567 lines
125 KiB
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567 lines
125 KiB
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<document id="C6E1360D516EC2860101B1CE8A9E767F" ID-CLB-Dataset="298857" ID-DOI="10.11646/zootaxa.5471.3.5" ID-GBIF-Dataset="0c4363d9-e2b8-4d39-9121-bc268e1fe3a6" ID-ISSN="1175-5326" ID-Zenodo-Dep="12190170" ID-ZooBank="C7E5ADC9-EBD7-455D-A0C3-7109FBC52AD3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718893521820" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2024" docId="03F80A59FFB7E92DFF2FFB26FD39FAAA" docLanguage="en" docName="zootaxa.5471.3.5.pdf" docOrigin="Zootaxa 5471 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5471.3.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Keratodegnathus javanicus Jaloszynski 2024, sp. n." docType="treatment" docVersion="6" lastPageNumber="363" masterDocId="FFC17221FFB6E925FFB8FFA0FF91FFD3" masterDocTitle="A new species of Keratodegnathus Pace in Indonesia (Coleoptera, Staphylinidae)" masterLastPageNumber="364" masterPageNumber="355" pageNumber="356" updateTime="1719239874310" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="16C8513992A4A8049B5E905997DA7D20">A new species of Keratodegnathus Pace in Indonesia (Coleoptera, Staphylinidae)</mods:title>
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<mods:namePart id="30E7225B193748838DD9B35BF69D4BAD">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="E37787731895C673C1659B23D1AD3B21">Zootaxa</mods:title>
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<mods:part id="6F66E780C7F078B14C2B575DC364377A">
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<mods:date id="FD6BE0031D0AA8FECC122333F70A3742">2024</mods:date>
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<mods:number id="B3BC0AD621A66378AE4A91017AB237EE">2024-06-19</mods:number>
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<mods:number id="37062E315D24B6309FAF13CB87BFF7B3">3</mods:number>
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<mods:url id="D8C1D0F8126173CB94F3925F77F2FD48">http://dx.doi.org/10.11646/zootaxa.5471.3.5</mods:url>
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<treatment id="03F80A59FFB7E92DFF2FFB26FD39FAAA" ID-DOI="http://doi.org/10.5281/zenodo.12190832" ID-GBIF-Taxon="233743931" ID-Zenodo-Dep="12190832" LSID="urn:lsid:plazi:treatment:03F80A59FFB7E92DFF2FFB26FD39FAAA" httpUri="http://treatment.plazi.org/id/03F80A59FFB7E92DFF2FFB26FD39FAAA" lastPageId="8" lastPageNumber="363" pageId="1" pageNumber="356">
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<subSubSection id="C34BE8C4FFB7E924FF2FFB26FD53FB72" box="[151,706,1158,1185]" pageId="1" pageNumber="356" type="nomenclature">
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<paragraph id="8BEEBB4FFFB7E924FF2FFB26FD53FB72" blockId="1.[151,706,1158,1221]" box="[151,706,1158,1185]" pageId="1" pageNumber="356">
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<heading id="D0A60C23FFB7E924FF2FFB26FD53FB72" bold="true" box="[151,706,1158,1185]" fontSize="11" level="1" pageId="1" pageNumber="356" reason="1">
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<emphasis id="B925675DFFB7E924FF2FFB26FD53FB72" bold="true" box="[151,706,1158,1185]" pageId="1" pageNumber="356">
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<taxonomicName id="4C51C0CCFFB7E924FF2FFB26FDE9FB72" authority="Jaloszynski" authorityName="Jaloszynski" authorityYear="2024" box="[151,632,1158,1185]" class="Insecta" family="Staphylinidae" genus="Keratodegnathus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="356" phylum="Arthropoda" rank="species" species="javanicus" status="sp. nov.">
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<emphasis id="B925675DFFB7E924FF2FFB26FE73FB72" bold="true" box="[151,482,1158,1185]" italics="true" pageId="1" pageNumber="356">Keratodegnathus javanicus</emphasis>
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Jałoszyński
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</taxonomicName>
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<taxonomicNameLabel id="A216DA26FFB7E924FDC7FB27FD53FB72" box="[639,706,1159,1185]" pageId="1" pageNumber="356" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34BE8C4FFB7E924FF2FFB0BFE88FB16" box="[151,281,1195,1221]" pageId="1" pageNumber="356" type="description">
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<paragraph id="8BEEBB4FFFB7E924FF2FFB0BFE88FB16" blockId="1.[151,706,1158,1221]" box="[151,281,1195,1221]" pageId="1" pageNumber="356">
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(
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<figureCitation id="136AA7CAFFB7E924FF27FB0BFE80FB16" box="[159,273,1195,1221]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2‒5" captionStart-2="FIGURES 6‒10" captionStart-3="FIGURE 11" captionStart-4="FIGURES 12‒18" captionStart-5="FIGURES 19‒24" captionStartId-0="2.[151,250,1651,1675]" captionStartId-1="3.[151,264,1776,1800]" captionStartId-2="4.[151,264,1598,1622]" captionStartId-3="5.[151,250,1645,1669]" captionStartId-4="6.[151,264,1873,1897]" captionStartId-5="7.[151,264,1649,1673]" captionTargetBox-0="[203,1391,314,1617]" captionTargetBox-1="[177,1411,453,1752]" captionTargetBox-2="[177,1411,181,1574]" captionTargetBox-3="[151,1435,273,1622]" captionTargetBox-4="[151,1435,273,1850]" captionTargetBox-5="[164,1417,188,1625]" captionTargetId-0="figure-88@2.[189,1398,306,1625]" captionTargetId-1="figure-149@3.[177,1411,453,1752]" captionTargetId-2="figure-19@4.[177,1411,181,1574]" captionTargetId-3="figure-62@5.[151,1435,273,1622]" captionTargetId-4="figure-67@6.[151,1435,273,1850]" captionTargetId-5="figure-15@7.[164,1423,181,1625]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionText-0="FIGURE 1. Keratodegnathus javanicus sp. n., holotype female, habitus in dorsal view." captionText-1="FIGURES 2‒5. Keratodegnathus javanicus sp. n., holotype female. Head and anterior region of thorax in lateral view (2); head in anterodorsal view (3); head, prothorax and elytral base in dorsal view (4; arrowheads indicate macrosetae); head and prothorax in ventral view (5). Abbreviations: acxc, anterior coxal carina; cl, clypeus; cmb, connecting membrane; cs, cervical sclerite; dmt, dorsal mandibular tooth; fr, frons; gal, galea; gen, gena; gp, gular plate; gr, genal carina; gs, gular suture; hr, hypostomal ridge; hs, hypostomal suture; hyp, hypomeron; hyr, hypomeral ridge; lac, lacinia; lbr, labrum; lpc, lateral pronotal carina; mn, mentum; mxp2‒4, maxillary palpomere 2‒4; nss, notosternal suture; ocr, occipital ridge; pd, pedicel; prl prelabium; pst, prosternum; ptp, posterior tentorial pit; sc, scape; smn, submentum; trt, trochantin; vt, vertex." captionText-2="FIGURES 6‒10. Keratodegnathus javanicus sp. n., holotype female. Head in anterodorsal view (6); mouthparts in dorsal view (7); anteromedian region of labrum in anterodorsal view (8); maxillo-labial complex in ventral view (9); distal region of prelabium in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; clp, clypeus; cmb, connecting membrane; dmt, dorsal mandibular tooth; fr, frons; gal, galea; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; md, mandible; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pmns, premental sclerite; prl, prelabium; smn, submentum." captionText-3="FIGURE 11. Keratodegnathus javanicus sp. n., holotype female, posterior region of thorax and abdomen in dorsal view. Abbreviations:asc, adsutural carina; bc, basal carina; go1‒3, glandular opening type 1‒3 (each indicated by different arrowhead); pt, paratergite; sp, spiracle; tIII‒X, tergite III‒X." captionText-4="FIGURES 12‒18. Keratodegnathus javanicus sp. n., holotype female. Thorax and abdomen in ventral view (12); protarsus in lateral (13) and lateroventral (14) views; mesotarsus in lateroventral view (15); metatarsus in lateral view (16); distal region of metatarsomere 4 in lateroventral view (17); details of claw insertion and empodium of metatarsus (18). Abbreviations: aest3, metanepisternum; es, empodial seta; mc, marginal carina; sIII‒IX, sternite III‒IX; v3, metaventrite." captionText-5="FIGURES 19‒24. Keratodegnathus javanicus sp. n., holotype female. Sternite VIII (19); tergite VIII (20); segment IX and X (21); mediobasal region of segments IX and X (22); spermatheca (23‒24). Abbreviations: tIX‒X, tergite IX‒X; vp, vaginal plate (sensu Maruyama (2006: fig. 16A))." figureDoi-0="http://doi.org/10.5281/zenodo.12190172" figureDoi-1=
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34BE8C4FFB7E924FF2FFB53FDCDFA86" pageId="1" pageNumber="356" type="materials_examined">
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<paragraph id="8BEEBB4FFFB7E924FF2FFB53FDCDFA86" blockId="1.[151,1437,1266,2013]" pageId="1" pageNumber="356">
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<emphasis id="B925675DFFB7E924FF2FFB53FED9FADF" bold="true" box="[151,328,1266,1293]" pageId="1" pageNumber="356">Type material.</emphasis>
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<materialsCitation id="3B39B112FFB7E924FEE9FB53FDCDFA86" ID-GBIF-Occurrence="4899229301" collectingDate="1997-05-24" collectingDateMax="1997-05-28" collectingDateMin="1997-05-24" collectionCode="MHNG" collectorName="S. Kurbatov" country="Indonesia" elevation="1450" location="Mt. Gede" pageId="1" pageNumber="356" specimenCount="1" specimenCount-female="1" stateProvince="West Java Province" typeStatus="holotype">
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<emphasis id="B925675DFFB7E924FEE9FB53FCCDFADF" bold="true" box="[337,860,1266,1293]" pageId="1" pageNumber="356">
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<typeStatus id="54EA05EDFFB7E924FEE9FB53FE53FADE" box="[337,450,1267,1293]" pageId="1" pageNumber="356" type="holotype">Holotype</typeStatus>
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:
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<collectingCountry id="F346FBDFFFB7E924FE6AFB52FDD6FADF" box="[466,583,1266,1292]" name="Indonesia" pageId="1" pageNumber="356">Indonesia</collectingCountry>
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(
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<collectingRegion id="499575ADFFB7E924FDEFFB52FCDEFADF" box="[599,847,1266,1293]" country="Indonesia" name="Jawa Barat" pageId="1" pageNumber="356">West Java Province</collectingRegion>
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):
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</emphasis>
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<specimenCount id="9D5770C6FFB7E924FCDEFB53FCEAFADE" box="[870,891,1267,1293]" count="1" pageId="1" pageNumber="356" type="female">♀</specimenCount>
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, two labels: “W Java,
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<location id="8E8EED94FFB7E924FB3DFB53FB63FADE" LSID="urn:lsid:plazi:treatment:03F80A59FFB7E92DFF2FFB26FD39FAAA:8E8EED94FFB7E924FB3DFB53FB63FADE" box="[1157,1266,1267,1293]" country="Indonesia" name="Mt. Gede" pageId="1" pageNumber="356" stateProvince="West Java Province">Mt. Gede</location>
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/
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<quantity id="4CA916AAFFB7E924FAB5FB52FA0DFADE" box="[1293,1436,1266,1293]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="1" pageNumber="356" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">
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<elevation id="007C5C7CFFB7E924FAB5FB52FA0DFADE" box="[1293,1436,1266,1293]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="1" pageNumber="356" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">1400-1500m</elevation>
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</quantity>
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/
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<date id="FFEF9D8FFFB7E924FF10FAB7FEA6FAE2" box="[168,311,1303,1329]" pageId="1" pageNumber="356" value="1997-05-24" valueMax="1997-05-28" valueMin="1997-05-24">
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<collectingDate id="EFAB6467FFB7E924FF10FAB7FEA6FAE2" box="[168,311,1303,1329]" pageId="1" pageNumber="356" value="1997-05-24" valueMax="1997-05-28" valueMin="1997-05-24">24-28. V. 97</collectingDate>
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</date>
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/ leg.
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<collectorName id="26A4DE99FFB7E924FE3CFAB7FD81FAE2" box="[388,528,1303,1329]" pageId="1" pageNumber="356">S. Kurbatov</collectorName>
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” [white, printed], “
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<taxonomicName id="4C51C0CCFFB7E924FD49FAB7FC6DFAE2" authorityName="Pace" authorityYear="2014" box="[753,1020,1303,1329]" class="Insecta" family="Staphylinidae" genus="Keratodegnathus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="356" phylum="Arthropoda" rank="genus">
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<emphasis id="B925675DFFB7E924FD49FAB7FC6DFAE2" box="[753,1020,1303,1329]" italics="true" pageId="1" pageNumber="356">KERATODEGNATHUS</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="4C51C0CCFFB7E924FBAEFAB7FB10FAE3" authorityName="Jaloszynski" authorityYear="2024" box="[1046,1153,1303,1328]" class="Insecta" family="Staphylinidae" genus="Keratodegnathus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="356" phylum="Arthropoda" rank="species" species="javanicus">
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<emphasis id="B925675DFFB7E924FBAEFAB7FB10FAE3" box="[1046,1153,1303,1328]" italics="true" pageId="1" pageNumber="356">javanicus</emphasis>
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</taxonomicName>
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m. / Jałoszyński, 2024 / HOLOTYPUS” [red, printed] (
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<collectionCode id="ED40238AFFB7E924FE4CFA9BFDC1FA86" box="[500,592,1339,1365]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1" pageNumber="356" type="Museum">MHNG</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34BE8C4FFB7E924FF7FFAFEFD1BFA36" pageId="1" pageNumber="356" type="diagnosis">
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<paragraph id="8BEEBB4FFFB7E924FF7FFAFEFD1BFA36" blockId="1.[151,1437,1266,2013]" pageId="1" pageNumber="356">
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<emphasis id="B925675DFFB7E924FF7FFAFEFED3FAAB" bold="true" box="[199,322,1374,1400]" pageId="1" pageNumber="356">Diagnosis.</emphasis>
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Female with dark brown head, elytra and distal region of abdomen, and with red pronotum and proximal region of abdomen (
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<figureCitation id="136AA7CAFFB7E924FE44FA23FDD3FA4E" box="[508,578,1411,1437]" captionStart="FIGURE 1" captionStartId="2.[151,250,1651,1675]" captionTargetBox="[203,1391,314,1617]" captionTargetId="figure-88@2.[189,1398,306,1625]" captionTargetPageId="2" captionText="FIGURE 1. Keratodegnathus javanicus sp. n., holotype female, habitus in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12190172" httpUri="https://zenodo.org/record/12190172/files/figure.png" pageId="1" pageNumber="356">Fig. 1</figureCitation>
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); antennomere 11 (
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<figureCitation id="136AA7CAFFB7E924FC95FA23FCE2FA4E" box="[813,883,1411,1437]" captionStart="FIGURE 1" captionStartId="2.[151,250,1651,1675]" captionTargetBox="[203,1391,314,1617]" captionTargetId="figure-88@2.[189,1398,306,1625]" captionTargetPageId="2" captionText="FIGURE 1. Keratodegnathus javanicus sp. n., holotype female, habitus in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12190172" httpUri="https://zenodo.org/record/12190172/files/figure.png" pageId="1" pageNumber="356">Fig. 1</figureCitation>
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) conspicuously elongate, twice as long as 10; spermatheca (
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<figureCitation id="136AA7CAFFB7E924FE8BFA07FE15FA12" box="[307,388,1447,1473]" captionStart="FIGURES 19‒24" captionStartId="7.[151,264,1649,1673]" captionTargetBox="[164,1417,188,1625]" captionTargetId="figure-15@7.[164,1423,181,1625]" captionTargetPageId="7" captionText="FIGURES 19‒24. Keratodegnathus javanicus sp. n., holotype female. Sternite VIII (19); tergite VIII (20); segment IX and X (21); mediobasal region of segments IX and X (22); spermatheca (23‒24). Abbreviations: tIX‒X, tergite IX‒X; vp, vaginal plate (sensu Maruyama (2006: fig. 16A))." figureDoi="http://doi.org/10.5281/zenodo.12190188" httpUri="https://zenodo.org/record/12190188/files/figure.png" pageId="1" pageNumber="356">Fig. 24</figureCitation>
|
|||
|
|
) with slender receptacle and a long duct tightly coiled around a large spheroid reservoir, duct coils form elongate, flat and ovoid structure.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C34BE8C4FFB7E92DFF7FFA4EFE14FC1A" lastPageId="8" lastPageNumber="363" pageId="1" pageNumber="356" type="description">
|
|||
|
|
<paragraph id="8BEEBB4FFFB7E924FF7FFA4EFEBDF9A6" blockId="1.[151,1437,1266,2013]" pageId="1" pageNumber="356">
|
|||
|
|
<emphasis id="B925675DFFB7E924FF7FFA4EFEC8F9DB" bold="true" box="[199,345,1518,1544]" pageId="1" pageNumber="356">Description.</emphasis>
|
|||
|
|
Body of female (
|
|||
|
|
<figureCitation id="136AA7CAFFB7E924FD9FFA4FFDFBF9DA" box="[551,618,1519,1545]" captionStart="FIGURE 1" captionStartId="2.[151,250,1651,1675]" captionTargetBox="[203,1391,314,1617]" captionTargetId="figure-88@2.[189,1398,306,1625]" captionTargetPageId="2" captionText="FIGURE 1. Keratodegnathus javanicus sp. n., holotype female, habitus in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12190172" httpUri="https://zenodo.org/record/12190172/files/figure.png" pageId="1" pageNumber="356">Fig. 1</figureCitation>
|
|||
|
|
) strongly flattened and slender, head, elytra, distal region of abdomen, antennomeres 1–9 and legs except tarsi dark brown with reddish hue, pronotum and proximal region of abdomen brownish-red, antennomeres 10 and 11 pale yellowish, palps and tarsi light brow; cuticle glossy; setae dark brown. BL
|
|||
|
|
<quantity id="4CA916AAFFB7E924FF79F9FAFEB9F9A6" box="[193,296,1626,1653]" metricMagnitude="-3" metricUnit="m" metricValue="5.05" pageId="1" pageNumber="356" unit="mm" value="5.05">5.05 mm</quantity>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BEEBB4FFFB7E927FF7FF9DFFAF7FECE" blockId="1.[151,1437,1266,2013]" lastBlockId="2.[151,1437,151,285]" lastPageId="2" lastPageNumber="357" pageId="1" pageNumber="356">
|
|||
|
|
Head (
|
|||
|
|
<figureCitation id="136AA7CAFFB7E924FEAAF9DFFEE6F94A" box="[274,375,1663,1689]" captionStart-0="FIGURES 2‒5" captionStart-1="FIGURES 6‒10" captionStartId-0="3.[151,264,1776,1800]" captionStartId-1="4.[151,264,1598,1622]" captionTargetBox-0="[177,1411,453,1752]" captionTargetBox-1="[177,1411,181,1574]" captionTargetId-0="figure-149@3.[177,1411,453,1752]" captionTargetId-1="figure-19@4.[177,1411,181,1574]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 2‒5. Keratodegnathus javanicus sp. n., holotype female. Head and anterior region of thorax in lateral view (2); head in anterodorsal view (3); head, prothorax and elytral base in dorsal view (4; arrowheads indicate macrosetae); head and prothorax in ventral view (5). Abbreviations: acxc, anterior coxal carina; cl, clypeus; cmb, connecting membrane; cs, cervical sclerite; dmt, dorsal mandibular tooth; fr, frons; gal, galea; gen, gena; gp, gular plate; gr, genal carina; gs, gular suture; hr, hypostomal ridge; hs, hypostomal suture; hyp, hypomeron; hyr, hypomeral ridge; lac, lacinia; lbr, labrum; lpc, lateral pronotal carina; mn, mentum; mxp2‒4, maxillary palpomere 2‒4; nss, notosternal suture; ocr, occipital ridge; pd, pedicel; prl prelabium; pst, prosternum; ptp, posterior tentorial pit; sc, scape; smn, submentum; trt, trochantin; vt, vertex." captionText-1="FIGURES 6‒10. Keratodegnathus javanicus sp. n., holotype female. Head in anterodorsal view (6); mouthparts in dorsal view (7); anteromedian region of labrum in anterodorsal view (8); maxillo-labial complex in ventral view (9); distal region of prelabium in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; clp, clypeus; cmb, connecting membrane; dmt, dorsal mandibular tooth; fr, frons; gal, galea; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; md, mandible; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pmns, premental sclerite; prl, prelabium; smn, submentum." figureDoi-0="http://doi.org/10.5281/zenodo.12190176" figureDoi-1="http://doi.org/10.5281/zenodo.12190180" httpUri-0="https://zenodo.org/record/12190176/files/figure.png" httpUri-1="https://zenodo.org/record/12190180/files/figure.png" pageId="1" pageNumber="356">Figs 2–6</figureCitation>
|
|||
|
|
) broadly subtrapezoidal, as broad as pronotum, broadest at eyes, HL
|
|||
|
|
<quantity id="4CA916AAFFB7E924FB30F9DFFB7FF94A" box="[1160,1262,1663,1689]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="1" pageNumber="356" unit="mm" value="0.63">0.63 mm</quantity>
|
|||
|
|
, HW 1.00 mm. Eyes large and strongly convex, finely faceted, strongly elongate oval. Clypeus (
|
|||
|
|
<figureCitation id="136AA7CAFFB7E924FB9DF903FBFEF96E" box="[1061,1135,1699,1725]" captionStart="FIGURES 2‒5" captionStartId="3.[151,264,1776,1800]" captionTargetBox="[177,1411,453,1752]" captionTargetId="figure-149@3.[177,1411,453,1752]" captionTargetPageId="3" captionText="FIGURES 2‒5. Keratodegnathus javanicus sp. n., holotype female. Head and anterior region of thorax in lateral view (2); head in anterodorsal view (3); head, prothorax and elytral base in dorsal view (4; arrowheads indicate macrosetae); head and prothorax in ventral view (5). Abbreviations: acxc, anterior coxal carina; cl, clypeus; cmb, connecting membrane; cs, cervical sclerite; dmt, dorsal mandibular tooth; fr, frons; gal, galea; gen, gena; gp, gular plate; gr, genal carina; gs, gular suture; hr, hypostomal ridge; hs, hypostomal suture; hyp, hypomeron; hyr, hypomeral ridge; lac, lacinia; lbr, labrum; lpc, lateral pronotal carina; mn, mentum; mxp2‒4, maxillary palpomere 2‒4; nss, notosternal suture; ocr, occipital ridge; pd, pedicel; prl prelabium; pst, prosternum; ptp, posterior tentorial pit; sc, scape; smn, submentum; trt, trochantin; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.12190176" httpUri="https://zenodo.org/record/12190176/files/figure.png" pageId="1" pageNumber="356">Figs 4</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="136AA7CAFFB7E924FBC4F903FB1DF96E" box="[1148,1164,1699,1725]" captionStart="FIGURES 6‒10" captionStartId="4.[151,264,1598,1622]" captionTargetBox="[177,1411,181,1574]" captionTargetId="figure-19@4.[177,1411,181,1574]" captionTargetPageId="4" captionText="FIGURES 6‒10. Keratodegnathus javanicus sp. n., holotype female. Head in anterodorsal view (6); mouthparts in dorsal view (7); anteromedian region of labrum in anterodorsal view (8); maxillo-labial complex in ventral view (9); distal region of prelabium in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; clp, clypeus; cmb, connecting membrane; dmt, dorsal mandibular tooth; fr, frons; gal, galea; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; md, mandible; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pmns, premental sclerite; prl, prelabium; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.12190180" httpUri="https://zenodo.org/record/12190180/files/figure.png" pageId="1" pageNumber="356">6</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFB7E924FB21F903FB3CF96E" box="[1177,1197,1699,1725]" italics="true" pageId="1" pageNumber="356">cl</emphasis>
|
|||
|
|
) nearly flat, strongly transverse and with nearly straight anterior margin; frons (
|
|||
|
|
<figureCitation id="136AA7CAFFB7E924FC94F967FCE4F932" box="[812,885,1735,1761]" captionStart="FIGURES 2‒5" captionStartId="3.[151,264,1776,1800]" captionTargetBox="[177,1411,453,1752]" captionTargetId="figure-149@3.[177,1411,453,1752]" captionTargetPageId="3" captionText="FIGURES 2‒5. Keratodegnathus javanicus sp. n., holotype female. Head and anterior region of thorax in lateral view (2); head in anterodorsal view (3); head, prothorax and elytral base in dorsal view (4; arrowheads indicate macrosetae); head and prothorax in ventral view (5). Abbreviations: acxc, anterior coxal carina; cl, clypeus; cmb, connecting membrane; cs, cervical sclerite; dmt, dorsal mandibular tooth; fr, frons; gal, galea; gen, gena; gp, gular plate; gr, genal carina; gs, gular suture; hr, hypostomal ridge; hs, hypostomal suture; hyp, hypomeron; hyr, hypomeral ridge; lac, lacinia; lbr, labrum; lpc, lateral pronotal carina; mn, mentum; mxp2‒4, maxillary palpomere 2‒4; nss, notosternal suture; ocr, occipital ridge; pd, pedicel; prl prelabium; pst, prosternum; ptp, posterior tentorial pit; sc, scape; smn, submentum; trt, trochantin; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.12190176" httpUri="https://zenodo.org/record/12190176/files/figure.png" pageId="1" pageNumber="356">Figs 4</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="136AA7CAFFB7E924FC3AF967FC03F932" box="[898,914,1735,1761]" captionStart="FIGURES 6‒10" captionStartId="4.[151,264,1598,1622]" captionTargetBox="[177,1411,181,1574]" captionTargetId="figure-19@4.[177,1411,181,1574]" captionTargetPageId="4" captionText="FIGURES 6‒10. Keratodegnathus javanicus sp. n., holotype female. Head in anterodorsal view (6); mouthparts in dorsal view (7); anteromedian region of labrum in anterodorsal view (8); maxillo-labial complex in ventral view (9); distal region of prelabium in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; clp, clypeus; cmb, connecting membrane; dmt, dorsal mandibular tooth; fr, frons; gal, galea; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; md, mandible; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pmns, premental sclerite; prl, prelabium; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.12190180" httpUri="https://zenodo.org/record/12190180/files/figure.png" pageId="1" pageNumber="356">6</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFB7E924FC18F967FC22F932" box="[928,947,1735,1761]" italics="true" pageId="1" pageNumber="356">fr</emphasis>
|
|||
|
|
) subtriangular, weakly convex, broadening posteriorly, where it is confluent with strongly transverse vertex (
|
|||
|
|
<figureCitation id="136AA7CAFFB7E924FC32F94BFC43F8D6" box="[906,978,1771,1797]" captionStart="FIGURES 2‒5" captionStartId="3.[151,264,1776,1800]" captionTargetBox="[177,1411,453,1752]" captionTargetId="figure-149@3.[177,1411,453,1752]" captionTargetPageId="3" captionText="FIGURES 2‒5. Keratodegnathus javanicus sp. n., holotype female. Head and anterior region of thorax in lateral view (2); head in anterodorsal view (3); head, prothorax and elytral base in dorsal view (4; arrowheads indicate macrosetae); head and prothorax in ventral view (5). Abbreviations: acxc, anterior coxal carina; cl, clypeus; cmb, connecting membrane; cs, cervical sclerite; dmt, dorsal mandibular tooth; fr, frons; gal, galea; gen, gena; gp, gular plate; gr, genal carina; gs, gular suture; hr, hypostomal ridge; hs, hypostomal suture; hyp, hypomeron; hyr, hypomeral ridge; lac, lacinia; lbr, labrum; lpc, lateral pronotal carina; mn, mentum; mxp2‒4, maxillary palpomere 2‒4; nss, notosternal suture; ocr, occipital ridge; pd, pedicel; prl prelabium; pst, prosternum; ptp, posterior tentorial pit; sc, scape; smn, submentum; trt, trochantin; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.12190176" httpUri="https://zenodo.org/record/12190176/files/figure.png" pageId="1" pageNumber="356">Fig. 4</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFB7E924FC58F94BFC65F8D7" box="[992,1012,1771,1796]" italics="true" pageId="1" pageNumber="356">vt</emphasis>
|
|||
|
|
) slightly impressed posteromedially; supraantennal tubercles lacking, oval antennal fossae exposed dorsally. Clypeolabral connecting membrane (
|
|||
|
|
<figureCitation id="136AA7CAFFB7E924FAD2F8AFFF36F89E" captionStart="FIGURES 2‒5" captionStartId="3.[151,264,1776,1800]" captionTargetBox="[177,1411,453,1752]" captionTargetId="figure-149@3.[177,1411,453,1752]" captionTargetPageId="3" captionText="FIGURES 2‒5. Keratodegnathus javanicus sp. n., holotype female. Head and anterior region of thorax in lateral view (2); head in anterodorsal view (3); head, prothorax and elytral base in dorsal view (4; arrowheads indicate macrosetae); head and prothorax in ventral view (5). Abbreviations: acxc, anterior coxal carina; cl, clypeus; cmb, connecting membrane; cs, cervical sclerite; dmt, dorsal mandibular tooth; fr, frons; gal, galea; gen, gena; gp, gular plate; gr, genal carina; gs, gular suture; hr, hypostomal ridge; hs, hypostomal suture; hyp, hypomeron; hyr, hypomeral ridge; lac, lacinia; lbr, labrum; lpc, lateral pronotal carina; mn, mentum; mxp2‒4, maxillary palpomere 2‒4; nss, notosternal suture; ocr, occipital ridge; pd, pedicel; prl prelabium; pst, prosternum; ptp, posterior tentorial pit; sc, scape; smn, submentum; trt, trochantin; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.12190176" httpUri="https://zenodo.org/record/12190176/files/figure.png" pageId="1" pageNumber="356">Figs 4</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="136AA7CAFFB7E924FF0EF893FF57F89E" box="[182,198,1843,1869]" captionStart="FIGURES 6‒10" captionStartId="4.[151,264,1598,1622]" captionTargetBox="[177,1411,181,1574]" captionTargetId="figure-19@4.[177,1411,181,1574]" captionTargetPageId="4" captionText="FIGURES 6‒10. Keratodegnathus javanicus sp. n., holotype female. Head in anterodorsal view (6); mouthparts in dorsal view (7); anteromedian region of labrum in anterodorsal view (8); maxillo-labial complex in ventral view (9); distal region of prelabium in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; clp, clypeus; cmb, connecting membrane; dmt, dorsal mandibular tooth; fr, frons; gal, galea; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; md, mandible; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pmns, premental sclerite; prl, prelabium; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.12190180" httpUri="https://zenodo.org/record/12190180/files/figure.png" pageId="1" pageNumber="356">6</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFB7E924FF6DF893FE95F89E" box="[213,260,1843,1869]" italics="true" pageId="1" pageNumber="356">cmb</emphasis>
|
|||
|
|
) about as long as labrum and subtrapezoidal. Tempora in dorsal view slightly shorter than half length of eye. Genae (
|
|||
|
|
<figureCitation id="136AA7CAFFB7E924FEF5F8F7FE07F8A2" box="[333,406,1878,1905]" captionStart="FIGURES 2‒5" captionStartId="3.[151,264,1776,1800]" captionTargetBox="[177,1411,453,1752]" captionTargetId="figure-149@3.[177,1411,453,1752]" captionTargetPageId="3" captionText="FIGURES 2‒5. Keratodegnathus javanicus sp. n., holotype female. Head and anterior region of thorax in lateral view (2); head in anterodorsal view (3); head, prothorax and elytral base in dorsal view (4; arrowheads indicate macrosetae); head and prothorax in ventral view (5). Abbreviations: acxc, anterior coxal carina; cl, clypeus; cmb, connecting membrane; cs, cervical sclerite; dmt, dorsal mandibular tooth; fr, frons; gal, galea; gen, gena; gp, gular plate; gr, genal carina; gs, gular suture; hr, hypostomal ridge; hs, hypostomal suture; hyp, hypomeron; hyr, hypomeral ridge; lac, lacinia; lbr, labrum; lpc, lateral pronotal carina; mn, mentum; mxp2‒4, maxillary palpomere 2‒4; nss, notosternal suture; ocr, occipital ridge; pd, pedicel; prl prelabium; pst, prosternum; ptp, posterior tentorial pit; sc, scape; smn, submentum; trt, trochantin; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.12190176" httpUri="https://zenodo.org/record/12190176/files/figure.png" pageId="1" pageNumber="356">Fig. 5</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFB7E924FE1EF8F7FE5FF8A3" box="[422,462,1879,1904]" italics="true" pageId="1" pageNumber="356">gen</emphasis>
|
|||
|
|
) with complete genal ridges extending from their posterior margins to the level of cardines. Gular plate (
|
|||
|
|
<figureCitation id="136AA7CAFFB7E924FE20F8DBFE4EF846" box="[408,479,1914,1941]" captionStart="FIGURES 2‒5" captionStartId="3.[151,264,1776,1800]" captionTargetBox="[177,1411,453,1752]" captionTargetId="figure-149@3.[177,1411,453,1752]" captionTargetPageId="3" captionText="FIGURES 2‒5. Keratodegnathus javanicus sp. n., holotype female. Head and anterior region of thorax in lateral view (2); head in anterodorsal view (3); head, prothorax and elytral base in dorsal view (4; arrowheads indicate macrosetae); head and prothorax in ventral view (5). Abbreviations: acxc, anterior coxal carina; cl, clypeus; cmb, connecting membrane; cs, cervical sclerite; dmt, dorsal mandibular tooth; fr, frons; gal, galea; gen, gena; gp, gular plate; gr, genal carina; gs, gular suture; hr, hypostomal ridge; hs, hypostomal suture; hyp, hypomeron; hyr, hypomeral ridge; lac, lacinia; lbr, labrum; lpc, lateral pronotal carina; mn, mentum; mxp2‒4, maxillary palpomere 2‒4; nss, notosternal suture; ocr, occipital ridge; pd, pedicel; prl prelabium; pst, prosternum; ptp, posterior tentorial pit; sc, scape; smn, submentum; trt, trochantin; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.12190176" httpUri="https://zenodo.org/record/12190176/files/figure.png" pageId="1" pageNumber="356">Fig. 5</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFB7E924FE57F8DBFD9AF847" box="[495,523,1915,1940]" italics="true" pageId="1" pageNumber="356">gp</emphasis>
|
|||
|
|
) and submentum (
|
|||
|
|
<figureCitation id="136AA7CAFFB7E924FD5DF8DBFCBDF846" box="[741,812,1914,1941]" captionStart="FIGURES 2‒5" captionStartId="3.[151,264,1776,1800]" captionTargetBox="[177,1411,453,1752]" captionTargetId="figure-149@3.[177,1411,453,1752]" captionTargetPageId="3" captionText="FIGURES 2‒5. Keratodegnathus javanicus sp. n., holotype female. Head and anterior region of thorax in lateral view (2); head in anterodorsal view (3); head, prothorax and elytral base in dorsal view (4; arrowheads indicate macrosetae); head and prothorax in ventral view (5). Abbreviations: acxc, anterior coxal carina; cl, clypeus; cmb, connecting membrane; cs, cervical sclerite; dmt, dorsal mandibular tooth; fr, frons; gal, galea; gen, gena; gp, gular plate; gr, genal carina; gs, gular suture; hr, hypostomal ridge; hs, hypostomal suture; hyp, hypomeron; hyr, hypomeral ridge; lac, lacinia; lbr, labrum; lpc, lateral pronotal carina; mn, mentum; mxp2‒4, maxillary palpomere 2‒4; nss, notosternal suture; ocr, occipital ridge; pd, pedicel; prl prelabium; pst, prosternum; ptp, posterior tentorial pit; sc, scape; smn, submentum; trt, trochantin; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.12190176" httpUri="https://zenodo.org/record/12190176/files/figure.png" pageId="1" pageNumber="356">Fig. 5</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFB7E924FC84F8DBFCF8F847" box="[828,873,1915,1940]" italics="true" pageId="1" pageNumber="356">smn</emphasis>
|
|||
|
|
) confluent and laterally delimited by continuous gular and hypostomal sutures (
|
|||
|
|
<figureCitation id="136AA7CAFFB7E924FE49F83FFDA7F86A" box="[497,566,1950,1977]" captionStart="FIGURES 2‒5" captionStartId="3.[151,264,1776,1800]" captionTargetBox="[177,1411,453,1752]" captionTargetId="figure-149@3.[177,1411,453,1752]" captionTargetPageId="3" captionText="FIGURES 2‒5. Keratodegnathus javanicus sp. n., holotype female. Head and anterior region of thorax in lateral view (2); head in anterodorsal view (3); head, prothorax and elytral base in dorsal view (4; arrowheads indicate macrosetae); head and prothorax in ventral view (5). Abbreviations: acxc, anterior coxal carina; cl, clypeus; cmb, connecting membrane; cs, cervical sclerite; dmt, dorsal mandibular tooth; fr, frons; gal, galea; gen, gena; gp, gular plate; gr, genal carina; gs, gular suture; hr, hypostomal ridge; hs, hypostomal suture; hyp, hypomeron; hyr, hypomeral ridge; lac, lacinia; lbr, labrum; lpc, lateral pronotal carina; mn, mentum; mxp2‒4, maxillary palpomere 2‒4; nss, notosternal suture; ocr, occipital ridge; pd, pedicel; prl prelabium; pst, prosternum; ptp, posterior tentorial pit; sc, scape; smn, submentum; trt, trochantin; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.12190176" httpUri="https://zenodo.org/record/12190176/files/figure.png" pageId="1" pageNumber="356">Fig. 5</figureCitation>
|
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|
;
|
|||
|
|
<emphasis id="B925675DFFB7E924FDFBF83FFDCDF86B" box="[579,604,1951,1976]" italics="true" pageId="1" pageNumber="356">gs</emphasis>
|
|||
|
|
,
|
|||
|
|
<emphasis id="B925675DFFB7E924FDD2F83FFD12F86A" box="[618,643,1951,1977]" italics="true" pageId="1" pageNumber="356">hs</emphasis>
|
|||
|
|
), narrowest site of gula situated just behind barely discernible, narrow and elongate posterior tentorial pits (
|
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|
|
<figureCitation id="136AA7CAFFB7E924FD86F863FD14F80E" box="[574,645,1986,2013]" captionStart="FIGURES 2‒5" captionStartId="3.[151,264,1776,1800]" captionTargetBox="[177,1411,453,1752]" captionTargetId="figure-149@3.[177,1411,453,1752]" captionTargetPageId="3" captionText="FIGURES 2‒5. Keratodegnathus javanicus sp. n., holotype female. Head and anterior region of thorax in lateral view (2); head in anterodorsal view (3); head, prothorax and elytral base in dorsal view (4; arrowheads indicate macrosetae); head and prothorax in ventral view (5). Abbreviations: acxc, anterior coxal carina; cl, clypeus; cmb, connecting membrane; cs, cervical sclerite; dmt, dorsal mandibular tooth; fr, frons; gal, galea; gen, gena; gp, gular plate; gr, genal carina; gs, gular suture; hr, hypostomal ridge; hs, hypostomal suture; hyp, hypomeron; hyr, hypomeral ridge; lac, lacinia; lbr, labrum; lpc, lateral pronotal carina; mn, mentum; mxp2‒4, maxillary palpomere 2‒4; nss, notosternal suture; ocr, occipital ridge; pd, pedicel; prl prelabium; pst, prosternum; ptp, posterior tentorial pit; sc, scape; smn, submentum; trt, trochantin; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.12190176" httpUri="https://zenodo.org/record/12190176/files/figure.png" pageId="1" pageNumber="356">Fig. 5</figureCitation>
|
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|
;
|
|||
|
|
<emphasis id="B925675DFFB7E924FD2CF863FD29F80F" box="[660,696,1987,2012]" italics="true" pageId="1" pageNumber="356">ptp</emphasis>
|
|||
|
|
). Hypostomal ridges (
|
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|
|
<figureCitation id="136AA7CAFFB7E924FC02F863FB91F80E" box="[954,1024,1986,2013]" captionStart="FIGURES 2‒5" captionStartId="3.[151,264,1776,1800]" captionTargetBox="[177,1411,453,1752]" captionTargetId="figure-149@3.[177,1411,453,1752]" captionTargetPageId="3" captionText="FIGURES 2‒5. Keratodegnathus javanicus sp. n., holotype female. Head and anterior region of thorax in lateral view (2); head in anterodorsal view (3); head, prothorax and elytral base in dorsal view (4; arrowheads indicate macrosetae); head and prothorax in ventral view (5). Abbreviations: acxc, anterior coxal carina; cl, clypeus; cmb, connecting membrane; cs, cervical sclerite; dmt, dorsal mandibular tooth; fr, frons; gal, galea; gen, gena; gp, gular plate; gr, genal carina; gs, gular suture; hr, hypostomal ridge; hs, hypostomal suture; hyp, hypomeron; hyr, hypomeral ridge; lac, lacinia; lbr, labrum; lpc, lateral pronotal carina; mn, mentum; mxp2‒4, maxillary palpomere 2‒4; nss, notosternal suture; ocr, occipital ridge; pd, pedicel; prl prelabium; pst, prosternum; ptp, posterior tentorial pit; sc, scape; smn, submentum; trt, trochantin; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.12190176" httpUri="https://zenodo.org/record/12190176/files/figure.png" pageId="1" pageNumber="356">Fig. 5</figureCitation>
|
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;
|
|||
|
|
<emphasis id="B925675DFFB7E924FBB7F863FBB9F80E" box="[1039,1064,1987,2013]" italics="true" pageId="1" pageNumber="356">hr</emphasis>
|
|||
|
|
) complete, each extending from posterior margin of submentum (marked by anterior ends of posterior tentorial pits) to the level of anterior margins of cardines. Head largely impunctate, small setiferous punctures present only mesad and posterad antennal fossae, on tempora and on genae (mainly along genal ridges, posterior margins and gular sutures); setae sparse, moderately long and suberect, those on posterior and mesal regions of genae much shorter than those on frons and vertex.
|
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|
|
</paragraph>
|
|||
|
|
<caption id="DF2EEBC7FFB4E927FF2FF9D3FB9FF958" ID-DOI="http://doi.org/10.5281/zenodo.12190172" ID-Zenodo-Dep="12190172" box="[151,1038,1651,1675]" httpUri="https://zenodo.org/record/12190172/files/figure.png" pageId="2" pageNumber="357" startId="2.[151,250,1651,1675]" targetBox="[203,1391,314,1617]" targetPageId="2" targetType="figure">
|
|||
|
|
<paragraph id="8BEEBB4FFFB4E927FF2FF9D3FB9FF958" blockId="2.[151,1038,1651,1675]" box="[151,1038,1651,1675]" pageId="2" pageNumber="357">
|
|||
|
|
<emphasis id="B925675DFFB4E927FF2FF9D3FE82F958" bold="true" box="[151,275,1651,1675]" pageId="2" pageNumber="357">FIGURE 1.</emphasis>
|
|||
|
|
<taxonomicName id="4C51C0CCFFB4E927FEA1F9D3FDBEF958" authority="Jałoszyński, 2024" authorityName="Jałoszyński" authorityYear="2024" box="[281,559,1651,1675]" class="Insecta" family="Staphylinidae" genus="Keratodegnathus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="357" phylum="Arthropoda" rank="species" species="javanicus" status="sp. nov.">
|
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|
|
<emphasis id="B925675DFFB4E927FEA1F9D3FDBEF958" box="[281,559,1651,1675]" italics="true" pageId="2" pageNumber="357">Keratodegnathus javanicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A216DA26FFB4E927FD8EF9D4FDFCF958" box="[566,621,1652,1675]" pageId="2" pageNumber="357" rank="species">sp. n.</taxonomicNameLabel>
|
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|
|
, holotype female, habitus in dorsal view.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BEEBB4FFFB4E926FF7FF97DFB99FE7E" blockId="2.[151,1436,1757,2036]" lastBlockId="3.[151,1437,151,429]" lastPageId="3" lastPageNumber="358" pageId="2" pageNumber="357">
|
|||
|
|
Labrum (
|
|||
|
|
<figureCitation id="136AA7CAFFB4E927FE96F97DFEE6F92B" box="[302,375,1757,1784]" captionStart="FIGURES 2‒5" captionStartId="3.[151,264,1776,1800]" captionTargetBox="[177,1411,453,1752]" captionTargetId="figure-149@3.[177,1411,453,1752]" captionTargetPageId="3" captionText="FIGURES 2‒5. Keratodegnathus javanicus sp. n., holotype female. Head and anterior region of thorax in lateral view (2); head in anterodorsal view (3); head, prothorax and elytral base in dorsal view (4; arrowheads indicate macrosetae); head and prothorax in ventral view (5). Abbreviations: acxc, anterior coxal carina; cl, clypeus; cmb, connecting membrane; cs, cervical sclerite; dmt, dorsal mandibular tooth; fr, frons; gal, galea; gen, gena; gp, gular plate; gr, genal carina; gs, gular suture; hr, hypostomal ridge; hs, hypostomal suture; hyp, hypomeron; hyr, hypomeral ridge; lac, lacinia; lbr, labrum; lpc, lateral pronotal carina; mn, mentum; mxp2‒4, maxillary palpomere 2‒4; nss, notosternal suture; ocr, occipital ridge; pd, pedicel; prl prelabium; pst, prosternum; ptp, posterior tentorial pit; sc, scape; smn, submentum; trt, trochantin; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.12190176" httpUri="https://zenodo.org/record/12190176/files/figure.png" pageId="2" pageNumber="357">Figs 4</figureCitation>
|
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|
|
,
|
|||
|
|
<figureCitation id="136AA7CAFFB4E927FE3BF97DFE21F924" box="[387,432,1757,1783]" captionStart="FIGURES 6‒10" captionStartId="4.[151,264,1598,1622]" captionTargetBox="[177,1411,181,1574]" captionTargetId="figure-19@4.[177,1411,181,1574]" captionTargetPageId="4" captionText="FIGURES 6‒10. Keratodegnathus javanicus sp. n., holotype female. Head in anterodorsal view (6); mouthparts in dorsal view (7); anteromedian region of labrum in anterodorsal view (8); maxillo-labial complex in ventral view (9); distal region of prelabium in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; clp, clypeus; cmb, connecting membrane; dmt, dorsal mandibular tooth; fr, frons; gal, galea; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; md, mandible; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pmns, premental sclerite; prl, prelabium; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.12190180" httpUri="https://zenodo.org/record/12190180/files/figure.png" pageId="2" pageNumber="357">6–7</figureCitation>
|
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|
|
;
|
|||
|
|
<emphasis id="B925675DFFB4E927FE03F97DFE4DF924" box="[443,476,1757,1783]" italics="true" pageId="2" pageNumber="357">lbr</emphasis>
|
|||
|
|
) strongly transverse, with evenly rounded anterior margin, except for shallow median emargination (
|
|||
|
|
<figureCitation id="136AA7CAFFB4E927FE84F8A1FE13F8CF" box="[316,386,1793,1820]" captionStart="FIGURES 6‒10" captionStartId="4.[151,264,1598,1622]" captionTargetBox="[177,1411,181,1574]" captionTargetId="figure-19@4.[177,1411,181,1574]" captionTargetPageId="4" captionText="FIGURES 6‒10. Keratodegnathus javanicus sp. n., holotype female. Head in anterodorsal view (6); mouthparts in dorsal view (7); anteromedian region of labrum in anterodorsal view (8); maxillo-labial complex in ventral view (9); distal region of prelabium in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; clp, clypeus; cmb, connecting membrane; dmt, dorsal mandibular tooth; fr, frons; gal, galea; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; md, mandible; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pmns, premental sclerite; prl, prelabium; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.12190180" httpUri="https://zenodo.org/record/12190180/files/figure.png" pageId="2" pageNumber="357">Fig. 8</figureCitation>
|
|||
|
|
), dorsal surface with two rows of symmetrically distributed long setae and asymmetrically, sparsely distributed minute setae. Mandibles (only partly exposed in studied specimen) long and slender, with pointed apices, each with dorsal tooth (
|
|||
|
|
<figureCitation id="136AA7CAFFB4E927FDECF8E9FD08F8B0" box="[596,665,1865,1891]" captionStart="FIGURES 6‒10" captionStartId="4.[151,264,1598,1622]" captionTargetBox="[177,1411,181,1574]" captionTargetId="figure-19@4.[177,1411,181,1574]" captionTargetPageId="4" captionText="FIGURES 6‒10. Keratodegnathus javanicus sp. n., holotype female. Head in anterodorsal view (6); mouthparts in dorsal view (7); anteromedian region of labrum in anterodorsal view (8); maxillo-labial complex in ventral view (9); distal region of prelabium in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; clp, clypeus; cmb, connecting membrane; dmt, dorsal mandibular tooth; fr, frons; gal, galea; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; md, mandible; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pmns, premental sclerite; prl, prelabium; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.12190180" httpUri="https://zenodo.org/record/12190180/files/figure.png" pageId="2" pageNumber="357">Fig. 6</figureCitation>
|
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|
|
;
|
|||
|
|
<emphasis id="B925675DFFB4E927FD1FF8E9FD40F8B0" box="[679,721,1865,1891]" italics="true" pageId="2" pageNumber="357">dmt</emphasis>
|
|||
|
|
). Cardo (
|
|||
|
|
<figureCitation id="136AA7CAFFB4E927FC86F8E9FC12F8B7" box="[830,899,1865,1892]" captionStart="FIGURES 2‒5" captionStartId="3.[151,264,1776,1800]" captionTargetBox="[177,1411,453,1752]" captionTargetId="figure-149@3.[177,1411,453,1752]" captionTargetPageId="3" captionText="FIGURES 2‒5. Keratodegnathus javanicus sp. n., holotype female. Head and anterior region of thorax in lateral view (2); head in anterodorsal view (3); head, prothorax and elytral base in dorsal view (4; arrowheads indicate macrosetae); head and prothorax in ventral view (5). Abbreviations: acxc, anterior coxal carina; cl, clypeus; cmb, connecting membrane; cs, cervical sclerite; dmt, dorsal mandibular tooth; fr, frons; gal, galea; gen, gena; gp, gular plate; gr, genal carina; gs, gular suture; hr, hypostomal ridge; hs, hypostomal suture; hyp, hypomeron; hyr, hypomeral ridge; lac, lacinia; lbr, labrum; lpc, lateral pronotal carina; mn, mentum; mxp2‒4, maxillary palpomere 2‒4; nss, notosternal suture; ocr, occipital ridge; pd, pedicel; prl prelabium; pst, prosternum; ptp, posterior tentorial pit; sc, scape; smn, submentum; trt, trochantin; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.12190176" httpUri="https://zenodo.org/record/12190176/files/figure.png" pageId="2" pageNumber="357">Fig. 5</figureCitation>
|
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|
|
;
|
|||
|
|
<emphasis id="B925675DFFB4E927FC29F8E9FC3AF8B0" box="[913,939,1865,1891]" italics="true" pageId="2" pageNumber="357">cd</emphasis>
|
|||
|
|
) strongly transverse; basistipes (
|
|||
|
|
<figureCitation id="136AA7CAFFB4E927FAA7F8E9FAF5F8B7" box="[1311,1380,1865,1892]" captionStart="FIGURES 6‒10" captionStartId="4.[151,264,1598,1622]" captionTargetBox="[177,1411,181,1574]" captionTargetId="figure-19@4.[177,1411,181,1574]" captionTargetPageId="4" captionText="FIGURES 6‒10. Keratodegnathus javanicus sp. n., holotype female. Head in anterodorsal view (6); mouthparts in dorsal view (7); anteromedian region of labrum in anterodorsal view (8); maxillo-labial complex in ventral view (9); distal region of prelabium in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; clp, clypeus; cmb, connecting membrane; dmt, dorsal mandibular tooth; fr, frons; gal, galea; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; md, mandible; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pmns, premental sclerite; prl, prelabium; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.12190180" httpUri="https://zenodo.org/record/12190180/files/figure.png" pageId="2" pageNumber="357">Fig. 9</figureCitation>
|
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|
|
;
|
|||
|
|
<emphasis id="B925675DFFB4E927FACAF8E9FA02F8B0" box="[1394,1427,1865,1891]" italics="true" pageId="2" pageNumber="357">bst</emphasis>
|
|||
|
|
) subtriangular and elongate; mediostipes (
|
|||
|
|
<figureCitation id="136AA7CAFFB4E927FDDAF8CDFD37F85B" box="[610,678,1901,1928]" captionStart="FIGURES 6‒10" captionStartId="4.[151,264,1598,1622]" captionTargetBox="[177,1411,181,1574]" captionTargetId="figure-19@4.[177,1411,181,1574]" captionTargetPageId="4" captionText="FIGURES 6‒10. Keratodegnathus javanicus sp. n., holotype female. Head in anterodorsal view (6); mouthparts in dorsal view (7); anteromedian region of labrum in anterodorsal view (8); maxillo-labial complex in ventral view (9); distal region of prelabium in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; clp, clypeus; cmb, connecting membrane; dmt, dorsal mandibular tooth; fr, frons; gal, galea; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; md, mandible; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pmns, premental sclerite; prl, prelabium; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.12190180" httpUri="https://zenodo.org/record/12190180/files/figure.png" pageId="2" pageNumber="357">Fig. 9</figureCitation>
|
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|
|
;
|
|||
|
|
<emphasis id="B925675DFFB4E927FD0AF8CEFD48F854" box="[690,729,1902,1927]" italics="true" pageId="2" pageNumber="357">mst</emphasis>
|
|||
|
|
) subrectangular; galea and lacinia (
|
|||
|
|
<figureCitation id="136AA7CAFFB4E927FBDDF8CDFB46F85B" box="[1125,1239,1901,1928]" captionStart="FIGURES 6‒10" captionStartId="4.[151,264,1598,1622]" captionTargetBox="[177,1411,181,1574]" captionTargetId="figure-19@4.[177,1411,181,1574]" captionTargetPageId="4" captionText="FIGURES 6‒10. Keratodegnathus javanicus sp. n., holotype female. Head in anterodorsal view (6); mouthparts in dorsal view (7); anteromedian region of labrum in anterodorsal view (8); maxillo-labial complex in ventral view (9); distal region of prelabium in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; clp, clypeus; cmb, connecting membrane; dmt, dorsal mandibular tooth; fr, frons; gal, galea; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; md, mandible; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pmns, premental sclerite; prl, prelabium; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.12190180" httpUri="https://zenodo.org/record/12190180/files/figure.png" pageId="2" pageNumber="357">Figs 9, 10</figureCitation>
|
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|
|
;
|
|||
|
|
<emphasis id="B925675DFFB4E927FB5AF8CDFA97F854" box="[1250,1286,1901,1927]" italics="true" pageId="2" pageNumber="357">gal</emphasis>
|
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|
|
,
|
|||
|
|
<emphasis id="B925675DFFB4E927FAACF8CDFAA7F854" box="[1300,1334,1901,1927]" italics="true" pageId="2" pageNumber="357">lac</emphasis>
|
|||
|
|
) slender, with dense mesal and distal setae, galea much longer than lacinia. Maxillary palp short and slender, palpomere 1 (
|
|||
|
|
<figureCitation id="136AA7CAFFB4E927FF27F815FF72F803" box="[159,227,1973,2000]" captionStart="FIGURES 6‒10" captionStartId="4.[151,264,1598,1622]" captionTargetBox="[177,1411,181,1574]" captionTargetId="figure-19@4.[177,1411,181,1574]" captionTargetPageId="4" captionText="FIGURES 6‒10. Keratodegnathus javanicus sp. n., holotype female. Head in anterodorsal view (6); mouthparts in dorsal view (7); anteromedian region of labrum in anterodorsal view (8); maxillo-labial complex in ventral view (9); distal region of prelabium in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; clp, clypeus; cmb, connecting membrane; dmt, dorsal mandibular tooth; fr, frons; gal, galea; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; md, mandible; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pmns, premental sclerite; prl, prelabium; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.12190180" httpUri="https://zenodo.org/record/12190180/files/figure.png" pageId="2" pageNumber="357">Fig. 9</figureCitation>
|
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|
|
;
|
|||
|
|
<emphasis id="B925675DFFB4E927FF57F816FEBDF803" box="[239,300,1974,2000]" italics="true" pageId="2" pageNumber="357">mxp1</emphasis>
|
|||
|
|
) short, annular, palpomere 2 (
|
|||
|
|
<figureCitation id="136AA7CAFFB4E927FDC2F815FD2FF803" box="[634,702,1973,2000]" captionStart="FIGURES 6‒10" captionStartId="4.[151,264,1598,1622]" captionTargetBox="[177,1411,181,1574]" captionTargetId="figure-19@4.[177,1411,181,1574]" captionTargetPageId="4" captionText="FIGURES 6‒10. Keratodegnathus javanicus sp. n., holotype female. Head in anterodorsal view (6); mouthparts in dorsal view (7); anteromedian region of labrum in anterodorsal view (8); maxillo-labial complex in ventral view (9); distal region of prelabium in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; clp, clypeus; cmb, connecting membrane; dmt, dorsal mandibular tooth; fr, frons; gal, galea; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; md, mandible; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pmns, premental sclerite; prl, prelabium; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.12190180" httpUri="https://zenodo.org/record/12190180/files/figure.png" pageId="2" pageNumber="357">Fig. 9</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFB4E927FD72F816FC96F803" box="[714,775,1974,2000]" italics="true" pageId="2" pageNumber="357">mxp2</emphasis>
|
|||
|
|
) strongly elongate, weakly curved and broadening distally; palpomere 4 (
|
|||
|
|
<figureCitation id="136AA7CAFFB4E927FE81F879FE11F827" box="[313,384,2009,2036]" captionStart="FIGURES 6‒10" captionStartId="4.[151,264,1598,1622]" captionTargetBox="[177,1411,181,1574]" captionTargetId="figure-19@4.[177,1411,181,1574]" captionTargetPageId="4" captionText="FIGURES 6‒10. Keratodegnathus javanicus sp. n., holotype female. Head in anterodorsal view (6); mouthparts in dorsal view (7); anteromedian region of labrum in anterodorsal view (8); maxillo-labial complex in ventral view (9); distal region of prelabium in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; clp, clypeus; cmb, connecting membrane; dmt, dorsal mandibular tooth; fr, frons; gal, galea; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; md, mandible; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pmns, premental sclerite; prl, prelabium; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.12190180" httpUri="https://zenodo.org/record/12190180/files/figure.png" pageId="2" pageNumber="357">Fig. 9</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFB4E927FE29F87AFE5FF827" box="[401,462,2010,2036]" italics="true" pageId="2" pageNumber="357">mxp4</emphasis>
|
|||
|
|
) strongly elongate, aciculate. Submentum (
|
|||
|
|
<figureCitation id="136AA7CAFFB4E927FC7FF879FB9EF827" box="[967,1039,2009,2036]" captionStart="FIGURES 6‒10" captionStartId="4.[151,264,1598,1622]" captionTargetBox="[177,1411,181,1574]" captionTargetId="figure-19@4.[177,1411,181,1574]" captionTargetPageId="4" captionText="FIGURES 6‒10. Keratodegnathus javanicus sp. n., holotype female. Head in anterodorsal view (6); mouthparts in dorsal view (7); anteromedian region of labrum in anterodorsal view (8); maxillo-labial complex in ventral view (9); distal region of prelabium in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; clp, clypeus; cmb, connecting membrane; dmt, dorsal mandibular tooth; fr, frons; gal, galea; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; md, mandible; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pmns, premental sclerite; prl, prelabium; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.12190180" httpUri="https://zenodo.org/record/12190180/files/figure.png" pageId="2" pageNumber="357">Fig. 9</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFB4E927FBA7F87AFBDDF820" box="[1055,1100,2010,2035]" italics="true" pageId="2" pageNumber="357">smn</emphasis>
|
|||
|
|
) subtrapezoidal and strongly transverse, its median region slightly impressed and with matte microsculpture contrasting with smooth lateral margins; mentum (
|
|||
|
|
<figureCitation id="136AA7CAFFB5E926FED7FF1BFE27FF06" box="[367,438,187,213]" captionStart="FIGURES 6‒10" captionStartId="4.[151,264,1598,1622]" captionTargetBox="[177,1411,181,1574]" captionTargetId="figure-19@4.[177,1411,181,1574]" captionTargetPageId="4" captionText="FIGURES 6‒10. Keratodegnathus javanicus sp. n., holotype female. Head in anterodorsal view (6); mouthparts in dorsal view (7); anteromedian region of labrum in anterodorsal view (8); maxillo-labial complex in ventral view (9); distal region of prelabium in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; clp, clypeus; cmb, connecting membrane; dmt, dorsal mandibular tooth; fr, frons; gal, galea; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; md, mandible; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pmns, premental sclerite; prl, prelabium; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.12190180" httpUri="https://zenodo.org/record/12190180/files/figure.png" pageId="3" pageNumber="358">Fig. 9</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFB5E926FE7DFF1BFE76FF07" box="[453,487,187,212]" italics="true" pageId="3" pageNumber="358">mn</emphasis>
|
|||
|
|
) subtrapezoidal, strongly transverse, covered with dense microgranules; prelabium (
|
|||
|
|
<figureCitation id="136AA7CAFFB5E926FF27FF7FFF76FF2A" box="[159,231,223,249]" captionStart="FIGURES 6‒10" captionStartId="4.[151,264,1598,1622]" captionTargetBox="[177,1411,181,1574]" captionTargetId="figure-19@4.[177,1411,181,1574]" captionTargetPageId="4" captionText="FIGURES 6‒10. Keratodegnathus javanicus sp. n., holotype female. Head in anterodorsal view (6); mouthparts in dorsal view (7); anteromedian region of labrum in anterodorsal view (8); maxillo-labial complex in ventral view (9); distal region of prelabium in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; clp, clypeus; cmb, connecting membrane; dmt, dorsal mandibular tooth; fr, frons; gal, galea; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; md, mandible; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pmns, premental sclerite; prl, prelabium; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.12190180" httpUri="https://zenodo.org/record/12190180/files/figure.png" pageId="3" pageNumber="358">Fig. 9</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFB5E926FF4FFF7FFE89FF2A" box="[247,280,223,249]" italics="true" pageId="3" pageNumber="358">prl</emphasis>
|
|||
|
|
) strongly elongate, with subtrapezoidal membranous proximal region and a pair of distal premental sclerites (
|
|||
|
|
<figureCitation id="136AA7CAFFB5E926FEBEFEA3FECCFECE" box="[262,349,259,285]" captionStart="FIGURES 6‒10" captionStartId="4.[151,264,1598,1622]" captionTargetBox="[177,1411,181,1574]" captionTargetId="figure-19@4.[177,1411,181,1574]" captionTargetPageId="4" captionText="FIGURES 6‒10. Keratodegnathus javanicus sp. n., holotype female. Head in anterodorsal view (6); mouthparts in dorsal view (7); anteromedian region of labrum in anterodorsal view (8); maxillo-labial complex in ventral view (9); distal region of prelabium in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; clp, clypeus; cmb, connecting membrane; dmt, dorsal mandibular tooth; fr, frons; gal, galea; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; md, mandible; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pmns, premental sclerite; prl, prelabium; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.12190180" httpUri="https://zenodo.org/record/12190180/files/figure.png" pageId="3" pageNumber="358">Fig. 10</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFB5E926FED5FEA3FE39FECF" box="[365,424,259,284]" italics="true" pageId="3" pageNumber="358">pmns</emphasis>
|
|||
|
|
) narrowly separated at middle, each bearing one short lateromedian seta and one long anterolateral seta; labial palps with palpomere 1 (
|
|||
|
|
<figureCitation id="136AA7CAFFB5E926FD7DFE87FC88FE92" box="[709,793,295,321]" captionStart="FIGURES 6‒10" captionStartId="4.[151,264,1598,1622]" captionTargetBox="[177,1411,181,1574]" captionTargetId="figure-19@4.[177,1411,181,1574]" captionTargetPageId="4" captionText="FIGURES 6‒10. Keratodegnathus javanicus sp. n., holotype female. Head in anterodorsal view (6); mouthparts in dorsal view (7); anteromedian region of labrum in anterodorsal view (8); maxillo-labial complex in ventral view (9); distal region of prelabium in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; clp, clypeus; cmb, connecting membrane; dmt, dorsal mandibular tooth; fr, frons; gal, galea; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; md, mandible; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pmns, premental sclerite; prl, prelabium; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.12190180" httpUri="https://zenodo.org/record/12190180/files/figure.png" pageId="3" pageNumber="358">Fig. 10</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFB5E926FC9EFE87FCD8FE92" box="[806,841,295,321]" italics="true" pageId="3" pageNumber="358">lp1</emphasis>
|
|||
|
|
) strongly elongate, palpomere 2 (
|
|||
|
|
<figureCitation id="136AA7CAFFB5E926FB70FE87FA8DFE92" box="[1224,1308,295,321]" captionStart="FIGURES 6‒10" captionStartId="4.[151,264,1598,1622]" captionTargetBox="[177,1411,181,1574]" captionTargetId="figure-19@4.[177,1411,181,1574]" captionTargetPageId="4" captionText="FIGURES 6‒10. Keratodegnathus javanicus sp. n., holotype female. Head in anterodorsal view (6); mouthparts in dorsal view (7); anteromedian region of labrum in anterodorsal view (8); maxillo-labial complex in ventral view (9); distal region of prelabium in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; clp, clypeus; cmb, connecting membrane; dmt, dorsal mandibular tooth; fr, frons; gal, galea; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; md, mandible; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pmns, premental sclerite; prl, prelabium; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.12190180" httpUri="https://zenodo.org/record/12190180/files/figure.png" pageId="3" pageNumber="358">Fig. 10</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFB5E926FA91FE87FADCFE92" box="[1321,1357,295,321]" italics="true" pageId="3" pageNumber="358">lp2</emphasis>
|
|||
|
|
) much narrower and shorter than lp1, strongly elongate, palpomere 3 (
|
|||
|
|
<figureCitation id="136AA7CAFFB5E926FCEBFEEBFC34FEB6" box="[851,933,331,357]" captionStart="FIGURES 6‒10" captionStartId="4.[151,264,1598,1622]" captionTargetBox="[177,1411,181,1574]" captionTargetId="figure-19@4.[177,1411,181,1574]" captionTargetPageId="4" captionText="FIGURES 6‒10. Keratodegnathus javanicus sp. n., holotype female. Head in anterodorsal view (6); mouthparts in dorsal view (7); anteromedian region of labrum in anterodorsal view (8); maxillo-labial complex in ventral view (9); distal region of prelabium in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; clp, clypeus; cmb, connecting membrane; dmt, dorsal mandibular tooth; fr, frons; gal, galea; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; md, mandible; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pmns, premental sclerite; prl, prelabium; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.12190180" httpUri="https://zenodo.org/record/12190180/files/figure.png" pageId="3" pageNumber="358">Fig. 10</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFB5E926FC08FEEBFC45FEB6" box="[944,980,331,357]" italics="true" pageId="3" pageNumber="358">lp3</emphasis>
|
|||
|
|
) narrow and long, weakly curved, nearly rod-like with rounded apex; ligula (
|
|||
|
|
<figureCitation id="136AA7CAFFB5E926FD90FECFFDEDFE5A" box="[552,636,367,393]" captionStart="FIGURES 6‒10" captionStartId="4.[151,264,1598,1622]" captionTargetBox="[177,1411,181,1574]" captionTargetId="figure-19@4.[177,1411,181,1574]" captionTargetPageId="4" captionText="FIGURES 6‒10. Keratodegnathus javanicus sp. n., holotype female. Head in anterodorsal view (6); mouthparts in dorsal view (7); anteromedian region of labrum in anterodorsal view (8); maxillo-labial complex in ventral view (9); distal region of prelabium in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; clp, clypeus; cmb, connecting membrane; dmt, dorsal mandibular tooth; fr, frons; gal, galea; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; md, mandible; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pmns, premental sclerite; prl, prelabium; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.12190180" httpUri="https://zenodo.org/record/12190180/files/figure.png" pageId="3" pageNumber="358">Fig. 10</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFB5E926FD30FECFFD37FE5A" box="[648,678,367,393]" italics="true" pageId="3" pageNumber="358">lig</emphasis>
|
|||
|
|
) bipartite, composed of two separate elongate processes, each with two apical setae; lateral lobes of hypopharynx (
|
|||
|
|
<figureCitation id="136AA7CAFFB5E926FD11FE33FD6CFE7E" box="[681,765,403,429]" captionStart="FIGURES 6‒10" captionStartId="4.[151,264,1598,1622]" captionTargetBox="[177,1411,181,1574]" captionTargetId="figure-19@4.[177,1411,181,1574]" captionTargetPageId="4" captionText="FIGURES 6‒10. Keratodegnathus javanicus sp. n., holotype female. Head in anterodorsal view (6); mouthparts in dorsal view (7); anteromedian region of labrum in anterodorsal view (8); maxillo-labial complex in ventral view (9); distal region of prelabium in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; clp, clypeus; cmb, connecting membrane; dmt, dorsal mandibular tooth; fr, frons; gal, galea; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; md, mandible; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pmns, premental sclerite; prl, prelabium; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.12190180" httpUri="https://zenodo.org/record/12190180/files/figure.png" pageId="3" pageNumber="358">Fig. 10</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFB5E926FCB0FE33FCB7FE7E" box="[776,806,403,429]" italics="true" pageId="3" pageNumber="358">llh</emphasis>
|
|||
|
|
) short and rounded.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF2EEBC7FFB5E926FF2FF950FBE2F80C" ID-DOI="http://doi.org/10.5281/zenodo.12190176" ID-Zenodo-Dep="12190176" httpUri="https://zenodo.org/record/12190176/files/figure.png" pageId="3" pageNumber="358" startId="3.[151,264,1776,1800]" targetBox="[177,1411,453,1752]" targetPageId="3" targetType="figure">
|
|||
|
|
<paragraph id="8BEEBB4FFFB5E926FF2FF950FBE2F80C" blockId="3.[151,1437,1776,2016]" pageId="3" pageNumber="358">
|
|||
|
|
<emphasis id="B925675DFFB5E926FF2FF950FEACF8DB" bold="true" box="[151,317,1776,1800]" pageId="3" pageNumber="358">FIGURES 2‒5.</emphasis>
|
|||
|
|
<taxonomicName id="4C51C0CCFFB5E926FEFEF950FDCFF8DB" authority="Jałoszyński, 2024" authorityName="Jałoszyński" authorityYear="2024" box="[326,606,1776,1800]" class="Insecta" family="Staphylinidae" genus="Keratodegnathus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="358" phylum="Arthropoda" rank="species" species="javanicus" status="sp. nov.">
|
|||
|
|
<emphasis id="B925675DFFB5E926FEFEF950FDCFF8DB" box="[326,606,1776,1800]" italics="true" pageId="3" pageNumber="358">Keratodegnathus javanicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A216DA26FFB5E926FDDFF950FD30F8D4" box="[615,673,1776,1799]" pageId="3" pageNumber="358" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
|
, holotype female. Head and anterior region of thorax in lateral view (
|
|||
|
|
<emphasis id="B925675DFFB5E926FA38F950FA1CF8DB" bold="true" box="[1408,1421,1776,1800]" pageId="3" pageNumber="358">2</emphasis>
|
|||
|
|
); head in anterodorsal view (
|
|||
|
|
<emphasis id="B925675DFFB5E926FE08F8B4FE2CF8FF" bold="true" box="[432,445,1812,1836]" pageId="3" pageNumber="358">3</emphasis>
|
|||
|
|
); head, prothorax and elytral base in dorsal view (
|
|||
|
|
<emphasis id="B925675DFFB5E926FC7FF8B4FC45F8FF" bold="true" box="[967,980,1812,1836]" pageId="3" pageNumber="358">4</emphasis>
|
|||
|
|
; arrowheads indicate macrosetae); head and prothorax in ventral view (
|
|||
|
|
<emphasis id="B925675DFFB5E926FE12F898FE26F883" bold="true" box="[426,439,1848,1872]" pageId="3" pageNumber="358">5</emphasis>
|
|||
|
|
). Abbreviations: acxc, anterior coxal carina; cl, clypeus; cmb, connecting membrane; cs, cervical sclerite; dmt, dorsal mandibular tooth; fr, frons; gal, galea; gen, gena; gp, gular plate; gr, genal carina; gs, gular suture; hr, hypostomal ridge; hs, hypostomal suture; hyp, hypomeron; hyr, hypomeral ridge; lac, lacinia; lbr, labrum; lpc, lateral pronotal carina; mn, mentum; mxp2‒4, maxillary palpomere 2‒4; nss, notosternal suture; ocr, occipital ridge; pd, pedicel; prl prelabium; pst, prosternum; ptp, posterior tentorial pit; sc, scape; smn, submentum; trt, trochantin; vt, vertex.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="DF2EEBC7FFB2E921FF2FF99EFE78F8D9" ID-DOI="http://doi.org/10.5281/zenodo.12190180" ID-Zenodo-Dep="12190180" httpUri="https://zenodo.org/record/12190180/files/figure.png" pageId="4" pageNumber="359" startId="4.[151,264,1598,1622]" targetBox="[177,1411,181,1574]" targetPageId="4" targetType="figure">
|
|||
|
|
<paragraph id="8BEEBB4FFFB2E921FF2FF99EFE78F8D9" blockId="4.[151,1436,1598,1802]" pageId="4" pageNumber="359">
|
|||
|
|
<emphasis id="B925675DFFB2E921FF2FF99EFEDAF985" bold="true" box="[151,331,1598,1622]" pageId="4" pageNumber="359">FIGURES 6‒10.</emphasis>
|
|||
|
|
<taxonomicName id="4C51C0CCFFB2E921FEECF99EFDFFF985" authority="Jałoszyński, 2024" authorityName="Jałoszyński" authorityYear="2024" box="[340,622,1598,1622]" class="Insecta" family="Staphylinidae" genus="Keratodegnathus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="359" phylum="Arthropoda" rank="species" species="javanicus" status="sp. nov.">
|
|||
|
|
<emphasis id="B925675DFFB2E921FEECF99EFDFFF985" box="[340,622,1598,1622]" italics="true" pageId="4" pageNumber="359">Keratodegnathus javanicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A216DA26FFB2E921FDCFF99EFD23F986" box="[631,690,1598,1621]" pageId="4" pageNumber="359" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
|
, holotype female. Head in anterodorsal view (
|
|||
|
|
<emphasis id="B925675DFFB2E921FB27F99EFB3DF985" bold="true" box="[1183,1196,1598,1622]" pageId="4" pageNumber="359">6</emphasis>
|
|||
|
|
); mouthparts in dorsal view (
|
|||
|
|
<emphasis id="B925675DFFB2E921FF60F9C2FF74F9A9" bold="true" box="[216,229,1634,1658]" pageId="4" pageNumber="359">7</emphasis>
|
|||
|
|
); anteromedian region of labrum in anterodorsal view (
|
|||
|
|
<emphasis id="B925675DFFB2E921FCA7F9C2FCBDF9A9" bold="true" box="[799,812,1634,1658]" pageId="4" pageNumber="359">8</emphasis>
|
|||
|
|
); maxillo-labial complex in ventral view (
|
|||
|
|
<emphasis id="B925675DFFB2E921FB66F9C2FB7AF9A9" bold="true" box="[1246,1259,1634,1658]" pageId="4" pageNumber="359">9</emphasis>
|
|||
|
|
); distal region of prelabium in ventral view (
|
|||
|
|
<emphasis id="B925675DFFB2E921FE14F926FE54F94D" bold="true" box="[428,453,1670,1694]" pageId="4" pageNumber="359">10</emphasis>
|
|||
|
|
). Abbreviations: bst, basistipes; cd, cardo; clp, clypeus; cmb, connecting membrane; dmt, dorsal mandibular tooth; fr, frons; gal, galea; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; md, mandible; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pmns, premental sclerite; prl, prelabium; smn, submentum.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BEEBB4FFFB2E920FF7FF8FCFB71FF2A" blockId="4.[151,1437,1884,2019]" lastBlockId="5.[151,1437,151,249]" lastPageId="5" lastPageNumber="360" pageId="4" pageNumber="359">
|
|||
|
|
Antennae (
|
|||
|
|
<figureCitation id="136AA7CAFFB2E921FEFBF8FCFE1DF8A5" box="[323,396,1884,1910]" captionStart="FIGURE 1" captionStartId="2.[151,250,1651,1675]" captionTargetBox="[203,1391,314,1617]" captionTargetId="figure-88@2.[189,1398,306,1625]" captionTargetPageId="2" captionText="FIGURE 1. Keratodegnathus javanicus sp. n., holotype female, habitus in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12190172" httpUri="https://zenodo.org/record/12190172/files/figure.png" pageId="4" pageNumber="359">Figs 1</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="136AA7CAFFB2E921FE24F8FCFE3DF8A5" box="[412,428,1884,1910]" captionStart="FIGURES 2‒5" captionStartId="3.[151,264,1776,1800]" captionTargetBox="[177,1411,453,1752]" captionTargetId="figure-149@3.[177,1411,453,1752]" captionTargetPageId="3" captionText="FIGURES 2‒5. Keratodegnathus javanicus sp. n., holotype female. Head and anterior region of thorax in lateral view (2); head in anterodorsal view (3); head, prothorax and elytral base in dorsal view (4; arrowheads indicate macrosetae); head and prothorax in ventral view (5). Abbreviations: acxc, anterior coxal carina; cl, clypeus; cmb, connecting membrane; cs, cervical sclerite; dmt, dorsal mandibular tooth; fr, frons; gal, galea; gen, gena; gp, gular plate; gr, genal carina; gs, gular suture; hr, hypostomal ridge; hs, hypostomal suture; hyp, hypomeron; hyr, hypomeral ridge; lac, lacinia; lbr, labrum; lpc, lateral pronotal carina; mn, mentum; mxp2‒4, maxillary palpomere 2‒4; nss, notosternal suture; ocr, occipital ridge; pd, pedicel; prl prelabium; pst, prosternum; ptp, posterior tentorial pit; sc, scape; smn, submentum; trt, trochantin; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.12190176" httpUri="https://zenodo.org/record/12190176/files/figure.png" pageId="4" pageNumber="359">4</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="136AA7CAFFB2E921FE02F8FCFE58F8A5" box="[442,457,1884,1910]" captionStart="FIGURES 6‒10" captionStartId="4.[151,264,1598,1622]" captionTargetBox="[177,1411,181,1574]" captionTargetId="figure-19@4.[177,1411,181,1574]" captionTargetPageId="4" captionText="FIGURES 6‒10. Keratodegnathus javanicus sp. n., holotype female. Head in anterodorsal view (6); mouthparts in dorsal view (7); anteromedian region of labrum in anterodorsal view (8); maxillo-labial complex in ventral view (9); distal region of prelabium in ventral view (10). Abbreviations: bst, basistipes; cd, cardo; clp, clypeus; cmb, connecting membrane; dmt, dorsal mandibular tooth; fr, frons; gal, galea; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; md, mandible; md, mandible; mn, mentum; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pmns, premental sclerite; prl, prelabium; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.12190180" httpUri="https://zenodo.org/record/12190180/files/figure.png" pageId="4" pageNumber="359">6</figureCitation>
|
|||
|
|
) slender, AnL
|
|||
|
|
<quantity id="4CA916AAFFB2E921FDCCF8FCFD4CF8A4" box="[628,733,1884,1911]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="359" unit="mm" value="2.7">2.70 mm</quantity>
|
|||
|
|
. Scape about 4 times as long as broad, clavate, broadest near distal third; pedicel only 1/3 as long as scape and distinctly narrower, about twice as long as broad; antennomere 3 conspicuously elongate, 3 × as long as broad and broadening distally; antennomeres 4–9 each 2.2–2.5 × as long as broad, each broadening distally, antennomere 10 slightly shorter and stouter than 9, weakly broadening distally, antennomere 11 conspicuously long, slightly longer than 9 and 10 combined, almost 4 × as long as broad, gradually narrowing from base to apex. Scape and pedicel glossy, remaining antennomeres matte, densely covered with fine microreticulation, antennomeres 10 and 11 with much sparser setae than remaining flagellomeres.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF2EEBC7FFB3E920FF2FF9CDFD13F91E" ID-DOI="http://doi.org/10.5281/zenodo.12190182" ID-Zenodo-Dep="12190182" httpUri="https://zenodo.org/record/12190182/files/figure.png" pageId="5" pageNumber="360" startId="5.[151,250,1645,1669]" targetBox="[151,1435,273,1622]" targetPageId="5" targetType="figure">
|
|||
|
|
<paragraph id="8BEEBB4FFFB3E920FF2FF9CDFD13F91E" blockId="5.[151,1437,1645,1742]" pageId="5" pageNumber="360">
|
|||
|
|
<emphasis id="B925675DFFB3E920FF2FF9CDFEB3F956" bold="true" box="[151,290,1645,1669]" pageId="5" pageNumber="360">FIGURE 11.</emphasis>
|
|||
|
|
<taxonomicName id="4C51C0CCFFB3E920FE94F9CEFDD7F956" authority="Jałoszyński, 2024" authorityName="Jałoszyński" authorityYear="2024" box="[300,582,1646,1670]" class="Insecta" family="Staphylinidae" genus="Keratodegnathus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="360" phylum="Arthropoda" rank="species" species="javanicus" status="sp. nov.">
|
|||
|
|
<emphasis id="B925675DFFB3E920FE94F9CEFDD7F956" box="[300,582,1646,1670]" italics="true" pageId="5" pageNumber="360">Keratodegnathus javanicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A216DA26FFB3E920FDE8F9CEFD1AF956" box="[592,651,1646,1669]" pageId="5" pageNumber="360" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
|
, holotype female, posterior region of thorax and abdomen in dorsal view. Abbreviations: asc, adsutural carina; bc, basal carina; go1‒3, glandular opening type 1‒3 (each indicated by different arrowhead); pt, paratergite; sp, spiracle; tIII‒X, tergite III‒X.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BEEBB4FFFB3E923FF7FF880FC31FF2A" blockId="5.[151,1437,1824,2030]" lastBlockId="6.[151,1436,150,249]" lastPageId="6" lastPageNumber="361" pageId="5" pageNumber="360">
|
|||
|
|
Pronotum (
|
|||
|
|
<figureCitation id="136AA7CAFFB3E920FEFCF880FE36F8E9" box="[324,423,1824,1850]" captionStart="FIGURES 2‒5" captionStartId="3.[151,264,1776,1800]" captionTargetBox="[177,1411,453,1752]" captionTargetId="figure-149@3.[177,1411,453,1752]" captionTargetPageId="3" captionText="FIGURES 2‒5. Keratodegnathus javanicus sp. n., holotype female. Head and anterior region of thorax in lateral view (2); head in anterodorsal view (3); head, prothorax and elytral base in dorsal view (4; arrowheads indicate macrosetae); head and prothorax in ventral view (5). Abbreviations: acxc, anterior coxal carina; cl, clypeus; cmb, connecting membrane; cs, cervical sclerite; dmt, dorsal mandibular tooth; fr, frons; gal, galea; gen, gena; gp, gular plate; gr, genal carina; gs, gular suture; hr, hypostomal ridge; hs, hypostomal suture; hyp, hypomeron; hyr, hypomeral ridge; lac, lacinia; lbr, labrum; lpc, lateral pronotal carina; mn, mentum; mxp2‒4, maxillary palpomere 2‒4; nss, notosternal suture; ocr, occipital ridge; pd, pedicel; prl prelabium; pst, prosternum; ptp, posterior tentorial pit; sc, scape; smn, submentum; trt, trochantin; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.12190176" httpUri="https://zenodo.org/record/12190176/files/figure.png" pageId="5" pageNumber="360">Figs 2, 4</figureCitation>
|
|||
|
|
) strongly convex, subtrapezoidal, broadest at level of anterior corners, strongly transverse; PL
|
|||
|
|
<quantity id="4CA916AAFFB3E920FF04F8E4FEB1F88D" box="[188,288,1860,1886]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="5" pageNumber="360" unit="mm" value="0.68">0.68 mm</quantity>
|
|||
|
|
, PW
|
|||
|
|
<quantity id="4CA916AAFFB3E920FEE0F8E4FE2CF88D" box="[344,445,1860,1887]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="5" pageNumber="360" unit="mm" value="0.95">0.95 mm</quantity>
|
|||
|
|
.Anterior pronotal margin weakly arcuate; anterior pronotal corners slightly obtuse-angled and with rounded apices; lateral margins straight; posterior corners strongly obtuse-angled and blunt; posterior margin weakly arcuate. Entire pronotal disc smooth and with sparse, fine setiferous punctures distributed in anterior half and along lateral margins; four macrosetae present along each lateral margin (
|
|||
|
|
<figureCitation id="136AA7CAFFB3E920FB93F810FBE1F819" box="[1067,1136,1968,1994]" captionStart="FIGURES 2‒5" captionStartId="3.[151,264,1776,1800]" captionTargetBox="[177,1411,453,1752]" captionTargetId="figure-149@3.[177,1411,453,1752]" captionTargetPageId="3" captionText="FIGURES 2‒5. Keratodegnathus javanicus sp. n., holotype female. Head and anterior region of thorax in lateral view (2); head in anterodorsal view (3); head, prothorax and elytral base in dorsal view (4; arrowheads indicate macrosetae); head and prothorax in ventral view (5). Abbreviations: acxc, anterior coxal carina; cl, clypeus; cmb, connecting membrane; cs, cervical sclerite; dmt, dorsal mandibular tooth; fr, frons; gal, galea; gen, gena; gp, gular plate; gr, genal carina; gs, gular suture; hr, hypostomal ridge; hs, hypostomal suture; hyp, hypomeron; hyr, hypomeral ridge; lac, lacinia; lbr, labrum; lpc, lateral pronotal carina; mn, mentum; mxp2‒4, maxillary palpomere 2‒4; nss, notosternal suture; ocr, occipital ridge; pd, pedicel; prl prelabium; pst, prosternum; ptp, posterior tentorial pit; sc, scape; smn, submentum; trt, trochantin; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.12190176" httpUri="https://zenodo.org/record/12190176/files/figure.png" pageId="5" pageNumber="360">Fig. 4</figureCitation>
|
|||
|
|
; indicated by arrowheads), sparse suberect setae distributed in anterior half of pronotal disc. Lateral pronotal carinae (
|
|||
|
|
<figureCitation id="136AA7CAFFB3E920FB03F874FA95F83D" box="[1211,1284,2004,2030]" captionStart="FIGURES 2‒5" captionStartId="3.[151,264,1776,1800]" captionTargetBox="[177,1411,453,1752]" captionTargetId="figure-149@3.[177,1411,453,1752]" captionTargetPageId="3" captionText="FIGURES 2‒5. Keratodegnathus javanicus sp. n., holotype female. Head and anterior region of thorax in lateral view (2); head in anterodorsal view (3); head, prothorax and elytral base in dorsal view (4; arrowheads indicate macrosetae); head and prothorax in ventral view (5). Abbreviations: acxc, anterior coxal carina; cl, clypeus; cmb, connecting membrane; cs, cervical sclerite; dmt, dorsal mandibular tooth; fr, frons; gal, galea; gen, gena; gp, gular plate; gr, genal carina; gs, gular suture; hr, hypostomal ridge; hs, hypostomal suture; hyp, hypomeron; hyr, hypomeral ridge; lac, lacinia; lbr, labrum; lpc, lateral pronotal carina; mn, mentum; mxp2‒4, maxillary palpomere 2‒4; nss, notosternal suture; ocr, occipital ridge; pd, pedicel; prl prelabium; pst, prosternum; ptp, posterior tentorial pit; sc, scape; smn, submentum; trt, trochantin; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.12190176" httpUri="https://zenodo.org/record/12190176/files/figure.png" pageId="5" pageNumber="360">Fig. 2</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFB3E920FAACF874FAA7F83D" box="[1300,1334,2004,2030]" italics="true" pageId="5" pageNumber="360">lpc</emphasis>
|
|||
|
|
) sharply marked and complete; hypomera (
|
|||
|
|
<figureCitation id="136AA7CAFFB0E923FDAAFF37FDE4FF62" box="[530,629,150,177]" captionStart="FIGURES 2‒5" captionStartId="3.[151,264,1776,1800]" captionTargetBox="[177,1411,453,1752]" captionTargetId="figure-149@3.[177,1411,453,1752]" captionTargetPageId="3" captionText="FIGURES 2‒5. Keratodegnathus javanicus sp. n., holotype female. Head and anterior region of thorax in lateral view (2); head in anterodorsal view (3); head, prothorax and elytral base in dorsal view (4; arrowheads indicate macrosetae); head and prothorax in ventral view (5). Abbreviations: acxc, anterior coxal carina; cl, clypeus; cmb, connecting membrane; cs, cervical sclerite; dmt, dorsal mandibular tooth; fr, frons; gal, galea; gen, gena; gp, gular plate; gr, genal carina; gs, gular suture; hr, hypostomal ridge; hs, hypostomal suture; hyp, hypomeron; hyr, hypomeral ridge; lac, lacinia; lbr, labrum; lpc, lateral pronotal carina; mn, mentum; mxp2‒4, maxillary palpomere 2‒4; nss, notosternal suture; ocr, occipital ridge; pd, pedicel; prl prelabium; pst, prosternum; ptp, posterior tentorial pit; sc, scape; smn, submentum; trt, trochantin; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.12190176" httpUri="https://zenodo.org/record/12190176/files/figure.png" pageId="6" pageNumber="361">Figs 2, 5</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFB0E923FD39FF37FD38FF62" box="[641,681,151,177]" italics="true" pageId="6" pageNumber="361">hyp</emphasis>
|
|||
|
|
) exposed in lateral view, impunctate and asetose. Hypomeral ridges (
|
|||
|
|
<figureCitation id="136AA7CAFFB0E923FF27FF1BFF74FF06" box="[159,229,186,213]" captionStart="FIGURES 2‒5" captionStartId="3.[151,264,1776,1800]" captionTargetBox="[177,1411,453,1752]" captionTargetId="figure-149@3.[177,1411,453,1752]" captionTargetPageId="3" captionText="FIGURES 2‒5. Keratodegnathus javanicus sp. n., holotype female. Head and anterior region of thorax in lateral view (2); head in anterodorsal view (3); head, prothorax and elytral base in dorsal view (4; arrowheads indicate macrosetae); head and prothorax in ventral view (5). Abbreviations: acxc, anterior coxal carina; cl, clypeus; cmb, connecting membrane; cs, cervical sclerite; dmt, dorsal mandibular tooth; fr, frons; gal, galea; gen, gena; gp, gular plate; gr, genal carina; gs, gular suture; hr, hypostomal ridge; hs, hypostomal suture; hyp, hypomeron; hyr, hypomeral ridge; lac, lacinia; lbr, labrum; lpc, lateral pronotal carina; mn, mentum; mxp2‒4, maxillary palpomere 2‒4; nss, notosternal suture; ocr, occipital ridge; pd, pedicel; prl prelabium; pst, prosternum; ptp, posterior tentorial pit; sc, scape; smn, submentum; trt, trochantin; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.12190176" httpUri="https://zenodo.org/record/12190176/files/figure.png" pageId="6" pageNumber="361">Fig. 5</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFB0E923FF4BFF1BFE89FF06" box="[243,280,187,213]" italics="true" pageId="6" pageNumber="361">hyr</emphasis>
|
|||
|
|
) present, each situated close to outer margin of procoxal cavity and anteriorly connecting with lateral pronotal carina. Prosternum (
|
|||
|
|
<figureCitation id="136AA7CAFFB0E923FE66FF7FFDB2FF2A" box="[478,547,222,249]" captionStart="FIGURES 2‒5" captionStartId="3.[151,264,1776,1800]" captionTargetBox="[177,1411,453,1752]" captionTargetId="figure-149@3.[177,1411,453,1752]" captionTargetPageId="3" captionText="FIGURES 2‒5. Keratodegnathus javanicus sp. n., holotype female. Head and anterior region of thorax in lateral view (2); head in anterodorsal view (3); head, prothorax and elytral base in dorsal view (4; arrowheads indicate macrosetae); head and prothorax in ventral view (5). Abbreviations: acxc, anterior coxal carina; cl, clypeus; cmb, connecting membrane; cs, cervical sclerite; dmt, dorsal mandibular tooth; fr, frons; gal, galea; gen, gena; gp, gular plate; gr, genal carina; gs, gular suture; hr, hypostomal ridge; hs, hypostomal suture; hyp, hypomeron; hyr, hypomeral ridge; lac, lacinia; lbr, labrum; lpc, lateral pronotal carina; mn, mentum; mxp2‒4, maxillary palpomere 2‒4; nss, notosternal suture; ocr, occipital ridge; pd, pedicel; prl prelabium; pst, prosternum; ptp, posterior tentorial pit; sc, scape; smn, submentum; trt, trochantin; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.12190176" httpUri="https://zenodo.org/record/12190176/files/figure.png" pageId="6" pageNumber="361">Fig. 5</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFB0E923FD97FF7FFDC1FF2B" box="[559,592,223,248]" italics="true" pageId="6" pageNumber="361">pst</emphasis>
|
|||
|
|
) also impunctate and asetose.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF2EEBC7FFB0E923FF2FF8F1FB16F805" ID-DOI="http://doi.org/10.5281/zenodo.12190184" ID-Zenodo-Dep="12190184" httpUri="https://zenodo.org/record/12190184/files/figure.png" pageId="6" pageNumber="361" startId="6.[151,264,1873,1897]" targetBox="[151,1435,273,1850]" targetPageId="6" targetType="figure">
|
|||
|
|
<paragraph id="8BEEBB4FFFB0E923FF2FF8F1FB16F805" blockId="6.[151,1437,1873,2006]" pageId="6" pageNumber="361">
|
|||
|
|
<emphasis id="B925675DFFB0E923FF2FF8F1FEC4F8B9" bold="true" box="[151,341,1873,1898]" pageId="6" pageNumber="361">FIGURES 12‒18.</emphasis>
|
|||
|
|
<taxonomicName id="4C51C0CCFFB0E923FEE3F8F2FDE3F8BA" authority="Jałoszyński, 2024" authorityName="Jałoszyński" authorityYear="2024" box="[347,626,1874,1898]" class="Insecta" family="Staphylinidae" genus="Keratodegnathus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="361" phylum="Arthropoda" rank="species" species="javanicus" status="sp. nov.">
|
|||
|
|
<emphasis id="B925675DFFB0E923FEE3F8F2FDE3F8BA" box="[347,626,1874,1898]" italics="true" pageId="6" pageNumber="361">Keratodegnathus javanicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A216DA26FFB0E923FDC1F8F2FD20F8BA" box="[633,689,1874,1897]" pageId="6" pageNumber="361" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
|
, holotype female. Thorax and abdomen in ventral view (
|
|||
|
|
<emphasis id="B925675DFFB0E923FB4DF8F2FA9FF8B9" bold="true" box="[1269,1294,1874,1898]" pageId="6" pageNumber="361">12</emphasis>
|
|||
|
|
); protarsus in lateral (
|
|||
|
|
<emphasis id="B925675DFFB0E923FF5DF8D6FF6FF85D" bold="true" box="[229,254,1910,1934]" pageId="6" pageNumber="361">13</emphasis>
|
|||
|
|
) and lateroventral (
|
|||
|
|
<emphasis id="B925675DFFB0E923FE70F8D6FE70F85D" bold="true" box="[456,481,1910,1934]" pageId="6" pageNumber="361">14</emphasis>
|
|||
|
|
) views; mesotarsus in lateroventral view (
|
|||
|
|
<emphasis id="B925675DFFB0E923FC29F8D6FC3BF85D" bold="true" box="[913,938,1910,1934]" pageId="6" pageNumber="361">15</emphasis>
|
|||
|
|
); metatarsus in lateral view (
|
|||
|
|
<emphasis id="B925675DFFB0E923FB6BF8D6FB7DF85D" bold="true" box="[1235,1260,1910,1934]" pageId="6" pageNumber="361">16</emphasis>
|
|||
|
|
); distal region of metatarsomere 4 in lateroventral view (
|
|||
|
|
<emphasis id="B925675DFFB0E923FD95F83AFDD7F861" bold="true" box="[557,582,1946,1970]" pageId="6" pageNumber="361">17</emphasis>
|
|||
|
|
); details of claw insertion and empodium of metatarsus (
|
|||
|
|
<emphasis id="B925675DFFB0E923FB2DF83AFB3FF861" bold="true" box="[1173,1198,1946,1970]" pageId="6" pageNumber="361">18</emphasis>
|
|||
|
|
). Abbreviations: aest3, metanepisternum; es, empodial seta; mc, marginal carina; sIII‒IX, sternite III‒IX; v3, metaventrite.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="DF2EEBC7FFB1E922FF2FF9D1FD96F902" ID-DOI="http://doi.org/10.5281/zenodo.12190188" ID-Zenodo-Dep="12190188" httpUri="https://zenodo.org/record/12190188/files/figure.png" pageId="7" pageNumber="362" startId="7.[151,264,1649,1673]" targetBox="[164,1417,188,1625]" targetPageId="7" targetType="figure">
|
|||
|
|
<paragraph id="8BEEBB4FFFB1E922FF2FF9D1FD96F902" blockId="7.[151,1436,1649,1746]" pageId="7" pageNumber="362">
|
|||
|
|
<emphasis id="B925675DFFB1E922FF2FF9D1FEC4F95A" bold="true" box="[151,341,1649,1673]" pageId="7" pageNumber="362">FIGURES 19‒24.</emphasis>
|
|||
|
|
<taxonomicName id="4C51C0CCFFB1E922FEE3F9D1FDE3F95A" authority="Jałoszyński, 2024" authorityName="Jałoszyński" authorityYear="2024" box="[347,626,1649,1673]" class="Insecta" family="Staphylinidae" genus="Keratodegnathus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="362" phylum="Arthropoda" rank="species" species="javanicus" status="sp. nov.">
|
|||
|
|
<emphasis id="B925675DFFB1E922FEE3F9D1FDE3F95A" box="[347,626,1649,1673]" italics="true" pageId="7" pageNumber="362">Keratodegnathus javanicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A216DA26FFB1E922FDC1F9D2FD23F95A" box="[633,690,1650,1673]" pageId="7" pageNumber="362" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
|
, holotype female. Sternite VIII (
|
|||
|
|
<emphasis id="B925675DFFB1E922FBB8F9D1FB88F95A" bold="true" box="[1024,1049,1649,1673]" pageId="7" pageNumber="362">19</emphasis>
|
|||
|
|
); tergite VIII (
|
|||
|
|
<emphasis id="B925675DFFB1E922FB09F9D1FB5BF95A" bold="true" box="[1201,1226,1649,1673]" pageId="7" pageNumber="362">20</emphasis>
|
|||
|
|
); segment IX and X (
|
|||
|
|
<emphasis id="B925675DFFB1E922FF18F935FF28F97E" bold="true" box="[160,185,1685,1709]" pageId="7" pageNumber="362">21</emphasis>
|
|||
|
|
); mediobasal region of segments IX and X (
|
|||
|
|
<emphasis id="B925675DFFB1E922FDCDF935FD1FF97E" bold="true" box="[629,654,1685,1709]" pageId="7" pageNumber="362">22</emphasis>
|
|||
|
|
); spermatheca (
|
|||
|
|
<emphasis id="B925675DFFB1E922FC88F935FCFEF97E" bold="true" box="[816,879,1685,1709]" pageId="7" pageNumber="362">23‒24</emphasis>
|
|||
|
|
). Abbreviations: tIX‒X, tergite IX‒X; vp, vaginal plate (sensu
|
|||
|
|
<bibRefCitation id="EFC0C6BEFFB1E922FF66F91AFE03F902" author="Maruyama, M." box="[222,402,1721,1746]" pageId="7" pageNumber="362" pagination="1 - 207" refId="ref3950" refString="Maruyama, M. (2006) Revision of the Palearctic species of the myrmecophilous genus Pella (Coleoptera, Staphylinidae, Aleocharinae). National Science Monographs, 32, 1 - 207." type="journal article" year="2006">Maruyama (2006</bibRefCitation>
|
|||
|
|
: fig. 16A)).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BEEBB4FFFB1E922FF7FF883FC1CF822" blockId="7.[151,1437,1827,2034]" pageId="7" pageNumber="362">
|
|||
|
|
Elytra (
|
|||
|
|
<figureCitation id="136AA7CAFFB1E922FEA6F883FEE1F8EE" box="[286,368,1827,1853]" captionStart="FIGURE 11" captionStartId="5.[151,250,1645,1669]" captionTargetBox="[151,1435,273,1622]" captionTargetId="figure-62@5.[151,1435,273,1622]" captionTargetPageId="5" captionText="FIGURE 11. Keratodegnathus javanicus sp. n., holotype female, posterior region of thorax and abdomen in dorsal view. Abbreviations:asc, adsutural carina; bc, basal carina; go1‒3, glandular opening type 1‒3 (each indicated by different arrowhead); pt, paratergite; sp, spiracle; tIII‒X, tergite III‒X." figureDoi="http://doi.org/10.5281/zenodo.12190182" httpUri="https://zenodo.org/record/12190182/files/figure.png" pageId="7" pageNumber="362">Fig. 11</figureCitation>
|
|||
|
|
) together subtrapezoidal, broadest at posterior corners; EL
|
|||
|
|
<quantity id="4CA916AAFFB1E922FB9AF883FB1AF8ED" box="[1058,1163,1827,1854]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="7" pageNumber="362" unit="mm" value="0.75">0.75 mm</quantity>
|
|||
|
|
, EW
|
|||
|
|
<quantity id="4CA916AAFFB1E922FB77F883FAA8F8ED" box="[1231,1337,1827,1854]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="362" unit="mm" value="1.3">1.30 mm</quantity>
|
|||
|
|
. Humeri obtuse-angled and not bulging, lateral elytral margins weakly rounded and diverging posteriorly, posterior elytral margins weakly rounded, posterolateral elytral corners blunt and nearly right-angled, adsutural posterior corners obtuse-angled and rounded, elytral suture accentuated by fine adsutural carina (
|
|||
|
|
<figureCitation id="136AA7CAFFB1E922FBBFF82FFBC8F87A" box="[1031,1113,1935,1961]" captionStart="FIGURE 11" captionStartId="5.[151,250,1645,1669]" captionTargetBox="[151,1435,273,1622]" captionTargetId="figure-62@5.[151,1435,273,1622]" captionTargetPageId="5" captionText="FIGURE 11. Keratodegnathus javanicus sp. n., holotype female, posterior region of thorax and abdomen in dorsal view. Abbreviations:asc, adsutural carina; bc, basal carina; go1‒3, glandular opening type 1‒3 (each indicated by different arrowhead); pt, paratergite; sp, spiracle; tIII‒X, tergite III‒X." figureDoi="http://doi.org/10.5281/zenodo.12190182" httpUri="https://zenodo.org/record/12190182/files/figure.png" pageId="7" pageNumber="362">Fig. 11</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFB1E922FBDBF830FB19F87A" box="[1123,1160,1936,1961]" italics="true" pageId="7" pageNumber="362">asc</emphasis>
|
|||
|
|
) on each elytron. Elytral disc sparsely punctate, covered with short and sparse suberect setae, each elytron with longitudinal row of four macrosetae along lateral margin (
|
|||
|
|
<figureCitation id="136AA7CAFFB1E922FDB4F877FDCEF822" box="[524,607,2007,2033]" captionStart="FIGURE 11" captionStartId="5.[151,250,1645,1669]" captionTargetBox="[151,1435,273,1622]" captionTargetId="figure-62@5.[151,1435,273,1622]" captionTargetPageId="5" captionText="FIGURE 11. Keratodegnathus javanicus sp. n., holotype female, posterior region of thorax and abdomen in dorsal view. Abbreviations:asc, adsutural carina; bc, basal carina; go1‒3, glandular opening type 1‒3 (each indicated by different arrowhead); pt, paratergite; sp, spiracle; tIII‒X, tergite III‒X." figureDoi="http://doi.org/10.5281/zenodo.12190182" httpUri="https://zenodo.org/record/12190182/files/figure.png" pageId="7" pageNumber="362">Fig. 11</figureCitation>
|
|||
|
|
; indicated by arrowheads).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BEEBB4FFFBEE92DFF7FFF37FE18FF62" blockId="8.[151,1437,151,1401]" box="[199,393,151,177]" pageId="8" pageNumber="363">Hind wings long.</paragraph>
|
|||
|
|
<paragraph id="8BEEBB4FFFBEE92DFF7FFF1BFE8EFF2A" blockId="8.[151,1437,151,1401]" pageId="8" pageNumber="363">
|
|||
|
|
Mesoscutellar shield largely exposed between elytral bases (
|
|||
|
|
<figureCitation id="136AA7CAFFBEE92DFCDFFF1BFC3AFF06" box="[871,939,187,213]" captionStart="FIGURES 2‒5" captionStartId="3.[151,264,1776,1800]" captionTargetBox="[177,1411,453,1752]" captionTargetId="figure-149@3.[177,1411,453,1752]" captionTargetPageId="3" captionText="FIGURES 2‒5. Keratodegnathus javanicus sp. n., holotype female. Head and anterior region of thorax in lateral view (2); head in anterodorsal view (3); head, prothorax and elytral base in dorsal view (4; arrowheads indicate macrosetae); head and prothorax in ventral view (5). Abbreviations: acxc, anterior coxal carina; cl, clypeus; cmb, connecting membrane; cs, cervical sclerite; dmt, dorsal mandibular tooth; fr, frons; gal, galea; gen, gena; gp, gular plate; gr, genal carina; gs, gular suture; hr, hypostomal ridge; hs, hypostomal suture; hyp, hypomeron; hyr, hypomeral ridge; lac, lacinia; lbr, labrum; lpc, lateral pronotal carina; mn, mentum; mxp2‒4, maxillary palpomere 2‒4; nss, notosternal suture; ocr, occipital ridge; pd, pedicel; prl prelabium; pst, prosternum; ptp, posterior tentorial pit; sc, scape; smn, submentum; trt, trochantin; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.12190176" httpUri="https://zenodo.org/record/12190176/files/figure.png" pageId="8" pageNumber="363">Fig. 4</figureCitation>
|
|||
|
|
), its visible region subtriangular, impunctate and asetose.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BEEBB4FFFBEE92DFF7FFEA3FB08FEB6" blockId="8.[151,1437,151,1401]" pageId="8" pageNumber="363">
|
|||
|
|
Legs (
|
|||
|
|
<figureCitation id="136AA7CAFFBEE92DFEB4FEA3FEDAFECE" box="[268,331,259,285]" captionStart="FIGURE 1" captionStartId="2.[151,250,1651,1675]" captionTargetBox="[203,1391,314,1617]" captionTargetId="figure-88@2.[189,1398,306,1625]" captionTargetPageId="2" captionText="FIGURE 1. Keratodegnathus javanicus sp. n., holotype female, habitus in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12190172" httpUri="https://zenodo.org/record/12190172/files/figure.png" pageId="8" pageNumber="363">Fig. 1</figureCitation>
|
|||
|
|
) long and slender, all tibiae straight, tarsi (
|
|||
|
|
<figureCitation id="136AA7CAFFBEE92DFC9DFEA3FC35FECE" box="[805,932,259,285]" captionStart="FIGURES 12‒18" captionStartId="6.[151,264,1873,1897]" captionTargetBox="[151,1435,273,1850]" captionTargetId="figure-67@6.[151,1435,273,1850]" captionTargetPageId="6" captionText="FIGURES 12‒18. Keratodegnathus javanicus sp. n., holotype female. Thorax and abdomen in ventral view (12); protarsus in lateral (13) and lateroventral (14) views; mesotarsus in lateroventral view (15); metatarsus in lateral view (16); distal region of metatarsomere 4 in lateroventral view (17); details of claw insertion and empodium of metatarsus (18). Abbreviations: aest3, metanepisternum; es, empodial seta; mc, marginal carina; sIII‒IX, sternite III‒IX; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.12190184" httpUri="https://zenodo.org/record/12190184/files/figure.png" pageId="8" pageNumber="363">Figs 13‒16</figureCitation>
|
|||
|
|
) 4-5-5, increasingly longer from fore to hind, each terminal tarsomere with distoventral set of pointed processes (
|
|||
|
|
<figureCitation id="136AA7CAFFBEE92DFC30FE87FC48FE92" box="[904,985,295,321]" captionStart="FIGURES 12‒18" captionStartId="6.[151,264,1873,1897]" captionTargetBox="[151,1435,273,1850]" captionTargetId="figure-67@6.[151,1435,273,1850]" captionTargetPageId="6" captionText="FIGURES 12‒18. Keratodegnathus javanicus sp. n., holotype female. Thorax and abdomen in ventral view (12); protarsus in lateral (13) and lateroventral (14) views; mesotarsus in lateroventral view (15); metatarsus in lateral view (16); distal region of metatarsomere 4 in lateroventral view (17); details of claw insertion and empodium of metatarsus (18). Abbreviations: aest3, metanepisternum; es, empodial seta; mc, marginal carina; sIII‒IX, sternite III‒IX; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.12190184" httpUri="https://zenodo.org/record/12190184/files/figure.png" pageId="8" pageNumber="363">Fig. 18</figureCitation>
|
|||
|
|
) and with single empodial seta (
|
|||
|
|
<figureCitation id="136AA7CAFFBEE92DFAFCFE87FA06FE92" box="[1348,1431,295,321]" captionStart="FIGURES 12‒18" captionStartId="6.[151,264,1873,1897]" captionTargetBox="[151,1435,273,1850]" captionTargetId="figure-67@6.[151,1435,273,1850]" captionTargetPageId="6" captionText="FIGURES 12‒18. Keratodegnathus javanicus sp. n., holotype female. Thorax and abdomen in ventral view (12); protarsus in lateral (13) and lateroventral (14) views; mesotarsus in lateroventral view (15); metatarsus in lateral view (16); distal region of metatarsomere 4 in lateroventral view (17); details of claw insertion and empodium of metatarsus (18). Abbreviations: aest3, metanepisternum; es, empodial seta; mc, marginal carina; sIII‒IX, sternite III‒IX; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.12190184" httpUri="https://zenodo.org/record/12190184/files/figure.png" pageId="8" pageNumber="363">Fig. 18</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFBEE92DFF2FFEEBFF3FFEB7" box="[151,174,331,356]" italics="true" pageId="8" pageNumber="363">es</emphasis>
|
|||
|
|
) inserted on elongate subcylindrical cuticular process bearing its own minute lateral seta.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BEEBB4FFFBEE92DFF7FFECFFCF8FD1E" blockId="8.[151,1437,151,1401]" pageId="8" pageNumber="363">
|
|||
|
|
Abdomen (
|
|||
|
|
<figureCitation id="136AA7CAFFBEE92DFEFFFECFFE01FE5A" box="[327,400,367,393]" captionStart="FIGURE 1" captionStartId="2.[151,250,1651,1675]" captionTargetBox="[203,1391,314,1617]" captionTargetId="figure-88@2.[189,1398,306,1625]" captionTargetPageId="2" captionText="FIGURE 1. Keratodegnathus javanicus sp. n., holotype female, habitus in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12190172" httpUri="https://zenodo.org/record/12190172/files/figure.png" pageId="8" pageNumber="363">Figs 1</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="136AA7CAFFBEE92DFE18FECFFE77FE5A" box="[416,486,367,393]" captionStart-0="FIGURE 11" captionStart-1="FIGURES 12‒18" captionStartId-0="5.[151,250,1645,1669]" captionStartId-1="6.[151,264,1873,1897]" captionTargetBox-0="[151,1435,273,1622]" captionTargetBox-1="[151,1435,273,1850]" captionTargetId-0="figure-62@5.[151,1435,273,1622]" captionTargetId-1="figure-67@6.[151,1435,273,1850]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 11. Keratodegnathus javanicus sp. n., holotype female, posterior region of thorax and abdomen in dorsal view. Abbreviations:asc, adsutural carina; bc, basal carina; go1‒3, glandular opening type 1‒3 (each indicated by different arrowhead); pt, paratergite; sp, spiracle; tIII‒X, tergite III‒X." captionText-1="FIGURES 12‒18. Keratodegnathus javanicus sp. n., holotype female. Thorax and abdomen in ventral view (12); protarsus in lateral (13) and lateroventral (14) views; mesotarsus in lateroventral view (15); metatarsus in lateral view (16); distal region of metatarsomere 4 in lateroventral view (17); details of claw insertion and empodium of metatarsus (18). Abbreviations: aest3, metanepisternum; es, empodial seta; mc, marginal carina; sIII‒IX, sternite III‒IX; v3, metaventrite." figureDoi-0="http://doi.org/10.5281/zenodo.12190182" figureDoi-1="http://doi.org/10.5281/zenodo.12190184" httpUri-0="https://zenodo.org/record/12190182/files/figure.png" httpUri-1="https://zenodo.org/record/12190184/files/figure.png" pageId="8" pageNumber="363">11‒12</figureCitation>
|
|||
|
|
) at base indistinctly narrower than elytra, slightly longer than head, pronotum and elytra combined; AbL 3.00 mm, AbW
|
|||
|
|
<quantity id="4CA916AAFFBEE92DFDE6FE33FD58FE7E" box="[606,713,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.28" pageId="8" pageNumber="363" unit="mm" value="1.28">1.28 mm</quantity>
|
|||
|
|
. Tergites III‒VI (
|
|||
|
|
<figureCitation id="136AA7CAFFBEE92DFC2EFE33FC7BFE7E" box="[918,1002,403,429]" captionStart="FIGURE 11" captionStartId="5.[151,250,1645,1669]" captionTargetBox="[151,1435,273,1622]" captionTargetId="figure-62@5.[151,1435,273,1622]" captionTargetPageId="5" captionText="FIGURE 11. Keratodegnathus javanicus sp. n., holotype female, posterior region of thorax and abdomen in dorsal view. Abbreviations:asc, adsutural carina; bc, basal carina; go1‒3, glandular opening type 1‒3 (each indicated by different arrowhead); pt, paratergite; sp, spiracle; tIII‒X, tergite III‒X." figureDoi="http://doi.org/10.5281/zenodo.12190182" httpUri="https://zenodo.org/record/12190182/files/figure.png" pageId="8" pageNumber="363">Fig. 11</figureCitation>
|
|||
|
|
) each with anterior transverse carina with its ends curved towards spiracle. Tergites III‒VI each with four setae along posterior margin and one pair of lateromedian setae; tergite VII with one pair of lateromedian setae and one pair of lateral setae in posterior third; tergite VIII (
|
|||
|
|
<figureCitation id="136AA7CAFFBEE92DFE9DFE5FFEE8FDCA" box="[293,377,511,537]" captionStart="FIGURES 19‒24" captionStartId="7.[151,264,1649,1673]" captionTargetBox="[164,1417,188,1625]" captionTargetId="figure-15@7.[164,1423,181,1625]" captionTargetPageId="7" captionText="FIGURES 19‒24. Keratodegnathus javanicus sp. n., holotype female. Sternite VIII (19); tergite VIII (20); segment IX and X (21); mediobasal region of segments IX and X (22); spermatheca (23‒24). Abbreviations: tIX‒X, tergite IX‒X; vp, vaginal plate (sensu Maruyama (2006: fig. 16A))." figureDoi="http://doi.org/10.5281/zenodo.12190188" httpUri="https://zenodo.org/record/12190188/files/figure.png" pageId="8" pageNumber="363">Fig. 20</figureCitation>
|
|||
|
|
, most macrosetae broken off) elongate subtrapezoidal with transverse rows of more numerous submedian and distal setae. Tergite VII (
|
|||
|
|
<figureCitation id="136AA7CAFFBEE92DFDE7FD83FD3EFDEE" box="[607,687,547,573]" captionStart="FIGURE 11" captionStartId="5.[151,250,1645,1669]" captionTargetBox="[151,1435,273,1622]" captionTargetId="figure-62@5.[151,1435,273,1622]" captionTargetPageId="5" captionText="FIGURE 11. Keratodegnathus javanicus sp. n., holotype female, posterior region of thorax and abdomen in dorsal view. Abbreviations:asc, adsutural carina; bc, basal carina; go1‒3, glandular opening type 1‒3 (each indicated by different arrowhead); pt, paratergite; sp, spiracle; tIII‒X, tergite III‒X." figureDoi="http://doi.org/10.5281/zenodo.12190182" httpUri="https://zenodo.org/record/12190182/files/figure.png" pageId="8" pageNumber="363">Fig. 11</figureCitation>
|
|||
|
|
) with posterior band of densely set microtrichia. Inner component of each paratergite on segments III‒VI with one minute distal seta, outer component with one long submedian seta and a group of minute distal setae. Sternites (
|
|||
|
|
<figureCitation id="136AA7CAFFBEE92DFD30FDCBFD49FD56" box="[648,728,619,645]" captionStart="FIGURES 12‒18" captionStartId="6.[151,264,1873,1897]" captionTargetBox="[151,1435,273,1850]" captionTargetId="figure-67@6.[151,1435,273,1850]" captionTargetPageId="6" captionText="FIGURES 12‒18. Keratodegnathus javanicus sp. n., holotype female. Thorax and abdomen in ventral view (12); protarsus in lateral (13) and lateroventral (14) views; mesotarsus in lateroventral view (15); metatarsus in lateral view (16); distal region of metatarsomere 4 in lateroventral view (17); details of claw insertion and empodium of metatarsus (18). Abbreviations: aest3, metanepisternum; es, empodial seta; mc, marginal carina; sIII‒IX, sternite III‒IX; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.12190184" httpUri="https://zenodo.org/record/12190184/files/figure.png" pageId="8" pageNumber="363">Fig. 12</figureCitation>
|
|||
|
|
) relatively densely setose; sternite VIII (
|
|||
|
|
<figureCitation id="136AA7CAFFBEE92DFB22FDCBFB7AFD56" box="[1178,1259,619,645]" captionStart="FIGURES 19‒24" captionStartId="7.[151,264,1649,1673]" captionTargetBox="[164,1417,188,1625]" captionTargetId="figure-15@7.[164,1423,181,1625]" captionTargetPageId="7" captionText="FIGURES 19‒24. Keratodegnathus javanicus sp. n., holotype female. Sternite VIII (19); tergite VIII (20); segment IX and X (21); mediobasal region of segments IX and X (22); spermatheca (23‒24). Abbreviations: tIX‒X, tergite IX‒X; vp, vaginal plate (sensu Maruyama (2006: fig. 16A))." figureDoi="http://doi.org/10.5281/zenodo.12190188" httpUri="https://zenodo.org/record/12190188/files/figure.png" pageId="8" pageNumber="363">Fig. 19</figureCitation>
|
|||
|
|
) subtrapezoidal with rounded posterior margin. Abdominal segments IX and X (
|
|||
|
|
<figureCitation id="136AA7CAFFBEE92DFCD0FD2FFC2BFD7A" box="[872,954,655,681]" captionStart="FIGURES 19‒24" captionStartId="7.[151,264,1649,1673]" captionTargetBox="[164,1417,188,1625]" captionTargetId="figure-15@7.[164,1423,181,1625]" captionTargetPageId="7" captionText="FIGURES 19‒24. Keratodegnathus javanicus sp. n., holotype female. Sternite VIII (19); tergite VIII (20); segment IX and X (21); mediobasal region of segments IX and X (22); spermatheca (23‒24). Abbreviations: tIX‒X, tergite IX‒X; vp, vaginal plate (sensu Maruyama (2006: fig. 16A))." figureDoi="http://doi.org/10.5281/zenodo.12190188" httpUri="https://zenodo.org/record/12190188/files/figure.png" pageId="8" pageNumber="363">Fig. 21</figureCitation>
|
|||
|
|
) with a pair of large sclerotized structures, possibly homologous with ‘vaginal plates’ of
|
|||
|
|
<bibRefCitation id="EFC0C6BEFFBEE92DFD2EFD13FCF4FD1E" author="Maruyama, M." box="[662,869,691,717]" pageId="8" pageNumber="363" pagination="1 - 207" refId="ref3950" refString="Maruyama, M. (2006) Revision of the Palearctic species of the myrmecophilous genus Pella (Coleoptera, Staphylinidae, Aleocharinae). National Science Monographs, 32, 1 - 207." type="journal article" year="2006">Maruyama (2006)</bibRefCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BEEBB4FFFBEE92DFF7FFD77FCF2FC8E" blockId="8.[151,1437,151,1401]" pageId="8" pageNumber="363">
|
|||
|
|
Abdominal glandular openings (
|
|||
|
|
<figureCitation id="136AA7CAFFBEE92DFD88FD77FD10FD22" box="[560,641,727,753]" captionStart="FIGURE 11" captionStartId="5.[151,250,1645,1669]" captionTargetBox="[151,1435,273,1622]" captionTargetId="figure-62@5.[151,1435,273,1622]" captionTargetPageId="5" captionText="FIGURE 11. Keratodegnathus javanicus sp. n., holotype female, posterior region of thorax and abdomen in dorsal view. Abbreviations:asc, adsutural carina; bc, basal carina; go1‒3, glandular opening type 1‒3 (each indicated by different arrowhead); pt, paratergite; sp, spiracle; tIII‒X, tergite III‒X." figureDoi="http://doi.org/10.5281/zenodo.12190182" httpUri="https://zenodo.org/record/12190182/files/figure.png" pageId="8" pageNumber="363">Fig. 11</figureCitation>
|
|||
|
|
). Tergite IV with minute round glandular opening (
|
|||
|
|
<figureCitation id="136AA7CAFFBEE92DFB05FD77FA9EFD22" box="[1213,1295,727,753]" captionStart="FIGURE 11" captionStartId="5.[151,250,1645,1669]" captionTargetBox="[151,1435,273,1622]" captionTargetId="figure-62@5.[151,1435,273,1622]" captionTargetPageId="5" captionText="FIGURE 11. Keratodegnathus javanicus sp. n., holotype female, posterior region of thorax and abdomen in dorsal view. Abbreviations:asc, adsutural carina; bc, basal carina; go1‒3, glandular opening type 1‒3 (each indicated by different arrowhead); pt, paratergite; sp, spiracle; tIII‒X, tergite III‒X." figureDoi="http://doi.org/10.5281/zenodo.12190182" httpUri="https://zenodo.org/record/12190182/files/figure.png" pageId="8" pageNumber="363">Fig. 11</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFBEE92DFAA3FD77FAD4FD22" box="[1307,1349,727,753]" italics="true" pageId="8" pageNumber="363">go1</emphasis>
|
|||
|
|
) mesad each spiracle. Sparse single openings surrounded by five cuticular processes (
|
|||
|
|
<figureCitation id="136AA7CAFFBEE92DFBA2FD5BFBFEFCC6" box="[1050,1135,763,789]" captionStart="FIGURE 11" captionStartId="5.[151,250,1645,1669]" captionTargetBox="[151,1435,273,1622]" captionTargetId="figure-62@5.[151,1435,273,1622]" captionTargetPageId="5" captionText="FIGURE 11. Keratodegnathus javanicus sp. n., holotype female, posterior region of thorax and abdomen in dorsal view. Abbreviations:asc, adsutural carina; bc, basal carina; go1‒3, glandular opening type 1‒3 (each indicated by different arrowhead); pt, paratergite; sp, spiracle; tIII‒X, tergite III‒X." figureDoi="http://doi.org/10.5281/zenodo.12190182" httpUri="https://zenodo.org/record/12190182/files/figure.png" pageId="8" pageNumber="363">Fig. 11</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFBEE92DFBC6FD5BFB36FCC6" box="[1150,1191,763,789]" italics="true" pageId="8" pageNumber="363">go2</emphasis>
|
|||
|
|
), found also only on tergite IV. Numerous minute pore plates (
|
|||
|
|
<figureCitation id="136AA7CAFFBEE92DFDD6FCBFFD50FCEA" box="[622,705,799,825]" captionStart="FIGURE 11" captionStartId="5.[151,250,1645,1669]" captionTargetBox="[151,1435,273,1622]" captionTargetId="figure-62@5.[151,1435,273,1622]" captionTargetPageId="5" captionText="FIGURE 11. Keratodegnathus javanicus sp. n., holotype female, posterior region of thorax and abdomen in dorsal view. Abbreviations:asc, adsutural carina; bc, basal carina; go1‒3, glandular opening type 1‒3 (each indicated by different arrowhead); pt, paratergite; sp, spiracle; tIII‒X, tergite III‒X." figureDoi="http://doi.org/10.5281/zenodo.12190182" httpUri="https://zenodo.org/record/12190182/files/figure.png" pageId="8" pageNumber="363">Fig. 11</figureCitation>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B925675DFFBEE92DFD76FCBFFD68FCEA" box="[718,761,799,825]" italics="true" pageId="8" pageNumber="363">go3</emphasis>
|
|||
|
|
) composed of 5‒6 oval or irregular openings present on all tergites, inside each opening 1‒3 microscopic pores are present.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BEEBB4FFFBEE92DFF7FFCC7FC46FC76" blockId="8.[151,1437,151,1401]" pageId="8" pageNumber="363">
|
|||
|
|
Spermatheca (
|
|||
|
|
<figureCitation id="136AA7CAFFBEE92DFED0FCC7FE78FC52" box="[360,489,871,897]" captionStart="FIGURES 19‒24" captionStartId="7.[151,264,1649,1673]" captionTargetBox="[164,1417,188,1625]" captionTargetId="figure-15@7.[164,1423,181,1625]" captionTargetPageId="7" captionText="FIGURES 19‒24. Keratodegnathus javanicus sp. n., holotype female. Sternite VIII (19); tergite VIII (20); segment IX and X (21); mediobasal region of segments IX and X (22); spermatheca (23‒24). Abbreviations: tIX‒X, tergite IX‒X; vp, vaginal plate (sensu Maruyama (2006: fig. 16A))." figureDoi="http://doi.org/10.5281/zenodo.12190188" httpUri="https://zenodo.org/record/12190188/files/figure.png" pageId="8" pageNumber="363">Figs 23‒24</figureCitation>
|
|||
|
|
)
|
|||
|
|
<quantity id="4CA916AAFFBEE92DFE41FCC7FDCFFC52" box="[505,606,871,897]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="8" pageNumber="363" unit="mm" value="0.48">0.48 mm</quantity>
|
|||
|
|
in length, with slender receptacle and a long duct tightly coiled around a large spheroid reservoir, duct coils form elongate, flat and ovoid structure.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BEEBB4FFFBEE92DFF7FFC0FFE14FC1A" blockId="8.[151,1437,151,1401]" box="[199,389,943,969]" pageId="8" pageNumber="363">Male. Unknown.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C34BE8C4FFBEE92DFF7FFC72FE30FC3E" box="[199,417,978,1005]" pageId="8" pageNumber="363" type="distribution">
|
|||
|
|
<paragraph id="8BEEBB4FFFBEE92DFF7FFC72FE30FC3E" blockId="8.[151,1437,151,1401]" box="[199,417,978,1005]" pageId="8" pageNumber="363">
|
|||
|
|
<emphasis id="B925675DFFBEE92DFF7FFC72FEF0FC3F" bold="true" box="[199,353,978,1004]" pageId="8" pageNumber="363">Distribution.</emphasis>
|
|||
|
|
<collectingRegion id="499575ADFFBEE92DFEDFFC73FE0DFC3E" box="[359,412,979,1005]" country="Indonesia" name="Jawa Timur" pageId="8" pageNumber="363">Java</collectingRegion>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C34BE8C4FFBEE92DFF7FFC57FC12FBC2" box="[199,899,1015,1041]" pageId="8" pageNumber="363" type="etymology">
|
|||
|
|
<paragraph id="8BEEBB4FFFBEE92DFF7FFC57FC12FBC2" blockId="8.[151,1437,151,1401]" box="[199,899,1015,1041]" pageId="8" pageNumber="363">
|
|||
|
|
<emphasis id="B925675DFFBEE92DFF7FFC57FEDFFBC2" bold="true" box="[199,334,1015,1041]" pageId="8" pageNumber="363">Etymology.</emphasis>
|
|||
|
|
The adjective
|
|||
|
|
<taxonomicName id="4C51C0CCFFBEE92DFE4BFC57FDCFFBC3" authorityName="Jałoszyński" authorityYear="2024" box="[499,606,1015,1040]" class="Insecta" family="Staphylinidae" genus="Keratodegnathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="363" phylum="Arthropoda" rank="species" species="javanicus">
|
|||
|
|
<emphasis id="B925675DFFBEE92DFE4BFC57FDCFFBC3" box="[499,606,1015,1040]" italics="true" pageId="8" pageNumber="363">javanicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
refers to the
|
|||
|
|
<typeStatus id="54EA05EDFFBEE92DFD4AFC57FCB3FBC2" box="[754,802,1015,1041]" pageId="8" pageNumber="363">type</typeStatus>
|
|||
|
|
locality.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C34BE8C4FFBEE92DFF7FFBBBFD39FAAA" pageId="8" pageNumber="363" type="discussion">
|
|||
|
|
<paragraph id="8BEEBB4FFFBEE92DFF7FFBBBFD39FAAA" blockId="8.[151,1437,151,1401]" pageId="8" pageNumber="363">
|
|||
|
|
<emphasis id="B925675DFFBEE92DFF7FFBBBFEAAFBE6" bold="true" box="[199,315,1051,1077]" pageId="8" pageNumber="363">Remarks.</emphasis>
|
|||
|
|
The female of
|
|||
|
|
<taxonomicName id="4C51C0CCFFBEE92DFE64FBBBFDEBFBE7" authorityName="Pace" authorityYear="2014" box="[476,634,1051,1076]" class="Insecta" family="Staphylinidae" genus="Keratodegnathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="363" phylum="Arthropoda" rank="species" species="rougemonti">
|
|||
|
|
<emphasis id="B925675DFFBEE92DFE64FBBBFDEBFBE7" box="[476,634,1051,1076]" italics="true" pageId="8" pageNumber="363">K. rougemonti</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(known to occur in
|
|||
|
|
<collectingRegion id="499575ADFFBEE92DFCE9FBBBFC09FBE6" box="[849,920,1051,1077]" country="Malaysia" name="Sabah" pageId="8" pageNumber="363">Sabah</collectingRegion>
|
|||
|
|
, Borneo) has antennomeres 10 and 11 subequal in length, whereas in
|
|||
|
|
<taxonomicName id="4C51C0CCFFBEE92DFE36FB9FFD8AFB8B" authorityName="Jałoszyński" authorityYear="2024" box="[398,539,1087,1112]" class="Insecta" family="Staphylinidae" genus="Keratodegnathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="363" phylum="Arthropoda" rank="species" species="javanicus">
|
|||
|
|
<emphasis id="B925675DFFBEE92DFE36FB9FFD8AFB8B" box="[398,539,1087,1112]" italics="true" pageId="8" pageNumber="363">K. javanicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
antennomere 11 is conspicuously elongate, twice as long as 10. Moreover, the abdomen of
|
|||
|
|
<taxonomicName id="4C51C0CCFFBEE92DFE9AFBC3FE53FBAF" authorityName="Pace" authorityYear="2014" box="[290,450,1123,1148]" class="Insecta" family="Staphylinidae" genus="Keratodegnathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="363" phylum="Arthropoda" rank="species" species="rougemonti">
|
|||
|
|
<emphasis id="B925675DFFBEE92DFE9AFBC3FE53FBAF" box="[290,450,1123,1148]" italics="true" pageId="8" pageNumber="363">K. rougemonti</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is uniformly reddish, and in
|
|||
|
|
<taxonomicName id="4C51C0CCFFBEE92DFCBCFBC3FC1EFBAF" authorityName="Jałoszyński" authorityYear="2024" box="[772,911,1123,1148]" class="Insecta" family="Staphylinidae" genus="Keratodegnathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="363" phylum="Arthropoda" rank="species" species="javanicus">
|
|||
|
|
<emphasis id="B925675DFFBEE92DFCBCFBC3FC1EFBAF" box="[772,911,1123,1148]" italics="true" pageId="8" pageNumber="363">K. javanicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
infuscate in posterior half. The spermatheca of
|
|||
|
|
<taxonomicName id="4C51C0CCFFBEE92DFF2FFB27FEA9FB73" authorityName="Pace" authorityYear="2014" box="[151,312,1159,1184]" class="Insecta" family="Staphylinidae" genus="Keratodegnathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="363" phylum="Arthropoda" rank="species" species="rougemonti">
|
|||
|
|
<emphasis id="B925675DFFBEE92DFF2FFB27FEA9FB73" box="[151,312,1159,1184]" italics="true" pageId="8" pageNumber="363">K. rougemonti</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
illustrated in
|
|||
|
|
<bibRefCitation id="EFC0C6BEFFBEE92DFE6CFB27FDC6FB72" author="Pace, R." box="[468,599,1159,1185]" pageId="8" pageNumber="363" pagination="727 - 794" refId="ref4031" refString="Pace, R. (2014) Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae). Linzer biologische Beitrage, 46 (1), 727 - 794." type="journal article" year="2014">Pace (2014</bibRefCitation>
|
|||
|
|
: fig. 130) is clearly incomplete, it seems that it is also partly damaged (the receptacle is not illustrated and the duct possibly distorted). The female of
|
|||
|
|
<taxonomicName id="4C51C0CCFFBEE92DFC6EFB0BFBC9FB17" authorityName="Assing" authorityYear="2019" box="[982,1112,1195,1220]" class="Insecta" family="Staphylinidae" genus="Keratodegnathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="363" phylum="Arthropoda" rank="species" species="cornutus">
|
|||
|
|
<emphasis id="B925675DFFBEE92DFC6EFB0BFBC9FB17" box="[982,1112,1195,1220]" italics="true" pageId="8" pageNumber="363">K. cornutus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(a species inhabiting
|
|||
|
|
<collectingRegion id="499575ADFFBEE92DFAF2FB0BFA0DFB16" box="[1354,1436,1195,1221]" country="Malaysia" name="Pahang" pageId="8" pageNumber="363">Pahang</collectingRegion>
|
|||
|
|
in peninsular
|
|||
|
|
<collectingCountry id="F346FBDFFFBEE92DFE8AFB6FFE0BFB3A" box="[306,410,1231,1257]" name="Malaysia" pageId="8" pageNumber="363">Malaysia</collectingCountry>
|
|||
|
|
) has the head, pronotum and elytra nearly black, while in
|
|||
|
|
<taxonomicName id="4C51C0CCFFBEE92DFB88FB6FFB2CFB3B" authorityName="Jałoszyński" authorityYear="2024" box="[1072,1213,1231,1256]" class="Insecta" family="Staphylinidae" genus="Keratodegnathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="363" phylum="Arthropoda" rank="species" species="javanicus">
|
|||
|
|
<emphasis id="B925675DFFBEE92DFB88FB6FFB2CFB3B" box="[1072,1213,1231,1256]" italics="true" pageId="8" pageNumber="363">K. javanicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
the head and elytra are dark brown, but the pronotum is contrastingly red. The spermatheca of
|
|||
|
|
<taxonomicName id="4C51C0CCFFBEE92DFC63FB53FBCCFADF" authorityName="Assing" authorityYear="2019" box="[987,1117,1267,1292]" class="Insecta" family="Staphylinidae" genus="Keratodegnathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="363" phylum="Arthropoda" rank="species" species="cornutus">
|
|||
|
|
<emphasis id="B925675DFFBEE92DFC63FB53FBCCFADF" box="[987,1117,1267,1292]" italics="true" pageId="8" pageNumber="363">K. cornutus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(illustrated in
|
|||
|
|
<bibRefCitation id="EFC0C6BEFFBEE92DFB47FB53FA09FADE" author="Assing, V." box="[1279,1432,1267,1293]" pageId="8" pageNumber="363" pagination="33 - 70" refId="ref3725" refString="Assing, V. (2019) On the Lomechusini fauna of the Palaearctic and Oriental regions. XXVI. New species, a new synonymy, and additional records (Coleoptera: Staphylinidae: Aleocharinae). Beitrage zur Entomologie, 69 (1), 33 - 70. https: // doi. org / 10.21248 / contrib. entomol. 69.1.033 - 070" type="journal article" year="2019">Assing (2019</bibRefCitation>
|
|||
|
|
: fig. 117)) is of a clearly different shape than that of
|
|||
|
|
<taxonomicName id="4C51C0CCFFBEE92DFD6BFAB7FCCFFAE3" authorityName="Jałoszyński" authorityYear="2024" box="[723,862,1303,1328]" class="Insecta" family="Staphylinidae" genus="Keratodegnathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="363" phylum="Arthropoda" rank="species" species="javanicus">
|
|||
|
|
<emphasis id="B925675DFFBEE92DFD6BFAB7FCCFFAE3" box="[723,862,1303,1328]" italics="true" pageId="8" pageNumber="363">K. javanicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; in addition to a different pattern and shape of coils around the large reservoir, there is a loop just proximal to the distal receptacle in
|
|||
|
|
<taxonomicName id="4C51C0CCFFBEE92DFB88FA9BFB22FA87" authorityName="Assing" authorityYear="2019" box="[1072,1203,1339,1364]" class="Insecta" family="Staphylinidae" genus="Keratodegnathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="363" phylum="Arthropoda" rank="species" species="cornutus">
|
|||
|
|
<emphasis id="B925675DFFBEE92DFB88FA9BFB22FA87" box="[1072,1203,1339,1364]" italics="true" pageId="8" pageNumber="363">K. cornutus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, whereas there is no such a loop in the spermatheca of
|
|||
|
|
<taxonomicName id="4C51C0CCFFBEE92DFDADFAFFFD30FAAB" authorityName="Jałoszyński" authorityYear="2024" box="[533,673,1375,1400]" class="Insecta" family="Staphylinidae" genus="Keratodegnathus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="363" phylum="Arthropoda" rank="species" species="javanicus">
|
|||
|
|
<emphasis id="B925675DFFBEE92DFDADFAFFFD30FAAB" box="[533,673,1375,1400]" italics="true" pageId="8" pageNumber="363">K. javanicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|