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<document id="B456FC24C5AFEA61AE9BFA5D871476F6" ID-DOI="10.5852/ejt.2024.967.2699" ID-ISSN="2118-9773" ID-Zenodo-Dep="13990819" ID-ZooBank="C08B8027-50CC-417E-BCD4-5183B9FF6738" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1729849508920" checkinUser="plazi" docAuthor="Zonstein, Sergei L." docDate="2024" docId="03A8B7235536FFA4FD6EE5D5FD49CAC0" docLanguage="en" docName="EJT.2024.967.1-185.pdf" docOrigin="European Journal of Taxonomy 967" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2699/12459" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Raveniola dolosa Zonstein, 2024, sp. nov." docType="treatment" docUuid="7EC137C9-9E8D-4B2C-8309-358072F8FF08" docUuidSource="ZooBank" docVersion="2" lastPageNumber="31" masterDocId="FF91CF5B552AFFBAFFEFE72AFFB2CE5B" masterDocTitle="A revision of the spider genus Raveniola (Araneae, Nemesiidae). II. Species from Central Asia" masterLastPageNumber="185" masterPageNumber="1" pageNumber="29" updateTime="1729850573173" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="E42BC9799A04DB56ED9C94735D23D8DC">A revision of the spider genus Raveniola (Araneae, Nemesiidae). II. Species from Central Asia</mods:title>
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<taxonomicName id="4C017DB65536FFA6FD6EE5D5FCE3CD41" authorityName="Zonstein" authorityYear="2024" box="[641,849,767,794]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="dolosa" status="sp. nov.">
<emphasis id="B975DA275536FFA6FD6EE5D5FCE3CD41" bold="true" box="[641,849,767,794]" italics="true" pageId="28" pageNumber="29">Raveniola dolosa</emphasis>
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<figureCitation id="133A1AB05536FFA6FF2EE46DFEBECD39" box="[193,268,839,866]" captionStart="Figs 1–9" captionStartId="107.[189,241,1848,1874]" captionTargetBox="[189,1398,268,1805]" captionTargetId="figure-84@107.[189,1398,265,1805]" captionTargetPageId="107" captionText="Figs 1–9.Raveniola spp., ♂♂, holotypes (except 3, 6), habitus, dorsal views. 1. R. caudataZonstein, 2009 (SMNH). 2. R. inopinata sp. nov. (SMNH). 3. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 4. R. alajensis sp. nov. (SMNH). 5. R. cucullata sp. nov. (SMNH). 6. R. cucullata, paratype from Shahob (SMNH). 7. R. dolosa sp. nov. (SMNH). 8. R. hirta sp. nov. (SMNH). 9. R. ignobilis sp. nov. (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990824" httpUri="https://zenodo.org/record/13990824/files/figure.png" pageId="28" pageNumber="29">Figs 7</figureCitation>
, 
<figureCitation id="133A1AB05536FFA6FEF6E46DFE8BCD3A" box="[281,313,839,865]" captionStart="Figs 36–44" captionStartId="111.[189,241,1837,1863]" captionTargetBox="[201,1398,270,1792]" captionTargetId="figure-83@111.[189,1398,265,1792]" captionTargetPageId="111" captionText="Figs 36–44. Raveniola spp., ♀♀, paratypes (except 36–37, 44), habitus, dorsal views. 36. R. redikorzevi (Spassky, 1937), from Kepele (SMNH). 37. R. afghana sp. nov., holotype (NHMD). 38. R. alajensis sp. nov. (SMNH). 39–40. R. cucullata sp. nov. from Shikildara (SMNH) and Childara (SMNH), respectively. 41. R. dolosa sp. nov. (SMNH). 42. R. hirta sp. nov. (SMNH). 43. R. ignobilis sp. nov. (SMNH). 44. R. karategensis sp. nov., holotype (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990834" httpUri="https://zenodo.org/record/13990834/files/figure.png" pageId="28" pageNumber="29">41</figureCitation>
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<figureCitation id="133A1AB05536FFA6FEAAE46DFED7CD3A" box="[325,357,839,865]" captionStart="Figs 69–75" captionStartId="115.[189,241,1831,1857]" captionTargetBox="[191,1400,265,1794]" captionTargetId="figure-66@115.[191,1400,265,1794]" captionTargetPageId="115" captionText="Figs 69–75. Raveniola spp., ♀♀ (69, 71–72, 74) and ♂♂ (70, 73, 75), live spiders; predominantly dorsal (69, 71–72, 74–75) and dorsolateral (70, 73) views. 69–70. R. cucullata sp. nov. (SMNH). 71. R. dolosa sp. nov. (SMNH).72. R. ornatasp. nov. (SMNH). 73–74. R. zyuzinisp. nov. (SMNH). 75. R. ferghanensis (Zonstein, 1984) (SMNH)." figureDoi="http://doi.org/10.5281/zenodo.13990842" httpUri="https://zenodo.org/record/13990842/files/figure.png" pageId="28" pageNumber="29">71</figureCitation>
, 
<figureCitation id="133A1AB05536FFA6FE9EE46DFE23CD3A" box="[369,401,839,865]" captionStart="Figs 82–90" captionStartId="117.[189,241,1853,1879]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-88@117.[189,1398,265,1816]" captionTargetPageId="117" captionText="Figs 82–90. Raveniola spp., ♂♂, holotypes (except 84, 88), cephalothorax, dorsal views. 82. R. caudata Zonstein, 2009 (SMNH). 83. R. inopinata sp. nov. (SMNH). 84. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 85. R. alajensis sp. nov. (SMNH). 86. R. concolor Zonstein, 2000 (MCSN AR 18). 87. R. cucullata sp. nov. (SMNH). 88. R. cucullata, paratype from Shahob (SMNH). 89. R. dolosa sp. nov. (SMNH). 90. R. hirta sp. nov. (SMNH). Scale bars: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990846" httpUri="https://zenodo.org/record/13990846/files/figure.png" pageId="28" pageNumber="29">89</figureCitation>
, 
<figureCitation id="133A1AB05536FFA6FE72E46DFE79CD3A" box="[413,459,839,865]" captionStart="Figs 109–117" captionStartId="120.[189,241,1835,1861]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-93@120.[189,1398,265,1816]" captionTargetPageId="120" captionText="Figs 109–117. Raveniola spp., ♂ (109) and ♀♀ (110–117), paratypes (except 109–111). Cephalothorax, dorsal views. 109. R. vulpina sp. nov., from Tereksai (SMNH). 110. R. redikorzevi (Spassky, 1937), ♀ from Kepele (SMNH). 111. R. afghana sp. nov., holotype (NHMD). 112. R. alajensis sp. nov. (SMNH). 113. R. concolor Zonstein, 2000 (MCSN AR 19). 114–115. R. cucullata sp. nov. from Mullokoni and Shahob, respectively (both SMNH). 116. R. dolosa sp. nov. (SMNH). 117. R. hirta sp. nov. (SMNH). Scale bars: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990858" httpUri="https://zenodo.org/record/13990858/files/figure.png" pageId="28" pageNumber="29">116</figureCitation>
, 
<figureCitation id="133A1AB05536FFA6FE39E46DFDB6CD3A" box="[470,516,839,865]" captionStart="Figs 136–147" captionStartId="123.[189,241,1725,1751]" captionTargetBox="[189,1398,265,1678]" captionTargetId="figure-112@123.[189,1398,265,1678]" captionTargetPageId="123" captionText="Figs 136–147. Raveniola spp., ♂♂, holotypes (except 138, 142), clypeus and eye tubercle, dorsal views. 136. R. caudata Zonstein, 2009 (SMNH). 137. R. inopinata sp. nov. (SMNH). 138. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 139. R. alajensis sp. nov. (SMNH). 140. R. concolor Zonstein, 2000 (MCSN AR 18). 141. R. cucullata sp. nov. (SMNH). 142. R. cucullata, paratype from Shahob (SMNH). 143. R. dolosa sp. nov. (SMNH). 144. R. hirta sp. nov. (SMNH). 145. R. ignobilis sp. nov. (SMNH). 146. R. insolita sp. nov. (ZISP). 147. R. ornata sp. nov. (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990866" httpUri="https://zenodo.org/record/13990866/files/figure.png" pageId="28" pageNumber="29">143</figureCitation>
, 
<figureCitation id="133A1AB05536FFA6FDFEE46DFDF2CD3A" box="[529,576,839,865]" captionStart="Figs 172–183" captionStartId="126.[189,241,1749,1775]" captionTargetBox="[189,1399,265,1701]" captionTargetId="figure-107@126.[189,1399,265,1701]" captionTargetPageId="126" captionText="Figs 172–183. Raveniola spp., ♀♀, paratypes (except 176, 181, 183), clypeus and eye tubercle, dorsal views. 172. R. dolosa sp. nov. (SMNH). 173–174. R. hirta sp. nov. (SMNH). 175. R. ignobilis sp. nov. (SMNH). 176. R. karategensis sp. nov., holotype (SMNH). 177. R. karategensis, paratype (SMNH). 178. R. ornata sp. nov., from Sanglok (SMNH). 179. R. ornatula sp. nov. (SMNH). 180. R. pamira sp. nov. (SMNH). 181. R. sororcula sp. nov., holotype (SMNH). 182. R. diluta sp. nov. (SMNH). 183. R. fedotovi (Charitonov, 1946), ♀ from Hojakurgan (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990872" httpUri="https://zenodo.org/record/13990872/files/figure.png" pageId="28" pageNumber="29">172</figureCitation>
, 
<figureCitation id="133A1AB05536FFA6FDA4E46DFDC8CD3A" box="[587,634,839,865]" captionStart="Figs 202–210" captionStartId="129.[189,241,1851,1877]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-88@129.[189,1398,265,1816]" captionTargetPageId="129" captionText="Figs 202–210. Raveniola spp., ♂♂, holotypes (except 204), sternum, labium and maxillae, ventral views. 202. R. caudata Zonstein, 2009 (SMNH). 203. R. inopinata sp. nov. (SMNH). 204. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 205. R. alajensis sp. nov. (SMNH). 206. R. concolor Zonstein, 2000 (MCSN AR 18). 207. R. cucullata sp. nov. (SMNH). 208. R. dolosa sp. nov. (SMNH). 209. R. hirta sp. nov. (SMNH). 210. R. ignobilis sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990878" httpUri="https://zenodo.org/record/13990878/files/figure.png" pageId="28" pageNumber="29">208</figureCitation>
, 
<figureCitation id="133A1AB05536FFA6FD69E46DFD07CD39" box="[646,693,839,866]" captionStart="Figs 229–237" captionStartId="132.[189,241,1827,1853]" captionTargetBox="[200,1385,265,1784]" captionTargetId="figure-90@132.[200,1385,265,1784]" captionTargetPageId="132" captionText="Figs 229–237. Raveniola spp., ♀♀, paratypes (except 229–230), sternum, labium and maxillae, ventral views. 229. R. redikorzevi (Spassky, 1937), ♀ from Kepele (SMNH). 230. R. afghana sp. nov., holotype (NHMD). 231. R. alajensis sp. nov. (SMNH). 232. R. concolor Zonstein, 2000 (MCSN AR 19). 233– 234. R. cucullata sp. nov. from Shikildara (SMNH) and Shahob (SMNH), respectively. 235. R. dolosa sp. nov.(SMNH). 236. R. hirta sp. nov. (SMNH). 237. R. ignobilis sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990884" httpUri="https://zenodo.org/record/13990884/files/figure.png" pageId="28" pageNumber="29">235</figureCitation>
, 
<figureCitation id="133A1AB05536FFA6FD2EE46DFD42CD39" box="[705,752,839,866]" captionStart="Figs 265–273" captionStartId="136.[189,241,1765,1791]" captionTargetBox="[189,1398,265,1731]" captionTargetId="figure-88@136.[189,1398,265,1731]" captionTargetPageId="136" captionText="Figs 265–273. Raveniola spp., ♂♂, holotypes (except 272), tibia and metatarsus I, retrolateral views. 265. R. dolosa sp. nov. (SMNH). 266. R. hirta sp. nov. (SMNH). 267. R. ignobilis sp. nov. (SMNH). 268. R. insolita sp. nov. (ZISP). 269. R. ornata sp. nov. (SMNH). 270. R. ornatula sp. nov. (SMNH). 271. R. pamira sp. nov. (SMNH). 272. R. pamira, paratype (SMNH). 273. R. diluta sp. nov. (SMNH). Modified hairs are indicated with arrows. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990895" httpUri="https://zenodo.org/record/13990895/files/figure.png" pageId="28" pageNumber="29">265</figureCitation>
, 
<figureCitation id="133A1AB05536FFA6FD14E46DFC98CD39" box="[763,810,839,866]" captionStart="Figs 290–309" captionStartId="139.[189,241,1678,1704]" captionTargetBox="[189,1398,265,1651]" captionTargetId="figure-180@139.[189,1398,265,1651]" captionTargetPageId="139" captionText="Figs 290–309. Raveniola spp., ♂♂, holotypes (except 290, 294, 300, 302, 308), metatarsus I, ventral views. 290. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 291. R. alajensis sp. nov. (SMNH). 292. R. concolor Zonstein, 2000 (MCSN AR 18). 293. R. cucullata sp. nov. (SMNH). 294. R. cucullata, paratype from Shahob (SMNH).295. R. dolosa sp. nov.(SMNH). 296. R. hirtasp. nov. (SMNH). 297. R. ignobilis sp. nov. (SMNH). 298. R. ornata sp. nov. (SMNH). 299. R. ornatula sp. nov. (SMNH). 300. R. ornatula, paratype (SMNH).301. R. pamirasp. nov. (SMNH). 302. R. pamira, paratype (SMNH). 303. R. diluta sp. nov. (SMNH). 304. R. kirgizica sp. nov. (SMNH). 305. R. kopetdaghensis (Fet, 1984) (ZISP). 306. R. mikhailovi Zonstein, 2021 (SMNH). 307. R. nenilini sp. nov. (SMNH). 308. R. virgata (Simon, 1891), ♂ from vicinity of Arslanbob (SMNH). 309. R. vulpina sp. nov. (SMNH). Modified hairs are indicated with arrows. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990903" httpUri="https://zenodo.org/record/13990903/files/figure.png" pageId="28" pageNumber="29">295</figureCitation>
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<figureCitation id="133A1AB05536FFA6FCD9E46DFCD7CD39" box="[822,869,839,866]" captionStart="Figs 349–363" captionStartId="143.[189,241,1727,1753]" captionTargetBox="[189,1390,270,1690]" captionTargetId="figure-139@143.[189,1398,265,1690]" captionTargetPageId="143" captionText="Figs 349–363. Raveniola spp., ♂♂, holotypes (except 351, 355, 363), palpal tibia, cymbium and copulatory bulb, retrolateral (349–358, 360–361, 363) and retroventral (359, 362) views.349. R. caudata Zonstein, 2009 (SMNH). 350. R. inopinata sp. nov. (SMNH). 351. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 352. R. alajensis sp. nov. (SMNH). 353. R. concolor Zonstein, 2000 (MCSN AR 18). 354. R. cucullata sp. nov. (SMNH). 355. R. cucullata, paratype from Shahob (SMNH). 356. R. dolosa sp. nov. (SMNH). 357. R. hirta sp. nov. (SMNH). 358–359. R. ignobilis sp. nov. (SMNH). 360. R. insolita sp. nov. (ZISP). 361. R. ornata sp. nov. (SMNH). 362. R. ornatula sp. nov. (SMNH). 363. R. pamira sp. nov., paratype (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990913" httpUri="https://zenodo.org/record/13990913/files/figure.png" pageId="28" pageNumber="29">356</figureCitation>
, 
<figureCitation id="133A1AB05536FFA6FC9EE46DFC68CD3A" box="[881,986,839,865]" captionStart-0="Figs 379–388" captionStart-1="Figs 389–399" captionStartId-0="145.[189,241,1844,1870]" captionStartId-1="146.[189,241,1854,1880]" captionTargetBox-0="[189,1398,265,1806]" captionTargetBox-1="[189,1398,265,1814]" captionTargetId-0="figure-75@145.[189,1398,265,1806]" captionTargetId-1="figure-68@146.[189,1398,265,1814]" captionTargetPageId-0="145" captionTargetPageId-1="146" captionText-0="Figs 379–388. Raveniola spp., ♂♂, holotypes, details of copulatoty bulb, retrolateral (379, 381, 383, 385, 388), ventral (380, 382, 384, 387) and retroventral (386) views. 379–380. R. caudata Zonstein, 2009 (SMNH). 381–382. R. inopinata sp. nov. (SMNH). 383–384. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH).385–387. R. alajensis sp. nov. (SMNH). 388. R. concolor Zonstein, 2000 (MCSN AR 18). Scale bars: 0.5 mm." captionText-1="Figs 389–399. Raveniola spp., ♂♂, holotypes (except 392–396), details of copulatoty bulb, retrolateral (389, 392, 395, 397), retroventral (390, 393, 398) and ventral (391, 394, 396, 399) views. 389– 391. R. cucullata sp. nov. (SMNH). 392–394. R. cucullata sp. nov., paratype from Shahob (SMNH). 395–396. R. cucullata, paratype from Sangdara (SMNH). 397–399. R. dolosa sp. nov. (SMNH). Scale bars: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13990917" figureDoi-1="http://doi.org/10.5281/zenodo.13990919" httpUri-0="https://zenodo.org/record/13990917/files/figure.png" httpUri-1="https://zenodo.org/record/13990919/files/figure.png" pageId="28" pageNumber="29">379–399</figureCitation>
, 
<figureCitation id="133A1AB05536FFA6FC09E46DFBFDCD39" box="[998,1103,839,866]" captionStart="Figs 487–503" captionStartId="156.[189,241,1695,1721]" captionTargetBox="[189,1398,265,1653]" captionTargetId="figure-102@156.[189,1398,265,1653]" captionTargetPageId="156" captionText="Figs 487–503. Raveniola spp., ♀♀, paratypes (except 487–489), spermathecae, dorsal views. 487–488. R. redikorzevi (Spassky, 1937), Kepele (SMNH). 489. R. afghana sp. nov. (NHMD). 490. Same, holotype. 491–493. R. alajensis sp. nov., Berksu. 494. R. concolor Zonstein, 2000 (MCSN AR 19). 495–496. R. cucullata sp. nov., Shikildara (SMNH). 497. R. cucullata, Shahob (SMNH). 498– 499. R. cucullata, Sangdara (SMNH). 500–503. R. dolosa sp. nov. (SMNH). Abbreviations: isb = inner spermathecal branch; osb = outer spermathecal branch; sb = spermathecal base. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990941" httpUri="https://zenodo.org/record/13990941/files/figure.png" pageId="28" pageNumber="29">500–503</figureCitation>
, 568–570, 643–650, 751
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE5536FFA6FF52E4A3FCC3CAD3" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="8BBE06355536FFA6FF52E4A3FE84CDF8" blockId="28.[189,310,905,931]" box="[189,310,905,931]" pageId="28" pageNumber="29">
<heading id="D0F6B1595536FFA6FF52E4A3FE84CDF8" bold="true" box="[189,310,905,931]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B975DA275536FFA6FF52E4A3FE84CDF8" bold="true" box="[189,310,905,931]" pageId="28" pageNumber="29">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE06355536FFA6FF52E490FCC3CAD3" blockId="28.[186,1399,953,1160]" pageId="28" pageNumber="29">
In the structure of the embolus and the spermathecae, 
<taxonomicName id="4C017DB65536FFA6FCD4E493FBBACD88" authorityName="Zonstein" authorityYear="2024" box="[827,1032,953,979]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="dolosa" status="sp. nov.">
<emphasis id="B975DA275536FFA6FCD4E493FBBACD88" box="[827,1032,953,979]" italics="true" pageId="28" pageNumber="29">Raveniola dolosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C5536FFA6FBFFE490FBD9CD8F" box="[1040,1131,954,980]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
resembles 
<taxonomicName id="4C017DB65536FFA6FB00E490FAC5CD88" authorityName="Zonstein" authorityYear="2024" box="[1263,1399,953,980]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="ignobilis" status="sp. nov.">
<emphasis id="B975DA275536FFA6FB00E490FAB2CD88" box="[1263,1280,954,979]" italics="true" pageId="28" pageNumber="29">R</emphasis>
. 
<emphasis id="B975DA275536FFA6FAFFE493FAC5CD88" box="[1296,1399,953,979]" italics="true" pageId="28" pageNumber="29">ignobilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C5536FFA6FF52E4F4FEA5CDA3" box="[189,279,990,1016]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
; it can be distinguished from the latter in having the embolus lacking a raised keel and in possessing noticeably shorter spermathecae (vs a clearly keeled embolus and longer branches of the spermathecae in 
<taxonomicName id="4C017DB65536FFA6FF31E30CFEDBCA64" authorityName="Zonstein" authorityYear="2024" box="[222,361,1061,1088]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="ignobilis">
<emphasis id="B975DA275536FFA6FF31E30CFF42CA64" box="[222,240,1062,1087]" italics="true" pageId="28" pageNumber="29">R</emphasis>
. 
<emphasis id="B975DA275536FFA6FEEDE30FFEDBCA64" box="[258,361,1061,1087]" italics="true" pageId="28" pageNumber="29">ignobilis</emphasis>
</taxonomicName>
; 
<figureCitation id="133A1AB05536FFA6FE94E30FFD90CA64" box="[379,546,1061,1087]" captionStart="Figs 389–399" captionStartId="146.[189,241,1854,1880]" captionTargetBox="[189,1398,265,1814]" captionTargetId="figure-68@146.[189,1398,265,1814]" captionTargetPageId="146" captionText="Figs 389–399. Raveniola spp., ♂♂, holotypes (except 392–396), details of copulatoty bulb, retrolateral (389, 392, 395, 397), retroventral (390, 393, 398) and ventral (391, 394, 396, 399) views. 389– 391. R. cucullata sp. nov. (SMNH). 392–394. R. cucullata sp. nov., paratype from Shahob (SMNH). 395–396. R. cucullata, paratype from Sangdara (SMNH). 397–399. R. dolosa sp. nov. (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990919" httpUri="https://zenodo.org/record/13990919/files/figure.png" pageId="28" pageNumber="29">Figs 397–399</figureCitation>
, 
<figureCitation id="133A1AB05536FFA6FDDEE30FFD2ACA1B" box="[561,664,1061,1088]" captionStart="Figs 487–503" captionStartId="156.[189,241,1695,1721]" captionTargetBox="[189,1398,265,1653]" captionTargetId="figure-102@156.[189,1398,265,1653]" captionTargetPageId="156" captionText="Figs 487–503. Raveniola spp., ♀♀, paratypes (except 487–489), spermathecae, dorsal views. 487–488. R. redikorzevi (Spassky, 1937), Kepele (SMNH). 489. R. afghana sp. nov. (NHMD). 490. Same, holotype. 491–493. R. alajensis sp. nov., Berksu. 494. R. concolor Zonstein, 2000 (MCSN AR 19). 495–496. R. cucullata sp. nov., Shikildara (SMNH). 497. R. cucullata, Shahob (SMNH). 498– 499. R. cucullata, Sangdara (SMNH). 500–503. R. dolosa sp. nov. (SMNH). Abbreviations: isb = inner spermathecal branch; osb = outer spermathecal branch; sb = spermathecal base. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990941" httpUri="https://zenodo.org/record/13990941/files/figure.png" pageId="28" pageNumber="29">500–503</figureCitation>
cf. 
<figureCitation id="133A1AB05536FFA6FD24E30FFCC3CA64" box="[715,881,1061,1087]" captionStart="Figs 400–408" captionStartId="147.[189,241,1853,1879]" captionTargetBox="[198,1398,265,1817]" captionTargetId="figure-50@147.[189,1398,265,1817]" captionTargetPageId="147" captionText="Figs 400–408. Raveniola spp., ♂♂, holotypes (except 406–408), details of copulatoty bulb; retrolateral (400, 403, 406–407), retroventral (401, 404) and ventral (402, 405, 408) views. 400–402. R. hirta sp. nov. (SMNH). 403–405. R. ignobilis sp. nov. (SMNH). 406–408. R. ignobilis, paratypes (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990921" httpUri="https://zenodo.org/record/13990921/files/figure.png" pageId="28" pageNumber="29">Figs 403–408</figureCitation>
, 
<figureCitation id="133A1AB05536FFA6FC6FE30FFC03CA1B" box="[896,945,1061,1088]" captionStart="Figs 504–521" captionStartId="157.[189,241,1719,1745]" captionTargetBox="[189,1398,265,1675]" captionTargetId="figure-86@157.[189,1398,265,1675]" captionTargetPageId="157" captionText="Figs 504–521. Raveniola spp., ♀♀, spermathecae, dorsal views. 504–508. R. hirta sp. nov., paratypes (SMNH). 509. R. ignobilis sp. nov., paratype (SMNH). 510. R. karategensis sp. nov., holotype (SMNH). 511–512. R. karategensis, paratypes (ZISP). 513–514. R. ornata sp. nov., paratypes from Ganjina (SMNH). 515–516. R. ornata, paratypes from Sanglok (SMNH). 517–519. R. ornatula sp. nov., paratypes (SMNH). 520–521. R. pamira sp. nov., paratypes (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990943" httpUri="https://zenodo.org/record/13990943/files/figure.png" pageId="28" pageNumber="29">509</figureCitation>
). Additionally, 
<taxonomicName id="4C017DB65536FFA6FB85E30CFB6ECA64" authorityName="Zonstein" authorityYear="2024" box="[1130,1244,1061,1088]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="dolosa">
<emphasis id="B975DA275536FFA6FB85E30CFBCECA64" box="[1130,1148,1062,1087]" italics="true" pageId="28" pageNumber="29">R</emphasis>
. 
<emphasis id="B975DA275536FFA6FB61E30FFB6ECA64" box="[1166,1244,1061,1087]" italics="true" pageId="28" pageNumber="29">dolosa</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C017DB65536FFA6FF52E360FEF4CA38" authorityName="Zonstein" authorityYear="2024" box="[189,326,1097,1124]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="ignobilis">
<emphasis id="B975DA275536FFA6FF52E360FF7DCA38" box="[189,207,1098,1123]" italics="true" pageId="28" pageNumber="29">R</emphasis>
. 
<emphasis id="B975DA275536FFA6FF30E363FEF4CA38" box="[223,326,1097,1123]" italics="true" pageId="28" pageNumber="29">ignobilis</emphasis>
</taxonomicName>
in having the posterior medium spinnerets strictly reduced in size (vs considerably larger PMS in the latter species; Figs 568–570 cf. Figs 573–574).
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE5536FFA6FF52E39AFE12CB1A" pageId="28" pageNumber="29" type="etymology">
<paragraph id="8BBE06355536FFA6FF52E39AFEF6CA91" blockId="28.[189,324,1200,1226]" box="[189,324,1200,1226]" pageId="28" pageNumber="29">
<heading id="D0F6B1595536FFA6FF52E39AFEF6CA91" bold="true" box="[189,324,1200,1226]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B975DA275536FFA6FF52E39AFEF6CA91" bold="true" box="[189,324,1200,1226]" pageId="28" pageNumber="29">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE06355536FFA6FF52E3F5FE12CB1A" blockId="28.[189,1399,1247,1345]" pageId="28" pageNumber="29">
The specific epithet is a Latin adjective ‘ 
<emphasis id="B975DA275536FFA6FD4FE3F5FCE2CAA2" box="[672,848,1247,1273]" italics="true" pageId="28" pageNumber="29">dolosus/-a/-um</emphasis>
’ (= deceptive) referring to a general similarity of this new species to the closest congeners, 
<taxonomicName id="4C017DB65536FFA6FD22E229FC01CB46" authorityName="Zonstein" authorityYear="2024" box="[717,947,1283,1309]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="ignobilis" status="sp. nov.">
<emphasis id="B975DA275536FFA6FD22E229FC01CB46" box="[717,947,1283,1309]" italics="true" pageId="28" pageNumber="29">Raveniola ignobilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C5536FFA6FC55E22EFBA7CB45" box="[954,1045,1284,1310]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C017DB65536FFA6FBBFE22EFB52CB46" authorityName="Zonstein" authorityYear="2024" box="[1104,1248,1283,1310]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="sororcula" status="sp. nov.">
<emphasis id="B975DA275536FFA6FBBFE22EFBD3CB46" box="[1104,1121,1284,1309]" italics="true" pageId="28" pageNumber="29">R</emphasis>
. 
<emphasis id="B975DA275536FFA6FB9FE229FB52CB46" box="[1136,1248,1283,1309]" italics="true" pageId="28" pageNumber="29">sororcula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C5536FFA6FB07E22EFAF4CB45" box="[1256,1350,1284,1310]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
; the gender is feminine.
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE5536FFA6FF52E243FBB8C8F1" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8BBE06355536FFA6FF52E243FE18CBD8" blockId="28.[189,426,1385,1411]" box="[189,426,1385,1411]" pageId="28" pageNumber="29">
<heading id="D0F6B1595536FFA6FF52E243FE18CBD8" bold="true" box="[189,426,1385,1411]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B975DA275536FFA6FF52E243FE18CBD8" bold="true" box="[189,426,1385,1411]" pageId="28" pageNumber="29">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE06355536FFA6FF05E2B3FEEECBE8" blockId="28.[189,1398,1433,1532]" box="[234,348,1433,1459]" pageId="28" pageNumber="29">
<emphasis id="B975DA275536FFA6FF05E2B3FEEECBE8" bold="true" box="[234,348,1433,1459]" pageId="28" pageNumber="29">
<typeStatus id="54BAB8975536FFA6FF05E2B3FEEECBE8" box="[234,348,1433,1459]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BBE06355536FFA6FF52E297FBA7CBA0" blockId="28.[189,1398,1433,1532]" pageId="28" pageNumber="29">
<materialsCitation id="3B690C685536FFA6FF52E297FBA7CBA0" collectingDate="1988-07-12" collectionCode="SMNH" collectorName="Peter I Mts &amp; S. Zonstein" country="Tajikistan" elevation="1900" latitude="38.85" location="Childara Canyon" longLatPrecision="1172" longitude="70.3" municipality="Shahobdara Gorge" pageId="28" pageNumber="29" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="F31646A55536FFA6FF52E297FED2CB8C" box="[189,352,1469,1495]" name="Tajikistan" pageId="28" pageNumber="29">TAJIKISTAN</collectingCountry>
• 
<specimenCount id="9D07CDBC5536FFA6FE92E294FE24CB83" box="[381,406,1470,1496]" count="1" pageId="28" pageNumber="29" type="male">♂</specimenCount>
; 
<collectorName id="26F463E35536FFA6FE4BE294FD9FCB83" box="[420,557,1470,1496]" pageId="28" pageNumber="29">Peter I Mts</collectorName>
, 
<location id="8EDE50EE5536FFA6FDD4E297FCB5CB8C" LSID="urn:lsid:plazi:treatment:03A8B7235536FFA4FD6EE5D5FD49CAC0:8EDE50EE5536FFA6FDD4E297FCB5CB8C" box="[571,775,1469,1495]" country="Tajikistan" latitude="38.85" longLatPrecision="1172" longitude="70.3" municipality="Shahobdara Gorge" name="Childara Canyon" pageId="28" pageNumber="29">Childara Canyon</location>
, 
<collectingMunicipality id="6BDA9C4F5536FFA6FCFAE297FC47CB8C" box="[789,1013,1469,1495]" pageId="28" pageNumber="29">Shahobdara Gorge</collectingMunicipality>
, 
<locationDeviation id="9A0C1F175536FFA6FBEDE297FAC0CB8C" box="[1026,1394,1469,1496]" location="Shahob Village" pageId="28" pageNumber="29">
<quantity id="4CF9ABD05536FFA6FBEDE297FB8DCB8C" box="[1026,1087,1469,1495]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="28" pageNumber="29" unit="km" value="4.0">4 km</quantity>
NNW of Shahob Village
</locationDeviation>
; 
<geoCoordinate id="EE3560F25536FFA6FF52E2CAFE9BCBA7" box="[189,297,1504,1532]" degrees="38" direction="north" minutes="51" orientation="latitude" pageId="28" pageNumber="29" precision="925" value="38.85">38°51′ N</geoCoordinate>
, 
<geoCoordinate id="EE3560F25536FFA6FEDAE2CAFE2CCBA7" box="[309,414,1504,1532]" degrees="70" direction="east" minutes="18" orientation="longitude" pageId="28" pageNumber="29" precision="925" value="70.3">70°18′ E</geoCoordinate>
; 
<elevation id="002CE1065536FFA6FE44E2CBFDF1CBA0" box="[427,579,1505,1532]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="28" pageNumber="29" unit="m" value="1900.0">
<quantity id="4CF9ABD05536FFA6FE44E2CBFDB6CBA7" box="[427,516,1505,1532]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="28" pageNumber="29" unit="m" value="1900.0">1900 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFBF20F55536FFA6FDBEE2CBFD56CBA0" box="[593,740,1505,1531]" pageId="28" pageNumber="29" value="1988-07-12">
<collectingDate id="EFFBD91D5536FFA6FDBEE2CBFD56CBA0" box="[593,740,1505,1531]" pageId="28" pageNumber="29" value="1988-07-12">12 Jul. 1988</collectingDate>
</date>
; 
<collectorName id="26F463E35536FFA6FD1FE2CBFCC4CBA0" box="[752,886,1505,1531]" pageId="28" pageNumber="29">S. Zonstein</collectorName>
leg.; 
<collectionCode id="ED109EF05536FFA6FC57E2CBFBA3CBA0" box="[952,1041,1505,1531]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="28" pageNumber="29">SMNH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BBE06355536FFA6FF05E10EFE66C865" blockId="28.[189,1399,1572,1706]" box="[234,468,1572,1598]" pageId="28" pageNumber="29">
<emphasis id="B975DA275536FFA6FF05E10EFED5C865" bold="true" box="[234,359,1572,1598]" pageId="28" pageNumber="29">
<typeStatus id="54BAB8975536FFA6FF05E10EFED5C865" box="[234,359,1572,1598]" pageId="28" pageNumber="29" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="9D07CDBC5536FFA6FE97E10EFE7AC865" box="[376,456,1572,1598]" count="13" pageId="28" pageNumber="29" type="female">13 ♀♀</specimenCount>
)
</paragraph>
<paragraph id="8BBE06355536FFA6FF52E162FBB8C8F1" blockId="28.[189,1399,1572,1706]" pageId="28" pageNumber="29">
<materialsCitation id="3B690C685536FFA6FF52E162FDE4C8DD" collectingDate="1988-07-12" collectingDateMax="1988-07-13" collectingDateMin="1988-07-12" collectionCode="V, SMNH" collectorName="S. Zonstein &amp; Ovchinnikov" country="Tajikistan" elevation="2000" latitude="38.85" location="Childara Canyon" longLatPrecision="1172" longitude="70.3" municipality="Shahobdara Gorge" pageId="28" pageNumber="29" specimenCount="11" specimenCount-female="11">
<collectingCountry id="F31646A55536FFA6FF52E162FED2C839" box="[189,352,1608,1634]" name="Tajikistan" pageId="28" pageNumber="29">TAJIKISTAN</collectingCountry>
• 
<specimenCount id="9D07CDBC5536FFA6FE97E162FE7BC839" box="[376,457,1608,1634]" count="11" pageId="28" pageNumber="29" type="female">11 ♀♀</specimenCount>
; same locality as for holotype; 
<elevation id="002CE1065536FFA6FCD7E162FBABC839" box="[824,1049,1608,1634]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="28" pageNumber="29" unit="m" value="2000.0" valueMax="2100.0" valueMin="1900.0">
<quantity id="4CF9ABD05536FFA6FCD7E162FC68C839" box="[824,986,1608,1634]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="28" pageNumber="29" unit="m" value="2000.0" valueMax="2100.0" valueMin="1900.0">1900–2100 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFBF20F55536FFA6FBC9E162FB57C839" box="[1062,1253,1608,1634]" pageId="28" pageNumber="29" value="1988-07-12" valueMax="1988-07-13" valueMin="1988-07-12">
<collectingDate id="EFFBD91D5536FFA6FBC9E162FB57C839" box="[1062,1253,1608,1634]" pageId="28" pageNumber="29" value="1988-07-12" valueMax="1988-07-13" valueMin="1988-07-12">12–13 Jul. 1988</collectingDate>
</date>
; 
<collectorName id="26F463E35536FFA6FB1EE162FAC5C839" box="[1265,1399,1608,1634]" pageId="28" pageNumber="29">S. Zonstein</collectorName>
and S. 
<collectionCode id="ED109EF05536FFA6FEECE146FEA9C8DD" box="[259,283,1644,1670]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="28" pageNumber="29" type="Museum">V</collectionCode>
. 
<collectorName id="26F463E35536FFA6FEC9E146FE0DC8DD" box="[294,447,1644,1670]" pageId="28" pageNumber="29">Ovchinnikov</collectorName>
leg.; 
<collectionCode id="ED109EF05536FFA6FDEEE146FDE4C8DD" box="[513,598,1644,1670]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="28" pageNumber="29">SMNH</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B690C685536FFA6FD81E146FBB6C8F1" collectingDate="2019-07-08" collectionCode="SMNH" collectorName="S. Zonstein" country="Tajikistan" elevation="1800" latitude="38.833332" location="Shahob Village" longLatPrecision="1173" longitude="70.316666" municipality="Shahobdara Gorge" pageId="28" pageNumber="29" specimenCount="2" specimenCount-female="2">
<specimenCount id="9D07CDBC5536FFA6FD81E146FD03C8DD" box="[622,689,1644,1670]" count="2" pageId="28" pageNumber="29" type="female">2 ♀♀</specimenCount>
; same locality as for holotype, 
<quantity id="4CF9ABD05536FFA6FBCFE146FBC1C8DD" box="[1056,1139,1644,1671]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="28" pageNumber="29" unit="km" value="2.5">2.5 km</quantity>
N of 
<location id="8EDE50EE5536FFA6FB58E146FAC0C8DD" LSID="urn:lsid:plazi:treatment:03A8B7235536FFA4FD6EE5D5FD49CAC0:8EDE50EE5536FFA6FB58E146FAC0C8DD" box="[1207,1394,1644,1670]" country="Tajikistan" latitude="38.833332" longLatPrecision="1173" longitude="70.316666" name="Shahob Village" pageId="28" pageNumber="29">Shahob Village</location>
; 
<geoCoordinate id="EE3560F25536FFA6FF52E1A4FE99C8F1" box="[189,299,1678,1706]" degrees="38" direction="north" minutes="50" orientation="latitude" pageId="28" pageNumber="29" precision="925" value="38.833332">38°50′ N</geoCoordinate>
, 
<geoCoordinate id="EE3560F25536FFA6FED7E1A4FE11C8F1" box="[312,419,1678,1706]" degrees="70" direction="east" minutes="19" orientation="longitude" pageId="28" pageNumber="29" precision="925" value="70.316666">70°19′ E</geoCoordinate>
; 
<elevation id="002CE1065536FFA6FE5FE1BAFDFBC8F1" box="[432,585,1680,1706]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="28" pageNumber="29" unit="m" value="1800.0">
<quantity id="4CF9ABD05536FFA6FE5FE1BAFDBBC8F1" box="[432,521,1680,1706]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="28" pageNumber="29" unit="m" value="1800.0">1800 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFBF20F55536FFA6FDB9E1BAFD69C8F1" box="[598,731,1680,1706]" pageId="28" pageNumber="29" value="2019-07-08">
<collectingDate id="EFFBD91D5536FFA6FDB9E1BAFD69C8F1" box="[598,731,1680,1706]" pageId="28" pageNumber="29" value="2019-07-08">8 Jul. 2019</collectingDate>
</date>
; 
<collectorName id="26F463E35536FFA6FD08E1BAFCDFC8F1" box="[743,877,1680,1706]" pageId="28" pageNumber="29">S. Zonstein</collectorName>
leg.; 
<collectionCode id="ED109EF05536FFA6FC40E1BAFBB6C8F1" box="[943,1028,1680,1706]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="28" pageNumber="29">SMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE5536FFA4FF52E1F8FD7BCDD5" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29" type="description">
<paragraph id="8BBE06355536FFA6FF52E1F8FEFCC8B7" blockId="28.[189,334,1746,1772]" box="[189,334,1746,1772]" pageId="28" pageNumber="29">
<heading id="D0F6B1595536FFA6FF52E1F8FEFCC8B7" bold="true" box="[189,334,1746,1772]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B975DA275536FFA6FF52E1F8FEFCC8B7" bold="true" box="[189,334,1746,1772]" pageId="28" pageNumber="29">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE06355536FFA6FF05E028FE19C940" blockId="28.[189,427,1793,1856]" box="[234,427,1793,1820]" pageId="28" pageNumber="29">
<emphasis id="B975DA275536FFA6FF05E028FE98C947" bold="true" box="[234,298,1794,1820]" pageId="28" pageNumber="29">Male</emphasis>
(
<typeStatus id="54BAB8975536FFA6FED6E02BFE11C940" box="[313,419,1793,1819]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
)
</paragraph>
<paragraph id="8BBE06355536FFA6FF52E00CFE1BC91B" blockId="28.[189,427,1793,1856]" box="[189,425,1830,1856]" pageId="28" pageNumber="29">
<smallCapsWord id="8D5890E95536FFA6FF52E00CFEAEC965" baselines="1849,1849" box="[189,284,1830,1856]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Habitus" pageId="28" pageNumber="29">HABITUS</smallCapsWord>
. See 
<figureCitation id="133A1AB05536FFA6FEB2E00CFE1BC91B" box="[349,425,1830,1856]" captionStart="Figs 1–9" captionStartId="107.[189,241,1848,1874]" captionTargetBox="[189,1398,268,1805]" captionTargetId="figure-84@107.[189,1398,265,1805]" captionTargetPageId="107" captionText="Figs 1–9.Raveniola spp., ♂♂, holotypes (except 3, 6), habitus, dorsal views. 1. R. caudataZonstein, 2009 (SMNH). 2. R. inopinata sp. nov. (SMNH). 3. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 4. R. alajensis sp. nov. (SMNH). 5. R. cucullata sp. nov. (SMNH). 6. R. cucullata, paratype from Shahob (SMNH). 7. R. dolosa sp. nov. (SMNH). 8. R. hirta sp. nov. (SMNH). 9. R. ignobilis sp. nov. (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990824" httpUri="https://zenodo.org/record/13990824/files/figure.png" pageId="28" pageNumber="29">Fig. 7.</figureCitation>
</paragraph>
<paragraph id="8BBE06355536FFA6FF52E042FB15C9D9" blockId="28.[189,1191,1896,1923]" box="[189,1191,1896,1923]" pageId="28" pageNumber="29">
<smallCapsWord id="8D5890E95536FFA6FF52E042FEDEC9DB" baselines="1915,1915" box="[189,364,1896,1922]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="28" pageNumber="29">MEASUREMENTS</smallCapsWord>
. TBL 12.55, CL 5.24, CW 4.97, LL 0.50, LW 0.87, SL 2.51, SW 2.41.
</paragraph>
<paragraph id="8BBE06355536FFA7FF52E080FDF3CF13" blockId="28.[189,1398,1962,2025]" lastBlockId="29.[189,1398,265,328]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<smallCapsWord id="8D5890E95536FFA6FF52E080FEAAC998" baselines="1981,1982" box="[189,280,1962,1988]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Colour" pageId="28" pageNumber="29">COLOUR</smallCapsWord>
. Carapace, chelicerae and leg I dull reddish brown; palps and legs II–IV lighter reddish brown; eye tubercle dark brown; sternum, labium and maxillae light yellowish brown; abdomen tan brown, with dark brown dorsal chevron-like pattern and a few small brown marks on ventral side; book-lungs and spinnerets pale yellowish brown.
</paragraph>
<paragraph id="8BBE06355537FFA7FF52E65BFE46CC1F" blockId="29.[187,1399,369,580]" pageId="29" pageNumber="30">
<smallCapsWord id="8D5890E95537FFA7FF52E65BFECACFD1" baselines="388,389" box="[189,376,369,395]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Cephalothorax" pageId="29" pageNumber="30">CEPHALOTHORAX</smallCapsWord>
. Carapace and chelicerae as shown in 
<figureCitation id="133A1AB05537FFA7FCDDE65BFC35CFD0" box="[818,903,369,395]" captionStart="Figs 82–90" captionStartId="117.[189,241,1853,1879]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-88@117.[189,1398,265,1816]" captionTargetPageId="117" captionText="Figs 82–90. Raveniola spp., ♂♂, holotypes (except 84, 88), cephalothorax, dorsal views. 82. R. caudata Zonstein, 2009 (SMNH). 83. R. inopinata sp. nov. (SMNH). 84. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 85. R. alajensis sp. nov. (SMNH). 86. R. concolor Zonstein, 2000 (MCSN AR 18). 87. R. cucullata sp. nov. (SMNH). 88. R. cucullata, paratype from Shahob (SMNH). 89. R. dolosa sp. nov. (SMNH). 90. R. hirta sp. nov. (SMNH). Scale bars: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990846" httpUri="https://zenodo.org/record/13990846/files/figure.png" pageId="29" pageNumber="30">Fig. 89</figureCitation>
. Clypeus and eye group as in 
<figureCitation id="133A1AB05537FFA7FB34E65BFA8CCFD0" box="[1243,1342,369,395]" captionStart="Figs 136–147" captionStartId="123.[189,241,1725,1751]" captionTargetBox="[189,1398,265,1678]" captionTargetId="figure-112@123.[189,1398,265,1678]" captionTargetPageId="123" captionText="Figs 136–147. Raveniola spp., ♂♂, holotypes (except 138, 142), clypeus and eye tubercle, dorsal views. 136. R. caudata Zonstein, 2009 (SMNH). 137. R. inopinata sp. nov. (SMNH). 138. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 139. R. alajensis sp. nov. (SMNH). 140. R. concolor Zonstein, 2000 (MCSN AR 18). 141. R. cucullata sp. nov. (SMNH). 142. R. cucullata, paratype from Shahob (SMNH). 143. R. dolosa sp. nov. (SMNH). 144. R. hirta sp. nov. (SMNH). 145. R. ignobilis sp. nov. (SMNH). 146. R. insolita sp. nov. (ZISP). 147. R. ornata sp. nov. (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990866" httpUri="https://zenodo.org/record/13990866/files/figure.png" pageId="29" pageNumber="30">Fig. 143</figureCitation>
. Eye diameters and interdistances: AME 0.14(0.20), ALE 0.26, PLE 0.16, PME 0.12; AME–AME 0.10(0.05), ALE–AME 0.11(0.08), ALE–PLE 0.10, PLE–PME 0.06, PME–PME 0.37. Anterior cheliceral edge with unmodified setae; rastellum not developed. Each cheliceral furrow with 9 promarginal teeth and 3–4 mesobasal denticles. MIT indiscernible. Sternum, labium and maxillae as shown in 
<figureCitation id="133A1AB05537FFA7FB73E52FFAB1CC7B" box="[1180,1283,517,544]" captionStart="Figs 202–210" captionStartId="129.[189,241,1851,1877]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-88@129.[189,1398,265,1816]" captionTargetPageId="129" captionText="Figs 202–210. Raveniola spp., ♂♂, holotypes (except 204), sternum, labium and maxillae, ventral views. 202. R. caudata Zonstein, 2009 (SMNH). 203. R. inopinata sp. nov. (SMNH). 204. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 205. R. alajensis sp. nov. (SMNH). 206. R. concolor Zonstein, 2000 (MCSN AR 18). 207. R. cucullata sp. nov. (SMNH). 208. R. dolosa sp. nov. (SMNH). 209. R. hirta sp. nov. (SMNH). 210. R. ignobilis sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990878" httpUri="https://zenodo.org/record/13990878/files/figure.png" pageId="29" pageNumber="30">Fig. 208</figureCitation>
. Maxillae with 36–39 cuspules each.
</paragraph>
<paragraph id="8BBE06355537FFA7FF52E544FD69CCAC" blockId="29.[189,1399,622,759]" pageId="29" pageNumber="30">
<smallCapsWord id="8D5890E95537FFA7FF52E544FF44CCDD" baselines="641,641" box="[189,246,622,648]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Legs" pageId="29" pageNumber="30">LEGS</smallCapsWord>
. Tibia and metatarsus I as in 
<figureCitation id="133A1AB05537FFA7FDBCE544FD0FCCD2" box="[595,701,622,649]" captionStart="Figs 265–273" captionStartId="136.[189,241,1765,1791]" captionTargetBox="[189,1398,265,1731]" captionTargetId="figure-88@136.[189,1398,265,1731]" captionTargetPageId="136" captionText="Figs 265–273. Raveniola spp., ♂♂, holotypes (except 272), tibia and metatarsus I, retrolateral views. 265. R. dolosa sp. nov. (SMNH). 266. R. hirta sp. nov. (SMNH). 267. R. ignobilis sp. nov. (SMNH). 268. R. insolita sp. nov. (ZISP). 269. R. ornata sp. nov. (SMNH). 270. R. ornatula sp. nov. (SMNH). 271. R. pamira sp. nov. (SMNH). 272. R. pamira, paratype (SMNH). 273. R. diluta sp. nov. (SMNH). Modified hairs are indicated with arrows. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990895" httpUri="https://zenodo.org/record/13990895/files/figure.png" pageId="29" pageNumber="30">Figs 265</figureCitation>
, 
<figureCitation id="133A1AB05537FFA7FD26E544FD4BCCD2" box="[713,761,622,649]" captionStart="Figs 290–309" captionStartId="139.[189,241,1678,1704]" captionTargetBox="[189,1398,265,1651]" captionTargetId="figure-180@139.[189,1398,265,1651]" captionTargetPageId="139" captionText="Figs 290–309. Raveniola spp., ♂♂, holotypes (except 290, 294, 300, 302, 308), metatarsus I, ventral views. 290. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 291. R. alajensis sp. nov. (SMNH). 292. R. concolor Zonstein, 2000 (MCSN AR 18). 293. R. cucullata sp. nov. (SMNH). 294. R. cucullata, paratype from Shahob (SMNH).295. R. dolosa sp. nov.(SMNH). 296. R. hirtasp. nov. (SMNH). 297. R. ignobilis sp. nov. (SMNH). 298. R. ornata sp. nov. (SMNH). 299. R. ornatula sp. nov. (SMNH). 300. R. ornatula, paratype (SMNH).301. R. pamirasp. nov. (SMNH). 302. R. pamira, paratype (SMNH). 303. R. diluta sp. nov. (SMNH). 304. R. kirgizica sp. nov. (SMNH). 305. R. kopetdaghensis (Fet, 1984) (ZISP). 306. R. mikhailovi Zonstein, 2021 (SMNH). 307. R. nenilini sp. nov. (SMNH). 308. R. virgata (Simon, 1891), ♂ from vicinity of Arslanbob (SMNH). 309. R. vulpina sp. nov. (SMNH). Modified hairs are indicated with arrows. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990903" httpUri="https://zenodo.org/record/13990903/files/figure.png" pageId="29" pageNumber="30">295</figureCitation>
. Scopula: entire and distal on metatarsi I–II; entire on tarsi I–II; narrowly divided by setae on tarsus III; widely divided on tarsus IV. Trichobothria: 2 rows of 8–9 each on tibiae, 12–14 on metatarsi, 10–11 on tarsi, 7–8 on cymbium. PTC I–IV with 9–10 and 7–8 teeth on inner and outer margins, respectively.
</paragraph>
<paragraph id="8BBE06355537FFA7FF52E40AFADCCD95" blockId="29.[189,1401,800,975]" pageId="29" pageNumber="30">
<smallCapsWord id="8D5890E95537FFA7FF52E40AFE9ECD62" baselines="819,820" box="[189,300,800,826]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Spination" pageId="29" pageNumber="30">SPINATION</smallCapsWord>
. All femora with 1–2 basodorsal spines and 3–4 dorsal bristles; metatarsus I and tarsi I–IV aspinose. Palp: femur pd2, rd1; patella pd1; tibia d1, p2, r3, v5; cymbium d4(5)+12–15 spikes. Leg I: femur pd3, rd3; patella p1; tibia p2, pv1, r2, rv2+ 
<specimenCount id="9D07CDBC5537FFA7FCD8E440FCD7CDDF" box="[823,869,874,900]" count="2" pageId="29" pageNumber="30" type="generic">2M.</specimenCount>
Leg II: femur pd3; patella p2; tibia p3, v7; metatarsus v5(4). Leg III: femur pd3, rd2; patella p2, r1; tibia d2(1), p4, r3, v7; metatarsus d3, p4, r3, v8. Leg IV: femur pd3(2), rd2(1); patella p1, r1; tibia d1, p3, r3, v9(7); metatarsus d3, p4(3), r5(4), v9.
</paragraph>
<paragraph id="8BBE06355537FFA7FF52E4D2FB31CA6C" blockId="29.[189,1398,1015,1079]" pageId="29" pageNumber="30">
<smallCapsWord id="8D5890E95537FFA7FF52E4D2FF46CA4B" baselines="1035,1035" box="[189,244,1016,1042]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Palp" pageId="29" pageNumber="30">PALP</smallCapsWord>
. Tibia, cymbium and copulatory bulb as shown in 
<figureCitation id="133A1AB05537FFA7FCBEE4D2FC0ACA49" box="[849,952,1015,1042]" captionStart="Figs 349–363" captionStartId="143.[189,241,1727,1753]" captionTargetBox="[189,1390,270,1690]" captionTargetId="figure-139@143.[189,1398,265,1690]" captionTargetPageId="143" captionText="Figs 349–363. Raveniola spp., ♂♂, holotypes (except 351, 355, 363), palpal tibia, cymbium and copulatory bulb, retrolateral (349–358, 360–361, 363) and retroventral (359, 362) views.349. R. caudata Zonstein, 2009 (SMNH). 350. R. inopinata sp. nov. (SMNH). 351. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 352. R. alajensis sp. nov. (SMNH). 353. R. concolor Zonstein, 2000 (MCSN AR 18). 354. R. cucullata sp. nov. (SMNH). 355. R. cucullata, paratype from Shahob (SMNH). 356. R. dolosa sp. nov. (SMNH). 357. R. hirta sp. nov. (SMNH). 358–359. R. ignobilis sp. nov. (SMNH). 360. R. insolita sp. nov. (ZISP). 361. R. ornata sp. nov. (SMNH). 362. R. ornatula sp. nov. (SMNH). 363. R. pamira sp. nov., paratype (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990913" httpUri="https://zenodo.org/record/13990913/files/figure.png" pageId="29" pageNumber="30">Fig. 356</figureCitation>
. Embolus with moderately long basal portion provided with very low keel and short, hooked apical part (
<figureCitation id="133A1AB05537FFA7FC20E337FBC5CA6C" box="[975,1143,1053,1079]" captionStart="Figs 389–399" captionStartId="146.[189,241,1854,1880]" captionTargetBox="[189,1398,265,1814]" captionTargetId="figure-68@146.[189,1398,265,1814]" captionTargetPageId="146" captionText="Figs 389–399. Raveniola spp., ♂♂, holotypes (except 392–396), details of copulatoty bulb, retrolateral (389, 392, 395, 397), retroventral (390, 393, 398) and ventral (391, 394, 396, 399) views. 389– 391. R. cucullata sp. nov. (SMNH). 392–394. R. cucullata sp. nov., paratype from Shahob (SMNH). 395–396. R. cucullata, paratype from Sangdara (SMNH). 397–399. R. dolosa sp. nov. (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990919" httpUri="https://zenodo.org/record/13990919/files/figure.png" pageId="29" pageNumber="30">Figs 397–399</figureCitation>
).
</paragraph>
<paragraph id="8BBE06355537FFA7FF52E34AFAA5CAC4" blockId="29.[189,1398,1120,1184]" pageId="29" pageNumber="30">
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. See Fig. 568. PMS: length 0.36, diameter 0.16. PLS: maximal diameter 0.49; length of basal, medial and apical segments 0.75, 0.43, 0.38; total length 1.56; apical segment triangular.
</paragraph>
<paragraph id="8BBE06355537FFA7FF52E3E3FE55CAB9" blockId="29.[189,487,1224,1251]" box="[189,487,1224,1251]" pageId="29" pageNumber="30">
<tableNote id="76E707BB5537FFA7FF52E3E3FE55CAB9" box="[189,487,1224,1251]" pageId="29" pageNumber="30" targetBox="[197,1267,1265,1469]" targetPageId="29">
<smallCapsWord id="8D5890E95537FFA7FF52E3E3FF58CABA" baselines="1244,1244" box="[189,234,1225,1251]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Leg" pageId="29" pageNumber="30">LEG</smallCapsWord>
<smallCapsWord id="8D5890E95537FFA7FF1DE3E4FE28CABA" baselines="1244" box="[242,410,1230,1249]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="measurements" pageId="29" pageNumber="30">MEASUREMENTS</smallCapsWord>
. ♂(♀)
</tableNote>
</paragraph>
<paragraph id="8BBE06355537FFA7FEBCE3DBFB41CBE6" pageId="29" pageNumber="30">
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<emphasis id="B975DA275537FFA7FEBCE3DBFE1ACB50" bold="true" box="[339,424,1265,1291]" pageId="29" pageNumber="30">Femur</emphasis>
</th>
<th id="76E06D0B55370045FE34E3DBFDEECB50" box="[475,604,1265,1291]" gridcol="2" gridrow="0" pageId="29" pageNumber="30">
<emphasis id="B975DA275537FFA7FE1FE3DBFDF4CB50" bold="true" box="[496,582,1265,1291]" pageId="29" pageNumber="30">Patella</emphasis>
</th>
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<emphasis id="B975DA275537FFA7FD76E3DBFD69CB50" bold="true" box="[665,731,1265,1291]" pageId="29" pageNumber="30">Tibia</emphasis>
</th>
<th id="76E06D0B55370045FCFEE3DBFC2DCB50" box="[785,927,1265,1291]" gridcol="4" gridrow="0" pageId="29" pageNumber="30">
<emphasis id="B975DA275537FFA7FCFEE3DBFC2DCB50" bold="true" box="[785,927,1265,1291]" pageId="29" pageNumber="30">Metatarsus</emphasis>
</th>
<th id="76E06D0B55370045FC59E3DBFB8ACB50" box="[950,1080,1265,1291]" gridcol="5" gridrow="0" pageId="29" pageNumber="30">
<emphasis id="B975DA275537FFA7FC22E3DBFB93CB50" bold="true" box="[973,1057,1265,1291]" pageId="29" pageNumber="30">Tarsus</emphasis>
</th>
<th id="76E06D0B55370045FBBBE3DBFB41CB50" box="[1108,1267,1265,1291]" gridcol="6" gridrow="0" pageId="29" pageNumber="30">
<emphasis id="B975DA275537FFA7FB6BE3DBFB76CB50" bold="true" box="[1156,1220,1265,1291]" pageId="29" pageNumber="30">Total</emphasis>
</th>
</tr>
<tr id="3531047755370045FF2AE23EFB41CB74" box="[197,1267,1300,1327]" gridrow="1" pageId="29" pageNumber="30">
<th id="76E06D0B55370045FF2AE23EFEAACB74" box="[197,280,1300,1327]" gridcol="0" gridrow="1" pageId="29" pageNumber="30">Palp</th>
<td id="76E06D0B55370045FED3E23EFE0FCB74" box="[316,445,1300,1327]" gridcol="1" gridrow="1" pageId="29" pageNumber="30">3.13 (2.79)</td>
<td id="76E06D0B55370045FE34E23EFDEECB74" box="[475,604,1300,1327]" gridcol="2" gridrow="1" pageId="29" pageNumber="30">1.61 (1.58)</td>
<td id="76E06D0B55370045FD96E23EFD49CB74" box="[633,763,1300,1327]" gridcol="3" gridrow="1" pageId="29" pageNumber="30">2.49 (1.89)</td>
<td id="76E06D0B55370045FCFEE23EFC2DCB74" box="[785,927,1300,1327]" gridcol="4" gridrow="1" pageId="29" pageNumber="30">–</td>
<td id="76E06D0B55370045FC59E23EFB8ACB74" box="[950,1080,1300,1327]" gridcol="5" gridrow="1" pageId="29" pageNumber="30">0.83 (1.92)</td>
<td id="76E06D0B55370045FBBBE23EFB41CB74" box="[1108,1267,1300,1327]" gridcol="6" gridrow="1" pageId="29" pageNumber="30">8.06 (8.18)</td>
</tr>
<tr id="3531047755370045FF2AE212FB41CB08" box="[197,1267,1336,1363]" gridrow="2" pageId="29" pageNumber="30">
<th id="76E06D0B55370045FF2AE212FEAACB08" box="[197,280,1336,1363]" gridcol="0" gridrow="2" pageId="29" pageNumber="30">Leg I</th>
<td id="76E06D0B55370045FED3E212FE0FCB08" box="[316,445,1336,1363]" gridcol="1" gridrow="2" pageId="29" pageNumber="30">5.65 (3.70)</td>
<td id="76E06D0B55370045FE34E212FDEECB08" box="[475,604,1336,1363]" gridcol="2" gridrow="2" pageId="29" pageNumber="30">2.94 (2.43)</td>
<td id="76E06D0B55370045FD96E212FD49CB08" box="[633,763,1336,1363]" gridcol="3" gridrow="2" pageId="29" pageNumber="30">4.32 (2.88)</td>
<td id="76E06D0B55370045FCFEE212FC2DCB08" box="[785,927,1336,1363]" gridcol="4" gridrow="2" pageId="29" pageNumber="30">4.07 (2.21)</td>
<td id="76E06D0B55370045FC59E212FB8ACB08" box="[950,1080,1336,1363]" gridcol="5" gridrow="2" pageId="29" pageNumber="30">2.43 (1.71)</td>
<td id="76E06D0B55370045FBBBE212FB41CB08" box="[1108,1267,1336,1363]" gridcol="6" gridrow="2" pageId="29" pageNumber="30">19.41 (12.93)</td>
</tr>
<tr id="3531047755370045FF2AE271FB41CB2D" box="[197,1267,1371,1398]" gridrow="3" pageId="29" pageNumber="30">
<th id="76E06D0B55370045FF2AE271FEAACB2D" box="[197,280,1371,1398]" gridcol="0" gridrow="3" pageId="29" pageNumber="30">Leg II</th>
<td id="76E06D0B55370045FED3E271FE0FCB2D" box="[316,445,1371,1398]" gridcol="1" gridrow="3" pageId="29" pageNumber="30">5.26 (3.36)</td>
<td id="76E06D0B55370045FE34E271FDEECB2D" box="[475,604,1371,1398]" gridcol="2" gridrow="3" pageId="29" pageNumber="30">2.58 (2.16)</td>
<td id="76E06D0B55370045FD96E271FD49CB2D" box="[633,763,1371,1398]" gridcol="3" gridrow="3" pageId="29" pageNumber="30">3.94 (2.56)</td>
<td id="76E06D0B55370045FCFEE271FC2DCB2D" box="[785,927,1371,1398]" gridcol="4" gridrow="3" pageId="29" pageNumber="30">3.93 (2.20)</td>
<td id="76E06D0B55370045FC59E271FB8ACB2D" box="[950,1080,1371,1398]" gridcol="5" gridrow="3" pageId="29" pageNumber="30">2.29 (1.79)</td>
<td id="76E06D0B55370045FBBBE271FB41CB2D" box="[1108,1267,1371,1398]" gridcol="6" gridrow="3" pageId="29" pageNumber="30">18.02 (12.07)</td>
</tr>
<tr id="3531047755370045FF2AE254FB41CBC2" box="[197,1267,1406,1433]" gridrow="4" pageId="29" pageNumber="30">
<th id="76E06D0B55370045FF2AE254FEAACBC2" box="[197,280,1406,1433]" gridcol="0" gridrow="4" pageId="29" pageNumber="30">Leg III</th>
<td id="76E06D0B55370045FED3E254FE0FCBC2" box="[316,445,1406,1433]" gridcol="1" gridrow="4" pageId="29" pageNumber="30">4.35 (3.09)</td>
<td id="76E06D0B55370045FE34E254FDEECBC2" box="[475,604,1406,1433]" gridcol="2" gridrow="4" pageId="29" pageNumber="30">2.07 (1.81)</td>
<td id="76E06D0B55370045FD96E254FD49CBC2" box="[633,763,1406,1433]" gridcol="3" gridrow="4" pageId="29" pageNumber="30">2.96 (2.04)</td>
<td id="76E06D0B55370045FCFEE254FC2DCBC2" box="[785,927,1406,1433]" gridcol="4" gridrow="4" pageId="29" pageNumber="30">4.14 (2.72)</td>
<td id="76E06D0B55370045FC59E254FB8ACBC2" box="[950,1080,1406,1433]" gridcol="5" gridrow="4" pageId="29" pageNumber="30">2.14 (1.87)</td>
<td id="76E06D0B55370045FBBBE254FB41CBC2" box="[1108,1267,1406,1433]" gridcol="6" gridrow="4" pageId="29" pageNumber="30">15.66 (11.53)</td>
</tr>
<tr id="3531047755370045FF2AE288FB41CBE6" box="[197,1267,1442,1469]" gridrow="5" pageId="29" pageNumber="30">
<th id="76E06D0B55370045FF2AE288FEAACBE6" box="[197,280,1442,1469]" gridcol="0" gridrow="5" pageId="29" pageNumber="30">Leg IV</th>
<td id="76E06D0B55370045FED3E288FE0FCBE6" box="[316,445,1442,1469]" gridcol="1" gridrow="5" pageId="29" pageNumber="30">5.46 (3.98)</td>
<td id="76E06D0B55370045FE34E288FDEECBE6" box="[475,604,1442,1469]" gridcol="2" gridrow="5" pageId="29" pageNumber="30">2.46 (2.18)</td>
<td id="76E06D0B55370045FD96E288FD49CBE6" box="[633,763,1442,1469]" gridcol="3" gridrow="5" pageId="29" pageNumber="30">4.08 (3.31)</td>
<td id="76E06D0B55370045FCFEE288FC2DCBE6" box="[785,927,1442,1469]" gridcol="4" gridrow="5" pageId="29" pageNumber="30">5.82 (3.66)</td>
<td id="76E06D0B55370045FC59E288FB8ACBE6" box="[950,1080,1442,1469]" gridcol="5" gridrow="5" pageId="29" pageNumber="30">2.78 (2.04)</td>
<td id="76E06D0B55370045FBBBE288FB41CBE6" box="[1108,1267,1442,1469]" gridcol="6" gridrow="5" pageId="29" pageNumber="30">20.60 (15.17)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BBE06355537FFA7FF05E2C9FE77CBA6" blockId="29.[189,453,1507,1571]" box="[234,453,1507,1533]" pageId="29" pageNumber="30">
<emphasis id="B975DA275537FFA7FF05E2C9FEF7CBA6" bold="true" box="[234,325,1507,1533]" pageId="29" pageNumber="30">Female</emphasis>
(
<typeStatus id="54BAB8975537FFA7FEBAE2C9FE0CCBA6" box="[341,446,1507,1533]" pageId="29" pageNumber="30" type="paratype">paratype</typeStatus>
)
</paragraph>
<paragraph id="8BBE06355537FFA7FF52E123FE05C879" blockId="29.[189,453,1507,1571]" box="[189,439,1544,1571]" pageId="29" pageNumber="30">
<smallCapsWord id="8D5890E95537FFA7FF52E123FEAEC87A" baselines="1564,1564" box="[189,284,1545,1571]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Habitus" pageId="29" pageNumber="30">HABITUS</smallCapsWord>
. See 
<figureCitation id="133A1AB05537FFA7FEB2E122FE05C879" box="[349,439,1544,1570]" captionStart="Figs 36–44" captionStartId="111.[189,241,1837,1863]" captionTargetBox="[201,1398,270,1792]" captionTargetId="figure-83@111.[189,1398,265,1792]" captionTargetPageId="111" captionText="Figs 36–44. Raveniola spp., ♀♀, paratypes (except 36–37, 44), habitus, dorsal views. 36. R. redikorzevi (Spassky, 1937), from Kepele (SMNH). 37. R. afghana sp. nov., holotype (NHMD). 38. R. alajensis sp. nov. (SMNH). 39–40. R. cucullata sp. nov. from Shikildara (SMNH) and Childara (SMNH), respectively. 41. R. dolosa sp. nov. (SMNH). 42. R. hirta sp. nov. (SMNH). 43. R. ignobilis sp. nov. (SMNH). 44. R. karategensis sp. nov., holotype (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990834" httpUri="https://zenodo.org/record/13990834/files/figure.png" pageId="29" pageNumber="30">Fig. 41.</figureCitation>
</paragraph>
<paragraph id="8BBE06355537FFA7FF52E166FB14C83E" blockId="29.[189,1190,1611,1638]" box="[189,1190,1611,1638]" pageId="29" pageNumber="30">
<smallCapsWord id="8D5890E95537FFA7FF52E166FEDEC83F" baselines="1631,1631" box="[189,364,1612,1638]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="29" pageNumber="30">MEASUREMENTS</smallCapsWord>
. TBL 15.60, CL 5.26, CW 4.53, LL 0.87, LW 1.11, SL 2.76, SW 2.48.
</paragraph>
<paragraph id="8BBE06355537FFA7FF52E1A5FC52C8F3" blockId="29.[189,992,1678,1705]" box="[189,992,1678,1705]" pageId="29" pageNumber="30">
<smallCapsWord id="8D5890E95537FFA7FF52E1A5FEAAC8FC" baselines="1698,1698" box="[189,280,1679,1705]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Colour" pageId="29" pageNumber="30">COLOUR</smallCapsWord>
. Similar to that of male, but carapace and legs slightly paler.
</paragraph>
<paragraph id="8BBE06355537FFA7FF52E1F8FC7BC9DB" blockId="29.[187,1399,1746,1921]" pageId="29" pageNumber="30">
<smallCapsWord id="8D5890E95537FFA7FF52E1F8FECAC8B1" baselines="1765,1765" box="[189,376,1746,1772]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Cephalothorax" pageId="29" pageNumber="30">CEPHALOTHORAX</smallCapsWord>
. Carapace and chelicerae as shown in 
<figureCitation id="133A1AB05537FFA7FCC3E1F8FC3CC8B7" box="[812,910,1746,1772]" captionStart="Figs 109–117" captionStartId="120.[189,241,1835,1861]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-93@120.[189,1398,265,1816]" captionTargetPageId="120" captionText="Figs 109–117. Raveniola spp., ♂ (109) and ♀♀ (110–117), paratypes (except 109–111). Cephalothorax, dorsal views. 109. R. vulpina sp. nov., from Tereksai (SMNH). 110. R. redikorzevi (Spassky, 1937), ♀ from Kepele (SMNH). 111. R. afghana sp. nov., holotype (NHMD). 112. R. alajensis sp. nov. (SMNH). 113. R. concolor Zonstein, 2000 (MCSN AR 19). 114–115. R. cucullata sp. nov. from Mullokoni and Shahob, respectively (both SMNH). 116. R. dolosa sp. nov. (SMNH). 117. R. hirta sp. nov. (SMNH). Scale bars: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990858" httpUri="https://zenodo.org/record/13990858/files/figure.png" pageId="29" pageNumber="30">Fig. 116</figureCitation>
. Clypeus and eye group as in 
<figureCitation id="133A1AB05537FFA7FB32E1F8FA8DC8B7" box="[1245,1343,1746,1772]" captionStart="Figs 172–183" captionStartId="126.[189,241,1749,1775]" captionTargetBox="[189,1399,265,1701]" captionTargetId="figure-107@126.[189,1399,265,1701]" captionTargetPageId="126" captionText="Figs 172–183. Raveniola spp., ♀♀, paratypes (except 176, 181, 183), clypeus and eye tubercle, dorsal views. 172. R. dolosa sp. nov. (SMNH). 173–174. R. hirta sp. nov. (SMNH). 175. R. ignobilis sp. nov. (SMNH). 176. R. karategensis sp. nov., holotype (SMNH). 177. R. karategensis, paratype (SMNH). 178. R. ornata sp. nov., from Sanglok (SMNH). 179. R. ornatula sp. nov. (SMNH). 180. R. pamira sp. nov. (SMNH). 181. R. sororcula sp. nov., holotype (SMNH). 182. R. diluta sp. nov. (SMNH). 183. R. fedotovi (Charitonov, 1946), ♀ from Hojakurgan (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990872" httpUri="https://zenodo.org/record/13990872/files/figure.png" pageId="29" pageNumber="30">Fig. 172</figureCitation>
. Eye diameters and interdistances: AME 0.13(0.18), ALE 0.25, PLE 0.18, PME 0.16; AME–AME 0.12(0.07), ALE–AME 0.10(0.07), ALE–PLE 0.09, PLE–PME 0.05, PME–PME 0.35. Cheliceral rastellum absent. Each cheliceral furrow with 9 promarginal teeth and 9–10 mesobasal denticles. Sternum, labium and maxillae as shown in 
<figureCitation id="133A1AB05537FFA7FE52E04CFD91C9DA" box="[445,547,1894,1921]" captionStart="Figs 229–237" captionStartId="132.[189,241,1827,1853]" captionTargetBox="[200,1385,265,1784]" captionTargetId="figure-90@132.[200,1385,265,1784]" captionTargetPageId="132" captionText="Figs 229–237. Raveniola spp., ♀♀, paratypes (except 229–230), sternum, labium and maxillae, ventral views. 229. R. redikorzevi (Spassky, 1937), ♀ from Kepele (SMNH). 230. R. afghana sp. nov., holotype (NHMD). 231. R. alajensis sp. nov. (SMNH). 232. R. concolor Zonstein, 2000 (MCSN AR 19). 233– 234. R. cucullata sp. nov. from Shikildara (SMNH) and Shahob (SMNH), respectively. 235. R. dolosa sp. nov.(SMNH). 236. R. hirta sp. nov. (SMNH). 237. R. ignobilis sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990884" httpUri="https://zenodo.org/record/13990884/files/figure.png" pageId="29" pageNumber="30">Fig. 235</figureCitation>
. Maxillae with ca 70 cuspules each.
</paragraph>
<paragraph id="8BBE06355537FFA7FF52E080FAC5C9B3" blockId="29.[189,1399,1961,2025]" pageId="29" pageNumber="30">
<smallCapsWord id="8D5890E95537FFA7FF52E080FF44C999" baselines="1981,1981" box="[189,246,1962,1988]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Legs" pageId="29" pageNumber="30">LEGS</smallCapsWord>
. Scopula: distal on metatarsi I–II; entire on palpal tarsus and tarsus I; widely divided by setae on tarsus II; rudimentary on tarsi III–IV. Trichobothria: 2 rows of 11–12 each on tibiae, 16–18 on metatarsi;
</paragraph>
<paragraph id="8BBE06355534FFA4FF52E623FD0BCF13" blockId="30.[189,1398,265,329]" pageId="30" pageNumber="31">13–15 on tarsi; 11 on palpal tarsus. Palpal claw with 5 promarginal teeth. PTC I–II and III–IV with 4–5 and 5–7 teeth on each margin, respectively.</paragraph>
<paragraph id="8BBE06355534FFA4FF52E658FEB4CC7A" blockId="30.[188,1400,370,545]" pageId="30" pageNumber="31">
<smallCapsWord id="8D5890E95534FFA4FF52E658FE9ECFD1" baselines="389,389" box="[189,300,370,396]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Spination" pageId="30" pageNumber="31">SPINATION</smallCapsWord>
. All femora with one basodorsal spine and 3 dorsal bristles; palpal patella, patellae I–II, and tarsi I–IV aspinose. Palp: femur pd1; tibia p1, v7; tarsus v3. Leg I: femur pd1; tibia v6; metatarsus v6. Leg II: femur pd1; tibia p2, v6; metatarsus v6. Leg III: femur pd2, rd2; patella p2, r1; tibia d1, p2, r2(1), v7; metatarsus d3(2), p3, r3, v9(8). Leg IV: femur rd1; patella r1; tibia p2, r2, v7; metatarsus p3, r3, v8(7).
</paragraph>
<paragraph id="8BBE06355534FFA4FF52E561FC04CCD0" blockId="30.[189,1398,587,651]" pageId="30" pageNumber="31">
<smallCapsWord id="8D5890E95534FFA4FF52E561FED0CC38" baselines="606,606" box="[189,354,587,613]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Spermathecae" pageId="30" pageNumber="31">SPERMATHECAE</smallCapsWord>
. Each of paired spermathecae V-shaped with relatively low and wide base carrying two densely located, short and weakly diverging branches (
<figureCitation id="133A1AB05534FFA4FCADE55AFC18CCD0" box="[834,938,624,651]" captionStart="Figs 487–503" captionStartId="156.[189,241,1695,1721]" captionTargetBox="[189,1398,265,1653]" captionTargetId="figure-102@156.[189,1398,265,1653]" captionTargetPageId="156" captionText="Figs 487–503. Raveniola spp., ♀♀, paratypes (except 487–489), spermathecae, dorsal views. 487–488. R. redikorzevi (Spassky, 1937), Kepele (SMNH). 489. R. afghana sp. nov. (NHMD). 490. Same, holotype. 491–493. R. alajensis sp. nov., Berksu. 494. R. concolor Zonstein, 2000 (MCSN AR 19). 495–496. R. cucullata sp. nov., Shikildara (SMNH). 497. R. cucullata, Shahob (SMNH). 498– 499. R. cucullata, Sangdara (SMNH). 500–503. R. dolosa sp. nov. (SMNH). Abbreviations: isb = inner spermathecal branch; osb = outer spermathecal branch; sb = spermathecal base. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990941" httpUri="https://zenodo.org/record/13990941/files/figure.png" pageId="30" pageNumber="31">Fig. 500</figureCitation>
).
</paragraph>
<paragraph id="8BBE06355534FFA4FF52E59EFAA4CCA8" blockId="30.[188,1398,692,756]" pageId="30" pageNumber="31">
<smallCapsWord id="8D5890E95534FFA4FF52E59EFE8ECC97" baselines="711,711" box="[189,316,692,718]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Spinnerets" pageId="30" pageNumber="31">SPINNERETS</smallCapsWord>
. See Figs 569–570. PMS: length 0.47, diameter 0.15. PLS: maximal diameter 0.57; length of basal, medial and apical segments 0.98, 0.42, 0.43; total length 1.83; apical segment triangular.
</paragraph>
<paragraph id="8BBE06355534FFA4FF52E436FE81CD6D" blockId="30.[189,307,796,822]" box="[189,307,796,822]" pageId="30" pageNumber="31">
<heading id="D0F6B1595534FFA4FF52E436FE81CD6D" bold="true" box="[189,307,796,822]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B975DA275534FFA4FF52E436FE81CD6D" bold="true" box="[189,307,796,822]" pageId="30" pageNumber="31">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE06355534FFA4FF52E464FD7BCDD5" blockId="30.[188,1399,845,910]" pageId="30" pageNumber="31">
Carapace length varies in females (n =9) from 4.30 to 6.17. Variations in the habitus and structure of the spermathecae as shown in 
<figureCitation id="133A1AB05534FFA4FE1AE459FDFDCDD6" box="[501,591,883,909]" captionStart="Figs 69–75" captionStartId="115.[189,241,1831,1857]" captionTargetBox="[191,1400,265,1794]" captionTargetId="figure-66@115.[191,1400,265,1794]" captionTargetPageId="115" captionText="Figs 69–75. Raveniola spp., ♀♀ (69, 71–72, 74) and ♂♂ (70, 73, 75), live spiders; predominantly dorsal (69, 71–72, 74–75) and dorsolateral (70, 73) views. 69–70. R. cucullata sp. nov. (SMNH). 71. R. dolosa sp. nov. (SMNH).72. R. ornatasp. nov. (SMNH). 73–74. R. zyuzinisp. nov. (SMNH). 75. R. ferghanensis (Zonstein, 1984) (SMNH)." figureDoi="http://doi.org/10.5281/zenodo.13990842" httpUri="https://zenodo.org/record/13990842/files/figure.png" pageId="30" pageNumber="31">Figs 71</figureCitation>
, 
<figureCitation id="133A1AB05534FFA4FDB4E459FD74CDD5" box="[603,710,883,910]" captionStart="Figs 487–503" captionStartId="156.[189,241,1695,1721]" captionTargetBox="[189,1398,265,1653]" captionTargetId="figure-102@156.[189,1398,265,1653]" captionTargetPageId="156" captionText="Figs 487–503. Raveniola spp., ♀♀, paratypes (except 487–489), spermathecae, dorsal views. 487–488. R. redikorzevi (Spassky, 1937), Kepele (SMNH). 489. R. afghana sp. nov. (NHMD). 490. Same, holotype. 491–493. R. alajensis sp. nov., Berksu. 494. R. concolor Zonstein, 2000 (MCSN AR 19). 495–496. R. cucullata sp. nov., Shikildara (SMNH). 497. R. cucullata, Shahob (SMNH). 498– 499. R. cucullata, Sangdara (SMNH). 500–503. R. dolosa sp. nov. (SMNH). Abbreviations: isb = inner spermathecal branch; osb = outer spermathecal branch; sb = spermathecal base. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990941" httpUri="https://zenodo.org/record/13990941/files/figure.png" pageId="30" pageNumber="31">501–503</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE5534FFA4FF52E49CFBA1CA7C" pageId="30" pageNumber="31" type="biology_ecology">
<paragraph id="8BBE06355534FFA4FF52E49CFE92CD8B" blockId="30.[189,288,950,976]" box="[189,288,950,976]" pageId="30" pageNumber="31">
<heading id="D0F6B1595534FFA4FF52E49CFE92CD8B" bold="true" box="[189,288,950,976]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B975DA275534FFA4FF52E49CFE92CD8B" bold="true" box="[189,288,950,976]" pageId="30" pageNumber="31">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE06355534FFA4FF52E4CDFBA1CA7C" blockId="30.[189,1398,999,1063]" pageId="30" pageNumber="31">
All spiders were found hiding in soil cavities under stones in riverside woodlands, dominated by 
<taxonomicName id="4C017DB65534FFA4FF52E327FE52CA7D" box="[189,480,1036,1062]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="seravschanica">
<emphasis id="B975DA275534FFA4FF52E327FE52CA7D" box="[189,480,1036,1062]" italics="true" pageId="30" pageNumber="31">Juniperus seravschanica</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C017DB65534FFA4FE01E326FD3CCA7D" box="[494,654,1036,1062]" class="Magnoliopsida" family="Juglandaceae" genus="Juglans" kingdom="Plantae" order="Fagales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="regia">
<emphasis id="B975DA275534FFA4FE01E326FD3CCA7D" box="[494,654,1036,1062]" italics="true" pageId="30" pageNumber="31">Juglans regia</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C017DB65534FFA4FD28E326FCE2CA7C" box="[711,848,1036,1063]" class="Magnoliopsida" family="Salicaceae" genus="Populus" kingdom="Plantae" order="Malpighiales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B975DA275534FFA4FD28E326FC95CA7D" box="[711,807,1036,1062]" italics="true" pageId="30" pageNumber="31">Populus</emphasis>
sp.
</taxonomicName>
(
<figureCitation id="133A1AB05534FFA4FCB0E326FBB5CA7C" box="[863,1031,1036,1063]" captionStart="Figs 643–650" captionStartId="170.[189,241,1816,1842]" captionTargetBox="[189,1399,265,1784]" captionTargetId="figure-37@170.[189,1399,265,1784]" captionTargetPageId="170" captionText="Figs 643–650. Raveniola spp., habitats of two sympatric congeners, R. cucullata sp. nov. and R. dolosa sp. nov. (643–646), and retreats used by representatives of the latter species (647–650). Tajikistan, Peter I Mts, surroundings of Shahob." figureDoi="http://doi.org/10.5281/zenodo.13990957" httpUri="https://zenodo.org/record/13990957/files/figure.png" pageId="30" pageNumber="31">Figs 643–650</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE5534FFA4FF52E37AFD49CAC0" box="[189,763,1104,1179]" pageId="30" pageNumber="31" type="distribution">
<paragraph id="8BBE06355534FFA4FF52E37AFEE4CA31" blockId="30.[189,342,1104,1130]" box="[189,342,1104,1130]" pageId="30" pageNumber="31">
<heading id="D0F6B1595534FFA4FF52E37AFEE4CA31" bold="true" box="[189,342,1104,1130]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B975DA275534FFA4FF52E37AFEE4CA31" bold="true" box="[189,342,1104,1130]" pageId="30" pageNumber="31">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE06355534FFA4FF52E3ABFD49CAC0" blockId="30.[189,763,1152,1179]" box="[189,763,1152,1179]" pageId="30" pageNumber="31">
Known only from the 
<typeStatus id="54BAB8975534FFA4FE2DE3ABFE47CAC0" box="[450,501,1153,1179]" pageId="30" pageNumber="31">type</typeStatus>
locality. See 
<figureCitation id="133A1AB05534FFA4FD7DE3ABFD46CAC0" box="[658,756,1152,1179]" captionStart="Figs 751–760" captionStartId="184.[189,241,1662,1688]" captionTargetBox="[258,1323,265,1629]" captionTargetId="figure-247@184.[258,1323,265,1629]" captionTargetPageId="184" captionText="Figs 751–760. Distribution records of Raveniola spp. 751. R. alajensis sp. nov. (), R. cucullata" figureDoi="http://doi.org/10.5281/zenodo.13990987" httpUri="https://zenodo.org/record/13990987/files/figure.png" pageId="30" pageNumber="31">Fig. 751</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>