treatments-xml/data/2C/B2/61/2CB2612F12885FC0BB61101462F92B7F.xml

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<mods:title id="7CAE6DF09E0A9CA54FA6B03E44D3F87C">Systematic revision of the snorkel snail genus Rhiostoma Benson, 1860 (Gastropoda, Caenogastropoda, Cyclophoridae) with descriptions of new species</mods:title>
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<mods:namePart id="0911853F9F975F825BCD702F2563FB76">Tongkerd, Piyoros</mods:namePart>
<mods:affiliation id="715E42CF056D856370588EE7C1D745F2">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="0FAB4D015FD7D6D9560B629068A0442A">Tumpeesuwan, Sakboworn</mods:namePart>
<mods:affiliation id="1A0925C8FAEFD93D81CEC704D9E9628A">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand &amp; Present address: Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai, Maha Sarakham 44150 Thailand</mods:affiliation>
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<mods:namePart id="2B0A8E01D72E9202B8E76EC39601F342">Inkhavilay, Khamla</mods:namePart>
<mods:affiliation id="0EEEF5941FAD973B8D4AB4D2F63F33F1">Department of Biology, Faculty of Natural Science, National University of Laos, P. O. Box 7322, Dongdok, Vientiane, Laos</mods:affiliation>
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<mods:namePart id="243AE129D3FC0DE957CE6C3F15C1DF3F">Prasankok, Pongpun</mods:namePart>
<mods:affiliation id="5195E4478B378C2B112634B5C9168AB0">School of Biology, Institute of Science, Suranaree University of Technology, Nakhon Ratchasima 30000, Thailand</mods:affiliation>
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<mods:namePart id="F9E7D9F47F99EA843152F07BAFE046E8">Jeratthitikul, Ekgachai</mods:namePart>
<mods:affiliation id="C4F8D2BCBED38CC44890732D4C43142D">Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
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<mods:namePart id="7167D95D868E88BACFF4FDBD3661D088">Panha, Somsak</mods:namePart>
<mods:affiliation id="21AA8CF3A6E42E6338C735DFD41474BA">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="937AACDFC223680DD672F1FEC881B2DE">Sutcharit, Chirasak</mods:namePart>
<mods:affiliation id="7122A5509E5021AADFB448EFBC30673E">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<taxonomicName id="BB0B7736A9932E6763D46BAC705620C1" ID-CoL="B3CKH" LSID="2CB2612F-1288-5FC0-BB61-101462F92B7F" authority="(Thach &amp; Huber, 2020)" baseAuthorityName="Thach &amp; Huber" baseAuthorityYear="2020" class="Gastropoda" family="Cyclophoridae" genus="Opisthoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisthoporus battambangensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="battambangensis">Opisthoporus battambangensis (Thach &amp; Huber, 2020)</taxonomicName>
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<figureCitation id="149082C15855F03EDB5DA8A59E01E95A" captionStart="Figure 57" captionStartId="F57" captionText="Figure 57. A Pterocyclos subalatus Sykes, 1903, holotype UMZC 1032 from Gunong Inas, Perak, Malay Peninsula B Pterocyclos spelaeotes (Tomlin, 1931), lectotype NMW 1955.158.01107 from Batu Cave, Kula Lumpur, Selangor C Pterocyclos umbraticus (Benthem Jutting, 1949), holotype ZMA 135622 from Maxwell's Hill, Perak, Malaysia D Cyclotus boxalli syntype NHMUK 1894.5.23.1 from Kina Balu E Opisthoporus battambangensis holotype MNHN-IM- 2000 - 35506 from Battambang, Cambodia F Opisthoporus grohi holotype MNHN-IM- 2000 - 35507 from Dak Nong, Vietnam. Photograph: H Wood, NMW (B), M Caballer, A Lardeur and V Heros, MNHN (E, F)." figureDoi="10.3897/zookeys.1142.90097.figure57" httpUri="https://binary.pensoft.net/fig/799915" pageId="0" pageNumber="1">Fig. 57E</figureCitation>
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<taxonomicName id="990853FCC7870D6391B8673A65A701E8" authorityName="Thach &amp; F. Huber" authorityYear="2020" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma battambangensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="battambangensis">Rhiostoma battambangensis</taxonomicName>
Thach &amp; Huber in Thach, 2020: 18, figs 153, 154. Type locality: Battambang City, Battambang Province, Cambodia.
</paragraph>
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<paragraph id="C0E23564427364DC409DC7A88ADEC86F" pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph id="9877C3B8F08D0E8DE568459A6EC90B1E" pageId="0" pageNumber="1">
<emphasis id="0E2A14B5A7315B1B293F4BB7D1402E2E" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
MNHN-IM-2000-35506 (Fig. 
<figureCitation id="71BF6217E4D182D0DE4C8900D96270D1" captionStart="Figure 57" captionStartId="F57" captionText="Figure 57. A Pterocyclos subalatus Sykes, 1903, holotype UMZC 1032 from Gunong Inas, Perak, Malay Peninsula B Pterocyclos spelaeotes (Tomlin, 1931), lectotype NMW 1955.158.01107 from Batu Cave, Kula Lumpur, Selangor C Pterocyclos umbraticus (Benthem Jutting, 1949), holotype ZMA 135622 from Maxwell's Hill, Perak, Malaysia D Cyclotus boxalli syntype NHMUK 1894.5.23.1 from Kina Balu E Opisthoporus battambangensis holotype MNHN-IM- 2000 - 35506 from Battambang, Cambodia F Opisthoporus grohi holotype MNHN-IM- 2000 - 35507 from Dak Nong, Vietnam. Photograph: H Wood, NMW (B), M Caballer, A Lardeur and V Heros, MNHN (E, F)." figureDoi="10.3897/zookeys.1142.90097.figure57" httpUri="https://binary.pensoft.net/fig/799915" pageId="0" pageNumber="1">57E</figureCitation>
) from Battambang City, Battambang Province.
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<paragraph id="F3ED41D40D062076D317299764CC9306" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="6CB21ADEA85E06040E9EEF6A095A2B47" pageId="0" pageNumber="1">
This species was described based on the holotype, and by three paratypes housed in the 
<normalizedToken id="D65D655B0CC356397DA2FB4C6B833001" originalValue="authors’">authors'</normalizedToken>
collections. The operculum is attached with the holotype and has a calcareous, multispiral, and plate-shaped operculum; the shell has a short detached whorl and yellowish brown periostracum. The outer peristome is slightly expanded with short tubular breathing device located on the suture area of the detached whorl, and the inner peristome has a shallow incision. These are the distinguishing characters of 
<taxonomicName id="A0501432D3673C9DCFAF5BAA8FAC2686" authorityName="Benson in Pfeiffer" authorityYear="1851" class="Gastropoda" family="Cyclophoridae" genus="Opisthoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisthoporus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="DA1A09AF9696061FE69C1988CF6E7232" italics="true" pageId="0" pageNumber="1">Opisthoporus</emphasis>
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; therefore, we have relocated this species into the genus 
<taxonomicName id="1B234E10673952D8A317EBDCA9E4F4D1" authorityName="Benson in Pfeiffer" authorityYear="1851" class="Gastropoda" family="Cyclophoridae" genus="Opisthoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisthoporus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="BE31F8CFF21602D2FA9FA8A95D12A417" italics="true" pageId="0" pageNumber="1">Opisthoporus</emphasis>
</taxonomicName>
.
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