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<document id="CE465E8F5DC051637BDCD2AFC84987DD" ID-DOI="10.5852/ejt.2019.526" ID-GBIF-Dataset="2da65c0a-9c48-43f5-ada3-e088de28dec1" ID-ISSN="2118-9773" ID-Zenodo-Dep="3152217" ID-ZooBank="3FFC527D-E9CA-4D9F-BF59-076FA60171AC" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="existingObjects,operationResults,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1558535825802" checkinUser="plazi" docAuthor="Nurinsiyah, Ayu Savitri, Neiber, Marco T. &amp; Hausdorf, Bernhard" docDate="2019" docId="03B1DB4C8B324E384C73FEF73962FE62" docLanguage="en" docName="ej-526_nurinsiyah_neiber_hausdorf.pdf.imf" docOrigin="European Journal of Taxonomy 526" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Landouria parahyangensis Nurinsiyah &amp; Neiber &amp; Hausdorf 2019, sp. nov." docType="treatment" docUuid="41EEB2F2-7781-4D19-9F62-34316F374749" docUuidSource="ZooBank" docVersion="15" lastPageNumber="34" masterDocId="FF88A3348B2D4E194E35FFFC3C0DFFDB" masterDocTitle="Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java" masterLastPageNumber="73" masterPageNumber="1" pageNumber="32" updateTime="1716316918745" updateUser="carolina">
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<mods:title id="FDC90FFF0EA5BE4DE0D41000C61A288C">Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java</mods:title>
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<mods:roleTerm id="BAC53A4F5C8DC80012DE55D8B7A6A2D4">Author</mods:roleTerm>
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<mods:namePart id="09DC071A23CCCCBB6FEFD66513118C39">Nurinsiyah, Ayu Savitri</mods:namePart>
<mods:nameIdentifier id="F76B0966C2D64805AE95B8804B55E4CB" type="LSID">5AFBFEFB-B243-49CC-AACC-248B2F007BC8</mods:nameIdentifier>
<mods:affiliation id="58FC768B6D6EAE443CEC2ABCBABC2077">Museum Zoologicum Bogoriense, Research Center for Biology, Indonesian Institute of Science, Cibinong, West Java, Indonesia.</mods:affiliation>
<mods:nameIdentifier id="88638B77DC7E43F84A2762E5F7C42202" type="email">ayu_nurinsiyah@yahoo.com &amp; ayus002@lipi.go.id</mods:nameIdentifier>
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<mods:namePart id="0B073748C04D4EBDE0E6F6434EEB3541">Neiber, Marco T.</mods:namePart>
<mods:nameIdentifier id="7F2D4EC98527EEBB525CB65D38C54C43" type="LSID">8F6006F5-D880-4841-AC32-B97E18D7BB5C</mods:nameIdentifier>
<mods:affiliation id="5A310512641ED1211F5A187CDC512193">Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
<mods:nameIdentifier id="4095E63C80A8BBA5CBD1BEA0EF73CD22" type="email">mneiber@hotmail.de</mods:nameIdentifier>
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<mods:namePart id="9600077D1CB04BE1CEFF266DDB0F2FE9">Hausdorf, Bernhard</mods:namePart>
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<mods:affiliation id="B12BD6376519C4BAB84805CBF94DD91D">Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany.</mods:affiliation>
<mods:nameIdentifier id="0BD5244EDE07BBE682928EA2821CE446" type="email">hausdorf@zoologie.uni-hamburg.de</mods:nameIdentifier>
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<mods:typeOfResource id="D05BE72D7BCA735E402B403F44ABA4A5">text</mods:typeOfResource>
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<mods:title id="69D4769CF0BF337234E177956405211A">European Journal of Taxonomy</mods:title>
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<mods:date id="DB98F81C589B12B3D8816991D09127F2">2019</mods:date>
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<mods:number id="E149E4A76E051E3ECC4D14ACE7AE6134">2019-05-21</mods:number>
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<mods:number id="5F797AF8FBD5408B9BAA0DAA7658BF36">526</mods:number>
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<treatment id="03B1DB4C8B324E384C73FEF73962FE62" ID-GBIF-Taxon="157239581" LSID="urn:lsid:zoobank.org:act:41EEB2F2-7781-4D19-9F62-34316F374749" httpUri="http://treatment.plazi.org/id/03B1DB4C8B324E384C73FEF73962FE62" lastPageId="33" lastPageNumber="34" pageId="31" pageNumber="32">
<subSubSection id="C30239D18B324E064C73FEF738BBFE93" pageId="31" pageNumber="32" type="nomenclature">
<paragraph id="8BA76A5A8B324E064C73FEF73FE3FEFD" blockId="31.[381,1206,267,365]" box="[582,1006,267,294]" pageId="31" pageNumber="32">
<heading id="D0EFDD368B324E064C73FEF73FE3FEFD" box="[582,1006,267,294]" centered="true" fontSize="11" level="2" pageId="31" pageNumber="32" reason="2">
<taxonomicName id="4C1811D98B324E064C73FEF73F80FEFD" ID-CoL="7RTHY" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[582,909,267,294]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="parahyangensis" status="sp. nov.">
<emphasis id="B96CB6488B324E064C73FEF73F80FEFD" bold="true" box="[582,909,267,294]" italics="true" pageId="31" pageNumber="32">Landouria parahyangensis</emphasis>
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<taxonomicNameLabel id="A25F0B338B324E064DA1FEF03FE3FEFD" box="[916,1006,268,294]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
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urn:lsid:zoobank.org:act:
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</paragraph>
</subSubSection>
<subSubSection id="C30239D18B324E064CA4FEAE3FAEFEB7" box="[657,931,338,365]" pageId="31" pageNumber="32" type="description">
<paragraph id="8BA76A5A8B324E064CA4FEAE3FAEFEB7" blockId="31.[381,1206,267,365]" box="[657,931,338,365]" pageId="31" pageNumber="32">
Figs
<figureCitation id="132376DF8B324E064CFEFEAE3EE6FEB7" box="[715,747,338,364]" captionStart="Figs 48–51" captionStartId="31.[189,241,1824,1850]" captionTargetBox="[280,1308,265,2019]" captionTargetId="figure@32.[280,1308,265,2019]" captionTargetPageId="32" captionText="Figs 48–51 (opposite page). Shells of species of Landouria from Java. 48. L. parahyangensis sp. nov., West Java, Karangnunggal, surroundings of Pasir Taraje Cave (holotype, MZB 20429). 49. L. nusakambangensis sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 412298a). 50. L. petrukensis sp. nov., Central Java, Karangbolong, way to Petruk Cave (holotype, MZB 19203). 51. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). Scale bar: 5 mm." httpUri="https://zenodo.org/record/3152259/files/figure.png" pageId="31" pageNumber="32">48</figureCitation>
,
<figureCitation id="132376DF8B324E064CC2FEAE3F15FEB6" box="[759,792,338,365]" captionStart="Figs 52–55" captionStartId="33.[189,241,1717,1743]" captionTargetBox="[189,1398,265,1993]" captionTargetId="figure@34.[189,1398,265,1993]" captionTargetPageId="34" captionText="Figs 52–55 (opposite page). Distribution of species of Landouria in Java (10′ grid). 52. L. winteriana (Pfeiffer, 1842), L. naggsi sp. nov., L. nusakambangensis sp. nov., L. petrukensis sp. nov., L. tholiformis sp. nov., L. tonywhitteni sp. nov., L. smimensis (Mousson, 1848). 53. L. conoidea (Leschke, 1914), L. abdidalem sp. nov., L. sewuensis sp. nov., L. sukoliloensis sp. nov., L. pacitanensis sp. nov., L. intumescens (Martens, 1867), L. madurensis sp. nov. 54. L. rotatoria (Pfeiffer, 1842), L. nodifera sp. nov., L. parahyangensis sp. nov., L. schepmani (Möllendorff, 1897), L. zonifera sp. nov., L. ciliocincta (Möllendorff, 1897), L. pakidulan sp. nov. 55. L. epiplatia (Möllendorff, 1897), L. monticola van Benthem Jutting, 1950, L. davini Dharma, 2015, L. menorehensis sp. nov., L. dharmai sp. nov., L. moussoniana (Martens, 1867)." httpUri="https://zenodo.org/record/3152265/files/figure.png" pageId="31" pageNumber="32">54</figureCitation>
;
<tableCitation id="C69A5FE18B324E064D16FEAE3FAEFEB7" box="[803,931,338,364]" captionStart-0="Table 3" captionStart-1="Table 4" captionStartId-0="13.[189,258,268,294]" captionStartId-1="22.[188,257,551,579]" captionTargetBox-0="[201,1370,481,1908]" captionTargetBox-1="[205,1929,661,1732]" captionTargetPageId-0="13" captionTargetPageId-1="22" captionText-0="Table 3 (continued on next 3 pages). Shell measurements for species of Landouria Godwin-Austen, 1918 (in mm). Abbreviations: D = shell diameter; D3W = diameter of the third whorl; da = diameter of the aperture; H = shell height; ha = height of the aperture; max = maximum; min = minimum; n = number of measured spec.; std = standard deviation; U = diameter of the umbilicus; W = number of whorls." captionText-1="Table 4 (continued on next 3 pages). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java. Characteristics most important for distinguishing similar taxa are printed in bold." httpUri-0="http://table.plazi.org/id/DF673AD28B204E144E88FEF03D19FE68" httpUri-1="http://table.plazi.org/id/DF673AD28B3B4E0F4E89FDDB3F73FDBC" pageId="31" pageNumber="32" tableUuid-0="DF673AD28B204E144E88FEF03D19FE68" tableUuid-1="DF673AD28B3B4E0F4E89FDDB3F73FDBC">Tables 3–4</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B324E064E88FE6F3DC4FE24" pageId="31" pageNumber="32" type="diagnosis">
<paragraph id="8BA76A5A8B324E064E88FE6F3D3BFE76" blockId="31.[189,310,403,429]" box="[189,310,403,429]" pageId="31" pageNumber="32">
<heading id="D0EFDD368B324E064E88FE6F3D3BFE76" bold="true" box="[189,310,403,429]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="B96CB6488B324E064E88FE6F3D3BFE76" bold="true" box="[189,310,403,429]" pageId="31" pageNumber="32">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B324E064E88FE3E3DC4FE24" blockId="31.[189,1398,450,511]" pageId="31" pageNumber="32">
<taxonomicName id="4C1811D98B324E064E88FE3E3DF4FE07" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[189,505,450,476]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="parahyangensis" status="sp. nov.">
<emphasis id="B96CB6488B324E064E88FE3E3DF4FE07" box="[189,505,450,476]" italics="true" pageId="31" pageNumber="32">Landouria parahyangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B324E064FCAFE3E3E54FE07" box="[511,601,450,476]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
is characterized by a sharply keeled, tightly coiled conical shell with a very wide umbilicus.
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B324E064E88FDDB3E98FD6D" pageId="31" pageNumber="32" type="etymology">
<paragraph id="8BA76A5A8B324E064E88FDDB3D49FD9A" blockId="31.[189,324,551,577]" box="[189,324,551,577]" pageId="31" pageNumber="32">
<heading id="D0EFDD368B324E064E88FDDB3D49FD9A" bold="true" box="[189,324,551,577]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="B96CB6488B324E064E88FDDB3D49FD9A" bold="true" box="[189,324,551,577]" pageId="31" pageNumber="32">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B324E064E88FDA93E98FD6D" blockId="31.[189,1399,597,694]" pageId="31" pageNumber="32">
<taxonomicName id="4C1811D98B324E064E88FDA93DF5FDB4" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[189,504,597,623]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="parahyangensis" status="sp. nov.">
<emphasis id="B96CB6488B324E064E88FDA93DF5FDB4" box="[189,504,597,623]" italics="true" pageId="31" pageNumber="32">Landouria parahyangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B324E064FC8FDA93E58FDB4" box="[509,597,597,623]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
is named after its occurrence in the mountainous region Parahyangan. In the Sundanese language, the name ʻParahyanganʼ means ʻthe abode of hyangs (gods)ʼ, which were supposed to reside in the mountain tops.
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B324E064E88FD21387DFB64" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="8BA76A5A8B324E064E88FD213DA7FD2C" blockId="31.[189,426,733,759]" box="[189,426,733,759]" pageId="31" pageNumber="32">
<heading id="D0EFDD368B324E064E88FD213DA7FD2C" bold="true" box="[189,426,733,759]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="B96CB6488B324E064E88FD213DA7FD2C" bold="true" box="[189,426,733,759]" pageId="31" pageNumber="32">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B324E064EDFFCF73D51FCFE" blockId="31.[189,1398,779,911]" box="[234,348,779,805]" pageId="31" pageNumber="32">
<typeStatus id="54A3D4F88B324E064EDFFCF73D51FCFE" box="[234,348,779,805]" pageId="31" pageNumber="32" type="holotype">
<emphasis id="B96CB6488B324E064EDFFCF73D51FCFE" bold="true" box="[234,348,779,805]" pageId="31" pageNumber="32">Holotype</emphasis>
</typeStatus>
</paragraph>
<paragraph id="8BA76A5A8B324E064E88FCD33DFEFC54" blockId="31.[189,1398,779,911]" pageId="31" pageNumber="32">
<materialsCitation id="3B7060078B324E064E88FCD33DFEFC54" ID-GBIF-Occurrence="2243137365" collectedFrom="agroforestry" collectingDate="2014-07-04" collectionCode="MZB" collectorName="A. S. Nurinsiyah &amp; T. Setiadi" country="Indonesia" elevation="243" latitude="-7.666111" location="surroundings of Pasir Taraje Cave" longLatPrecision="21" longitude="108.11694" municipality="Karangnunggal" pageId="31" pageNumber="32" specimenCode="20429" specimenCount="1" stateProvince="West Java" typeStatus="holotype">
<collectingCountry id="F30F2ACA8B324E064E88FCD33D56FC92" box="[189,347,815,841]" name="Indonesia" pageId="31" pageNumber="32">INDONESIA</collectingCountry>
•
<typeStatus id="54A3D4F88B324E064F42FCD23DECFC93" box="[375,481,814,840]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
;
<collectingRegion id="49DCA4B88B324E064FDBFCD33E65FC92" box="[494,616,815,841]" country="Indonesia" name="Jawa Barat" pageId="31" pageNumber="32">West Java</collectingRegion>
,
<collectingMunicipality id="6BC3F0208B324E064C40FCD23F20FC93" box="[629,813,814,840]" pageId="31" pageNumber="32">Karangnunggal</collectingMunicipality>
,
<location id="8EC73C818B324E064D0FFCD238DCFC92" LSID="urn:lsid:plazi:treatment:03B1DB4C8B324E384C73FEF73962FE62:8EC73C818B324E064D0FFCD238DCFC92" box="[826,1233,814,841]" country="Indonesia" latitude="-7.666111" longLatPrecision="21" longitude="108.11694" municipality="Karangnunggal" name="surroundings of Pasir Taraje Cave" pageId="31" pageNumber="32" stateProvince="West Java">surroundings of Pasir Taraje Cave</location>
,
<collectedFrom id="380429A78B324E064AEAFCD2397FFC93" box="[1247,1394,814,840]" pageId="31" pageNumber="32">agroforestry</collectedFrom>
;
<geoCoordinate id="EE2C0C9D8B324E064E88FCAC3D32FCB7" box="[189,319,848,876]" degrees="7" direction="south" minutes="39" orientation="latitude" pageId="31" pageNumber="32" precision="15" seconds="58" value="-7.666111">7°39′58″ S</geoCoordinate>
,
<geoCoordinate id="EE2C0C9D8B324E064F79FCAC3DE3FCB7" box="[332,494,848,876]" degrees="108" direction="east" minutes="07" orientation="longitude" pageId="31" pageNumber="32" precision="15" seconds="01" value="108.11694">108°07′01″ E</geoCoordinate>
;
<elevation id="00358D698B324E064FCEFCAE3E8BFCB7" box="[507,646,850,876]" metricMagnitude="2" metricUnit="m" metricValue="2.43" pageId="31" pageNumber="32" unit="m" value="243.0">
<quantity id="4CE0C7BF8B324E064FCEFCAE3E4BFCB7" box="[507,582,850,876]" metricMagnitude="2" metricUnit="m" metricValue="2.43" pageId="31" pageNumber="32" unit="m" value="243.0">243 m</quantity>
a.s.l.
</elevation>
;
<collectingDate id="EFE2B5728B324E064CA1FCAE3F17FCB7" box="[660,794,850,876]" pageId="31" pageNumber="32" value="2014-07-04">
<date id="FFA64C9A8B324E064CA1FCAE3F17FCB7" box="[660,794,850,876]" pageId="31" pageNumber="32" value="2014-07-04">4 Jul. 2014</date>
</collectingDate>
;
<collectorName id="26ED0F8C8B324E064D10FCAE3FEEFCB7" box="[805,995,850,876]" pageId="31" pageNumber="32">A.S. Nurinsiyah</collectorName>
and
<collectorName id="26ED0F8C8B324E064A28FCAE3880FCB7" box="[1053,1165,850,876]" pageId="31" pageNumber="32">T. Setiadi</collectorName>
leg.; D = 15.0 mm, H =
<quantity id="4CE0C7BF8B324E064EC4FC893D43FC54" box="[241,334,885,911]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="31" pageNumber="32" unit="mm" value="8.4">8.4 mm</quantity>
;
<collectionCode id="ED09F29F8B324E064F6FFC893D97FC54" box="[346,410,885,911]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="31" pageNumber="32" type="Museum">MZB</collectionCode>
<specimenCode id="DBBEC2218B324E064F94FC893DE3FC54" box="[417,494,885,911]" pageId="31" pageNumber="32">20429</specimenCode>
.
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</paragraph>
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<paragraph id="8BA76A5A8B324E064EDFFC533D6AFC12" blockId="31.[189,1398,943,1216]" box="[234,359,943,969]" pageId="31" pageNumber="32">
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<emphasis id="B96CB6488B324E064EDFFC533D6AFC12" bold="true" box="[234,359,943,969]" pageId="31" pageNumber="32">Paratypes</emphasis>
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<paragraph id="8BA76A5A8B324E064E88FC2E387DFB64" blockId="31.[189,1398,943,1216]" pageId="31" pageNumber="32">
<collectingCountry id="F30F2ACA8B324E064E88FC2E3D56FC37" box="[189,347,978,1004]" name="Indonesia" pageId="31" pageNumber="32">INDONESIA</collectingCountry>
–
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<emphasis id="B96CB6488B324E064F43FC2E3DFBFC37" bold="true" box="[374,502,978,1004]" pageId="31" pageNumber="32">West Java</emphasis>
</collectingRegion>
•
<specimenCount id="9D1EA1D38B324E064C39FC2E3E6DFC37" box="[524,608,978,1004]" count="3" pageId="31" pageNumber="32" type="generic">3 spec.</specimenCount>
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,
<collectedFrom id="380429A78B324E064AEAFC2E397FFC37" box="[1247,1394,978,1004]" pageId="31" pageNumber="32">agroforestry</collectedFrom>
;
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,
<geoCoordinate id="EE2C0C9D8B324E064F7FFC0F3DE1FBCB" box="[330,492,1011,1040]" degrees="108" direction="east" minutes="06" orientation="longitude" pageId="31" pageNumber="32" precision="15" seconds="55" value="108.11527">108°06′55″ E</geoCoordinate>
;
<elevation id="00358D698B324E064FCDFC093E8CFBD4" box="[504,641,1013,1040]" metricMagnitude="2" metricUnit="m" metricValue="2.34" pageId="31" pageNumber="32" unit="m" value="234.0">
<quantity id="4CE0C7BF8B324E064FCDFC093E4FFBCB" box="[504,578,1013,1040]" metricMagnitude="2" metricUnit="m" metricValue="2.34" pageId="31" pageNumber="32" unit="m" value="234.0">234 m</quantity>
a.s.l.
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;
<collectionCode id="ED09F29F8B324E064CBBFC0A3EC3FBCB" box="[654,718,1014,1040]" collectionName="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="31" pageNumber="32" type="Museum">MZB</collectionCode>
<specimenCode id="DBBEC2218B324E064CE0FC093F13FBD4" box="[725,798,1013,1039]" pageId="31" pageNumber="32">20431</specimenCode>
•
<materialsCitation id="3B7060078B324E064D02FC09386FFBE9" ID-GBIF-Occurrence="2243137533" collectedFrom="agroforestry and bamboo" collectionCode="MZB" country="Indonesia" elevation="237" latitude="-7.6666665" location="near Sela Dower Cave" longLatPrecision="21" longitude="108.11667" municipality="Karangnunggal" pageId="31" pageNumber="32" specimenCode="20428" specimenCount="10" stateProvince="Jawa Barat" typeStatus="paratype">
<specimenCount id="9D1EA1D38B324E064D02FC093F97FBCB" box="[823,922,1013,1040]" count="10" pageId="31" pageNumber="32" type="generic">10 spec.</specimenCount>
;
<collectingMunicipality id="6BC3F0208B324E064D93FC093853FBD4" box="[934,1118,1013,1039]" pageId="31" pageNumber="32">Karangnunggal</collectingMunicipality>
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<location id="8EC73C818B324E064A5CFC0A397FFBD4" LSID="urn:lsid:plazi:treatment:03B1DB4C8B324E384C73FEF73962FE62:8EC73C818B324E064A5CFC0A397FFBD4" box="[1129,1394,1013,1040]" country="Indonesia" latitude="-7.6666665" longLatPrecision="21" longitude="108.11667" municipality="Karangnunggal" name="near Sela Dower Cave" pageId="31" pageNumber="32" stateProvince="Jawa Barat">near Sela Dower Cave</location>
,
<collectedFrom id="380429A78B324E064E88FBE43DE4FBE9" box="[189,489,1048,1074]" pageId="31" pageNumber="32">agroforestry and bamboo</collectedFrom>
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,
<geoCoordinate id="EE2C0C9D8B324E064CB3FBEA3F24FBE8" box="[646,809,1046,1075]" degrees="108" direction="east" minutes="07" orientation="longitude" pageId="31" pageNumber="32" precision="15" seconds="00" value="108.11667">108°07′00″ E</geoCoordinate>
;
<elevation id="00358D698B324E064D03FBE43FCFFBE9" box="[822,962,1048,1075]" metricMagnitude="2" metricUnit="m" metricValue="2.37" pageId="31" pageNumber="32" unit="m" value="237.0">
<quantity id="4CE0C7BF8B324E064D03FBE43F8FFBE8" box="[822,898,1048,1075]" metricMagnitude="2" metricUnit="m" metricValue="2.37" pageId="31" pageNumber="32" unit="m" value="237.0">237 m</quantity>
a.s.l.
</elevation>
;
<collectionCode id="ED09F29F8B324E064DE4FBE5381CFBE8" box="[977,1041,1049,1075]" collectionName="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="31" pageNumber="32" type="Museum">MZB</collectionCode>
<specimenCode id="DBBEC2218B324E064A2CFBE4386FFBE9" box="[1049,1122,1048,1074]" pageId="31" pageNumber="32">20428</specimenCode>
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•
<materialsCitation id="3B7060078B324E064A48FBE43E05FB8E" ID-GBIF-Occurrence="2243137452" collectionCode="ZMH" country="Indonesia" elevation="237" latitude="-7.6666665" location="Sela Dower Cave" longLatPrecision="21" longitude="108.11667" municipality="Karangnunggal" pageId="31" pageNumber="32" specimenCode="ZMH 148186" specimenCount="5" stateProvince="Jawa Barat" typeStatus="paratype">
<specimenCount id="9D1EA1D38B324E064A48FBE438DEFBE8" box="[1149,1235,1048,1075]" count="5" pageId="31" pageNumber="32" type="generic">5 spec.</specimenCount>
; same data as for preceding;
<specimenCode id="DBBEC2218B324E064F5DFBC03E05FB8E" box="[360,520,1083,1110]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="31" pageNumber="32" type="Museum">ZMH 148186</specimenCode>
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•
<materialsCitation id="3B7060078B324E064C2AFBC03849FB8D" ID-GBIF-Occurrence="2243137375" box="[543,1092,1083,1110]" collectingDate="2014-07-04" collectionCode="MZB" collectorName="A. S. Nurinsiyah &amp; T. Setiadi" country="Indonesia" elevation="243" latitude="-7.666111" location="Pasir Taraje Cave" longLatPrecision="21" longitude="108.11694" municipality="Karangnunggal" pageId="31" pageNumber="32" specimenCode="20430" specimenCount="3" stateProvince="Jawa Barat" typeStatus="paratype">
<specimenCount id="9D1EA1D38B324E064C2AFBC03E7EFB8D" box="[543,627,1084,1110]" count="3" pageId="31" pageNumber="32" type="generic">3 spec.</specimenCount>
; same data as for holotype;
<collectionCode id="ED09F29F8B324E064D80FBC03FF8FB8D" box="[949,1013,1084,1110]" collectionName="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="31" pageNumber="32" type="Museum">MZB</collectionCode>
<specimenCode id="DBBEC2218B324E064DCEFBC03849FB8D" box="[1019,1092,1084,1110]" pageId="31" pageNumber="32">20430</specimenCode>
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•
<materialsCitation id="3B7060078B324E064A69FBC03DD9FBA2" ID-GBIF-Occurrence="2243137453" collectingDate="2014-07-04" collectionCode="ZMH" collectorName="A. S. Nurinsiyah &amp; T. Setiadi" country="Indonesia" elevation="243" latitude="-7.666111" location="Pasir Taraje Cave" longLatPrecision="21" longitude="108.11694" municipality="Karangnunggal" pageId="31" pageNumber="32" specimenCode="ZMH 148187" specimenCount="3" stateProvince="Jawa Barat" typeStatus="paratype">
<specimenCount id="9D1EA1D38B324E064A69FBC038BDFB8D" box="[1116,1200,1084,1110]" count="3" pageId="31" pageNumber="32" type="generic">3 spec.</specimenCount>
; same data as for holotype;
<specimenCode id="DBBEC2218B324E064F06FBA33DD9FBA2" box="[307,468,1119,1145]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="31" pageNumber="32" type="Museum">ZMH 148187</specimenCode>
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•
<materialsCitation id="3B7060078B324E064FDBFBA33F23FB46" ID-GBIF-Occurrence="2243137467" collectedFrom="agroforestry and shrubs" collectionCode="ZMH" country="Indonesia" elevation="237" latitude="-7.6669445" location="surroundings of Liang Cai" longLatPrecision="21" longitude="108.1175" municipality="Karangnunggal" pageId="31" pageNumber="32" specimenCode="ZMH 148185" specimenCount="1" stateProvince="Jawa Barat" typeStatus="paratype">
<specimenCount id="9D1EA1D38B324E064FDBFBA33E4EFBA2" box="[494,579,1119,1145]" count="1" pageId="31" pageNumber="32" type="generic">1 spec.</specimenCount>
;
<collectingMunicipality id="6BC3F0208B324E064C65FBA33F05FBA2" box="[592,776,1119,1145]" pageId="31" pageNumber="32">Karangnunggal</collectingMunicipality>
,
<location id="8EC73C818B324E064D21FBA33841FBA2" LSID="urn:lsid:plazi:treatment:03B1DB4C8B324E384C73FEF73962FE62:8EC73C818B324E064D21FBA33841FBA2" box="[788,1100,1119,1145]" country="Indonesia" latitude="-7.6669445" longLatPrecision="21" longitude="108.1175" municipality="Karangnunggal" name="surroundings of Liang Cai" pageId="31" pageNumber="32" stateProvince="Jawa Barat">surroundings of Liang Cai</location>
,
<collectedFrom id="380429A78B324E064A6CFBA3397FFBA2" box="[1113,1394,1119,1145]" pageId="31" pageNumber="32">agroforestry and shrubs</collectedFrom>
;
<geoCoordinate id="EE2C0C9D8B324E064E88FB7C3D33FB47" box="[189,318,1152,1180]" degrees="7" direction="south" minutes="40" orientation="latitude" pageId="31" pageNumber="32" precision="15" seconds="01" value="-7.6669445">7°40′01″ S</geoCoordinate>
,
<geoCoordinate id="EE2C0C9D8B324E064F7FFB7C3DE6FB47" box="[330,491,1152,1180]" degrees="108" direction="east" minutes="07" orientation="longitude" pageId="31" pageNumber="32" precision="15" seconds="03" value="108.1175">108°07′03″ E</geoCoordinate>
;
<elevation id="00358D698B324E064FCDFB7E3E8CFB47" box="[504,641,1154,1180]" metricMagnitude="2" metricUnit="m" metricValue="2.37" pageId="31" pageNumber="32" unit="m" value="237.0">
<quantity id="4CE0C7BF8B324E064FCDFB7E3E4FFB47" box="[504,578,1154,1180]" metricMagnitude="2" metricUnit="m" metricValue="2.37" pageId="31" pageNumber="32" unit="m" value="237.0">237 m</quantity>
a.s.l.
</elevation>
;
<specimenCode id="DBBEC2218B324E064CBBFB7E3F23FB46" box="[654,814,1154,1181]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="31" pageNumber="32" type="Museum">ZMH 148185</specimenCode>
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•
<materialsCitation id="3B7060078B324E064D72FB7E3861FB64" ID-GBIF-Occurrence="2243137366" collectedFrom="agroforestry" collectionCode="MZB" country="Indonesia" county="Tasikmalaya" elevation="240" latitude="-7.67" location="mouth of Bojong Cave" longLatPrecision="21" longitude="108.12" municipality="Karangnunggal" pageId="31" pageNumber="32" specimenCode="20427" specimenCount="1" stateProvince="Jawa Barat" typeStatus="paratype">
<specimenCount id="9D1EA1D38B324E064D72FB7E3F96FB47" box="[839,923,1154,1180]" count="1" pageId="31" pageNumber="32" type="generic">1 spec.</specimenCount>
;
<collectingCounty id="62C612D68B324E064D92FB7E3833FB47" box="[935,1086,1154,1180]" pageId="31" pageNumber="32">Tasikmalaya</collectingCounty>
,
<collectingMunicipality id="6BC3F0208B324E064A7CFB7E390CFB47" box="[1097,1281,1154,1180]" pageId="31" pageNumber="32">Karangnunggal</collectingMunicipality>
,
<location id="8EC73C818B324E064B39FB7E3D57FB64" LSID="urn:lsid:plazi:treatment:03B1DB4C8B324E384C73FEF73962FE62:8EC73C818B324E064B39FB7E3D57FB64" country="Indonesia" county="Tasikmalaya" latitude="-7.67" longLatPrecision="21" longitude="108.12" municipality="Karangnunggal" name="mouth of Bojong Cave" pageId="31" pageNumber="32" stateProvince="Jawa Barat">mouth of Bojong Cave</location>
,
<collectedFrom id="380429A78B324E064F53FB593DF4FB64" box="[358,505,1189,1215]" pageId="31" pageNumber="32">agroforestry</collectedFrom>
;
<geoCoordinate id="EE2C0C9D8B324E064C31FB5F3E8BFB64" box="[516,646,1187,1215]" degrees="7" direction="south" minutes="40" orientation="latitude" pageId="31" pageNumber="32" precision="15" seconds="12" value="-7.67">7°40′12″ S</geoCoordinate>
,
<geoCoordinate id="EE2C0C9D8B324E064CA6FB5F3F39FB1B" box="[659,820,1187,1216]" degrees="108" direction="east" minutes="07" orientation="longitude" pageId="31" pageNumber="32" precision="15" seconds="12" value="108.12">108°07′12″ E</geoCoordinate>
;
<elevation id="00358D698B324E064D75FB593FC7FB64" box="[832,970,1189,1216]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="31" pageNumber="32" unit="m" value="240.0">
<quantity id="4CE0C7BF8B324E064D75FB593F86FB1B" box="[832,907,1189,1216]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="31" pageNumber="32" unit="m" value="240.0">240 m</quantity>
a.s.l.
</elevation>
;
<collectionCode id="ED09F29F8B324E064DEDFB5A3815FB1B" box="[984,1048,1190,1216]" collectionName="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="31" pageNumber="32" type="Museum">MZB</collectionCode>
<specimenCode id="DBBEC2218B324E064A2AFB593861FB64" box="[1055,1132,1189,1215]" pageId="31" pageNumber="32">20427</specimenCode>
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.
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="C30239D18B324E064E88FB1A3DB6F9C2" pageId="31" pageNumber="32" type="description">
<paragraph id="8BA76A5A8B324E064E88FB1A3D43FADB" blockId="31.[189,334,1254,1280]" box="[189,334,1254,1280]" pageId="31" pageNumber="32">
<heading id="D0EFDD368B324E064E88FB1A3D43FADB" bold="true" box="[189,334,1254,1280]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="B96CB6488B324E064E88FB1A3D43FADB" bold="true" box="[189,334,1254,1280]" pageId="31" pageNumber="32">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B324E064E88FAE93E1AFA04" blockId="31.[189,1399,1301,1503]" pageId="31" pageNumber="32">
SHELL (Fig. 48;
<tableCitation id="C69A5FE18B324E064F4AFAE93E0BFAF4" box="[383,518,1301,1327]" captionStart-0="Table 3" captionStart-1="Table 4" captionStartId-0="13.[189,258,268,294]" captionStartId-1="22.[188,257,551,579]" captionTargetBox-0="[201,1370,481,1908]" captionTargetBox-1="[205,1929,661,1732]" captionTargetPageId-0="13" captionTargetPageId-1="22" captionText-0="Table 3 (continued on next 3 pages). Shell measurements for species of Landouria Godwin-Austen, 1918 (in mm). Abbreviations: D = shell diameter; D3W = diameter of the third whorl; da = diameter of the aperture; H = shell height; ha = height of the aperture; max = maximum; min = minimum; n = number of measured spec.; std = standard deviation; U = diameter of the umbilicus; W = number of whorls." captionText-1="Table 4 (continued on next 3 pages). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java. Characteristics most important for distinguishing similar taxa are printed in bold." httpUri-0="http://table.plazi.org/id/DF673AD28B204E144E88FEF03D19FE68" httpUri-1="http://table.plazi.org/id/DF673AD28B3B4E0F4E89FDDB3F73FDBC" pageId="31" pageNumber="32" tableUuid-0="DF673AD28B204E144E88FEF03D19FE68" tableUuid-1="DF673AD28B3B4E0F4E89FDDB3F73FDBC">Tables 3–4</tableCitation>
). Conical, with 6.5–7 hardly convex whorls; protoconch almost smooth; teleoconch with fine, irregular wrinkles; without distinct incised spiral lines; scaly processes sparse, only at periphery; tuberculate around umbilicus; brownish-corneous; body whorl sharply keeled; aperture rounded rhombic; upper insertion of peristome slightly descending; peristome expanded, reflexed and slightly thickened; umbilicus hardly eccentric, wide, comprising 39–45% of shell diameter, hardly obscured by columellar edge.
</paragraph>
<paragraph id="8BA76A5A8B324E064E88FA033DB6F9C2" blockId="31.[189,443,1535,1562]" box="[189,443,1535,1562]" pageId="31" pageNumber="32">GENITALIA. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C30239D18B324E384E88F9BD3F52FE93" lastPageId="33" lastPageNumber="34" pageId="31" pageNumber="32" type="discussion">
<paragraph id="8BA76A5A8B324E064E88F9BD3D22F980" blockId="31.[189,303,1601,1627]" box="[189,303,1601,1627]" pageId="31" pageNumber="32">
<heading id="D0EFDD368B324E064E88F9BD3D22F980" bold="true" box="[189,303,1601,1627]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="B96CB6488B324E064E88F9BD3D22F980" bold="true" box="[189,303,1601,1627]" pageId="31" pageNumber="32">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B324E064E88F993397AF90B" blockId="31.[189,1399,1647,1744]" pageId="31" pageNumber="32">
<taxonomicName id="4C1811D98B324E064E88F9933E0CF952" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[189,513,1647,1673]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="parahyangensis" status="sp. nov.">
<emphasis id="B96CB6488B324E064E88F9933E0CF952" box="[189,513,1647,1673]" italics="true" pageId="31" pageNumber="32">Landouria parahyangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B324E064C3AF9933E7CF952" box="[527,625,1647,1673]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
resembles
<taxonomicName id="4C1811D98B324E064D37F98C3F9DF952" baseAuthorityName="Mollendorff" baseAuthorityYear="1897" box="[770,912,1647,1673]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="epiplatia">
<emphasis id="B96CB6488B324E064D37F98C3F9DF952" box="[770,912,1647,1673]" italics="true" pageId="31" pageNumber="32">L. epiplatia</emphasis>
</taxonomicName>
(Figs 26–28) with regard to the wide umbilicus. It differs from
<taxonomicName id="4C1811D98B324E064FC1F96F3E70F977" baseAuthorityName="Mollendorff" baseAuthorityYear="1897" box="[500,637,1682,1708]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="epiplatia">
<emphasis id="B96CB6488B324E064FC1F96F3E70F977" box="[500,637,1682,1708]" italics="true" pageId="31" pageNumber="32">L. epiplatia</emphasis>
</taxonomicName>
in the less sharply keeled, higher, conical shell (H
<quantity id="4CE0C7BF8B324E064AD2F96E397AF976" box="[1255,1399,1682,1709]" metricMagnitude="-3" metricUnit="m" metricValue="8.85" metricValueMax="9.3" metricValueMin="8.4" pageId="31" pageNumber="32" unit="mm" value="8.85" valueMax="9.3" valueMin="8.4">8.4–9.3 mm</quantity>
vs
<quantity id="4CE0C7BF8B324E064ED5F9493D7DF90B" box="[224,368,1717,1744]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" metricValueMax="6.9" metricValueMin="5.7" pageId="31" pageNumber="32" unit="mm" value="6.3" valueMax="6.9" valueMin="5.7">5.7–6.9 mm</quantity>
in
<taxonomicName id="4C1811D98B324E064FACF94A3E2FF914" baseAuthorityName="Mollendorff" baseAuthorityYear="1897" box="[409,546,1717,1743]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="epiplatia">
<emphasis id="B96CB6488B324E064FACF94A3E2FF914" box="[409,546,1717,1743]" italics="true" pageId="31" pageNumber="32">L. epiplatia</emphasis>
</taxonomicName>
; D/H 1.69–1.94 vs
<quantity id="4CE0C7BF8B324E064D27F9493FA5F914" box="[786,936,1717,1743]" metricMagnitude="-2" metricUnit="m" metricValue="6.3119" metricValueMax="7.4676" metricValueMin="5.156199999999999" pageId="31" pageNumber="32" unit="in" value="2.485" valueMax="2.94" valueMin="2.03">2.03–2.94 in</quantity>
<taxonomicName id="4C1811D98B324E064D84F94A3836F914" baseAuthorityName="Mollendorff" baseAuthorityYear="1897" box="[945,1083,1717,1743]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="epiplatia">
<emphasis id="B96CB6488B324E064D84F94A3836F914" box="[945,1083,1717,1743]" italics="true" pageId="31" pageNumber="32">L. epiplatia</emphasis>
</taxonomicName>
) with more whorls (6.5–7
</paragraph>
<caption id="DF673AD28B324E064E88F8DC3EF0F830" httpUri="https://zenodo.org/record/3152259/files/figure.png" pageId="31" pageNumber="32" targetBox="[280,1308,265,2019]" targetPageId="32">
<paragraph id="8BA76A5A8B324E064E88F8DC3EF0F830" blockId="31.[189,1399,1824,2027]" pageId="31" pageNumber="32">
<emphasis id="B96CB6488B324E064E88F8DC3D47F8E1" bold="true" box="[189,330,1824,1850]" pageId="31" pageNumber="32">Figs 48–51</emphasis>
(opposite page). Shells of species of
<taxonomicName id="4C1811D98B324E064D0AF8DD3FB6F8E0" authorityName="Godwin-Austen" authorityYear="1918" box="[831,955,1825,1851]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="genus">
<emphasis id="B96CB6488B324E064D0AF8DD3FB6F8E0" box="[831,955,1825,1851]" italics="true" pageId="31" pageNumber="32">Landouria</emphasis>
</taxonomicName>
from
<collectingRegion id="49DCA4B88B324E064A21F8DD3841F8E0" box="[1044,1100,1825,1851]" country="Indonesia" name="Jawa Timur" pageId="31" pageNumber="32">Java</collectingRegion>
.
<emphasis id="B96CB6488B324E064A55F8DC3870F8E1" bold="true" box="[1120,1149,1824,1850]" pageId="31" pageNumber="32">48</emphasis>
.
<taxonomicName id="4C1811D98B324E064AA0F8DD397AF8E0" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[1173,1399,1825,1851]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="parahyangensis" status="sp. nov.">
<emphasis id="B96CB6488B324E064AA0F8DD397AF8E0" box="[1173,1399,1825,1851]" italics="true" pageId="31" pageNumber="32">L. parahyangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B324E064E88F8B83D2CF885" box="[189,289,1860,1886]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
,
<collectingRegion id="49DCA4B88B324E064F06F8B83DBFF885" box="[307,434,1860,1886]" country="Indonesia" name="Jawa Barat" pageId="31" pageNumber="32">West Java</collectingRegion>
, Karangnunggal, surroundings of Pasir Taraje Cave (
<typeStatus id="54A3D4F88B324E064A65F8B838B6F885" box="[1104,1211,1860,1886]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
, MZB 20429).
<emphasis id="B96CB6488B324E064E88F89B3CD7F85A" bold="true" box="[189,218,1895,1921]" pageId="31" pageNumber="32">49</emphasis>
.
<taxonomicName id="4C1811D98B324E064ED8F8943DE2F85A" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[237,495,1895,1921]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="nusakambangensis" status="sp. nov.">
<emphasis id="B96CB6488B324E064ED8F8943DE2F85A" box="[237,495,1895,1921]" italics="true" pageId="31" pageNumber="32">L. nusakambangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B324E064FCFF89B3E51F85A" box="[506,604,1895,1921]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
,
<collectingRegion id="49DCA4B88B324E064C59F89B3F08F85A" box="[620,773,1895,1921]" country="Indonesia" name="Jawa Tengah" pageId="31" pageNumber="32">Central Java</collectingRegion>
, Nusa Kambangan Island, nature monument Gligir (
<typeStatus id="54A3D4F88B324E064EF0F8763D3DF87F" box="[197,304,1930,1956]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
, ZMA 412298a).
<emphasis id="B96CB6488B324E064FCAF8763E11F87F" bold="true" box="[511,540,1930,1956]" pageId="31" pageNumber="32">50</emphasis>
.
<taxonomicName id="4C1811D98B324E064C1FF8773EC3F87F" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[554,718,1930,1956]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="petrukensis" status="sp. nov.">
<emphasis id="B96CB6488B324E064C1FF8773EC3F87F" box="[554,718,1930,1956]" italics="true" pageId="31" pageNumber="32">L. petrukensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B324E064CE0F8773F3CF87E" box="[725,817,1931,1957]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
,
<collectingRegion id="49DCA4B88B324E064D09F8763FDCF87E" box="[828,977,1930,1957]" country="Indonesia" name="Jawa Tengah" pageId="31" pageNumber="32">Central Java</collectingRegion>
, Karangbolong, way to Petruk Cave (
<typeStatus id="54A3D4F88B324E064EF0F8513D3DF81C" box="[197,304,1965,1991]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
, MZB 19203).
<emphasis id="B96CB6488B324E064FD6F8513E0DF81C" bold="true" box="[483,512,1965,1991]" pageId="31" pageNumber="32">51</emphasis>
.
<taxonomicName id="4C1811D98B324E064C3BF8523EA3F81C" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[526,686,1965,1991]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="tholiformis" status="sp. nov.">
<emphasis id="B96CB6488B324E064C3BF8523EA3F81C" box="[526,686,1965,1991]" italics="true" pageId="31" pageNumber="32">L. tholiformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B324E064C81F8523F1DF813" box="[692,784,1966,1992]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
,
<collectingRegion id="49DCA4B88B324E064D2FF8513FAEF81C" box="[794,931,1965,1991]" country="Indonesia" name="Yogyakarta" pageId="31" pageNumber="32">Yogyakarta</collectingRegion>
, Gunung Kidul, Tepus, near Poktunggal Beach (
<typeStatus id="54A3D4F88B324E064F23F82C3D8CF831" box="[278,385,2000,2026]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
, MZB 17999). Scale bar:
<quantity id="4CE0C7BF8B324E064C84F82C3EF5F830" box="[689,760,2000,2027]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="31" pageNumber="32" unit="mm" value="5.0">5 mm</quantity>
.
</paragraph>
</caption>
<paragraph id="8BA76A5A8B0C4E384E88FEF03F52FE93" blockId="33.[189,1399,267,329]" pageId="33" pageNumber="34">
vs
<quantity id="4CE0C7BF8B0C4E384ED7FEF73D77FEFE" box="[226,378,267,294]" metricMagnitude="-1" metricUnit="m" metricValue="1.4605" metricValueMax="1.5875" metricValueMin="1.3335" pageId="33" pageNumber="34" unit="in" value="5.75" valueMax="6.25" valueMin="5.25">5.25–6.25 in</quantity>
<taxonomicName id="4C1811D98B0C4E384FB0FEF03E1DFEFE" baseAuthorityName="Mollendorff" baseAuthorityYear="1897" box="[389,528,267,293]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="epiplatia">
<emphasis id="B96CB6488B0C4E384FB0FEF03E1DFEFE" box="[389,528,267,293]" italics="true" pageId="33" pageNumber="34">L. epiplatia</emphasis>
</taxonomicName>
).
<taxonomicName id="4C1811D98B0C4E384C19FEF73F60FEFE" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[556,877,267,293]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="parahyangensis" status="sp. nov.">
<emphasis id="B96CB6488B0C4E384C19FEF73F60FEFE" box="[556,877,267,293]" italics="true" pageId="33" pageNumber="34">Landouria parahyangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B0C4E384D4DFEF03FDAFEFD" box="[888,983,268,294]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
also resembles
<taxonomicName id="4C1811D98B0C4E384AA9FEF03900FEFE" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[1180,1293,268,293]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="naggsi" status="sp. nov.">
<emphasis id="B96CB6488B0C4E384AA9FEF03900FEFE" box="[1180,1293,268,293]" italics="true" pageId="33" pageNumber="34">L. naggsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B0C4E384B2DFEF0397AFEFD" box="[1304,1399,268,294]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
(Fig. 25). For differences, see remarks under that species.
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B0C4E384E88FE8C3962FE62" box="[189,1391,368,441]" pageId="33" pageNumber="34" type="distribution">
<paragraph id="8BA76A5A8B0C4E384E88FE8C3D5BFE51" blockId="33.[189,342,368,394]" box="[189,342,368,394]" pageId="33" pageNumber="34">
<heading id="D0EFDD368B0C4E384E88FE8C3D5BFE51" bold="true" box="[189,342,368,394]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="B96CB6488B0C4E384E88FE8C3D5BFE51" bold="true" box="[189,342,368,394]" pageId="33" pageNumber="34">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B0C4E384E88FE633962FE62" blockId="33.[189,1391,414,441]" box="[189,1391,414,441]" pageId="33" pageNumber="34">
<taxonomicName id="4C1811D98B0C4E384E88FE633DF7FE62" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[189,506,415,441]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="parahyangensis" status="sp. nov.">
<emphasis id="B96CB6488B0C4E384E88FE633DF7FE62" box="[189,506,415,441]" italics="true" pageId="33" pageNumber="34">Landouria parahyangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B0C4E384C34FE633E51FE62" box="[513,604,415,441]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
is endemic to the Karangnunggal karst area in
<collectingRegion id="49DCA4B88B0C4E384AB1FE6338F4FE62" box="[1156,1273,415,441]" country="Indonesia" name="Jawa Barat" pageId="33" pageNumber="34">West Java</collectingRegion>
(Fig. 54).
</paragraph>
</subSubSection>
</treatment>
</document>
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