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164 lines
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<document ID-DOI="10.11646/zootaxa.5264.1.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="7836172" ID-ZooBank="47E141C8-F69A-422E-B7A7-E1AF4AD201BE" approvalRequired="6" approvalRequired_for_taxonomicNames="3" approvalRequired_for_textStreams="3" checkinTime="1681726002599" checkinUser="plazi" docAuthor="Nozaki, Tsubasa" docDate="2023" docId="03AB87F4FFBF8B26FF21FCFBFA653552" docLanguage="en" docName="zootaxa.5264.1.4.pdf" docOrigin="Zootaxa 5264 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5264.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Andrikothelyna Pace 2000" docType="key" docVersion="2" lastPageNumber="75" masterDocId="FF92FF8CFFB48B2DFFB6FFCEFFF93055" masterDocTitle="Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan" masterLastPageNumber="76" masterPageNumber="64" pageNumber="75" updateTime="1681726413705" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Nozaki, Tsubasa</mods:namePart>
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<mods:affiliation>Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka 819 - 0395, Japan & The Kyushu University Museum, Fukuoka 812 - 8581, Japan</mods:affiliation>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2023</mods:date>
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<mods:number>2023-04-12</mods:number>
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<mods:number>5264</mods:number>
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<mods:identifier type="DOI">10.11646/zootaxa.5264.1.4</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:03AB87F4FFBF8B26FF21FCFBFA653552" httpUri="http://treatment.plazi.org/id/03AB87F4FFBF8B26FF21FCFBFA653552" lastPageNumber="75" pageId="11" pageNumber="75">
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Key to species of
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<taxonomicName ID-CoL="8XJKK" authorityName="Pace" authorityYear="2000" box="[366,555,820,847]" class="Insecta" family="Staphylinidae" genus="Andrikothelyna" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="75" phylum="Arthropoda" rank="genus">
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<emphasis bold="true" box="[366,555,820,847]" italics="true" pageId="11" pageNumber="75">Andrikothelyna</emphasis>
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</emphasis>
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</heading>
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<keyLead pageId="11" pageNumber="75">
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1 Pronotum color reddish yellow to orange; elytra with low of granules along suture and carina near lateral margin weak (
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<figureCitation captionStart="FIGURES 1–8" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[214,1377,193,1897]" captionTargetId="figure-32@3.[206,1381,181,1920]" captionTargetPageId="3" captionText="FIGURES 1–8. Habitus of Andrikothelyna spp. 1, 5―Holotype of Andrikothelyna rubiginosa sp. nov., female; 2, 6―Holotype of Andrikothelyna limbata sp. nov., male (Kyushu); 3, 7―Andrikothelyna limbata sp. nov. male (Ishigaki Is.); 4, 8―Holotype of Andrikothelyna naomichii sp. nov. male." figureDoi="http://doi.org/10.5281/zenodo.7836176" httpUri="https://zenodo.org/record/7836176/files/figure.png" pageId="11" pageNumber="75">Fig. 8</figureCitation>
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)................................................................................................. 2
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</keyLead>
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</paragraph>
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<paragraph blockId="11.[151,1437,884,1287]" pageId="11" pageNumber="75">
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<keyLead pageId="11" pageNumber="75">
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- Pronotum color reddish brown to black; elytra with low of granules along suture and carina near lateral margin distinct (
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<figureCitation captionStart="FIGURES 1–8" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[214,1377,193,1897]" captionTargetId="figure-32@3.[206,1381,181,1920]" captionTargetPageId="3" captionText="FIGURES 1–8. Habitus of Andrikothelyna spp. 1, 5―Holotype of Andrikothelyna rubiginosa sp. nov., female; 2, 6―Holotype of Andrikothelyna limbata sp. nov., male (Kyushu); 3, 7―Andrikothelyna limbata sp. nov. male (Ishigaki Is.); 4, 8―Holotype of Andrikothelyna naomichii sp. nov. male." figureDoi="http://doi.org/10.5281/zenodo.7836176" httpUri="https://zenodo.org/record/7836176/files/figure.png" pageId="11" pageNumber="75">Figs 5–7</figureCitation>
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)............................................................................................... 3
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep pageId="11" pageNumber="75">
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<paragraph blockId="11.[151,1437,884,1287]" pageId="11" pageNumber="75">
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<keyLead pageId="11" pageNumber="75">
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2 Pronotum having posterior medial depression (fig.
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<bibRefCitation author="Pace, R." box="[773,876,1002,1023]" pageId="11" pageNumber="75" pagination="109 - 163" refId="ref7639" refString="Pace, R. (2000) Aleocharinae di Papua-Nuova Guinea (Coleoptera, Staphylinidae). Staphylinidae). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Entomologie 70, 109 - 163." type="journal article" year="2000">Pace 2000</bibRefCitation>
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); abdominal tergite VIII having two lateral large teeth and two medial short teeth at posterior margin (fig.
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<bibRefCitation author="Pace, R." box="[735,837,1031,1053]" pageId="11" pageNumber="75" pagination="109 - 163" refId="ref7639" refString="Pace, R. (2000) Aleocharinae di Papua-Nuova Guinea (Coleoptera, Staphylinidae). Staphylinidae). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Entomologie 70, 109 - 163." type="journal article" year="2000">Pace 2000</bibRefCitation>
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)......................................
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<taxonomicName authorityName="Pace" authorityYear="2000" box="[1326,1436,1031,1053]" class="Insecta" family="Staphylinidae" genus="Andrikothelyna" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="75" phylum="Arthropoda" rank="species" species="papuana">
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<emphasis box="[1326,1341,1031,1052]" italics="true" pageId="11" pageNumber="75">A</emphasis>
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.
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<emphasis box="[1352,1436,1032,1053]" italics="true" pageId="11" pageNumber="75">papuana</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph blockId="11.[151,1437,884,1287]" pageId="11" pageNumber="75">
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<keyLead pageId="11" pageNumber="75">
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- Pronotum without posterior medial depression (
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<figureCitation box="[693,783,1060,1082]" captionStart="FIGURES 1–8" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[214,1377,193,1897]" captionTargetId="figure-32@3.[206,1381,181,1920]" captionTargetPageId="3" captionText="FIGURES 1–8. Habitus of Andrikothelyna spp. 1, 5―Holotype of Andrikothelyna rubiginosa sp. nov., female; 2, 6―Holotype of Andrikothelyna limbata sp. nov., male (Kyushu); 3, 7―Andrikothelyna limbata sp. nov. male (Ishigaki Is.); 4, 8―Holotype of Andrikothelyna naomichii sp. nov. male." figureDoi="http://doi.org/10.5281/zenodo.7836176" httpUri="https://zenodo.org/record/7836176/files/figure.png" pageId="11" pageNumber="75">Figs 4, 8</figureCitation>
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); abdominal tergite VIII having four to six short teeth at posterior margin (
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<figureCitation box="[307,376,1090,1112]" captionStart="FIGURES 31–36" captionStartId="10.[151,264,1933,1957]" captionTargetBox="[180,1409,189,1896]" captionTargetId="figure-18@10.[167,1420,181,1909]" captionTargetPageId="10" captionText="FIGURES 31–36. Andrikothelyna naomichii sp. nov. 31―abdominal tergite VIII, male; 32―abdominal sternite VIII male; 33―abdominal sternite VIII female; 34―median lobe of male aedeagus; 35―Paramere of male aedeagus; 36―female spermatheca." figureDoi="http://doi.org/10.5281/zenodo.7836195" httpUri="https://zenodo.org/record/7836195/files/figure.png" pageId="11" pageNumber="75">Fig. 31</figureCitation>
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).....................................................................
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<taxonomicName authority="Nozaki, 2023" authorityName="Nozaki" authorityYear="2023" box="[1231,1354,1090,1111]" class="Insecta" family="Staphylinidae" genus="Andrikothelyna" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="75" phylum="Arthropoda" rank="species" species="naomichii" status="sp. nov.">
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<emphasis box="[1231,1246,1090,1111]" italics="true" pageId="11" pageNumber="75">A</emphasis>
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.
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<emphasis box="[1258,1354,1090,1111]" italics="true" pageId="11" pageNumber="75">naomichii</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[1360,1436,1090,1111]" pageId="11" pageNumber="75">
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<taxonomicNameLabel box="[1360,1436,1090,1111]" pageId="11" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</keyLead>
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</paragraph>
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<keyLead box="[151,1436,1119,1141]" pageId="11" pageNumber="75">
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3 Posterior margin of each elytron having a pair of yellow semicircular maculation (
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<figureCitation box="[1015,1100,1119,1141]" captionStart="FIGURES 1–8" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[214,1377,193,1897]" captionTargetId="figure-32@3.[206,1381,181,1920]" captionTargetPageId="3" captionText="FIGURES 1–8. Habitus of Andrikothelyna spp. 1, 5―Holotype of Andrikothelyna rubiginosa sp. nov., female; 2, 6―Holotype of Andrikothelyna limbata sp. nov., male (Kyushu); 3, 7―Andrikothelyna limbata sp. nov. male (Ishigaki Is.); 4, 8―Holotype of Andrikothelyna naomichii sp. nov. male." figureDoi="http://doi.org/10.5281/zenodo.7836176" httpUri="https://zenodo.org/record/7836176/files/figure.png" pageId="11" pageNumber="75">Figs 2–3</figureCitation>
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)..........
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<taxonomicName authority="Nozaki, 2023" authorityName="Nozaki" authorityYear="2023" box="[1254,1353,1119,1141]" class="Insecta" family="Staphylinidae" genus="Andrikothelyna" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="75" phylum="Arthropoda" rank="species" species="limbata" status="sp. nov.">
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<emphasis box="[1254,1269,1119,1140]" italics="true" pageId="11" pageNumber="75">A</emphasis>
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.
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<emphasis box="[1280,1353,1119,1140]" italics="true" pageId="11" pageNumber="75">limbata</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[1360,1436,1119,1140]" pageId="11" pageNumber="75">
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<taxonomicNameLabel box="[1360,1436,1119,1140]" pageId="11" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</keyLead>
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</paragraph>
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<paragraph blockId="11.[151,1437,884,1287]" box="[151,1436,1148,1170]" pageId="11" pageNumber="75">
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<keyLead box="[151,1436,1148,1170]" pageId="11" pageNumber="75">- Posterior margin of each elytron without yellow semicircular maculation......................................... 3</keyLead>
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4 Abdominal tergite VIII having six teeth (four major teeth and two tiny teeth) at posterior margin (
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<figureCitation box="[1162,1232,1178,1200]" captionStart="FIGURES 16–19" captionStartId="6.[151,264,1900,1924]" captionTargetBox="[197,1383,622,1849]" captionTargetId="figure-191@6.[151,1436,597,1877]" captionTargetPageId="6" captionText="FIGURES 16–19. Andrikothelyna rubiginosa sp. nov. female. 16―Meso- and metaventrite; 17―abdominal tergite VIII; 18― abdominal sternite VIII; 19―spermatheca." figureDoi="http://doi.org/10.5281/zenodo.7836182" httpUri="https://zenodo.org/record/7836182/files/figure.png" pageId="11" pageNumber="75">Fig. 17</figureCitation>
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).................................................................................................
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<taxonomicName authority="Nozaki, 2023" authorityName="Nozaki" authorityYear="2023" box="[1223,1353,1207,1229]" class="Insecta" family="Staphylinidae" genus="Andrikothelyna" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="75" phylum="Arthropoda" rank="species" species="rubiginosa" status="sp. nov.">
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<emphasis box="[1223,1238,1207,1228]" italics="true" pageId="11" pageNumber="75">A</emphasis>
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.
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<emphasis box="[1249,1353,1207,1228]" italics="true" pageId="11" pageNumber="75">rubiginosa</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[1360,1436,1207,1228]" pageId="11" pageNumber="75">
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<taxonomicNameLabel box="[1360,1436,1207,1228]" pageId="11" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</keyLead>
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</paragraph>
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<paragraph blockId="11.[151,1437,884,1287]" pageId="11" pageNumber="75">
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<keyLead pageId="11" pageNumber="75">
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- Abdominal tergite VIII having four teeth at posterior margin, two medial teeth very short (fig.
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<bibRefCitation author="Pace, R." box="[1188,1288,1237,1258]" pageId="11" pageNumber="75" pagination="33 - 63" refId="ref7746" refString="Pace, R. (2013) New distributional data, New Species and Three New Genera of Aleocharinae from Malaysia, Vietnam and Taiwan (Coleoptera: Staphylinidae). Tropical Zoology, 26 (1), 33 - 63. https: // doi. org / 10.1080 / 03946975.2013.775832" type="journal article" year="2013">Pace 2013</bibRefCitation>
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).............................................................................................
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<taxonomicName authorityName="Nozaki" authorityYear="2023" baseAuthorityName="Pace" baseAuthorityYear="2013" box="[1220,1320,1266,1287]" class="Insecta" family="Staphylinidae" genus="Andrikothelyna" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="75" phylum="Arthropoda" rank="species" species="orientis" status="comb. nov.">
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<emphasis box="[1220,1235,1266,1287]" italics="true" pageId="11" pageNumber="75">A</emphasis>
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.
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<emphasis box="[1247,1320,1266,1287]" italics="true" pageId="11" pageNumber="75">orientis</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[1326,1436,1266,1287]" pageId="11" pageNumber="75">
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<taxonomicNameLabel box="[1326,1436,1266,1287]" pageId="11" pageNumber="75" rank="species">comb. nov.</taxonomicNameLabel>
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</emphasis>
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</keyLead>
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</paragraph>
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</treatment>
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