137 lines
15 KiB
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137 lines
15 KiB
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<document id="A3D6C37A209E3A4B13198D317D0AC635" ID-DOI="10.11646/zootaxa.3700.3.7" ID-GBIF-Dataset="371f30c1-ff9d-4fde-9429-fc9d284caf03" ID-ISSN="1175-5326" ID-Zenodo-Dep="249376" ID-ZooBank="E3FA2E4B-DDF4-4FDB-8CF1-FFBA215D40CA" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460663374311" checkinUser="plazi" docAuthor="Massa, Bruno" docDate="2013" docId="03C187FAFFA7B341FF64D01C59C9FAA2" docLanguage="en" docName="zt03700p475.pdf" docOrigin="Zootaxa 3700 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tmethis maroccanus Bolivar 1908" docType="treatment" docVersion="13" lastPageNumber="440" masterDocId="FFF8FF82FFA2B344FFF3D33D5D23FFDA" masterDocTitle="Pamphagidae (Orthoptera: Caelifera) of North Africa: key to genera and the annotated check-list of species" masterLastPageNumber="475" masterPageNumber="435" pageNumber="440" updateTime="1698347717602" updateUser="plazi">
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<mods:mods id="5DC7A79A5BCA0F6780F6E810F25ED3C4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="FB0F407CC4A9DF6A306D3E914A02B605">
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<mods:title id="DD265774B42E0A3D05C46EC32CEB757C">Pamphagidae (Orthoptera: Caelifera) of North Africa: key to genera and the annotated check-list of species</mods:title>
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</mods:titleInfo>
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<mods:name id="EDA898D254A359719A3B881603F2E0A2" type="personal">
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<mods:role id="0A73A439B83A2465426969AFBB8DDAD7">
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<mods:roleTerm id="974E49B0F851EF81C917F0199B14BAD0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B11C2572A7B170ADE7A9BAEC9BF8EA4A">Massa, Bruno</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="4AB543CD28464C52D5723E5B178EB5E1">text</mods:typeOfResource>
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<mods:titleInfo id="479CC4DDB7742539DAA51844F5B42212">
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<mods:title id="90EC695E54B12588E9F4D90ED3FC9562">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="465DB23297D8E77B83CB7DA94BA4A888">
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<mods:date id="82A95B18B526EEDCD8375510264EA9BA">2013</mods:date>
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<mods:detail id="420129996B4FA8FABB592AF2D94C69CD" type="volume">
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<mods:number id="DD523E4BFCC52C6BFEF91B3B7828E9DE">3700</mods:number>
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</mods:detail>
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<mods:detail id="29D13D68318FD934B68125C463D80DF6" type="issue">
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<mods:number id="5B39B153526DA7E6B90F4371B16D32FA">3</mods:number>
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</mods:detail>
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<mods:extent id="637410BFA4446F93D551F914FD76534A" unit="page">
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<mods:start id="6E9B819C0921CBA81356C7C923629AD1">435</mods:start>
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<mods:end id="4687C8FF124C9418109865B31A92E88B">475</mods:end>
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<mods:classification id="DD5DD795FD320B61FAD95D3C0CF6C0A7">journal article</mods:classification>
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<mods:identifier id="FAC60975CB20FF52EE978C5195609FCE" type="DOI">10.11646/zootaxa.3700.3.7</mods:identifier>
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<mods:identifier id="0BD37F4DE21C1AA186410037F5E17545" type="GBIF-Dataset">371f30c1-ff9d-4fde-9429-fc9d284caf03</mods:identifier>
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<mods:identifier id="61ABEE396B588EB13768D5EB00DE46E3" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="BF86E2D3F1FC0E2AB3861A2F61B7A711" type="Zenodo-Dep">249376</mods:identifier>
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<mods:identifier id="F49BBB3D9B59CA24E2361FB7EC2BB258" type="ZooBank">E3FA2E4B-DDF4-4FDB-8CF1-FFBA215D40CA</mods:identifier>
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</mods:mods>
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<treatment id="03C187FAFFA7B341FF64D01C59C9FAA2" ID-DOI="http://doi.org/10.5281/zenodo.5697468" ID-GBIF-Taxon="119574861" ID-Zenodo-Dep="5697468" LSID="urn:lsid:plazi:treatment:03C187FAFFA7B341FF64D01C59C9FAA2" httpUri="http://treatment.plazi.org/id/03C187FAFFA7B341FF64D01C59C9FAA2" lastPageNumber="440" pageId="5" pageNumber="440">
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<subSubSection id="C3726567FFA7B341FF64D01C5F1CFCE1" box="[151,575,801,827]" pageId="5" pageNumber="440" type="nomenclature">
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<paragraph id="8BD736ECFFA7B341FF64D01C5F1CFCE1" blockId="5.[151,575,801,827]" box="[151,575,801,827]" pageId="5" pageNumber="440">
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<heading id="D09F8180FFA7B341FF64D01C5F1CFCE1" bold="true" box="[151,575,801,827]" fontSize="11" level="1" pageId="5" pageNumber="440" reason="1">
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<taxonomicName id="4C684D6FFFA7B341FF64D01C5F1CFCE1" ID-CoL="7CDZX" authority="Bolivar, 1908" authorityName="Bolivar" authorityYear="1908" box="[151,575,801,827]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="maroccanus">
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<emphasis id="B91CEAFEFFA7B341FF64D01C5F1CFCE1" bold="true" box="[151,575,801,827]" pageId="5" pageNumber="440">
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<emphasis id="B91CEAFEFFA7B341FF64D01C5CB7FCE1" bold="true" box="[151,404,801,827]" italics="true" pageId="5" pageNumber="440">Tmethis maroccanus</emphasis>
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Bolivar, 1908
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3726567FFA7B341FF64D05A59C9FAA2" pageId="5" pageNumber="440" type="materials_examined">
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<paragraph id="8BD736ECFFA7B341FF64D05A5E2AFC7E" blockId="5.[151,1437,871,1400]" pageId="5" pageNumber="440">
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<emphasis id="B91CEAFEFFA7B341FF64D05A5C59FC5A" bold="true" box="[151,378,871,896]" pageId="5" pageNumber="440">Material examined</emphasis>
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.
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<collectingCountry id="F37F767CFFA7B341FE79D0555CD1FCA5" box="[394,498,872,895]" name="Morocco" pageId="5" pageNumber="440">Morocco</collectingCountry>
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, Marrakech,
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<specimenCount id="9D6EFD65FFA7B341FD75D0555F88FCA5" box="[646,683,872,895]" pageId="5" pageNumber="440" type="generic">7♂</specimenCount>
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, 4♀
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<typeStatus id="54D3884EFFA7B341FD10D0555E66FC5A" box="[739,837,872,896]" pageId="5" pageNumber="440" type="syntype">syntypes</typeStatus>
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of
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<taxonomicName id="4C684D6FFFA7B341FC9ED0545931FC5A" box="[877,1042,873,896]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="maroccanus">
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<emphasis id="B91CEAFEFFA7B341FC9ED0545931FC5A" box="[877,1042,873,896]" italics="true" pageId="5" pageNumber="440">T. maroccanus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C684D6FFFA7B341FBB8D05459CCFC5A" box="[1099,1263,873,896]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="maroccanus">
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<emphasis id="B91CEAFEFFA7B341FBB8D05459CCFC5A" box="[1099,1263,873,896]" italics="true" pageId="5" pageNumber="440">T. maroccanus</emphasis>
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</taxonomicName>
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<taxonomicName id="4C684D6FFFA7B341FB04D05558B6FC5A" box="[1271,1429,872,896]" pageId="5" pageNumber="440" rank="variety" variety="incristata">
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var.
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<emphasis id="B91CEAFEFFA7B341FADAD05458B6FC5A" box="[1321,1429,873,896]" italics="true" pageId="5" pageNumber="440">incristata</emphasis>
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</taxonomicName>
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; El Glaui,
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<specimenCount id="9D6EFD65FFA7B341FEF0D0B15C04FC7E" box="[259,295,908,932]" pageId="5" pageNumber="440" type="generic">1♂</specimenCount>
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, 2♀; Bou Lourane 15.
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<date id="FFD6102CFFA7B341FDD3D0B15FA0FC7E" box="[544,643,908,933]" pageId="5" pageNumber="440" value="1949-06-02">VI.49, 2</date>
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♀ (MNCN).
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</paragraph>
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<paragraph id="8BD736ECFFA7B341FF34D09259C9FAA2" blockId="5.[151,1437,871,1400]" pageId="5" pageNumber="440">
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The description was based mainly on the smooth integuments, with rounded tubercles and on the peculiar colour of hind wings, with an isolated brown spot (Bolivar 1908); these characters were also reported by Chopard (1943). Only Uvarov (1943) showed the characteristics of this taxon, that consent to separate it from
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<taxonomicName id="4C684D6FFFA7B341FAF5D0C45873FBCA" box="[1286,1360,1017,1040]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="cisti">
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<emphasis id="B91CEAFEFFA7B341FAF5D0C45873FBCA" box="[1286,1360,1017,1040]" italics="true" pageId="5" pageNumber="440">T. cisti</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C684D6FFFA7B341FA74D0C45C12FBEF" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="pulchripennis">
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<emphasis id="B91CEAFEFFA7B341FA74D0C45C12FBEF" italics="true" pageId="5" pageNumber="440">T. pulchripennis</emphasis>
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</taxonomicName>
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: the shape of the pronotum is broadly parabolic, with convex margins and low keel (
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<figureCitation id="13532A69FFA7B341FB13D7215844FBEF" box="[1248,1383,1052,1077]" captionStart="FIGURES 32 – 39" captionStartId="28.[151,264,1715,1737]" captionTargetBox="[300,1286,184,1690]" captionTargetId="figure@28.[300,1287,183,1693]" captionTargetPageId="28" captionText="FIGURES 32 – 39. Lateral view of pronotum of some taxa described in the genus Tmethis, here revised." httpUri="https://zenodo.org/record/249379/files/figure.png" pageId="5" pageNumber="440">Figs. 36–37</figureCitation>
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,
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<figureCitation id="13532A69FFA7B341FA81D7215D95FB82" captionStart="FIGURES 40 – 53" captionStartId="29.[151,264,2001,2023]" captionTargetBox="[220,1366,182,1957]" captionTargetId="figure@29.[220,1366,165,1979]" captionTargetPageId="29" captionText="FIGURES 40 – 53. Lateral view of males and females of some taxa described in the genus Tmethis, here revised." httpUri="https://zenodo.org/record/249380/files/figure.png" pageId="5" pageNumber="440">46– 49</figureCitation>
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,
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<figureCitation id="13532A69FFA7B341FF31D7025DC1FB82" box="[194,226,1087,1112]" captionStart="FIGURES 54 – 59" captionStartId="30.[151,264,1827,1849]" captionTargetBox="[290,1301,181,1799]" captionTargetId="figure@30.[286,1301,171,1806]" captionTargetPageId="30" captionText="FIGURES 54 – 59. Some of the taxa described in the genus Tmethis, view from above to show the pattern of wing colour pattern and the ratio between the length of wings and the body. Figs. 58 a and 58 b show the pattern of pronotum and wings of male and female of T. harterti n. status. Figs. 54 a and 55 a show the apical lobes, large and broadly separated, of epiphallus of T. cisti, while Figs. 59 b and 59 c show the relatively small and narrowly separated lobes of epiphallus of T. pulchripennis (after Uvarov 1943). Figs 54 b and 59 a show the differences in the spurs of hind tibiae, long and slender, with the first tarsal joint long, and its second pulvillus longer than the first or third in T. cisti; shorter and stouter, with first tarsal joint short, its second pulvillus not longer than the first or the third in T. pulchripennis (after Uvarov 1943)." httpUri="https://zenodo.org/record/249381/files/figure.png" pageId="5" pageNumber="440">57</figureCitation>
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).
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<collectingCountry id="F37F767CFFA7B341FF05D7025C0DFB82" box="[246,302,1087,1112]" name="India" pageId="5" pageNumber="440">Hind</collectingCountry>
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tibiae are yellow, hind wings are shorter than tegmina, and in the female tegmina hardly reach hind knees. The epiphallus is similar to that of
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<taxonomicName id="4C684D6FFFA7B341FD9BD7585E3EFBA7" box="[616,797,1124,1149]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="pulchripennis">
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<emphasis id="B91CEAFEFFA7B341FD9BD7585E3EFBA7" box="[616,797,1124,1149]" italics="true" pageId="5" pageNumber="440">T. pulchripennis</emphasis>
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</taxonomicName>
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, with hind lobes relatively small and narrowly separated. When Bolivar (1908) described this species from Marrakech, from the same locality described also the
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<taxonomicName id="4C684D6FFFA7B341FA80D7B55C21FB1E" pageId="5" pageNumber="440" rank="variety" variety="incristata">
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var.
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<emphasis id="B91CEAFEFFA7B341FF64D7905C21FB1E" box="[151,258,1197,1220]" italics="true" pageId="5" pageNumber="440">incristata</emphasis>
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</taxonomicName>
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, which is an extreme case of specimens without pronotum keel and of course has no taxonomical value. It covers an inner area of
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<collectingCountry id="F37F767CFFA7B341FE38D7ED5F13FB3D" box="[459,560,1232,1255]" name="Morocco" pageId="5" pageNumber="440">Morocco</collectingCountry>
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(Bolivar 1908, Werner 1932). Concerning the record from Colomb-Bechar (
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<collectingCountry id="F37F767CFFA7B341FF6DD7C95DD6FAD7" box="[158,245,1268,1293]" name="Algeria" pageId="5" pageNumber="440">Algeria</collectingCountry>
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) by Chopard (1943) see the subspecies
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<emphasis id="B91CEAFEFFA7B341FD3ED7C95E2DFAD7" box="[717,782,1268,1293]" italics="true" pageId="5" pageNumber="440">hirtus</emphasis>
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. In addition, considering the high variability of
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<taxonomicName id="4C684D6FFFA7B341FAB4D7C85C13FAEA" authority="Massa (2009)" authorityName="Massa" authorityYear="2009" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="cisti">
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<emphasis id="B91CEAFEFFA7B341FAB4D7C858B6FAD6" box="[1351,1429,1269,1292]" italics="true" pageId="5" pageNumber="440">T. cisti</emphasis>
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, Massa (2009)
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</taxonomicName>
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considered unlikely also the record by Usmani (2007) from South
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<collectingCountry id="F37F767CFFA7B341FBEFD62A597EFAEA" box="[1052,1117,1303,1328]" name="Libya" pageId="5" pageNumber="440">Libya</collectingCountry>
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(Fezzan), that in the light of characters above reported for
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<taxonomicName id="4C684D6FFFA7B341FE11D6005F71FA8F" box="[482,594,1340,1365]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="harterti">
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<emphasis id="B91CEAFEFFA7B341FE11D6005F71FA8F" box="[482,594,1340,1365]" italics="true" pageId="5" pageNumber="440">T. harterti</emphasis>
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</taxonomicName>
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, could belong to this taxon (see also
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<taxonomicName id="4C684D6FFFA7B341FC1ED60159C5FA8F" box="[1005,1254,1340,1365]" class="Insecta" family="Pamphagidae" genus="Neotmethis" higherTaxonomySource="CoL" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="bidentatus">
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<emphasis id="B91CEAFEFFA7B341FC1ED60159C5FA8F" box="[1005,1254,1340,1365]" italics="true" pageId="5" pageNumber="440">Neotmethis bidentatus</emphasis>
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</taxonomicName>
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). Consequently,
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<taxonomicName id="4C684D6FFFA7B341FF64D65C5C19FAA2" box="[151,314,1377,1400]" class="Insecta" family="Pamphagidae" genus="Tmethis" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="maroccanus">
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<emphasis id="B91CEAFEFFA7B341FF64D65C5C19FAA2" box="[151,314,1377,1400]" italics="true" pageId="5" pageNumber="440">T. maroccanus</emphasis>
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</taxonomicName>
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should cover only the inner western area of
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<collectingCountry id="F37F767CFFA7B341FCDFD65D5EB2FAAD" box="[812,913,1376,1399]" name="Morocco" pageId="5" pageNumber="440">Morocco</collectingCountry>
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and West
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<collectingCountry id="F37F767CFFA7B341FBF5D6625979FAA2" box="[1030,1114,1375,1400]" name="Algeria" pageId="5" pageNumber="440">Algeria</collectingCountry>
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(see below).
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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