127C3F3" box="[1084,1130,1184,1208]" pageId="71" pageNumber="110">type</typeStatus>
and subsequent descriptions of the species by
<bibRefCitation id="EFDD4B51FF9EA254FC68FB14A1EEC392" author="De Castro, P." box="[1006,1187,1217,1241]" pageId="71" pageNumber="110" pagination="317 - 372" refId="ref78519" refString="De Castro, P., 1965. Su alcune Soritidae (Foraminiferida) del Cretacico della Campania. Societa dei Naturalisti in Napoli, 74: 317 - 372." type="journal article" year="1965">De Castro (1965</bibRefCitation>
,
<bibRefCitation id="EFDD4B51FF9EA254FB28FB17A1A8C392" author="De Castro, P." box="[1198,1253,1217,1241]" pageId="71" pageNumber="110" pagination="86 - 97" refId="ref78728" refString="De Castro, P., 1985. Peneroplis parvus, Pseudorhapydionina dubia, P. laurinensis &amp; Pseudorhipidionina casertana. In: Schroeder, R. and Neumann, M., (eds.) Les Grands foraminiferes du Cretace moyen de la region mediterraneenne. Geobios Memoire special, 7: 86 - 97." type="journal article" year="1985">1985</bibRefCitation>
) are comprehensive and allow for confident identification (see also
<bibRefCitation id="EFDD4B51FF9EA254FCB6FAD5A169C250" author="Calonge-Garcia, A." box="[816,1060,1283,1307]" pageId="71" pageNumber="110" pagination="25 - 45" refId="ref76128" refString="Calonge-Garcia, A., 1996. Soritids of Cretaceous from Iberian Range (Spain). Coloquios de Paleontologia, 48: 25 - 45." type="journal article" year="1996">Calonge-Garcia, 1996</bibRefCitation>
; p. 38, pl. 2, figs. 8-13). A peneropolid with a porcelaneous, imperforate wall and multiple cribrate apertures. 5-6 chambers in the first whorl,
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the last (second) whorl. May become flabelliform. The species is very similar to the extant
<taxonomicName id="4C4C4D23FF9EA254FAAFFA51A11DC28A" authority="(Fitchell &amp; Moll)" baseAuthorityName="Fitchell &amp; Moll" class="Globothalamea" family="Charentiidae" genus="Peneroplis" kingdom="Chromista" order="Loftusiida" pageId="71" pageNumber="110" phylum="Foraminifera" rank="species" species="planatus">
<emphasis id="B938EAB2FF9EA254FAAFFA51A6C1C28A" italics="true" pageId="71" pageNumber="110">Peneroplis planatus</emphasis>
(Fitchell &amp; Moll)
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, but with a smaller size (equatorial diameter c.
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), plus more adult chambers per mm (11.5-17.5). See the Species Key Chart (Appendix) for diagnostic and other characteristics.
</paragraph>
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<taxonomicName id="4C4C4D23FF9EA254FCB6F9F8A6D4C10E" authorityName="De Castro" authorityYear="1965" box="[816,921,1582,1605]" class="Globothalamea" family="Charentiidae" genus="Peneroplis" kingdom="Chromista" order="Loftusiida" pageId="71" pageNumber="110" phylum="Foraminifera" rank="species" species="parvus">
<emphasis id="B938EAB2FF9EA254FCB6F9F8A6D4C10E" box="[816,921,1582,1605]" italics="true" pageId="71" pageNumber="110">P. parvus</emphasis>
</taxonomicName>
is amongst the oldest known representatives of
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<emphasis id="B938EAB2FF9EA254FCB6F998A6E9C12D" box="[816,932,1614,1638]" italics="true" pageId="71" pageNumber="110">Peneroplis</emphasis>
</taxonomicName>
, a genus often regarded as restricted to the Cenozoic and Recent (
<bibRefCitation id="EFDD4B51FF9EA254FBB6F9B9A019C1CC" author="Loeblich Jnr., A. R. &amp; Tappan, H." box="[1072,1364,1647,1671]" pageId="71" pageNumber="110" refId="ref87900" refString="Loeblich Jnr., A. R. &amp; Tappan, H., 1988. Foraminiferal genera and their classification. Van Nostrand Reinhold, 2 vols." type="book" year="1988">Loeblich &amp; Tappan, 1988</bibRefCitation>
;
<bibRefCitation id="EFDD4B51FF9EA254FAEEF9A6A154C1E3" author="BouDagher-Fadel, M. K." pageId="71" pageNumber="110" pagination="1 - 37" refId="ref75051" refString="BouDagher-Fadel, M. K., 2008. Biology and evolutionary history of larger benthic foraminifera. Developments in Palaeontology and Stratigraphy, 21: 1 - 37." type="journal article" year="2008">BouDagher-Fadel, 2008</bibRefCitation>
).
<bibRefCitation id="EFDD4B51FF9EA254FBBFF946A05BC1E3" author="Chiocchini, M." box="[1081,1302,1680,1704]" pageId="71" pageNumber="110" pagination="171 - 202" refId="ref76776" refString="Chiocchini, M., 2008 a. New benthic foraminifers (Miliolacea and Soritacea) from the Cenomanian and Upper Turonian of the Monte Cairo (Southern Latium, Central Italy). Memorie Descrittive della Carta Geologica d'Italia, 84: 171 - 202." type="journal article" year="2008">Chiocchini (2008a)</bibRefCitation>
introduced
<taxonomicName id="4C4C4D23FF9EA254FCB6F967A153C182" authorityName="Chiocchini" authorityYear="2008" box="[816,1054,1713,1737]" class="Globothalamea" family="Charentiidae" genus="Peneroplis" kingdom="Chromista" order="Loftusiida" pageId="71" pageNumber="110" phylum="Foraminifera" rank="species" species="cairoensis">
<emphasis id="B938EAB2FF9EA254FCB6F967A153C182" box="[816,1054,1713,1737]" italics="true" pageId="71" pageNumber="110">Peneroplis cairoensis</emphasis>
</taxonomicName>
from the initial part of their late Cenomanian (=intra-middle Cenomanian) of central
<collectingCountry id="F35B7630FF9EA254FAE3F904A0D4C1A1" box="[1381,1433,1746,1770]" name="Italy" pageId="71" pageNumber="110">Italy</collectingCountry>
. This species is slightly larger than
<taxonomicName id="4C4C4D23FF9EA254FB35F923A056C047" authorityName="De Castro" authorityYear="1965" box="[1203,1307,1781,1804]" class="Globothalamea" family="Charentiidae" genus="Peneroplis" kingdom="Chromista" order="Loftusiida" pageId="71" pageNumber="110" phylum="Foraminifera" rank="species" species="parvus">
<emphasis id="B938EAB2FF9EA254FB35F923A056C047" box="[1203,1307,1781,1804]" italics="true" pageId="71" pageNumber="110">P. parvus</emphasis>
</taxonomicName>
(c.
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diameter), has a greater number of chambers in the uncoiled stage and a larger diameter of apertural pores. It has not been recorded from outside its
<typeStatus id="54F78802FF9EA254FB6FF88EA05AC03B" box="[1257,1303,1880,1904]" pageId="71" pageNumber="110">type</typeStatus>
area and is not considered further.
<taxonomicName id="4C4C4D23FF9EA254FBB2F8AEA07EC0DB" baseAuthorityName="Peybernes" baseAuthorityYear="1984" box="[1076,1331,1912,1936]" class="Globothalamea" family="Charentiidae" genus="Peneroplis" kingdom="Chromista" order="Loftusiida" pageId="71" pageNumber="110" phylum="Foraminifera" rank="species" species="aragonensis">
<emphasis id="B938EAB2FF9EA254FBB2F8AEA07EC0DB" box="[1076,1331,1912,1936]" italics="true" pageId="71" pageNumber="110">Peneroplis aragonensis</emphasis>
</taxonomicName>
is a species introduced from the late Albian of the Spanish Pyrenees by
<bibRefCitation id="EFDD4B51FF9EA254FC16F86CA102C099" author="Peybernes, B." box="[912,1103,1978,2002]" pageId="71" pageNumber="110" pagination="491 - 499" refId="ref93070" refString="Peybernes, B., 1984. Foraminiferes benthiques nouveaux de l'Albien superieur du Massif du Turbon (Pyrenees espagnole). Benthos ' 83; 2 nd lnternational Symposium Benthic Foraminifera (Pau): 491 - 499." type="journal article" year="1984">Peybernès (1984)</bibRefCitation>
as
<taxonomicName id="4C4C4D23FF9EA254FBFCF86CA0D1C099" authorityName="Peybernes" authorityYear="1984" box="[1146,1436,1978,2002]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="71" pageNumber="110" phylum="Foraminifera" rank="species" species="aragonensis">
<emphasis id="B938EAB2FF9EA254FBFCF86CA0D1C099" box="[1146,1436,1978,2002]" italics="true" pageId="71" pageNumber="110">Broeckinella? aragonensis</emphasis>
</taxonomicName>
and subsequently transferred to
<taxonomicName id="4C4C4D23FF9EA254FB09F80DA04EC0B8" authorityName="De Montfort" authorityYear="1808" box="[1167,1283,2011,2035]" class="Globothalamea" family="Charentiidae" genus="Peneroplis" kingdom="Chromista" order="Loftusiida" pageId="71" pageNumber="110" phylum="Foraminifera" rank="genus">
<emphasis id="B938EAB2FF9EA254FB09F80DA04EC0B8" box="[1167,1283,2011,2035]" italics="true" pageId="71" pageNumber="110">Peneroplis</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFDD4B51FF9EA254FAA9F80DA16ECF5F" author="Schlagintweit, F. &amp; Rashidi, K." pageId="71" pageNumber="110" pagination="57 - 67" refId="ref96972" refString="Schlagintweit, F. &amp; Rashidi, K., 2020. Broeckinella hensoni n. sp., a new larger benthic foraminifera from the upper Maastrichtian of Iran and a revision of the genus Broeckinella Henson, 1948. Acta Palaeontologica Romaniae, 16 (2): 57 - 67." type="journal article" year="2020">Schlagintweit &amp; Rashidi (2020)</bibRefCitation>
. This species is much larger than
<taxonomicName id="4C4C4D23FF90A25AFF11FF49A5B3C7FD" authorityName="De Castro" authorityYear="1965" box="[151,254,159,182]" class="Globothalamea" family="Charentiidae" genus="Peneroplis" kingdom="Chromista" order="Loftusiida" pageId="73" pageNumber="112" phylum="Foraminifera" rank="species" species="parvus">
<emphasis id="B938EAB2FF90A25AFF11FF49A5B3C7FD" box="[151,254,159,182]" italics="true" pageId="73" pageNumber="112">P. parvus</emphasis>
</taxonomicName>
(diameter c.
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) with a tendency to become sub-annular in macrospheric forms. It too has not been recorded outside its
<typeStatus id="54F78802FF90A25AFEFEFF37A4EBC7B2" box="[376,422,225,249]" pageId="73" pageNumber="112">type</typeStatus>
area and is not considered further.
<bibRefCitation id="EFDD4B51FF90A25AFF53FED7A4E3C652" author="Saint-Marc, P." box="[213,430,257,281]" pageId="73" pageNumber="112" pagination="1 - 342" refId="ref96009" refString="Saint-Marc, P., 1974 a. Etude stratigraphique et micropaleontologique de l'Albien, du Cenomanien et du Turonien du Liban. Notes et Memoires sur le Moyen- Orient, XIII, Museum National d'Histoire Naturelle, Paris, pp. 1 - 342." type="book chapter" year="1974">Saint-Marc (1974a)</bibRefCitation>
illustrated a “
<taxonomicName id="4C4C4D23FF90A25AFDDEFED7A7B5C652" box="[600,760,257,281]" class="Globothalamea" family="Charentiidae" genus="Peneroplis" kingdom="Chromista" order="Loftusiida" pageId="73" pageNumber="112" phylum="Foraminifera" rank="species" species="undetermined">
<emphasis id="B938EAB2FF90A25AFDDEFED7A781C652" box="[600,716,257,281]" italics="true" pageId="73" pageNumber="112">Peneroplis</emphasis>
sp.
</taxonomicName>
” from the late Albian of
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. This is a rapidly uncoiling form, broader and more rounded than
<taxonomicName id="4C4C4D23FF90A25AFDFEFE92A7AEC610" authorityName="De Castro" authorityYear="1965" box="[632,739,324,347]" class="Globothalamea" family="Charentiidae" genus="Peneroplis" kingdom="Chromista" order="Loftusiida" pageId="73" pageNumber="112" phylum="Foraminifera" rank="species" species="parvus">
<emphasis id="B938EAB2FF90A25AFDFEFE92A7AEC610" box="[632,739,324,347]" italics="true" pageId="73" pageNumber="112">P. parvus</emphasis>
</taxonomicName>
in axial section, and slightly larger. Consorti (in Consorti et al., 2018) introduced a new genus and species,
<taxonomicName id="4C4C4D23FF90A25AFD2AFE53A4CDC6F4" ID-CoL="83DRG" class="Tubothalamea" family="Soritidae" genus="Pseudopeneroplis" kingdom="Chromista" order="Miliolida" pageId="73" pageNumber="112" phylum="Foraminifera" rank="species" species="oyonensis">
<emphasis id="B938EAB2FF90A25AFD2AFE53A4CDC6F4" italics="true" pageId="73" pageNumber="112">Pseudopeneroplis oyonensis</emphasis>
</taxonomicName>
, from the upper Cenomanian of
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. In contrast to
<taxonomicName id="4C4C4D23FF90A25AFEEBFE1EA4ADC6AB" authorityName="De Montfort" authorityYear="1808" box="[365,480,456,480]" class="Globothalamea" family="Charentiidae" genus="Peneroplis" kingdom="Chromista" order="Loftusiida" pageId="73" pageNumber="112" phylum="Foraminifera" rank="genus">
<emphasis id="B938EAB2FF90A25AFEEBFE1EA4ADC6AB" box="[365,480,456,480]" italics="true" pageId="73" pageNumber="112">Peneroplis</emphasis>
</taxonomicName>
this taxon develops subdivisions in the marginal area of the chamber lumen but is otherwise similar.
</paragraph>
<caption id="DF336628FF91A25BFE59FCA3A1F1C4C0" ID-DOI="http://doi.org/10.5281/zenodo.10834303" ID-Zenodo-Dep="10834303" box="[479,1212,885,907]" httpUri="https://zenodo.org/record/10834303/files/figure.png" pageId="72" pageNumber="111" startId="72.[479,512,885,906]" targetBox="[252,1437,166,876]" targetPageId="72" targetType="figure">
<paragraph id="8BF336A0FF91A25BFE59FCA3A1F1C4C0" blockId="72.[479,1212,885,907]" box="[479,1212,885,907]" pageId="72" pageNumber="111">
<emphasis id="B938EAB2FF91A25BFE59FCA3A768C4C1" bold="true" box="[479,549,885,906]" pageId="72" pageNumber="111">Fig. 57</emphasis>
Cenomanian paleogeographic distribution of
<taxonomicName id="4C4C4D23FF91A25BFC58FCA3A1FBC4C0" authorityName="Luperto-Sinni" authorityYear="1968" box="[990,1206,885,907]" class="Globothalamea" family="Nautiloculinidae" genus="Nummofallotia" kingdom="Chromista" order="Lituolida" pageId="72" pageNumber="111" phylum="Foraminifera" rank="species" species="apula">Nummofallotia? apula</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF336628FF91A25BFF64F8B1A66FC0F3" ID-DOI="http://doi.org/10.5281/zenodo.10834305" ID-Zenodo-Dep="10834305" httpUri="https://zenodo.org/record/10834305/files/figure.png" pageId="72" pageNumber="111" startId="72.[226,259,1895,1916]" targetBox="[239,1454,982,1875]" targetPageId="72" targetType="figure">
<paragraph id="8BF336A0FF91A25BFF64F8B1A66FC0F3" blockId="72.[226,1437,1895,1977]" pageId="72" pageNumber="111">
<emphasis id="B938EAB2FF91A25BFF64F8B1A46AC037" bold="true" box="[226,295,1895,1916]" pageId="72" pageNumber="111">Fig. 58</emphasis>
Representative illustrations of
<taxonomicName id="4C4C4D23FF91A25BFDD5F8B1A64FC036" authorityName="De Castro" authorityYear="1965" box="[595,770,1895,1917]" class="Globothalamea" family="Charentiidae" genus="Peneroplis" kingdom="Chromista" order="Loftusiida" pageId="72" pageNumber="111" phylum="Foraminifera" rank="species" species="parvus">
<emphasis id="B938EAB2FF91A25BFDD5F8B1A64FC036" box="[595,770,1895,1917]" italics="true" pageId="72" pageNumber="111">Peneroplis parvus</emphasis>
</taxonomicName>
:
<emphasis id="B938EAB2FF91A25BFC96F8B1A651C037" bold="true" box="[784,796,1895,1916]" pageId="72" pageNumber="111">a</emphasis>
Equatorial section, De Castro in
<bibRefCitation id="EFDD4B51FF91A25BFBDCF8B1A035C036" author="Schroeder, R. &amp; Neumann, M." box="[1114,1400,1895,1917]" pageId="72" pageNumber="111" pagination="1 - 161" refId="ref98244" refString="Schroeder, R. &amp; Neumann, M., 1985. Les Grands foraminiferes du Cretace moyen de la region mediterraneenne. Geobios Memoire special, 7: 1 - 161." type="journal article" year="1985">Schroeder &amp; Neumann (1985</bibRefCitation>
, pl. 39, fig. 1, Italy);
<emphasis id="B938EAB2FF91A25BFE03F853A4DFC0D1" bold="true" box="[389,402,1925,1946]" pageId="72" pageNumber="111">b</emphasis>
Axial section, De Castro in
<bibRefCitation id="EFDD4B51FF91A25BFD25F853A68CC0D0" author="Schroeder, R. &amp; Neumann, M." box="[675,961,1925,1947]" pageId="72" pageNumber="111" pagination="1 - 161" refId="ref98244" refString="Schroeder, R. &amp; Neumann, M., 1985. Les Grands foraminiferes du Cretace moyen de la region mediterraneenne. Geobios Memoire special, 7: 1 - 161." type="journal article" year="1985">Schroeder &amp; Neumann (1985</bibRefCitation>
, pl. 39, fig. 13, Italy);
<emphasis id="B938EAB2FF91A25BFB1FF853A1E9C0D1" bold="true" box="[1177,1188,1925,1946]" pageId="72" pageNumber="111">c</emphasis>
Oblique axial section, De Castro in
<bibRefCitation id="EFDD4B51FF91A25BFEB9F875A710C0F2" author="Schroeder, R. &amp; Neumann, M." box="[319,605,1955,1977]" pageId="72" pageNumber="111" pagination="1 - 161" refId="ref98244" refString="Schroeder, R. &amp; Neumann, M., 1985. Les Grands foraminiferes du Cretace moyen de la region mediterraneenne. Geobios Memoire special, 7: 1 - 161." type="journal article" year="1985">Schroeder &amp; Neumann (1985</bibRefCitation>
, pl. 39, fig. 5, Italy).
</paragraph>
</caption>
<paragraph id="8BF336A0FF90A25AFF11FDFDA7C6C378" blockId="73.[151,772,158,1075]" pageId="73" pageNumber="112">
A potential confusion species is
<taxonomicName id="4C4C4D23FF90A25AFE7FFDFDA4A1C52F" authority="Said &amp; Kenawy. Originally" authorityName="Said &amp; Kenawy. Originally" class="Globothalamea" family="Nezzazatidae" genus="Neodubrovkinella" kingdom="Chromista" order="Lituolida" pageId="73" pageNumber="112" phylum="Foraminifera" rank="species" species="turonica">
<emphasis id="B938EAB2FF90A25AFE7FFDFDA5FAC52F" italics="true" pageId="73" pageNumber="112">Neodubrovkinella turonica</emphasis>
Said &amp; Kenawy. Originally
</taxonomicName>
described as a species of
<taxonomicName id="4C4C4D23FF90A25AFF11FDBBA447C5CE" authorityName="De Montfort" authorityYear="1808" box="[151,266,621,645]" class="Globothalamea" family="Charentiidae" genus="Peneroplis" kingdom="Chromista" order="Loftusiida" pageId="73" pageNumber="112" phylum="Foraminifera" rank="genus">
<emphasis id="B938EAB2FF90A25AFF11FDBBA447C5CE" box="[151,266,621,645]" italics="true" pageId="73" pageNumber="112">Peneroplis</emphasis>
</taxonomicName>
(“
<taxonomicName id="4C4C4D23FF90A25AFEAEFDBBA747C5CE" baseAuthorityName="Schlagintweit &amp; Yazdi-Moghadam" baseAuthorityYear="2022" box="[296,522,621,645]" class="Globothalamea" family="Charentiidae" genus="Peneroplis" kingdom="Chromista" order="Loftusiida" pageId="73" pageNumber="112" phylum="Foraminifera" rank="species" species="turonicus">
<emphasis id="B938EAB2FF90A25AFEAEFDBBA747C5CE" box="[296,522,621,645]" italics="true" pageId="73" pageNumber="112">Peneroplis turonicus</emphasis>
</taxonomicName>
”) from
<collectingCountry id="F35B7630FF90A25AFDE2FDB8A7E9C5CD" box="[612,676,622,646]" name="Egypt" pageId="73" pageNumber="112">Egypt</collectingCountry>
(
<bibRefCitation id="EFDD4B51FF90A25AFD32FDBBA475C5ED" author="Said, R. &amp; Kenawy, A." pageId="73" pageNumber="112" pagination="77 - 86" refId="ref95802" refString="Said, R. &amp; Kenawy, A., 1957. Foraminifera from the Turonian rocks of Abu-Roash, Egypt. Contributions from the Cushman Foundation for Foraminiferal Research, 8 (2): 77 - 86." type="journal article" year="1957">Said &amp; Kenawy, 1957</bibRefCitation>
), this taxon has been shown by
<bibRefCitation id="EFDD4B51FF90A25AFD13FD58A749C58C" author="Schlagintweit, F. &amp; Yazdi-Moghadam, M." pageId="73" pageNumber="112" refId="ref97579" refString="Schlagintweit, F. &amp; Yazdi-Moghadam, M., 2022 a." type="book" year="2022">Schlagintweit &amp; Yazdi-Moghadam (2022a)</bibRefCitation>
to be a biokovinid (and Cenomanian, not Turonian). In well-preserved material the finely agglutinating wall with a pseudo-keriothecal structure is visible, excluding it from the
<taxonomicName id="4C4C4D23FF90A25AFDF1FCC5A790C460" ID-CoL="7NG82" box="[631,733,787,811]" class="Tubothalamea" kingdom="Chromista" order="Miliolida" pageId="73" pageNumber="112" phylum="Foraminifera" rank="order">Miliolida</taxonomicName>
. In specimens where the wall structure is not clear, a large proloculus in megalospheric forms, a tendency to rapidly uncoil with chambers enlarging rapidly, are sufficient to distinguish
<taxonomicName id="4C4C4D23FF90A25AFE90FC4EA4DEC4E4" box="[278,403,920,943]" class="Globothalamea" family="Nezzazatidae" genus="Neodubrovkinella" kingdom="Chromista" order="Lituolida" pageId="73" pageNumber="112" phylum="Foraminifera" rank="species" species="turonica">
<emphasis id="B938EAB2FF90A25AFE90FC4EA4DEC4E4" box="[278,403,920,943]" italics="true" pageId="73" pageNumber="112">N. turonica</emphasis>
</taxonomicName>
from
<emphasis id="B938EAB2FF90A25AFE5AFC4EA701C4E4" box="[476,588,920,943]" italics="true" pageId="73" pageNumber="112">
<taxonomicName id="4C4C4D23FF90A25AFE5AFC4EA705C4E4" authorityName="De Castro" authorityYear="1965" box="[476,584,920,943]" class="Globothalamea" family="Charentiidae" genus="Peneroplis" kingdom="Chromista" order="Loftusiida" pageId="73" pageNumber="112" phylum="Foraminifera" rank="species" species="parvus">P. parvus</taxonomicName>
,
</emphasis>
as is occasional streptospiral coiling in early chambers. It is worth noting that in many thin-sections, and even illustrations of three-dimensional specimens, the porecellanous nature of the wall of
<taxonomicName id="4C4C4D23FF90A25AFF6FFBCAA403C378" authorityName="De Castro" authorityYear="1965" box="[233,334,1052,1075]" class="Globothalamea" family="Charentiidae" genus="Peneroplis" kingdom="Chromista" order="Loftusiida" pageId="73" pageNumber="112" phylum="Foraminifera" rank="species" species="parvus">
<emphasis id="B938EAB2FF90A25AFF6FFBCAA403C378" box="[233,334,1052,1075]" italics="true" pageId="73" pageNumber="112">P. parvus</emphasis>
</taxonomicName>
can be difficult to determine.
</paragraph>
</subSubSection>
<subSubSection id="C356652BFF90A25AFF11FB8BA7FBC3F3" pageId="73" pageNumber="112" type="distribution">
<paragraph id="8BF336A0FF90A25AFF11FB8BA4E5C33E" blockId="73.[151,424,1117,1141]" box="[151,424,1117,1141]" pageId="73" pageNumber="112">Stratigraphic Distribution</paragraph>
<paragraph id="8BF336A0FF90A25AFF11FB76A7FBC3F3" blockId="73.[151,772,1184,2068]" box="[151,694,1184,1208]" pageId="73" pageNumber="112">Late Albian/early Cenomanian late Cenomanian.</paragraph>
</subSubSection>
<subSubSection id="C356652BFF90A25AFF11FB17A147C460" pageId="73" pageNumber="112" type="description">
<paragraph id="8BF336A0FF90A25AFF11FB17A46BC1A1" blockId="73.[151,772,1184,2068]" pageId="73" pageNumber="112">
The stratigraphic range of
<taxonomicName id="4C4C4D23FF90A25AFE3FFB14A76DC392" authorityName="De Castro" authorityYear="1965" box="[441,544,1218,1241]" class="Globothalamea" family="Charentiidae" genus="Peneroplis" kingdom="Chromista" order="Loftusiida" pageId="73" pageNumber="112" phylum="Foraminifera" rank="species" species="parvus">
<emphasis id="B938EAB2FF90A25AFE3FFB14A76DC392" box="[441,544,1218,1241]" italics="true" pageId="73" pageNumber="112">P. parvus</emphasis>
</taxonomicName>
has been the subject of a range of opinions in the literature. In his original description,
<bibRefCitation id="EFDD4B51FF90A25AFE99FAD2A4ACC250" author="De Castro, P." box="[287,481,1283,1307]" pageId="73" pageNumber="112" pagination="317 - 372" refId="ref78519" refString="De Castro, P., 1965. Su alcune Soritidae (Foraminiferida) del Cretacico della Campania. Societa dei Naturalisti in Napoli, 74: 317 - 372." type="journal article" year="1965">De Castro (1965)</bibRefCitation>
thought the
<typeStatus id="54F78802FF90A25AFDF5FAD2A7ECC257" box="[627,673,1284,1308]" pageId="73" pageNumber="112">type</typeStatus>
material (from central
<collectingCountry id="F35B7630FF90A25AFEADFAF2A410C277" box="[299,349,1316,1340]" name="Italy" pageId="73" pageNumber="112">Italy</collectingCountry>
) to be probably middle Cenomanian in age, a view maintained by him in 1985 (De Castro in
<bibRefCitation id="EFDD4B51FF90A25AFF11FAB0A4ABC235" author="Schroeder, R. &amp; Neumann, M." box="[151,486,1382,1406]" pageId="73" pageNumber="112" pagination="1 - 161" refId="ref98244" refString="Schroeder, R. &amp; Neumann, M., 1985. Les Grands foraminiferes du Cretace moyen de la region mediterraneenne. Geobios Memoire special, 7: 1 - 161." type="journal article" year="1985">Schroeder &amp; Neumann, 1985</bibRefCitation>
), although he considered the overall range to be latest Albian to intra-middle Cenomanian based on his assessment of known occurrences to that time. However, in 1991 he considered that in the broad
<typeStatus id="54F78802FF90A25AFEA5FA3DA41CC148" box="[291,337,1515,1539]" pageId="73" pageNumber="112">type</typeStatus>
area an upper Albian lower Cenomanian biozone could be introduced based on the total range of the species. Similarly,
<bibRefCitation id="EFDD4B51FF90A25AFE31F9FBA78CC10E" author="Chiocchini, M. &amp; Pampaloni, M. L. &amp; Pichezzi, R. M." box="[439,705,1581,1605]" pageId="73" pageNumber="112" refId="ref77109" refString="Chiocchini, M., Pampaloni, M. L., &amp; Pichezzi, R. M., 2012. Microfacies e microfossili delle successioni carbonatiche mesozoiche del Lazio e dell &quot; Abruzzo (Italia centrale) - Cretacico. Memorie per servire alla descrizione della Carta Geologica d'Italia. ISPRA, Servizio Geologico d'Italia, Dipartimento Difesa del Suolo, Rome." type="book" year="2012">Chiocchini et al. (2012)</bibRefCitation>
illustrated the species and limited its range to the around the Albian Cenomanian boundary. By contrast
<bibRefCitation id="EFDD4B51FF90A25AFD24F9B9A463C1E3" author="Calonge-Garcia, A." pageId="73" pageNumber="112" pagination="25 - 45" refId="ref76128" refString="Calonge-Garcia, A., 1996. Soritids of Cretaceous from Iberian Range (Spain). Coloquios de Paleontologia, 48: 25 - 45." type="journal article" year="1996">Calonge-Garcia (1996)</bibRefCitation>
and
<bibRefCitation id="EFDD4B51FF90A25AFEE3F946A4B0C1E3" author="Bilotte, M." box="[357,509,1680,1704]" pageId="73" pageNumber="112" pagination="5" refId="ref74518" refString="Bilotte, M., 1998. Larger benthic foraminifera. In: Hardenbol, J., Jacquin, T., Farley, M. B., de Graciansky, P. - C., Vail, P. (eds.) Cretaceous Biochronostratigraphy: Mesozoic and Cenozoic Sequence Chronostratigraphic Framework of European Basins. SEPM Spec. Publ. 60, chart 5." type="book chapter" year="1998">Bilotte (1998)</bibRefCitation>
restricted the species to the lower Cenomanian, and
<bibRefCitation id="EFDD4B51FF90A25AFE5AF967A727C182" author="Velic, I." box="[476,618,1713,1737]" pageId="73" pageNumber="112" pagination="1 - 113" refId="ref101499" refString="Velic, I., 2007. Stratigraphy and Palaeobiogeography of Mesozoic Benthic Foraminifera of the Karst Dinarides (SE Europe). Geologia Croatica, 60 (1): 1 - 113." type="journal article" year="2007">Velić (2007)</bibRefCitation>
to the upper Cenomanian.
</paragraph>
<paragraph id="8BF336A0FF90A25AFF11F922A02BC6F4" blockId="73.[151,772,1184,2068]" lastBlockId="73.[816,1437,158,811]" pageId="73" pageNumber="112">
Determining the range of this species is hampered by a lack of unequivocal illustrations in the literature coupled with a lack of independent age calibration.
<bibRefCitation id="EFDD4B51FF90A25AFDE9F8E1A457C024" author="De Lapparent, A. F. &amp; Philip, J. &amp; Marie, P. &amp; Dufaure, P." pageId="73" pageNumber="112" pagination="311 - 318" refId="ref78987" refString="De Lapparent, A. F., Philip, J., Marie, P. &amp; Dufaure, P., 1974. La montagne de Parandaz et les transgressions cretacees en Afghanistan central. Bulletin de la Societe Geologique de France, XVI (7): 311 - 318." type="journal article" year="1974">De Lapparent et al. (1974)</bibRefCitation>
provided an unillustrated record of
<taxonomicName id="4C4C4D23FF90A25AFD1BF88EA649C024" authorityName="De Castro" authorityYear="1965" box="[669,772,1880,1903]" class="Globothalamea" family="Charentiidae" genus="Peneroplis" kingdom="Chromista" order="Loftusiida" pageId="73" pageNumber="112" phylum="Foraminifera" rank="species" species="parvus">
<emphasis id="B938EAB2FF90A25AFD1BF88EA649C024" box="[669,772,1880,1903]" italics="true" pageId="73" pageNumber="112">P. parvus</emphasis>
</taxonomicName>
from
<collectingCountry id="F35B7630FF90A25AFF50F8AEA415C0DB" box="[214,344,1912,1936]" name="Afghanistan" pageId="73" pageNumber="112">Afghanistan</collectingCountry>
and also points out the difficulties in establishing the difference between the Cenomanian and Turonian in sections devoid of ammonites. Upper Cenomanian records can be supported by the illustrated record from
<collectingCountry id="F35B7630FF90A25AFF52F82AA46FCF5F" box="[212,290,2044,2068]" name="Croatia" pageId="73" pageNumber="112">Croatia</collectingCountry>
of
<bibRefCitation id="EFDD4B51FF90A25AFECDF82AA72CCF5F" author="Velic, I. &amp; Vlahovic, I." box="[331,609,2044,2068]" pageId="73" pageNumber="112" pagination="25 - 43" refId="ref101422" refString="Velic, I. &amp; Vlahovic, I., 1994. Foraminiferal assemblages in the Cenomanian of the Buzet-Savudrija area (northwestern Istria, Croatia). Geologia Croatica, 47 (1): 25 - 43." type="journal article" year="1994">Velić &amp; Vlahović (1994)</bibRefCitation>
, the illustrated occurrence from undifferentiated middle upper Cenomanian of central
<collectingRegion id="4988F842FF90A25AFC7DFF69A13CC79C" box="[1019,1137,191,215]" country="Mexico" name="Distrito Federal" pageId="73" pageNumber="112">Mexico by</collectingRegion>
<bibRefCitation id="EFDD4B51FF90A25AFBFCFF69A01FC79C" author="Omana, L. &amp; Lopez-Doncel, R. &amp; Torres-Hernandez, J. R." box="[1146,1362,191,215]" pageId="73" pageNumber="112" pagination="1 - 94" refId="ref91696" refString="Omana, L., Lopez-Doncel, R., Torres-Hernandez, J. R., 2013, Late Cretaceous (Cenomanian-Santonian) foraminiferal biostratigraphy from W Valles-San Luis Platform, Mexico: 9 th Symposium of Cretaceous System, Ankara, Turkey, pp. 1 - 94." type="book chapter" year="2013">Omaña et al. (2013</bibRefCitation>
,
<bibRefCitation id="EFDD4B51FF90A25AFAE6FF69A0DAC79C" author="Omana, L. &amp; Torres, J. R. &amp; Lopez-Doncel, R. &amp; Alencaster, G. &amp; Caballero, I. L." box="[1376,1431,191,215]" pageId="73" pageNumber="112" pagination="28 - 44" refId="ref91802" refString="Omana, L., Torres, J. R., Lopez-Doncel, R., Alencaster, G. &amp; Caballero, I. L., 2014. A pithonellid bloom in the Cenomanian-Turonian boundary interval from Cerritos in the western Valles - San Luis Potosi platform, Mexico: paleoenvironmental significance. Revista Mexicana de Ciencias Geologicas, 31 (1): 28 - 44." type="journal article" year="2014">2014</bibRefCitation>
,
<bibRefCitation id="EFDD4B51FF90A25AFCB6FF36A624C7B3" author="Omana, L. &amp; Lopez-Doncel, R. &amp; Torres, J. R. &amp; Alencaster, G. &amp; Lopez-Caballero, I." box="[816,873,224,248]" pageId="73" pageNumber="112" pagination="691 - 725" refId="ref91622" refString="Omana, L., Lopez-Doncel, R., Torres, J. R., Alencaster, G. &amp; Lopez-Caballero, I. 2019. Mid - late Cenomanian larger benthic foraminifers from the El Abra Formation W Valles-San Luis Potosi Platform, central - eastern Mexico: Taxonomy, biostratigraphy and paleoenvironmental implications. Boletin de la Sociedad Geologica Mexicana, 71 (3): 691 - 725." type="journal article" year="2019">2019</bibRefCitation>
), and the illustrated record of Shahin &amp; El Baz (2010, 2013) from Sinai. An additional illustrated record from southeast
<collectingCountry id="F35B7630FF90A25AFC5AFEF4A163C671" box="[988,1070,290,314]" name="Mexico" pageId="73" pageNumber="112">Mexico</collectingCountry>
is that of
<bibRefCitation id="EFDD4B51FF90A25AFB20FEF4A6D7C610" author="Rosales-Dominguez, M. &amp; Bermudez-Santana, J. C. &amp; Aguilar-Pina, M." pageId="73" pageNumber="112" pagination="697 - 712" refId="ref95499" refString="Rosales-Dominguez, M. del C., Bermudez-Santana, J. C. &amp; Aguilar-Pina, M., 1997. Mid and Upper Cretaceous foraminiferal assemblages from the Sierra de Chiapas, southeastern Mexico. Cretaceous Research, 18: 697 - 712." type="journal article" year="1997">Rosales-Dominguez et al. (1997)</bibRefCitation>
but their assessment that it is “post Cenomanian” in age is based partly on circular reasoning and is incorrect. The unusual Turonian record of
<bibRefCitation id="EFDD4B51FF90A25AFB7CFE53A635C6F4" author="Orabi, O. H. &amp; Khalil, H. M." pageId="73" pageNumber="112" pagination="615 - 633" refId="ref92238" refString="Orabi, O. H. &amp; Khalil, H. M., 2001. Low-oxygen tolerant benthic foraminifera from the Cenomanian-Turonian transition of Gebel Farsh El Ghozlan section, west-central Sinai, Egypt. Second International Conference of the Geology of Africa, 1: 615 - 633." type="journal article" year="2001">Orabi &amp; Khalil (2001)</bibRefCitation>
from Sinai is not substantiated by illustration.
</paragraph>
<paragraph id="8BF336A0FF90A25AFCB6FE1EA147C460" blockId="73.[816,1437,158,811]" pageId="73" pageNumber="112">
Other than Chiocchini et al., (2012), late Albian and early Cenomanian records of the species lack substantiation by illustration but include Scott (2002) from
<collectingCountry id="F35B7630FF90A25AFAC5FDDCA0D5C569" box="[1347,1432,522,546]" name="Mexico" pageId="73" pageNumber="112">Mexico</collectingCountry>
; Calonge et al., (2002);
<bibRefCitation id="EFDD4B51FF90A25AFBB4FDFDA1B6C508" author="Caus, E. &amp; Bernaus, J. M. &amp; Calonge, E. &amp; Martin-Chivelet, M." box="[1074,1275,555,579]" pageId="73" pageNumber="112" pagination="172 - 181" refId="ref76249" refString="Caus, E., Bernaus, J. M., Calonge, E. &amp; Martin-Chivelet, M., 2009. Mid-Cenomanian separation of Atlantic and Tethyan domains in Iberia by a land-bridge: The origin of larger foraminifera provinces? Palaeogeography, Palaeoclimatology, Palaeoecology, 283 (3 - 4): 172 - 181." type="journal article" year="2009">Caus et al. (2009)</bibRefCitation>
; Vicedo et al., (2011) and Consorti et al., (2016b) from
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;
<bibRefCitation id="EFDD4B51FF90A25AFADBFD9AA6B8C5CE" author="Bachmann, M. &amp; Bassiouni, M. A. A. &amp; Kuss, J." pageId="73" pageNumber="112" pagination="131 - 162" refId="ref73048" refString="Bachmann, M., Bassiouni, M. A. A. &amp; Kuss, J., 2003. Timing of mid-Cretaceous carbonate platform depositional cycles, northern Sinai, Egypt. Palaeogeography, Palaeoclimatology, Palaeoecology, 200: 131 - 162." type="journal article" year="2003">Bachmann et al. (2003)</bibRefCitation>
from Sinai; and
<bibRefCitation id="EFDD4B51FF90A25AFB21FDBBA0D1C5CE" author="Ilavsky, J. &amp; Salaj, J." box="[1191,1436,621,645]" pageId="73" pageNumber="112" pagination="17 - 24" refId="ref84778" refString="Ilavsky, J. &amp; Salaj, J., 1969. Microbiostratigraphie du Cretace de la region de Kasserine, Tunisie Centrale. Notes du Service Geologique de Tunisie, 30: 17 - 24." type="journal article" year="1969">Ilavsky &amp; Salaj (1969)</bibRefCitation>
from
<collectingCountry id="F35B7630FF90A25AFCEBFD58A68DC5ED" box="[877,960,654,678]" name="Tunisia" pageId="73" pageNumber="112">Tunisia</collectingCountry>
. Although not illustrated, the middle Cenomanian record from
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of
<bibRefCitation id="EFDD4B51FF90A25AFB36FD79A030C58C" author="Saint-Marc, P." box="[1200,1405,687,711]" pageId="73" pageNumber="112" pagination="103 - 131" refId="ref96200" refString="Saint-Marc, P., 1981. Lebanon. In: Reyment, R. A. &amp; Bengtson, P. (eds.) Aspects of Mid-Cretaceous Regional Geology. IGCP Projects 58, Academic Press, pp. 103 - 131." type="book chapter" year="1981">Saint-Marc (1981)</bibRefCitation>
is important as the co-occurrence with the ammonite
<taxonomicName id="4C4C4D23FF90A25AFAE4FD07A124C441" authority="(Brongniart)" baseAuthorityName="Brongniart" baseAuthorityYear="1822" class="Cephalopoda" family="Acanthoceratidae" genus="Calycoceras" kingdom="Animalia" pageId="73" pageNumber="112" phylum="Mollusca" rank="species" species="gentoni">
<emphasis id="B938EAB2FF90A25AFAE4FD07A691C441" italics="true" pageId="73" pageNumber="112">Calycoceras gentoni</emphasis>
(Brongniart)
</taxonomicName>
supports a middle Cenomanian age calibration.
</paragraph>
</subSubSection>
<subSubSection id="C356652BFF90A25AFCB6FC83A04DC4E4" pageId="73" pageNumber="112" type="distribution">
<paragraph id="8BF336A0FF90A25AFCB6FC83A1BAC426" blockId="73.[816,1271,853,877]" box="[816,1271,853,877]" pageId="73" pageNumber="112">Cenomanian Paleogeographic Distribution</paragraph>
<paragraph id="8BF336A0FF90A25AFCB6FC41A04DC4E4" blockId="73.[816,1437,919,1439]" box="[816,1280,919,943]" pageId="73" pageNumber="112">
Caribbean Neotethys (and?
<collectingCountry id="F35B7630FF90A25AFBEAFC41A1B8C4E4" box="[1132,1269,919,943]" name="Afghanistan" pageId="73" pageNumber="112">Afghanistan</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C356652BFF90A25AFCB6FC6FA680C2D4" pageId="73" pageNumber="112" type="description">
<paragraph id="8BF336A0FF90A25AFCB6FC6FA125C3DC" blockId="73.[816,1437,919,1439]" pageId="73" pageNumber="112">
In addition to the records above, the species as been recorded from other parts of southern
<collectingCountry id="F35B7630FF90A25AFB2FFC0FA197C4BA" box="[1193,1242,985,1009]" name="Italy" pageId="73" pageNumber="112">Italy</collectingCountry>
(
<bibRefCitation id="EFDD4B51FF90A25AFB6EFC0FA6BEC359" author="Luperto-Sinni, E. &amp; Borgomano, J." pageId="73" pageNumber="112" pagination="95 - 136" refId="ref88178" refString="Luperto-Sinni, E. &amp; Borgomano, J., 1989. Le Cretace superieur des Murges sud-orientales (Italie meridionale): stratigraphie et evolution des paleoenvironnements. Rivista Italiana di Paleontologia e Stratigrafia, 95: 95 - 136." type="journal article" year="1989">Luperto-Sinni &amp; Borgomano, 1989</bibRefCitation>
(late Cenomanian); Borghi &amp; Pignatti, 2006 (late Cenomanian); Di Stefano &amp; Ruberti, 2000, all without illustration) and the Iranian Zagros (
<bibRefCitation id="EFDD4B51FF90A25AFA96FBEAA6DCC33E" author="Jamalpour, M. &amp; Hamdi, B. &amp; Armoon, A." pageId="73" pageNumber="112" pagination="101 - 110" refId="ref85076" refString="Jamalpour, M., Hamdi, B. &amp; Armoon, A., 2018. Lithostratigraphy and biostratigraphy of well numbers 9 and 17 in Binak Oilfield in the southwest of Iran. Journal of the Palaeontological Society of India, 63 (1): 101 - 110." type="journal article" year="2018">Jamalpour et al., 2018</bibRefCitation>
(illustrated), (but not
<bibRefCitation id="EFDD4B51FF90A25AFB08FB8BA02DC33E" author="Omidi, R. &amp; Sadeghi, A. &amp; Hosseini &amp; Barzi, M. A. &amp; Akbar Bas Kelayeh, N." box="[1166,1376,1117,1141]" pageId="73" pageNumber="112" pagination="75 - 92" refId="ref91903" refString="Omidi, R., Sadeghi, A., Hosseini, Barzi, M. A. &amp; Akbar Bas Kelayeh, N. 2018. Biostratigraphy of the Sarvak and Ilam formations in Ab-Teymour Oil Field (Wells no. 1 and 14), Iranian Journal of Geology, 12 (46): 75 - 92. [in Persian]." type="journal article" year="2018">Omidi et al., 2018</bibRefCitation>
;
<bibRefCitation id="EFDD4B51FF90A25AFAE8FB88A6A2C3DD" author="Mohajer, M. D. &amp; Afghah, M. &amp; Dehghanian, M. &amp; Zakariaii, S. J. S." pageId="73" pageNumber="112" pagination="1647 - 1667" refId="ref90286" refString="Mohajer, M. D., Afghah, M., Dehghanian, M., &amp; Zakariaii, S. J. S., 2021 a. Biostratigraphy, Microfacies and Depositional Environment of the Sarvak Formation at Pyun Anticline (Zagros Basin, Southwest of Iran). Acta Geologica Sinica, 95 (5): 1647 - 1667." type="journal article" year="2021">Mohajer et al., 2021a</bibRefCitation>
,
<bibRefCitation id="EFDD4B51FF90A25AFC7AFBA9A10DC3DD" author="Mohajer, M. D. &amp; Afghah, M. &amp; Dehghanian, M. &amp; Zakariaii, S. J. S." box="[1020,1088,1151,1175]" pageId="73" pageNumber="112" pagination="877 - 902" refId="ref90404" refString="Mohajer, M. D., Afghah, M., Dehghanian, M., &amp; Zakariaii, S. J. S., 2022 a. Biozonation of Foraminifera and Oligosteginids of Late Albian-Middle Turonian at south flank of Khumi Anticline (East of Izeh Zone). Advanced Applications in Geology, 11 (4): 877 - 902." type="journal article" year="2022">2022a</bibRefCitation>
, b).
</paragraph>
<paragraph id="8BF336A0FF90A25AFCB6FB76A680C2D4" blockId="73.[816,1437,919,1439]" pageId="73" pageNumber="112">
The records from
<collectingCountry id="F35B7630FF90A25AFC71FB76A10FC3F3" box="[1015,1090,1184,1208]" name="Greece" pageId="73" pageNumber="112">Greece</collectingCountry>
(
<bibRefCitation id="EFDD4B51FF90A25AFBD5FB76A192C3F3" author="Fleury, J. - J." box="[1107,1247,1184,1208]" pageId="73" pageNumber="112" pagination="479 - 651" refId="ref81096" refString="Fleury, J. - J., 1980. Les zones de Gavrovo-Tripolitza et de Pinde-Olonos (Grece continentale et Peloponnese du Nord); Evolution d'une plate- forme et d'un basin dans leur cadre alpin. Societe Geologique du Nord, 4: 479 - 651." type="journal article" year="1980">Fleury, 1980</bibRefCitation>
(also 1971, unillustrated) = possible
<taxonomicName id="4C4C4D23FF90A25AFB9CFB17A06CC392" box="[1050,1313,1217,1241]" class="Tubothalamea" family="Soritidae" genus="Pseudorhapydionina" kingdom="Chromista" order="Miliolida" pageId="73" pageNumber="112" phylum="Foraminifera" rank="species" species="undetermined">
<emphasis id="B938EAB2FF90A25AFB9CFB17A06CC392" box="[1050,1313,1217,1241]" italics="true" pageId="73" pageNumber="112">Pseudorhapydionina sp.</emphasis>
</taxonomicName>
) and
<collectingCountry id="F35B7630FF90A25AFAE2FB17A0D1C392" box="[1380,1436,1217,1241]" name="Syria" pageId="73" pageNumber="112">Syria</collectingCountry>
(
<bibRefCitation id="EFDD4B51FF90A25AFCBEFB34A175C3B1" author="Ghanem, H. &amp; Kuss, J." box="[824,1080,1250,1274]" pageId="73" pageNumber="112" pagination="85 - 132" refId="ref82188" refString="Ghanem, H. &amp; Kuss, J., 2013. Stratigraphic control of the Aptian - Early Turonian sequences of the Levant Platform, Coastal Range, northwest Syria. GeoArabia, 18 (4): 85 - 132." type="journal article" year="2013">Ghanem &amp; Kuss, 2013</bibRefCitation>
= possible peneropolid, but not
<taxonomicName id="4C4C4D23FF90A25AFCB6FAD2A6D0C250" authorityName="De Castro" authorityYear="1965" box="[816,925,1284,1307]" class="Globothalamea" family="Charentiidae" genus="Peneroplis" kingdom="Chromista" order="Loftusiida" pageId="73" pageNumber="112" phylum="Foraminifera" rank="species" species="parvus">
<emphasis id="B938EAB2FF90A25AFCB6FAD2A6D0C250" box="[816,925,1284,1307]" italics="true" pageId="73" pageNumber="112">P. parvus</emphasis>
</taxonomicName>
) have illustrations which are not compatible with this species, whilst those from southern
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(
<bibRefCitation id="EFDD4B51FF90A25AFCBEFA93A6B3C216" author="Tasli, K. &amp; Ozer, E. &amp; Koc, H." box="[824,1022,1349,1373]" pageId="73" pageNumber="112" pagination="521 - 533" refId="ref100610" refString="Tasli, K., Ozer, E. &amp; Koc, H., 2006. Benthic foraminiferal assemblages of the Cretaceous platform carbonate succession in the Yavca area (Bolkar Mountains, S Turkey): biostratigraphy and paleoenvironments. Geobios, 39: 521 - 533." type="journal article" year="2006">Tasli et al., 2006</bibRefCitation>
;
<bibRefCitation id="EFDD4B51FF90A25AFB8BFA93A188C216" author="Sari, B. &amp; Tasli, K. &amp; Ozer, S." box="[1037,1221,1349,1373]" pageId="73" pageNumber="112" pagination="393 - 425" refId="ref96523" refString="Sari, B., Tasli, K. &amp; Ozer, S., 2009. Benthonic Foraminiferal Biostratigraphy of the Upper Cretaceous (Middle Cenomanian-Coniacian) Sequences of the Bey Daglari Carbonate Platform, Western Taurides, Turkey. Turkish Journal of Earth Sciences, 18: 393 - 425." type="journal article" year="2009">Sari et al., 2009</bibRefCitation>
) and southern
<collectingCountry id="F35B7630FF90A25AFAF4FA90A0D0C216" box="[1394,1437,1350,1373]" name="Iraq" pageId="73" pageNumber="112">Iraq</collectingCountry>
(
<bibRefCitation id="EFDD4B51FF90A25AFCBEFAB0A117C235" author="Al-Salihi, A. &amp; Ibrahim, Y. K." box="[824,1114,1382,1406]" pageId="73" pageNumber="112" pagination="196 - 207" refId="ref71294" refString="Al-Salihi, A. &amp; Ibrahim, Y. K., 2023. Larger Foraminifera of Middle and Late Cenomanian (Biostrome Association), Southern Iraq. Iraqi Geological Journal, 56 (1 A): 196 - 207." type="journal article" year="2023">Al-Salihi &amp; Ibrahim, 2023</bibRefCitation>
) are tentative or unconfirmed by illustration.
</paragraph>
</subSubSection>
</treatment>
</document>