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<document id="F7DD22939640E818B0EE8874E1C5BFDA" ID-DOI="10.37520/aemnp.2021.029" ID-ISSN="1804-6487" ID-Zenodo-Dep="5821215" ID-ZooBank="70C7FD52-9D69-4401-ADE8-0D043D39BB2C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1641390478763" checkinUser="plazi" docAuthor="Zettel, Herbert &amp; Laciny, Alice" docDate="2021" docId="D02C87B81F79FF9F74C50CC1FB52A9B5" docLanguage="en" docName="ActaEntMusNatPra.61.2.517-527.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 61 (2)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Thirumalaia Zettel &amp; Laciny 2021, gen. nov." docType="treatment" docVersion="3" lastPageNumber="518" masterDocId="2C15FFC01F78FF9E77340C2CFFD2AF08" masterDocTitle="The taxonomy of some unusual Microveliinae (Hemiptera: Heteroptera: Veliidae) from India" masterLastPageNumber="527" masterPageNumber="517" pageNumber="518" updateTime="1699285328636" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="C6E66A81F96DBE1AB6075ADF5F9BA860">The taxonomy of some unusual Microveliinae (Hemiptera: Heteroptera: Veliidae) from India</mods:title>
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<mods:namePart id="E276ED1CA20B83BE0F607191C7481470">Zettel, Herbert</mods:namePart>
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<mods:namePart id="323510BF42B9450F300542FB4B8A5403">Laciny, Alice</mods:namePart>
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<mods:date id="EBDB4FEECE9519122090C2971A290021">2021</mods:date>
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<mods:number id="1DA2D92DDA35FBD6E87CF742BCA06A50">2021-12-31</mods:number>
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<mods:identifier id="5790E8521C2F0117745DBCE1BE8F3D0E" type="DOI">10.37520/aemnp.2021.029</mods:identifier>
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<treatment id="D02C87B81F79FF9F74C50CC1FB52A9B5" ID-DOI="http://doi.org/10.5281/zenodo.5821255" ID-Zenodo-Dep="5821255" LSID="urn:lsid:plazi:treatment:D02C87B81F79FF9F74C50CC1FB52A9B5" httpUri="http://treatment.plazi.org/id/D02C87B81F79FF9F74C50CC1FB52A9B5" lastPageNumber="518" pageId="1" pageNumber="518">
<subSubSection id="109F65251F79FF9F74C50CC1FB35AE0C" box="[1009,1255,237,260]" pageId="1" pageNumber="518" type="nomenclature">
<paragraph id="583A36AE1F79FF9F74C50CC1FB35AE0C" blockId="1.[1009,1255,237,260]" box="[1009,1255,237,260]" pageId="1" pageNumber="518">
<heading id="037281C21F79FF9F74C50CC1FB35AE0C" box="[1009,1255,237,260]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="518" reason="3">
<emphasis id="6AF1EABC1F79FF9F74C50CC1FB35AE0C" bold="true" box="[1009,1255,237,260]" pageId="1" pageNumber="518">
<taxonomicName id="9F854D2D1F79FF9F74C50CC1FBAEAE0C" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[1009,1148,237,260]" class="Insecta" family="Veliidae" genus="Thirumalaia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="6AF1EABC1F79FF9F74C50CC1FBAEAE0C" bold="true" box="[1009,1148,237,260]" italics="true" pageId="1" pageNumber="518">Thirumalaia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F79FF9F73B70CC1FB35AE0C" box="[1155,1255,237,260]" pageId="1" pageNumber="518" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="109F65251F79FF9F73110D27FB60AE16" box="[1061,1202,267,286]" pageId="1" pageNumber="518" type="description">
<paragraph id="583A36AE1F79FF9F73110D27FB60AE16" blockId="1.[1061,1202,267,286]" box="[1061,1202,267,286]" pageId="1" pageNumber="518">
(
<figureCitation id="C0BE2A2B1F79FF9F731F0D27FBAAAE16" box="[1067,1144,267,286]" captionStart="Figs 1–4" captionStartId="2.[151,188,1717,1736]" captionTargetBox="[151,1436,189,1698]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 1–4. Habitus of Thirumalaia ocularis gen. nov. &amp; sp. nov. (1 – apterous male, 2 – apterous female), and of Eyarinella robusta gen. nov. &amp; sp. nov. (3 – apterous male, 4 – apterous female)." figureDoi="http://doi.org/10.5281/zenodo.5821217" httpUri="https://zenodo.org/record/5821217/files/figure.png" pageId="1" pageNumber="518">Figs 1, 2</figureCitation>
,
<figureCitation id="C0BE2A2B1F79FF9F73B50D27FB7EAE16" box="[1153,1196,267,286]" captionStart="Figs 9–15" captionStartId="5.[151,188,930,949]" captionTargetBox="[150,1431,193,906]" captionTargetId="figure-588@5.[150,1435,189,908]" captionTargetPageId="5" captionText="Figs 9–15. Structural details of Thirumalaia ocularis gen. nov. &amp; sp. nov.: 9 – head, first antennomere, and pronotum, dorsal; 10 – foreleg of male (arrow points at proximal end of grasping comb); 11 – abdominal segments 7 and 8, and proctiger of female, dorsal; 12 – abdominal segment 8 of male, ventral; 13 – pygophore of male, ventral; 14 – left paramere, lateral; 15 – proctiger of male, dorsal. Pilosity omitted, except in Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.5821221" httpUri="https://zenodo.org/record/5821221/files/figure.png" pageId="1" pageNumber="518">9–15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="109F65251F79FF9F74080D1AFAC1AE45" box="[828,1299,310,333]" pageId="1" pageNumber="518" type="type_taxon">
<paragraph id="583A36AE1F79FF9F74080D1AFAC1AE45" blockId="1.[828,1299,310,333]" box="[828,1299,310,333]" pageId="1" pageNumber="518">
<emphasis id="6AF1EABC1F79FF9F74080D1AFC02AE45" bold="true" box="[828,976,310,333]" pageId="1" pageNumber="518">
<typeStatus id="873E880C1F79FF9F74080D1AFCA6AE45" box="[828,884,310,333]" pageId="1" pageNumber="518">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="9F854D2D1F79FF9F74E20D1AFB6BAE45" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[982,1209,310,333]" class="Insecta" family="Veliidae" genus="Thirumalaia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="species" species="ocularis" status="sp. nov.">
<emphasis id="6AF1EABC1F79FF9F74E20D1AFB6BAE45" box="[982,1209,310,333]" italics="true" pageId="1" pageNumber="518">Thirumalaia ocularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F79FF9F73F40D1AFAC1AE45" box="[1216,1299,310,333]" pageId="1" pageNumber="518" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="109F65251F79FF9F74080D4AFB52A9B5" pageId="1" pageNumber="518" type="diagnosis">
<paragraph id="583A36AE1F79FF9F74080D4AFBFBACF5" blockId="1.[828,1437,358,1725]" pageId="1" pageNumber="518">
<emphasis id="6AF1EABC1F79FF9F74080D4AFC63AE75" bold="true" box="[828,945,358,381]" pageId="1" pageNumber="518">Diagnosis.</emphasis>
Body of male moderately stout (
<figureCitation id="C0BE2A2B1F79FF9F721D0D4AFAB8AE75" box="[1321,1386,358,381]" captionStart="Figs 1–4" captionStartId="2.[151,188,1717,1736]" captionTargetBox="[151,1436,189,1698]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 1–4. Habitus of Thirumalaia ocularis gen. nov. &amp; sp. nov. (1 – apterous male, 2 – apterous female), and of Eyarinella robusta gen. nov. &amp; sp. nov. (3 – apterous male, 4 – apterous female)." figureDoi="http://doi.org/10.5281/zenodo.5821217" httpUri="https://zenodo.org/record/5821217/files/figure.png" pageId="1" pageNumber="518">Fig. 1</figureCitation>
), of female much stouter (
<figureCitation id="C0BE2A2B1F79FF9F731F0DAAFBBDAE95" box="[1067,1135,390,413]" captionStart="Figs 1–4" captionStartId="2.[151,188,1717,1736]" captionTargetBox="[151,1436,189,1698]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 1–4. Habitus of Thirumalaia ocularis gen. nov. &amp; sp. nov. (1 – apterous male, 2 – apterous female), and of Eyarinella robusta gen. nov. &amp; sp. nov. (3 – apterous male, 4 – apterous female)." figureDoi="http://doi.org/10.5281/zenodo.5821217" httpUri="https://zenodo.org/record/5821217/files/figure.png" pageId="1" pageNumber="518">Fig. 2</figureCitation>
). Postocular region of head deeply inserted in anterior part of pronotum (
<figureCitation id="C0BE2A2B1F79FF9F72730D8AFA5DAEB5" box="[1351,1423,422,445]" captionStart="Figs 9–15" captionStartId="5.[151,188,930,949]" captionTargetBox="[150,1431,193,906]" captionTargetId="figure-588@5.[150,1435,189,908]" captionTargetPageId="5" captionText="Figs 9–15. Structural details of Thirumalaia ocularis gen. nov. &amp; sp. nov.: 9 – head, first antennomere, and pronotum, dorsal; 10 – foreleg of male (arrow points at proximal end of grasping comb); 11 – abdominal segments 7 and 8, and proctiger of female, dorsal; 12 – abdominal segment 8 of male, ventral; 13 – pygophore of male, ventral; 14 – left paramere, lateral; 15 – proctiger of male, dorsal. Pilosity omitted, except in Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.5821221" httpUri="https://zenodo.org/record/5821221/files/figure.png" pageId="1" pageNumber="518">Fig. 9</figureCitation>
). Eyes very large, synthlipsis 0.28–0.34 (
<figureCitation id="C0BE2A2B1F79FF9F73DE0DEAFAFCAED5" box="[1258,1326,454,477]" captionStart="Figs 9–15" captionStartId="5.[151,188,930,949]" captionTargetBox="[150,1431,193,906]" captionTargetId="figure-588@5.[150,1435,189,908]" captionTargetPageId="5" captionText="Figs 9–15. Structural details of Thirumalaia ocularis gen. nov. &amp; sp. nov.: 9 – head, first antennomere, and pronotum, dorsal; 10 – foreleg of male (arrow points at proximal end of grasping comb); 11 – abdominal segments 7 and 8, and proctiger of female, dorsal; 12 – abdominal segment 8 of male, ventral; 13 – pygophore of male, ventral; 14 – left paramere, lateral; 15 – proctiger of male, dorsal. Pilosity omitted, except in Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.5821221" httpUri="https://zenodo.org/record/5821221/files/figure.png" pageId="1" pageNumber="518">Fig. 9</figureCitation>
). Antenna very long and slender, about two thirds of body length (
<figureCitation id="C0BE2A2B1F79FF9F74770E2AFC76AD15" box="[835,932,518,541]" captionStart="Figs 1–4" captionStartId="2.[151,188,1717,1736]" captionTargetBox="[151,1436,189,1698]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 1–4. Habitus of Thirumalaia ocularis gen. nov. &amp; sp. nov. (1 – apterous male, 2 – apterous female), and of Eyarinella robusta gen. nov. &amp; sp. nov. (3 – apterous male, 4 – apterous female)." figureDoi="http://doi.org/10.5281/zenodo.5821217" httpUri="https://zenodo.org/record/5821217/files/figure.png" pageId="1" pageNumber="518">Figs 1, 2</figureCitation>
); antennomere I approximately as long as head length (in male longer than in female); relative lengths of antennomeres: IV ≧ III&gt; I&gt; II. Pronotum convexly rounded posteriorly (
<figureCitation id="C0BE2A2B1F79FF9F732A0E4AFBB3AD75" box="[1054,1121,614,637]" captionStart="Figs 9–15" captionStartId="5.[151,188,930,949]" captionTargetBox="[150,1431,193,906]" captionTargetId="figure-588@5.[150,1435,189,908]" captionTargetPageId="5" captionText="Figs 9–15. Structural details of Thirumalaia ocularis gen. nov. &amp; sp. nov.: 9 – head, first antennomere, and pronotum, dorsal; 10 – foreleg of male (arrow points at proximal end of grasping comb); 11 – abdominal segments 7 and 8, and proctiger of female, dorsal; 12 – abdominal segment 8 of male, ventral; 13 – pygophore of male, ventral; 14 – left paramere, lateral; 15 – proctiger of male, dorsal. Pilosity omitted, except in Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.5821221" httpUri="https://zenodo.org/record/5821221/files/figure.png" pageId="1" pageNumber="518">Fig. 9</figureCitation>
). Lateral evaporatorium situated approximately in middle of metapleuron. Legs long and slender. Claws preterminal. – Male: Propleuron unmodified. Femora unmodified. Protibia unmodified; protibial grasping comb long, about one third of tibia length (
<figureCitation id="C0BE2A2B1F79FF9F74770F2AFC42AC15" box="[835,912,774,797]" captionStart="Figs 9–15" captionStartId="5.[151,188,930,949]" captionTargetBox="[150,1431,193,906]" captionTargetId="figure-588@5.[150,1435,189,908]" captionTargetPageId="5" captionText="Figs 9–15. Structural details of Thirumalaia ocularis gen. nov. &amp; sp. nov.: 9 – head, first antennomere, and pronotum, dorsal; 10 – foreleg of male (arrow points at proximal end of grasping comb); 11 – abdominal segments 7 and 8, and proctiger of female, dorsal; 12 – abdominal segment 8 of male, ventral; 13 – pygophore of male, ventral; 14 – left paramere, lateral; 15 – proctiger of male, dorsal. Pilosity omitted, except in Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.5821221" httpUri="https://zenodo.org/record/5821221/files/figure.png" pageId="1" pageNumber="518">Fig. 10</figureCitation>
). Mesotibia without grasping comb. Claws simple, minimally surpassing apex of tarsus. Abdomen without projections. Segment 8 elongated. Pygophore pear-shaped, narrow at base, with truncated posterior margin (
<figureCitation id="C0BE2A2B1F79FF9F72750F4AFA5DAC75" box="[1345,1423,870,893]" captionStart="Figs 9–15" captionStartId="5.[151,188,930,949]" captionTargetBox="[150,1431,193,906]" captionTargetId="figure-588@5.[150,1435,189,908]" captionTargetPageId="5" captionText="Figs 9–15. Structural details of Thirumalaia ocularis gen. nov. &amp; sp. nov.: 9 – head, first antennomere, and pronotum, dorsal; 10 – foreleg of male (arrow points at proximal end of grasping comb); 11 – abdominal segments 7 and 8, and proctiger of female, dorsal; 12 – abdominal segment 8 of male, ventral; 13 – pygophore of male, ventral; 14 – left paramere, lateral; 15 – proctiger of male, dorsal. Pilosity omitted, except in Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.5821221" httpUri="https://zenodo.org/record/5821221/files/figure.png" pageId="1" pageNumber="518">Fig. 13</figureCitation>
). Proctiger very slender (
<figureCitation id="C0BE2A2B1F79FF9F73030FAAFB55AC95" box="[1079,1159,902,925]" captionStart="Figs 9–15" captionStartId="5.[151,188,930,949]" captionTargetBox="[150,1431,193,906]" captionTargetId="figure-588@5.[150,1435,189,908]" captionTargetPageId="5" captionText="Figs 9–15. Structural details of Thirumalaia ocularis gen. nov. &amp; sp. nov.: 9 – head, first antennomere, and pronotum, dorsal; 10 – foreleg of male (arrow points at proximal end of grasping comb); 11 – abdominal segments 7 and 8, and proctiger of female, dorsal; 12 – abdominal segment 8 of male, ventral; 13 – pygophore of male, ventral; 14 – left paramere, lateral; 15 – proctiger of male, dorsal. Pilosity omitted, except in Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.5821221" httpUri="https://zenodo.org/record/5821221/files/figure.png" pageId="1" pageNumber="518">Fig. 15</figureCitation>
). Parameres symmetrical, medium-sized, of characteristic shape as shown in
<figureCitation id="C0BE2A2B1F79FF9F72630F8AFC88ACD5" captionStart="Figs 9–15" captionStartId="5.[151,188,930,949]" captionTargetBox="[150,1431,193,906]" captionTargetId="figure-588@5.[150,1435,189,908]" captionTargetPageId="5" captionText="Figs 9–15. Structural details of Thirumalaia ocularis gen. nov. &amp; sp. nov.: 9 – head, first antennomere, and pronotum, dorsal; 10 – foreleg of male (arrow points at proximal end of grasping comb); 11 – abdominal segments 7 and 8, and proctiger of female, dorsal; 12 – abdominal segment 8 of male, ventral; 13 – pygophore of male, ventral; 14 – left paramere, lateral; 15 – proctiger of male, dorsal. Pilosity omitted, except in Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.5821221" httpUri="https://zenodo.org/record/5821221/files/figure.png" pageId="1" pageNumber="518">Figure 14</figureCitation>
. – Female: Genitalia situated in a strictly posterior opening of segment 7.
</paragraph>
<paragraph id="583A36AE1F79FF9F7408082AFB52A9B5" blockId="1.[828,1437,358,1725]" pageId="1" pageNumber="518">
<emphasis id="6AF1EABC1F79FF9F7408082AFBCDAB15" bold="true" box="[828,1055,1030,1053]" pageId="1" pageNumber="518">Comparative notes.</emphasis>
<taxonomicName id="9F854D2D1F79FF9F731D082AFB60AB15" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[1065,1202,1030,1053]" class="Insecta" family="Veliidae" genus="Thirumalaia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="6AF1EABC1F79FF9F731D082AFB60AB15" box="[1065,1202,1030,1053]" italics="true" pageId="1" pageNumber="518">Thirumalaia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F79FF9F7389082AFAF7AB15" box="[1213,1317,1030,1053]" pageId="1" pageNumber="518" rank="genus">gen. nov.</taxonomicNameLabel>
resembles some species of
<taxonomicName id="9F854D2D1F79FF9F74CC080AFB21AB35" authority="Distant, 1903" authorityName="Distant" authorityYear="1903" box="[1016,1267,1062,1085]" class="Insecta" family="Veliidae" genus="Baptista" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="genus">
<emphasis id="6AF1EABC1F79FF9F74CC080AFB81AB35" box="[1016,1107,1062,1085]" italics="true" pageId="1" pageNumber="518">Baptista</emphasis>
<bibRefCitation id="3C144B5F1F79FF9F7369080AFB21AB35" author="DISTANT W. L." box="[1117,1267,1062,1085]" pageId="1" pageNumber="518" pagination="219 - 274" refId="ref7875" refString="DISTANT W. L. 1903: Report on the Rhynchota. Part I. Heteroptera. Fascicula Malayenses Zoology 2: 219 - 274." type="journal article" year="1903">
Distant,
<quantity id="9F7D9B4B1F79FF9F7389080AFB21AB35" box="[1213,1267,1062,1085]" metricMagnitude="1" metricUnit="m" metricValue="4.83362" pageId="1" pageNumber="518" unit="in" value="1903.0">1903</quantity>
</bibRefCitation>
</taxonomicName>
in habitus and shares with this genus some other characters like the long protibial grasping comb, structure of antenna, and a concave anterior margin of the pronotum. Most species of
<taxonomicName id="9F854D2D1F79FF9F74B9088AFC3AABB5" authorityName="Distant" authorityYear="1903" box="[909,1000,1190,1213]" class="Insecta" family="Veliidae" genus="Baptista" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="genus">
<emphasis id="6AF1EABC1F79FF9F74B9088AFC3AABB5" box="[909,1000,1190,1213]" italics="true" pageId="1" pageNumber="518">Baptista</emphasis>
</taxonomicName>
, including those hitherto reported from
<collectingCountry id="2092763E1F79FF9F740808EAFCA5ABD5" box="[828,887,1222,1245]" name="India" pageId="1" pageNumber="518">India</collectingCountry>
, differ from
<taxonomicName id="9F854D2D1F79FF9F74CA08EAFB54ABD5" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[1022,1158,1222,1245]" class="Insecta" family="Veliidae" genus="Thirumalaia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="6AF1EABC1F79FF9F74CA08EAFB54ABD5" box="[1022,1158,1222,1245]" italics="true" pageId="1" pageNumber="518">Thirumalaia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F79FF9F73B908EAFB22ABD5" box="[1165,1264,1222,1245]" pageId="1" pageNumber="518" rank="genus">gen. nov.</taxonomicNameLabel>
by strong modifications of foreleg and abdominal segments 6–7 of the males (=
<taxonomicName id="9F854D2D1F79FF9F74AB092AFBB5AA15" authorityName="Andersen" authorityYear="1989" box="[927,1127,1286,1309]" class="Insecta" family="Veliidae" genus="Baptista" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="6AF1EABC1F79FF9F74AB092AFBB5AA15" box="[927,1127,1286,1309]" italics="true" pageId="1" pageNumber="518">Baptista femoralis</emphasis>
</taxonomicName>
group sensu Kඈඏൺർ &amp; YൺඇG 2000). Four species of
<taxonomicName id="9F854D2D1F79FF9F7307090AFB5CAA35" authorityName="Distant" authorityYear="1903" box="[1075,1166,1318,1341]" class="Insecta" family="Veliidae" genus="Baptista" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="genus">
<emphasis id="6AF1EABC1F79FF9F7307090AFB5CAA35" box="[1075,1166,1318,1341]" italics="true" pageId="1" pageNumber="518">Baptista</emphasis>
</taxonomicName>
from Southeast Asia and Southern
<collectingCountry id="2092763E1F79FF9F7497096AFC31AA55" box="[931,995,1350,1373]" name="China" pageId="1" pageNumber="518">China</collectingCountry>
form the
<taxonomicName id="9F854D2D1F79FF9F737C096AFB29AA55" baseAuthorityName="Andersen" baseAuthorityYear="1989" box="[1096,1275,1350,1373]" class="Insecta" family="Veliidae" genus="Baptista" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="species" species="collaris">
<emphasis id="6AF1EABC1F79FF9F737C096AFB29AA55" box="[1096,1275,1350,1373]" italics="true" pageId="1" pageNumber="518">Baptista collaris</emphasis>
</taxonomicName>
group and also lack such modifications (Zൾඍඍൾඅ 2004, Yൾ et al. 2014b). However,
<taxonomicName id="9F854D2D1F79FF9F749A09AAFBE4AA95" authority="Zettel &amp; Laciny, 2021" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[942,1078,1414,1437]" class="Insecta" family="Veliidae" genus="Thirumalaia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="6AF1EABC1F79FF9F749A09AAFBE4AA95" box="[942,1078,1414,1437]" italics="true" pageId="1" pageNumber="518">Thirumalaia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F79FF9F730B09AAFB76AA95" box="[1087,1188,1414,1437]" pageId="1" pageNumber="518" rank="genus">gen. nov.</taxonomicNameLabel>
possesses none of the characters defining this group either (described in detail by Zൾඍඍൾඅ 2004). The sexually dimorphic body shape, the enlarged compound eyes, and the characteristic parameres are the most important characteristics of
<taxonomicName id="9F854D2D1F79FF9F72270A2AFA4EA915" authority="For" authorityName="Zettel &amp; Laciny" authorityYear="2021" box="[1299,1436,1542,1565]" class="Insecta" family="Veliidae" genus="Thirumalaia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="6AF1EABC1F79FF9F72270A2AFA4EA915" box="[1299,1436,1542,1565]" italics="true" pageId="1" pageNumber="518">Thirumalaia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="71C257C71F79FF9F74080A0AFC4DA935" box="[828,927,1574,1597]" pageId="1" pageNumber="518" rank="genus">gen. nov.</taxonomicNameLabel>
For distinction from other genera see the key.
<emphasis id="6AF1EABC1F79FF9F74080A6AFC6BA955" bold="true" box="[828,953,1606,1629]" pageId="1" pageNumber="518">Etymology.</emphasis>
Dedicated to the memory of the Scientist of the Zoological Survey of
<collectingCountry id="2092763E1F79FF9F73110A4AFB8EA975" box="[1061,1116,1638,1661]" name="India" pageId="1" pageNumber="518">India</collectingCountry>
Dr. G. Thirumalai, honouring his significant contribution to the knowledge of Indian water bugs. Gender: feminine.
</paragraph>
</subSubSection>
</treatment>
</document>
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