treatments-xml/data/03/A3/CA/03A3CA13FFCDFFD5FF36967AFF41E15F.xml

<document id="7CC85BCC3210EF72C4098C0D529CA332" ID-CLB-Dataset="26538" ID-DOI="10.11646/zootaxa.4614.2.4" ID-GBIF-Dataset="57b938b5-ff1c-4df8-9d94-416d8a3668dc" ID-ISSN="1175-5326" ID-Zenodo-Dep="3242436" ID-ZooBank="FDC07BFF-38D1-4A0E-8599-F1DCB29E2112" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1560166616111" checkinUser="plazi" docAuthor="Davis, Hayden R., Bauer, Aaron M., Jackman, Todd R., Nashriq, Izneil &amp; Das, Indraneil" docDate="2019" docId="03A3CA13FFCDFFD5FF36967AFF41E15F" docLanguage="en" docName="zootaxa.4614.2.4.pdf" docOrigin="Zootaxa 4614 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyrtodactylus limajalur Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das 2019, sp. nov." docType="treatment" docUuid="E114F921-E406-4332-BABD-22C7BEDD6F99" docUuidSource="ZooBank" docVersion="10" lastPageNumber="347" masterDocId="FF9AB26BFFC7FFC5FFA1957FFFAEE731" masterDocTitle="Uncovering karst endemism within Borneo: two new Cyrtodactylus species from Sarawak, Malaysia" masterLastPageNumber="352" masterPageNumber="331" pageNumber="341" updateTime="1698733839631" updateUser="ExternalLinkService">
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<mods:title id="2D1D53CDC8E369C2F9E1C00B216C54F7">Uncovering karst endemism within Borneo: two new Cyrtodactylus species from Sarawak, Malaysia</mods:title>
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<mods:namePart id="F5C803E31999F3225D53C272E650945B">Davis, Hayden R.</mods:namePart>
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<mods:namePart id="772D6F640642033BE84E34328CE6624A">Bauer, Aaron M.</mods:namePart>
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<mods:namePart id="C16BAA642195C01FD7CAB59B74B8FE28">Jackman, Todd R.</mods:namePart>
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<mods:namePart id="5A9CCC21D6C9C6C3B3721FAF3D7B50CA">Nashriq, Izneil</mods:namePart>
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<taxonomicName id="4C0A0086FFCDFFCFFF36967AFE1AE42E" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[151,436,773,799]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="341" phylum="Chordata" rank="species" species="limajalur" status="sp. nov.">
<emphasis id="B97EA717FFCDFFCFFF36967AFE1AE42E" bold="true" box="[151,436,773,799]" italics="true" pageId="10" pageNumber="341">Cyrtodactylus limajalur</emphasis>
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<taxonomicNameLabel id="A24D1A6CFFCDFFCFFE1D967AFDB4E42E" box="[444,538,773,799]" pageId="10" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
<paragraph id="8BB57B05FFCDFFCFFF369658FE46E471" blockId="10.[151,538,773,832]" box="[151,488,807,832]" pageId="10" pageNumber="341">Five-banded Bent-toed Gecko</paragraph>
</subSubSection>
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<paragraph id="8BB57B05FFCDFFCFFF369610FDA4E4E1" blockId="10.[151,1437,879,2021]" pageId="10" pageNumber="341">
<materialsCitation id="3B627158FFCDFFCFFF369610FDA4E4E1" ID-GBIF-Occurrence="2251813346" collectingDate="2017-08-06" collectionCode="CAS, WGS" country="Malaysia" county="East" elevation="88" latitude="1.2965611" location="Adult" longLatPrecision="1" longitude="110.4425" municipality="Serian" pageId="10" pageNumber="341" specimenCode="CAS 262848, WGS 1984" specimenCount="1" typeStatus="holotype">
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<emphasis id="B97EA717FFCDFFCFFF369610FEAAE4B9" bold="true" box="[151,260,879,904]" pageId="10" pageNumber="341">Holotype</emphasis>
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(
<figureCitation id="13316780FFCDFFCFFEB99610FECFE4B9" box="[280,353,879,904]" captionStart="FIGURE 5" captionStartId="15.[151,250,1914,1936]" captionTargetBox="[161,1425,438,1779]" captionTargetId="figure@15.[161,1426,428,1817]" captionTargetPageId="15" captionText="FIGURE 5. Preserved type series of Cyrtodactylus limajalur sp. nov.; A) type series displaying dorsal side of body (from left to right: CAS 262849; CAS 262848; CAS 262847; and CAS 262846); and B) type series displaying ventral side of body (specimens arranged in the same order as figure 5A)." httpUri="https://zenodo.org/record/3242446/files/figure.png" pageId="10" pageNumber="341">Fig. 5</figureCitation>
). 
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male, 
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collected near 
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, 
<collectingMunicipality id="6BD1E17FFFCDFFCFFB3F9610FB46E4B9" box="[1182,1256,879,904]" pageId="10" pageNumber="341">Serian</collectingMunicipality>
, Sarawak, 
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<collectingCountry id="F31D3B95FFCDFFCFFF3696EBFEAFE49C" box="[151,257,916,941]" name="Malaysia" pageId="10" pageNumber="341">Malaysia</collectingCountry>
. (
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; 
<geoCoordinate id="EE3E1DC2FFCDFFCFFE6B96EBFDD0E49C" box="[458,638,916,941]" degrees="110" direction="east" minutes="26" orientation="longitude" pageId="10" pageNumber="341" precision="1" seconds="33.00" value="110.4425">110°26'33.00&quot;E</geoCoordinate>
; 
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<quantity id="4CF2D6E0FFCDFFCFFD2B96EBFD68E49D" box="[650,710,916,940]" metricMagnitude="1" metricUnit="m" metricValue="8.8" pageId="10" pageNumber="341" unit="m" value="88.0">88 m</quantity>
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; 
<specimenCode id="DBACD37EFFCDFFCFFD7096EBFCFEE49D" box="[721,848,916,940]" collectionCode="WGS" pageId="10" pageNumber="341">WGS 1984</specimenCode>
) by Izneil Nashriq and Hayden Davis on 
<collectingDate id="EFF0A42DFFCDFFCFFA8096EBFF61E4E1" pageId="10" pageNumber="341" value="2017-08-06">
<date id="FFB45DC5FFCDFFCFFA8096EBFF61E4E1" pageId="10" pageNumber="341" value="2017-08-06">August 06, 2017</date>
</collectingDate>
at 20:00–22:00 hrs (
<figureCitation id="13316780FFCDFFCFFE1696C8FE57E4E1" box="[439,505,951,976]" captionStart="FIGURE 1" captionStartId="3.[151,250,1124,1146]" captionTargetBox="[317,1278,184,1110]" captionTargetId="figure@3.[305,1282,176,1120]" captionTargetPageId="3" captionText="FIGURE 1. Preserved type series of Cyrtodactylus muluensis sp. nov. A) type series displaying dorsal side of body (from left to right: CAS 262995; CAS 262986; CAS 262983; CAS 262997; CAS 262996; and CAS 262984; CAS 262985 not shown); and B) type series displaying ventral side of body (specimens arranged in the same order as figure 1A)." httpUri="https://zenodo.org/record/3242438/files/figure.png" pageId="10" pageNumber="341">Fig. 1</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="8BB57B05FFCDFFCFFF6696A3FE84E329" blockId="10.[151,1437,879,2021]" pageId="10" pageNumber="341">
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<emphasis id="B97EA717FFCDFFCFFF6696A3FE91E4C4" bold="true" box="[199,319,988,1013]" pageId="10" pageNumber="341">Paratypes</emphasis>
</typeStatus>
. 
<materialsCitation id="3B627158FFCDFFCFFEEC96A2FE88E329" ID-GBIF-Occurrence="2251813309" collectingDate="2017-08-06" collectionCode="CAS" country="Malaysia" county="East" elevation="88" latitude="1.2965611" location="Adult" longLatPrecision="1" longitude="110.4425" municipality="Serian" pageId="10" pageNumber="341" specimenCode="CAS 262846, CAS 262847, CAS262849" specimenCount="1" typeStatus="paratype">
<typeStatus id="54B1C5A7FFCDFFCFFEEC96A2FE14E4C4" box="[333,442,989,1013]" pageId="10" pageNumber="341" type="paratype">Paratypes</typeStatus>
(
<specimenCode id="DBACD37EFFCDFFCFFE6896A3FDCEE4C4" box="[457,608,988,1013]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="10" pageNumber="341" type="Museum">CAS 262846</specimenCode>
; 
<specimenCode id="DBACD37EFFCDFFCFFDCA96A3FCAFE4C4" box="[619,769,988,1013]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="10" pageNumber="341" type="Museum">CAS 262847</specimenCode>
; and 
<specimenCode id="DBACD37EFFCDFFCFFC9D96A3FC66E4C4" box="[828,968,988,1013]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="10" pageNumber="341" type="Museum">CAS262849</specimenCode>
) (
<figureCitation id="13316780FFCDFFCFFC7E96A3FB8CE4C4" box="[991,1058,988,1013]" captionStart="FIGURE 5" captionStartId="15.[151,250,1914,1936]" captionTargetBox="[161,1425,438,1779]" captionTargetId="figure@15.[161,1426,428,1817]" captionTargetPageId="15" captionText="FIGURE 5. Preserved type series of Cyrtodactylus limajalur sp. nov.; A) type series displaying dorsal side of body (from left to right: CAS 262849; CAS 262848; CAS 262847; and CAS 262846); and B) type series displaying ventral side of body (specimens arranged in the same order as figure 5A)." httpUri="https://zenodo.org/record/3242446/files/figure.png" pageId="10" pageNumber="341">Fig. 5</figureCitation>
) have the same collection data as the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C310288EFFCDFFCFFF66915BFDA4E26D" pageId="10" pageNumber="341" type="diagnosis">
<paragraph id="8BB57B05FFCDFFCFFF66915BFDA4E26D" blockId="10.[151,1437,879,2021]" pageId="10" pageNumber="341">
<emphasis id="B97EA717FFCDFFCFFF66915BFE94E30C" bold="true" box="[199,314,1060,1085]" pageId="10" pageNumber="341">Diagnosis</emphasis>
. 
<taxonomicName id="4C0A0086FFCDFFCFFEEB915BFDF6E30D" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[330,600,1060,1084]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="341" phylum="Chordata" rank="species" species="limajalur" status="sp. nov.">
<emphasis id="B97EA717FFCDFFCFFEEB915BFDF6E30D" box="[330,600,1060,1084]" italics="true" pageId="10" pageNumber="341">Cyrtodactylus limajalur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFCDFFCFFDC0915BFD13E30C" box="[609,701,1060,1085]" pageId="10" pageNumber="341" rank="species">
<emphasis id="B97EA717FFCDFFCFFDC0915BFD13E30C" bold="true" box="[609,701,1060,1085]" pageId="10" pageNumber="341">sp. nov.</emphasis>
</taxonomicNameLabel>
can be differentiated from all other species of 
<taxonomicName id="4C0A0086FFCDFFCFFB7B915BFAD9E30D" box="[1242,1399,1060,1084]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="341" phylum="Chordata" rank="genus">
<emphasis id="B97EA717FFCDFFCFFB7B915BFAD9E30D" box="[1242,1399,1060,1084]" italics="true" pageId="10" pageNumber="341">Cyrtodactylus</emphasis>
</taxonomicName>
by the combination of the following characters: maximum SVL of at least 
<quantity id="4CF2D6E0FFCDFFCFFBA89137FBCCE36E" box="[1033,1122,1096,1120]" metricMagnitude="-2" metricUnit="m" metricValue="9.4" pageId="10" pageNumber="341" unit="mm" value="94.0">94 mm</quantity>
; 10–12 supralabials; 9–11 infralabials; weak tuberculation on body; no tubercles on ventral surface of forelimbs, gular region, or ventrolateral folds; 34–38 paravertebral tubercles; 11–13 longitudinal tubercle rows; 33–42 ventral scales; 19–22 subdigital lamellae on fourth toe; 5–6 enlarged femoral scales; no femoral pores; 7–8 precloacal pores; precloacal pit in males; enlarged median row of transverse scales; 5 dark dorsal body bands; body bands more than two times the width of interspaces; rostral chevron just posterior to orbitals; no white line edging the body bands and nuchal loop; and no scattered white tubercles on dorsum. These characters are scored across all currently recorded Bornean 
<taxonomicName id="4C0A0086FFCDFFCFFF36903BFE9AE26D" box="[151,308,1348,1372]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="341" phylum="Chordata" rank="genus">
<emphasis id="B97EA717FFCDFFCFFF36903BFE9AE26D" box="[151,308,1348,1372]" italics="true" pageId="10" pageNumber="341">Cyrtodactylus</emphasis>
</taxonomicName>
species in Table 3.
</paragraph>
</subSubSection>
<subSubSection id="C310288EFFCDFFCEFF669018FE80E009" lastPageId="11" lastPageNumber="342" pageId="10" pageNumber="341" type="description">
<paragraph id="8BB57B05FFCDFFCFFF669018FD8EE0F0" blockId="10.[151,1437,879,2021]" pageId="10" pageNumber="341">
<emphasis id="B97EA717FFCDFFCFFF669018FE72E2B1" bold="true" box="[199,476,1383,1408]" pageId="10" pageNumber="341">
Description of 
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</emphasis>
. Adult male; 
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SVL; head large, moderate in length (HL/SVL 0.28), relatively narrow (HW/HL 0.46), moderately flattened (HD/HL 0.36), distinct from neck, triangular in dorsal profile; loreal scales slightly concave posteriorly, flat anteriorly; frontal and prefrontal regions concave; canthus rostralis rounded; snout elongate (ES/HL 0.44), rounded in dorsal profile, laterally constricted; eye large (ED/HL 0.27); ear opening elliptical, moderate in size (EL/HL 0.10), obliquely oriented; eye to ear distance less than diameter of eye; rostral scale rectangular, partially divided dorsally by line extending from the posteriormost portion of the rostral scale to approximately half the distance anteriorly, two postnasal scales, one medial postrostralis (internasal), bordered laterally by first supralabials; external nares bordered anteriorly by rostral, ventrally by first supralabial; 12/10 (L/R) rectangular supralabials extending to the upturn of the labial margin, tapering abruptly directly below midpoint of eye; first supralabial largest; 9/11 (L/R) infralabials extending to the upturn of the labial margin, tapering abruptly directly below midpoint of eye; loreal scales weakly raised, same size as scales on top of head, occiput, and canthus rostralis; no tubercles on occiput or interorbital region; bony ridge bordering the orbital rim; transverse frontoparietal ridge; 38/34 (L/R) supracilliary scales, elongate, smooth, largest anteriorly; mental triangular, bordered laterally by first infralabials and posteriorly by left and right rectangular postmentals which contact medially for 40% of their length, forming a Y-shape; single row of slightly enlarged, elongate chinshields extending posteriorly to sixth infralabial scale; small, flat gular scales gradually grading posteriorly into larger, flat, smooth pectoral and ventral scales.
</paragraph>
<paragraph id="8BB57B05FFCDFFCEFF6692B3FAC7E671" blockId="10.[151,1437,879,2021]" lastBlockId="11.[151,1436,151,789]" lastPageId="11" lastPageNumber="342" pageId="10" pageNumber="341">Body with distinct, non-tuberculate ventrolateral folds; dorsal scales small, granular interspersed with low, regularly arranged smooth tubercles; small intervening tubercles occasionally present; tubercles extend from top of head to caudal constriction, and onto anterior one-fifth of tail; tubercles on occiput and nape small, those on posterior portion of body largest; approximately 13 longitudinal rows of tubercles slightly posterior of midbody; 38 paravertebral tubercles; 42 flat imbricate ventral scales between ventrolateral body folds; ventral scales larger than dorsal scales; precloacal scales with 5/3 (L/R) pores, slightly larger than ventral scales; shallow precloacal pit.</paragraph>
<paragraph id="8BB57B05FFCCFFCEFF669433FAD4E599" blockId="11.[151,1436,151,789]" pageId="11" pageNumber="342">Forelimbs relatively short (FL/SVL 0.17); scales on preaxial surface of forelimbs small, tubercles absent; scales on postaxial surface flat, tubercles absent; palmar scales weakly rounded; digits well developed, inflected at basal interphalangeal joints; 22/21 (L/R) subdigital lamellae on fourth toe rectangular, broadly expanded proximal to joint inflection, slightly expanded immediately distal to joint becoming gradually more expanded near the claw; claws well-developed, relatively short; hind limbs more robust than forelimbs, moderate in length (TBL/SVL 0.20); postaxial thigh scales flat, smooth, slightly larger than dorsal granular scales; ventral tibial scales flat, smooth; 8/6 (L/R) expanded femoral scales; expanded precloacal scales; 7 pore-bearing precloacal scales; precloacal scales expanded, forming pit; plantar scales slightly raised; digits well developed, inflected at basal, interphalangeal joints; 20/21 (L/R) subdigital lamellae on fourth toe rectangular, broadly expanded proximal to joint inflection, slightly expanded immediately distal to joint becoming gradually more expanded near the claw.</paragraph>
<paragraph id="8BB57B05FFCCFFCEFF6697CBFCB4E425" blockId="11.[151,1436,151,789]" pageId="11" pageNumber="342">
Tail regenerated from base, 114.0 mm long, 
<quantity id="4CF2D6E0FFCCFFCEFD1497CBFCA4E5FD" box="[693,778,692,717]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="11" pageNumber="342" unit="mm" value="6.6">6.6 mm</quantity>
wide at base, tapering to a point distally; dorsal scales flat, squarish; subcaudal region bearing enlarged median row of transverse scales; no caudal furrow; base of tail forming hemipenal swelling; and 3/2 (L/R) cloacal spurs.
</paragraph>
<caption id="DF752B8DFFCCFFCEFF369032FBF9E28E" httpUri="https://zenodo.org/record/3242444/files/figure.png" pageId="11" pageNumber="342" startId="11.[151,250,1357,1379]" targetBox="[163,1436,836,1342]" targetPageId="11">
<paragraph id="8BB57B05FFCCFFCEFF369032FBF9E28E" blockId="11.[151,1436,1357,1471]" pageId="11" pageNumber="342">
<emphasis id="B97EA717FFCCFFCEFF369032FEBCE253" bold="true" box="[151,274,1357,1379]" pageId="11" pageNumber="342">FIGURE 4</emphasis>
. A) Mulu karst formation demonstrating 
<taxonomicName id="4C0A0086FFCCFFCEFD759032FC7DE252" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[724,979,1357,1379]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="342" phylum="Chordata" rank="species" species="muluensis" status="sp. nov.">
<emphasis id="B97EA717FFCCFFCEFD759032FC7DE252" box="[724,979,1357,1379]" italics="true" pageId="11" pageNumber="342">Cyrtodactylus muluensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFCCFFCEFC7E9032FB9BE253" box="[991,1077,1357,1378]" pageId="11" pageNumber="342" rank="species">
<emphasis id="B97EA717FFCCFFCEFC7E9032FB9BE253" bold="true" box="[991,1077,1357,1378]" pageId="11" pageNumber="342">sp. nov.</emphasis>
</taxonomicNameLabel>
microhabitat; B) type locality of 
<taxonomicName id="4C0A0086FFCCFFCEFF369013FE24E2B3" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[151,394,1388,1410]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="342" phylum="Chordata" rank="species" species="limajalur" status="sp. nov.">
<emphasis id="B97EA717FFCCFFCEFF369013FE24E2B3" box="[151,394,1388,1410]" italics="true" pageId="11" pageNumber="342">Cyrtodactylus limajalur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFCCFFCEFE309012FE4FE2B3" box="[401,481,1389,1410]" pageId="11" pageNumber="342" rank="species">sp. nov.</taxonomicNameLabel>
The type locality of 
<taxonomicName id="4C0A0086FFCCFFCEFD199013FC95E2B3" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[696,827,1388,1410]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="342" phylum="Chordata" rank="species" species="muluensis" status="sp. nov.">
<emphasis id="B97EA717FFCCFFCEFD199013FC95E2B3" box="[696,827,1388,1410]" italics="true" pageId="11" pageNumber="342">C. muluensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFCCFFCEFCE39013FC3AE2B0" box="[834,916,1388,1409]" pageId="11" pageNumber="342" rank="species">
<emphasis id="B97EA717FFCCFFCEFCE39013FC3AE2B0" bold="true" box="[834,916,1388,1409]" pageId="11" pageNumber="342">sp. nov.</emphasis>
</taxonomicNameLabel>
is primarily old growth with low-lying plants near the karst walls; the type locality of 
<taxonomicName id="4C0A0086FFCCFFCEFDA990F4FD28E291" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[520,646,1418,1440]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="342" phylum="Chordata" rank="species" species="limajalur" status="sp. nov.">
<emphasis id="B97EA717FFCCFFCEFDA990F4FD28E291" box="[520,646,1418,1440]" italics="true" pageId="11" pageNumber="342">C. limajalur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFCCFFCEFD2F90F4FD4FE291" box="[654,737,1419,1440]" pageId="11" pageNumber="342" rank="species">
<emphasis id="B97EA717FFCCFFCEFD2F90F4FD4FE291" bold="true" box="[654,737,1419,1440]" pageId="11" pageNumber="342">sp. nov.</emphasis>
</taxonomicNameLabel>
is on the edge of a local palm oil plantation with some old growth remaining at the base of the karst wall and on the portions of the formation at higher elevation.
</paragraph>
</caption>
<paragraph id="8BB57B05FFCCFFCEFF6690A3FB79E199" blockId="11.[151,1437,1500,2029]" pageId="11" pageNumber="342">
<emphasis id="B97EA717FFCCFFCEFF6690A3FE3CE2C4" bold="true" box="[199,402,1500,1525]" pageId="11" pageNumber="342">Coloration in life</emphasis>
. Dorsum of head, body, limbs, and tail greyish-brown; V-shaped line on rostrum; wide darkbrown nuchal loop that extends to the tip of the snout, edged by white line; five dark-brown bands between nuchal loop and the posterior portion of the limb insertion, edged anteriorly and posteriorly by thin dark-brown lines; body bands at least two times wider than interspaces; front limbs with light-brown blotch pattern; hind limbs with two dark bands separated by white interspaces; ventral portion of body bearing uniform light cream color; tail regenerated bearing four white spots on the anterior half, light grey dorsal and ventral coloration.
</paragraph>
<paragraph id="8BB57B05FFCCFFCEFF6693CBFE80E009" blockId="11.[151,1437,1500,2029]" pageId="11" pageNumber="342">
<emphasis id="B97EA717FFCCFFCEFF6693CBFE9BE1FC" bold="true" box="[199,309,1716,1741]" pageId="11" pageNumber="342">Variation</emphasis>
. The 
<typeStatus id="54B1C5A7FFCCFFCEFEDB93CAFE4DE1FC" box="[378,483,1717,1741]" pageId="11" pageNumber="342" type="paratype">paratypes</typeStatus>
are very similar to the 
<typeStatus id="54B1C5A7FFCCFFCEFD4993CBFCE6E1FC" box="[744,840,1716,1741]" pageId="11" pageNumber="342" type="holotype">holotype</typeStatus>
in coloration and pattern (Table 4). 
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and
</materialsCitation>
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</materialsCitation>
displays reduced dorsal tuberculation, particularly on the anterior half of the body; 
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</materialsCitation>
has one less precloacal pore with 4/3 (L/R), a less well-defined rostral V-pattern, and no chevron posterior to the orbitals (
<figureCitation id="13316780FFCCFFCEFF3E9260FF42E009" box="[159,236,1823,1848]" captionStart="FIGURE 5" captionStartId="15.[151,250,1914,1936]" captionTargetBox="[161,1425,438,1779]" captionTargetId="figure@15.[161,1426,428,1817]" captionTargetPageId="15" captionText="FIGURE 5. Preserved type series of Cyrtodactylus limajalur sp. nov.; A) type series displaying dorsal side of body (from left to right: CAS 262849; CAS 262848; CAS 262847; and CAS 262846); and B) type series displaying ventral side of body (specimens arranged in the same order as figure 5A)." httpUri="https://zenodo.org/record/3242446/files/figure.png" pageId="11" pageNumber="342">Figs. 5</figureCitation>
&amp; 
<figureCitation id="13316780FFCCFFCEFEB19260FE8EE009" box="[272,288,1823,1848]" captionStart="FIGURE 6" captionStartId="16.[151,250,835,857]" captionTargetBox="[151,1436,179,814]" captionTargetId="figure@16.[151,1436,154,814]" captionTargetPageId="16" captionText="FIGURE 6. Photograph of live paratype of Cyrtodactylus limajalur sp. nov. (CAS 262846)." httpUri="https://zenodo.org/record/3242448/files/figure.png" pageId="11" pageNumber="342">6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C310288EFFCCFFCEFF66923BFCCBE0DD" pageId="11" pageNumber="342" type="distribution">
<paragraph id="8BB57B05FFCCFFCEFF66923BFCCBE0DD" blockId="11.[151,1437,1500,2029]" pageId="11" pageNumber="342">
<emphasis id="B97EA717FFCCFFCEFF66923BFEF7E06C" bold="true" box="[199,345,1860,1885]" pageId="11" pageNumber="342">Distribution</emphasis>
. 
<taxonomicName id="4C0A0086FFCCFFCEFECA923BFDD5E06D" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[363,635,1860,1884]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="342" phylum="Chordata" rank="species" species="limajalur" status="sp. nov.">
<emphasis id="B97EA717FFCCFFCEFECA923BFDD5E06D" box="[363,635,1860,1884]" italics="true" pageId="11" pageNumber="342">Cyrtodactylus limajalur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFCCFFCEFD27923BFD4AE06C" box="[646,740,1860,1885]" pageId="11" pageNumber="342" rank="species">
<emphasis id="B97EA717FFCCFFCEFD27923BFD4AE06C" bold="true" box="[646,740,1860,1885]" pageId="11" pageNumber="342">sp. nov.</emphasis>
</taxonomicNameLabel>
is only known from an isolated karst massif in the greater Serian area in southwestern 
<collectingCountry id="F31D3B95FFCCFFCEFE409218FDEBE0B1" box="[481,581,1895,1920]" name="Malaysia" pageId="11" pageNumber="342">Sarawak</collectingCountry>
. The Serian area has a high concentration of large, isolated karst massifs (
<bibRefCitation id="EF9B06F4FFCCFFCEFF3F92F3FEEDE094" author="Wilford, G. E." box="[158,323,1932,1957]" pageId="11" pageNumber="342" refId="ref12262" refString="Wilford, G. E. (1964) The Geology of Sarawak and Sabah Caves. Geological Survey, Borneo Region, 181 pp." type="book" year="1964">Wilford, 1964</bibRefCitation>
). 
<taxonomicName id="4C0A0086FFCCFFCEFEF992F3FDCBE095" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[344,613,1932,1956]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="342" phylum="Chordata" rank="species" species="limajalur" status="sp. nov.">
<emphasis id="B97EA717FFCCFFCEFEF992F3FDCBE095" box="[344,613,1932,1956]" italics="true" pageId="11" pageNumber="342">Cyrtodactylus limajalur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFCCFFCEFDCC92F3FD67E094" box="[621,713,1932,1957]" pageId="11" pageNumber="342" rank="species">
<emphasis id="B97EA717FFCCFFCEFDCC92F3FD67E094" bold="true" box="[621,713,1932,1957]" pageId="11" pageNumber="342">sp. nov.</emphasis>
</taxonomicNameLabel>
was collected from a karst massif approximately one kilometer from a main road. Surrounding formations were not sampled due to inaccessibility, indicating the possibility that this species inhabits other surrounding karst formations (
<figureCitation id="13316780FFCCFFCEFCB292ABFCF6E0DD" box="[787,856,2004,2029]" captionStart="FIGURE 4" captionStartId="11.[151,250,1357,1379]" captionTargetBox="[163,1436,836,1342]" captionTargetId="figure@11.[163,1448,836,1342]" captionTargetPageId="11" captionText="FIGURE 4. A) Mulu karst formation demonstrating Cyrtodactylus muluensis sp. nov. microhabitat; B) type locality of Cyrtodactylus limajalur sp. nov. The type locality of C. muluensis sp. nov. is primarily old growth with low-lying plants near the karst walls; the type locality of C. limajalur sp. nov. is on the edge of a local palm oil plantation with some old growth remaining at the base of the karst wall and on the portions of the formation at higher elevation." httpUri="https://zenodo.org/record/3242444/files/figure.png" pageId="11" pageNumber="342">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C310288EFFC9FFCAFF6695E8FC48E7C6" lastPageId="15" lastPageNumber="346" pageId="14" pageNumber="345" type="etymology">
<paragraph id="8BB57B05FFC9FFCBFF6695E8FCF5E62C" blockId="14.[151,1437,151,648]" pageId="14" pageNumber="345">
<emphasis id="B97EA717FFC9FFCBFF6695E8FEE9E781" bold="true" box="[199,327,151,176]" pageId="14" pageNumber="345">Etymology</emphasis>
. The specific epithet 
<taxonomicName id="4C0A0086FFC9FFCBFDE195E7FD06E781" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[576,680,152,176]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="345" phylum="Chordata" rank="species" species="limajalur">
<emphasis id="B97EA717FFC9FFCBFDE195E7FD06E781" box="[576,680,152,176]" italics="true" pageId="14" pageNumber="345">limajalur</emphasis>
</taxonomicName>
is in reference to the banding pattern of the species. “Lima jalur” loosely translated from Malay is five-banded. 
<taxonomicName id="4C0A0086FFC9FFCBFD0195C3FC03E7E5" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[672,941,188,212]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="345" phylum="Chordata" rank="species" species="limajalur" status="sp. nov.">
<emphasis id="B97EA717FFC9FFCBFD0195C3FC03E7E5" box="[672,941,188,212]" italics="true" pageId="14" pageNumber="345">Cyrtodactylus limajalur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFC9FFCBFC1495C3FBA1E7E4" box="[949,1039,188,213]" pageId="14" pageNumber="345" rank="species">
<emphasis id="B97EA717FFC9FFCBFC1495C3FBA1E7E4" bold="true" box="[949,1039,188,213]" pageId="14" pageNumber="345">sp. nov.</emphasis>
</taxonomicNameLabel>
has a very distinct banding pattern that is consistently comprised of five bands. This species is the first Bornean 
<taxonomicName id="4C0A0086FFC9FFCBFC5F959FFB35E7C9" box="[1022,1179,224,248]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="345" phylum="Chordata" rank="genus">
<emphasis id="B97EA717FFC9FFCBFC5F959FFB35E7C9" box="[1022,1179,224,248]" italics="true" pageId="14" pageNumber="345">Cyrtodactylus</emphasis>
</taxonomicName>
species that displays a consistent banding pattern, without blotches or other markings.
</paragraph>
<paragraph id="8BB57B05FFC9FFCBFF669458FD0CE6E1" blockId="14.[151,1437,151,648]" pageId="14" pageNumber="345">
<emphasis id="B97EA717FFC9FFCBFF669458FED1E671" bold="true" box="[199,383,295,320]" pageId="14" pageNumber="345">Natural history</emphasis>
. All specimens of 
<taxonomicName id="4C0A0086FFC9FFCBFDF69457FCCBE671" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[599,869,296,320]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="345" phylum="Chordata" rank="species" species="limajalur" status="sp. nov.">
<emphasis id="B97EA717FFC9FFCBFDF69457FCCBE671" box="[599,869,296,320]" italics="true" pageId="14" pageNumber="345">Cyrtodactylus limajalur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFC9FFCBFCCC9458FC66E671" box="[877,968,295,320]" pageId="14" pageNumber="345" rank="species">
<emphasis id="B97EA717FFC9FFCBFCCC9458FC66E671" bold="true" box="[877,968,295,320]" pageId="14" pageNumber="345">sp. nov.</emphasis>
</taxonomicNameLabel>
were collected between 19:00 and 22:00 hours on large karst boulders at the base of a large limestone face. No female specimens were gravid. They occur sympatrically with 
<taxonomicName id="4C0A0086FFC9FFCBFEDD940FFD1EE6B9" baseAuthorityName="Peters" baseAuthorityYear="1871" box="[380,688,368,392]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="345" phylum="Chordata" rank="species" species="consobrinus">
<emphasis id="B97EA717FFC9FFCBFEDD940FFD1EE6B9" box="[380,688,368,392]" italics="true" pageId="14" pageNumber="345">Cyrtodactylus consobrinus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0A0086FFC9FFCBFD52940FFC3CE6B9" authorityName="Inger" authorityYear="1958" box="[755,914,368,392]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="345" phylum="Chordata" rank="species" species="pubisulcus">
<emphasis id="B97EA717FFC9FFCBFD52940FFC3CE6B9" box="[755,914,368,392]" italics="true" pageId="14" pageNumber="345">C. pubisulcus</emphasis>
</taxonomicName>
, however, 
<taxonomicName id="4C0A0086FFC9FFCBFBBA940FFB65E6B9" baseAuthorityName="Peters" baseAuthorityYear="1871" box="[1051,1227,368,392]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="345" phylum="Chordata" rank="species" species="consobrinus">
<emphasis id="B97EA717FFC9FFCBFBBA940FFB65E6B9" box="[1051,1227,368,392]" italics="true" pageId="14" pageNumber="345">C. consobrinus</emphasis>
</taxonomicName>
was seen almost exclusively on large tree trunks and 
<taxonomicName id="4C0A0086FFC9FFCBFD9D94EBFD76E69D" authorityName="Inger" authorityYear="1958" box="[572,728,404,428]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="345" phylum="Chordata" rank="species" species="pubisulcus">
<emphasis id="B97EA717FFC9FFCBFD9D94EBFD76E69D" box="[572,728,404,428]" italics="true" pageId="14" pageNumber="345">C. pubisulcus</emphasis>
</taxonomicName>
was only found on low-lying vegetation, indicating that they may not be syntopic with 
<taxonomicName id="4C0A0086FFC9FFCBFE1894C7FDEFE6E1" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[441,577,440,464]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="345" phylum="Chordata" rank="species" species="limajalur" status="sp. nov.">
<emphasis id="B97EA717FFC9FFCBFE1894C7FDEFE6E1" box="[441,577,440,464]" italics="true" pageId="14" pageNumber="345">C. limajalur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFC9FFCBFDE894C8FD0CE6E1" box="[585,674,439,464]" pageId="14" pageNumber="345" rank="species">
<emphasis id="B97EA717FFC9FFCBFDE894C8FD0CE6E1" bold="true" box="[585,674,439,464]" pageId="14" pageNumber="345">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BB57B05FFC9FFCBFF6694A3FEF9E50E" blockId="14.[151,1437,151,648]" pageId="14" pageNumber="345">
There were no noticeable cave entrances on the karst massif. All specimens were collected within approximately ~ 
<quantity id="4CF2D6E0FFC9FFCBFEF1977DFED4E52B" box="[336,378,514,539]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="345" unit="m" value="5.0">5 m</quantity>
<superScript id="7C7FD64DFFC9FFCBFEDA9480FE2DE53C" attach="left" box="[379,387,511,525]" fontSize="6" pageId="14" pageNumber="345">2</superScript>
of each other, however, there did not appear to be anything different or significant about that specific location.
</paragraph>
<paragraph id="8BB57B05FFC9FFCBFF669735FAB7E5B6" blockId="14.[151,1437,151,648]" pageId="14" pageNumber="345">
All of the specimens comprising the 
<typeStatus id="54B1C5A7FFC9FFCBFD279734FD18E552" box="[646,694,587,611]" pageId="14" pageNumber="345">type</typeStatus>
series had regenerated tails. However, the tails were all fully regenerated indicating that the species may experience higher rates of predation at earlier stages in life.
</paragraph>
<caption id="DF752B8DFFC9FFCBFF3697C9FA88E5DC" ID-Table-UUID="DF752B8DFFC9FFCBFF3697C9FA88E5DC" httpUri="http://table.plazi.org/id/DF752B8DFFC9FFCBFF3697C9FA88E5DC" pageId="14" pageNumber="345" targetBox="[159,1303,777,1574]" targetIsTable="true" targetPageId="14">
<paragraph id="8BB57B05FFC9FFCBFF3697C9FA88E5DC" blockId="14.[151,1436,693,749]" pageId="14" pageNumber="345">
<emphasis id="B97EA717FFC9FFCBFF3697C9FEA3E5FC" bold="true" box="[151,269,694,717]" pageId="14" pageNumber="345">TABLE 5.</emphasis>
Meristic, color, and measurement data of the type series for 
<taxonomicName id="4C0A0086FFC9FFCBFC0B97CAFB05E5FC" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[938,1195,693,717]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="345" phylum="Chordata" rank="species" species="limajalur" status="sp. nov.">
<emphasis id="B97EA717FFC9FFCBFC0B97CAFB05E5FC" box="[938,1195,693,717]" italics="true" pageId="14" pageNumber="345">Cyrtodactylus limajalur</emphasis>
</taxonomicName>
<emphasis id="B97EA717FFC9FFCBFB1597C9FAA2E5FC" bold="true" box="[1204,1292,694,717]" pageId="14" pageNumber="345">
<taxonomicNameLabel id="A24D1A6CFFC9FFCBFB1597C9FAA2E5FC" box="[1204,1292,694,717]" pageId="14" pageNumber="345" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from Serian, Sarawak, Malaysia. Key to abbreviations in Materials and Methods. ‘R’ indicates regeneration in the TL row.
</paragraph>
</caption>
<paragraph id="8BB57B05FFC9FFCBFF3E9676FB14E117" pageId="14" pageNumber="345">
<table id="F90A89A5FFC9003AFF3E9676FAB9E117" box="[159,1303,777,1574]" gridcols="5" gridrows="19" pageId="14" pageNumber="345">
<tr id="353A7947FFC9003AFF3E9676FAB9E40F" box="[159,1303,777,830]" gridrow="0" pageId="14" pageNumber="345">
<th id="76EB103BFFC9003AFF3E9676FE29E40F" box="[159,391,777,830]" gridcol="0" gridrow="0" pageId="14" pageNumber="345">Character</th>
<th id="76EB103BFFC9003AFE4B9676FDC6E40F" box="[490,616,777,830]" gridcol="1" gridrow="0" pageId="14" pageNumber="345">CAS262848 Holotype</th>
<th id="76EB103BFFC9003AFD4A9676FCC6E40F" box="[747,872,777,830]" gridcol="2" gridrow="0" pageId="14" pageNumber="345">CAS262849 Paratype</th>
<th id="76EB103BFFC9003AFC6F9676FBE5E40F" box="[974,1099,777,830]" gridcol="3" gridrow="0" pageId="14" pageNumber="345">CAS262846 Paratype</th>
<th id="76EB103BFFC9003AFB3B9676FAB9E40F" box="[1178,1303,777,830]" gridcol="4" gridrow="0" pageId="14" pageNumber="345">CAS262847 Paratype</th>
</tr>
<tr id="353A7947FFC9003AFF3E962EFAB9E456" box="[159,1303,849,871]" gridrow="1" pageId="14" pageNumber="345">
<th id="76EB103BFFC9003AFF3E962EFE29E456" box="[159,391,849,871]" gridcol="0" gridrow="1" pageId="14" pageNumber="345">Sex</th>
<td id="76EB103BFFC9003AFE4B962EFDC6E456" box="[490,616,849,871]" gridcol="1" gridrow="1" pageId="14" pageNumber="345">M</td>
<td id="76EB103BFFC9003AFD4A962EFCC6E456" box="[747,872,849,871]" gridcol="2" gridrow="1" pageId="14" pageNumber="345">M</td>
<td id="76EB103BFFC9003AFC6F962EFBE5E456" box="[974,1099,849,871]" gridcol="3" gridrow="1" pageId="14" pageNumber="345">F</td>
<td id="76EB103BFFC9003AFB3B962EFAB9E456" box="[1178,1303,849,871]" gridcol="4" gridrow="1" pageId="14" pageNumber="345">F</td>
</tr>
<tr id="353A7947FFC9003AFF3E9604FAB9E4A0" box="[159,1303,891,913]" gridrow="2" pageId="14" pageNumber="345">
<th id="76EB103BFFC9003AFF3E9604FE29E4A0" box="[159,391,891,913]" gridcol="0" gridrow="2" pageId="14" pageNumber="345">Supralabials</th>
<td id="76EB103BFFC9003AFE4B9604FDC6E4A0" box="[490,616,891,913]" gridcol="1" gridrow="2" pageId="14" pageNumber="345">12</td>
<td id="76EB103BFFC9003AFD4A9604FCC6E4A0" box="[747,872,891,913]" gridcol="2" gridrow="2" pageId="14" pageNumber="345">12</td>
<td id="76EB103BFFC9003AFC6F9604FBE5E4A0" box="[974,1099,891,913]" gridcol="3" gridrow="2" pageId="14" pageNumber="345">13</td>
<td id="76EB103BFFC9003AFB3B9604FAB9E4A0" box="[1178,1303,891,913]" gridcol="4" gridrow="2" pageId="14" pageNumber="345">12</td>
</tr>
<tr id="353A7947FFC9003AFF3E96DBFAB9E48B" box="[159,1303,932,954]" gridrow="3" pageId="14" pageNumber="345">
<th id="76EB103BFFC9003AFF3E96DBFE29E48B" box="[159,391,932,954]" gridcol="0" gridrow="3" pageId="14" pageNumber="345">Infralabials</th>
<td id="76EB103BFFC9003AFE4B96DBFDC6E48B" box="[490,616,932,954]" gridcol="1" gridrow="3" pageId="14" pageNumber="345">9</td>
<td id="76EB103BFFC9003AFD4A96DBFCC6E48B" box="[747,872,932,954]" gridcol="2" gridrow="3" pageId="14" pageNumber="345">9</td>
<td id="76EB103BFFC9003AFC6F96DBFBE5E48B" box="[974,1099,932,954]" gridcol="3" gridrow="3" pageId="14" pageNumber="345">9</td>
<td id="76EB103BFFC9003AFB3B96DBFAB9E48B" box="[1178,1303,932,954]" gridcol="4" gridrow="3" pageId="14" pageNumber="345">10</td>
</tr>
<tr id="353A7947FFC9003AFF3E96B2FAB9E4D2" box="[159,1303,973,995]" gridrow="4" pageId="14" pageNumber="345">
<th id="76EB103BFFC9003AFF3E96B2FE29E4D2" box="[159,391,973,995]" gridcol="0" gridrow="4" pageId="14" pageNumber="345">Paravertebral tubercles</th>
<td id="76EB103BFFC9003AFE4B96B2FDC6E4D2" box="[490,616,973,995]" gridcol="1" gridrow="4" pageId="14" pageNumber="345">38</td>
<td id="76EB103BFFC9003AFD4A96B2FCC6E4D2" box="[747,872,973,995]" gridcol="2" gridrow="4" pageId="14" pageNumber="345">34</td>
<td id="76EB103BFFC9003AFC6F96B2FBE5E4D2" box="[974,1099,973,995]" gridcol="3" gridrow="4" pageId="14" pageNumber="345">38</td>
<td id="76EB103BFFC9003AFB3B96B2FAB9E4D2" box="[1178,1303,973,995]" gridcol="4" gridrow="4" pageId="14" pageNumber="345">36</td>
</tr>
<tr id="353A7947FFC9003AFF3E9689FAB9E33D" box="[159,1303,1014,1036]" gridrow="5" pageId="14" pageNumber="345">
<th id="76EB103BFFC9003AFF3E9689FE29E33D" box="[159,391,1014,1036]" gridcol="0" gridrow="5" pageId="14" pageNumber="345">Ventral Scales</th>
<td id="76EB103BFFC9003AFE4B9689FDC6E33D" box="[490,616,1014,1036]" gridcol="1" gridrow="5" pageId="14" pageNumber="345">42</td>
<td id="76EB103BFFC9003AFD4A9689FCC6E33D" box="[747,872,1014,1036]" gridcol="2" gridrow="5" pageId="14" pageNumber="345">33</td>
<td id="76EB103BFFC9003AFC6F9689FBE5E33D" box="[974,1099,1014,1036]" gridcol="3" gridrow="5" pageId="14" pageNumber="345">35</td>
<td id="76EB103BFFC9003AFB3B9689FAB9E33D" box="[1178,1303,1014,1036]" gridcol="4" gridrow="5" pageId="14" pageNumber="345">42</td>
</tr>
<tr id="353A7947FFC9003AFF3E915FFAB9E307" box="[159,1303,1056,1078]" gridrow="6" pageId="14" pageNumber="345">
<th id="76EB103BFFC9003AFF3E915FFE29E307" box="[159,391,1056,1078]" gridcol="0" gridrow="6" pageId="14" pageNumber="345">Lamellae on 4th toe</th>
<td id="76EB103BFFC9003AFE4B915FFDC6E307" box="[490,616,1056,1078]" gridcol="1" gridrow="6" pageId="14" pageNumber="345">19</td>
<td id="76EB103BFFC9003AFD4A915FFCC6E307" box="[747,872,1056,1078]" gridcol="2" gridrow="6" pageId="14" pageNumber="345">20</td>
<td id="76EB103BFFC9003AFC6F915FFBE5E307" box="[974,1099,1056,1078]" gridcol="3" gridrow="6" pageId="14" pageNumber="345">22</td>
<td id="76EB103BFFC9003AFB3B915FFAB9E307" box="[1178,1303,1056,1078]" gridcol="4" gridrow="6" pageId="14" pageNumber="345">22</td>
</tr>
<tr id="353A7947FFC9003AFF3E9136FAB9E36E" box="[159,1303,1097,1119]" gridrow="7" pageId="14" pageNumber="345">
<th id="76EB103BFFC9003AFF3E9136FE29E36E" box="[159,391,1097,1119]" gridcol="0" gridrow="7" pageId="14" pageNumber="345">Body Bands</th>
<td id="76EB103BFFC9003AFE4B9136FDC6E36E" box="[490,616,1097,1119]" gridcol="1" gridrow="7" pageId="14" pageNumber="345">5</td>
<td id="76EB103BFFC9003AFD4A9136FCC6E36E" box="[747,872,1097,1119]" gridcol="2" gridrow="7" pageId="14" pageNumber="345">5</td>
<td id="76EB103BFFC9003AFC6F9136FBE5E36E" box="[974,1099,1097,1119]" gridcol="3" gridrow="7" pageId="14" pageNumber="345">5</td>
<td id="76EB103BFFC9003AFB3B9136FAB9E36E" box="[1178,1303,1097,1119]" gridcol="4" gridrow="7" pageId="14" pageNumber="345">5</td>
</tr>
<tr id="353A7947FFC9003AFF3E910CFAB9E3B8" box="[159,1303,1139,1161]" gridrow="8" pageId="14" pageNumber="345">
<th id="76EB103BFFC9003AFF3E910CFE29E3B8" box="[159,391,1139,1161]" gridcol="0" gridrow="8" pageId="14" pageNumber="345">SVL</th>
<td id="76EB103BFFC9003AFE4B910CFDC6E3B8" box="[490,616,1139,1161]" gridcol="1" gridrow="8" pageId="14" pageNumber="345">90.8</td>
<td id="76EB103BFFC9003AFD4A910CFCC6E3B8" box="[747,872,1139,1161]" gridcol="2" gridrow="8" pageId="14" pageNumber="345">94.3</td>
<td id="76EB103BFFC9003AFC6F910CFBE5E3B8" box="[974,1099,1139,1161]" gridcol="3" gridrow="8" pageId="14" pageNumber="345">72.5</td>
<td id="76EB103BFFC9003AFB3B910CFAB9E3B8" box="[1178,1303,1139,1161]" gridcol="4" gridrow="8" pageId="14" pageNumber="345">89.0</td>
</tr>
<tr id="353A7947FFC9003AFF3E91E3FAB9E383" box="[159,1303,1180,1202]" gridrow="9" pageId="14" pageNumber="345">
<th id="76EB103BFFC9003AFF3E91E3FE29E383" box="[159,391,1180,1202]" gridcol="0" gridrow="9" pageId="14" pageNumber="345">TL</th>
<td id="76EB103BFFC9003AFE4B91E3FDC6E383" box="[490,616,1180,1202]" gridcol="1" gridrow="9" pageId="14" pageNumber="345">114R</td>
<td id="76EB103BFFC9003AFD4A91E3FCC6E383" box="[747,872,1180,1202]" gridcol="2" gridrow="9" pageId="14" pageNumber="345">78.3R</td>
<td id="76EB103BFFC9003AFC6F91E3FBE5E383" box="[974,1099,1180,1202]" gridcol="3" gridrow="9" pageId="14" pageNumber="345">99.4</td>
<td id="76EB103BFFC9003AFB3B91E3FAB9E383" box="[1178,1303,1180,1202]" gridcol="4" gridrow="9" pageId="14" pageNumber="345">112.8R</td>
</tr>
<tr id="353A7947FFC9003AFF3E91BAFAB9E3EA" box="[159,1303,1221,1243]" gridrow="10" pageId="14" pageNumber="345">
<th id="76EB103BFFC9003AFF3E91BAFE29E3EA" box="[159,391,1221,1243]" gridcol="0" gridrow="10" pageId="14" pageNumber="345">TW</th>
<td id="76EB103BFFC9003AFE4B91BAFDC6E3EA" box="[490,616,1221,1243]" gridcol="1" gridrow="10" pageId="14" pageNumber="345">6.6</td>
<td id="76EB103BFFC9003AFD4A91BAFCC6E3EA" box="[747,872,1221,1243]" gridcol="2" gridrow="10" pageId="14" pageNumber="345">12.1</td>
<td id="76EB103BFFC9003AFC6F91BAFBE5E3EA" box="[974,1099,1221,1243]" gridcol="3" gridrow="10" pageId="14" pageNumber="345">5</td>
<td id="76EB103BFFC9003AFB3B91BAFAB9E3EA" box="[1178,1303,1221,1243]" gridcol="4" gridrow="10" pageId="14" pageNumber="345">5.9</td>
</tr>
<tr id="353A7947FFC9003AFF3E9191FAB9E235" box="[159,1303,1262,1284]" gridrow="11" pageId="14" pageNumber="345">
<th id="76EB103BFFC9003AFF3E9191FE29E235" box="[159,391,1262,1284]" gridcol="0" gridrow="11" pageId="14" pageNumber="345">HL</th>
<td id="76EB103BFFC9003AFE4B9191FDC6E235" box="[490,616,1262,1284]" gridcol="1" gridrow="11" pageId="14" pageNumber="345">25.2</td>
<td id="76EB103BFFC9003AFD4A9191FCC6E235" box="[747,872,1262,1284]" gridcol="2" gridrow="11" pageId="14" pageNumber="345">26.9</td>
<td id="76EB103BFFC9003AFC6F9191FBE5E235" box="[974,1099,1262,1284]" gridcol="3" gridrow="11" pageId="14" pageNumber="345">21.3</td>
<td id="76EB103BFFC9003AFB3B9191FAB9E235" box="[1178,1303,1262,1284]" gridcol="4" gridrow="11" pageId="14" pageNumber="345">25.0</td>
</tr>
<tr id="353A7947FFC9003AFF3E9067FAB9E21F" box="[159,1303,1304,1326]" gridrow="12" pageId="14" pageNumber="345">
<th id="76EB103BFFC9003AFF3E9067FE29E21F" box="[159,391,1304,1326]" gridcol="0" gridrow="12" pageId="14" pageNumber="345">HW</th>
<td id="76EB103BFFC9003AFE4B9067FDC6E21F" box="[490,616,1304,1326]" gridcol="1" gridrow="12" pageId="14" pageNumber="345">11.7</td>
<td id="76EB103BFFC9003AFD4A9067FCC6E21F" box="[747,872,1304,1326]" gridcol="2" gridrow="12" pageId="14" pageNumber="345">16.1</td>
<td id="76EB103BFFC9003AFC6F9067FBE5E21F" box="[974,1099,1304,1326]" gridcol="3" gridrow="12" pageId="14" pageNumber="345">13.6</td>
<td id="76EB103BFFC9003AFB3B9067FAB9E21F" box="[1178,1303,1304,1326]" gridcol="4" gridrow="12" pageId="14" pageNumber="345">15.5</td>
</tr>
<tr id="353A7947FFC9003AFF3E903EFAB9E266" box="[159,1303,1345,1367]" gridrow="13" pageId="14" pageNumber="345">
<th id="76EB103BFFC9003AFF3E903EFE29E266" box="[159,391,1345,1367]" gridcol="0" gridrow="13" pageId="14" pageNumber="345">HD</th>
<td id="76EB103BFFC9003AFE4B903EFDC6E266" box="[490,616,1345,1367]" gridcol="1" gridrow="13" pageId="14" pageNumber="345">9.0</td>
<td id="76EB103BFFC9003AFD4A903EFCC6E266" box="[747,872,1345,1367]" gridcol="2" gridrow="13" pageId="14" pageNumber="345">9.5</td>
<td id="76EB103BFFC9003AFC6F903EFBE5E266" box="[974,1099,1345,1367]" gridcol="3" gridrow="13" pageId="14" pageNumber="345">7.6</td>
<td id="76EB103BFFC9003AFB3B903EFAB9E266" box="[1178,1303,1345,1367]" gridcol="4" gridrow="13" pageId="14" pageNumber="345">8.9</td>
</tr>
<tr id="353A7947FFC9003AFF3E9014FAB9E2B0" box="[159,1303,1387,1409]" gridrow="14" pageId="14" pageNumber="345">
<th id="76EB103BFFC9003AFF3E9014FE29E2B0" box="[159,391,1387,1409]" gridcol="0" gridrow="14" pageId="14" pageNumber="345">EE</th>
<td id="76EB103BFFC9003AFE4B9014FDC6E2B0" box="[490,616,1387,1409]" gridcol="1" gridrow="14" pageId="14" pageNumber="345">5.9</td>
<td id="76EB103BFFC9003AFD4A9014FCC6E2B0" box="[747,872,1387,1409]" gridcol="2" gridrow="14" pageId="14" pageNumber="345">6.2</td>
<td id="76EB103BFFC9003AFC6F9014FBE5E2B0" box="[974,1099,1387,1409]" gridcol="3" gridrow="14" pageId="14" pageNumber="345">6.0</td>
<td id="76EB103BFFC9003AFB3B9014FAB9E2B0" box="[1178,1303,1387,1409]" gridcol="4" gridrow="14" pageId="14" pageNumber="345">12.0</td>
</tr>
<tr id="353A7947FFC9003AFF3E90EBFAB9E29B" box="[159,1303,1428,1450]" gridrow="15" pageId="14" pageNumber="345">
<th id="76EB103BFFC9003AFF3E90EBFE29E29B" box="[159,391,1428,1450]" gridcol="0" gridrow="15" pageId="14" pageNumber="345">ES</th>
<td id="76EB103BFFC9003AFE4B90EBFDC6E29B" box="[490,616,1428,1450]" gridcol="1" gridrow="15" pageId="14" pageNumber="345">11.0</td>
<td id="76EB103BFFC9003AFD4A90EBFCC6E29B" box="[747,872,1428,1450]" gridcol="2" gridrow="15" pageId="14" pageNumber="345">11.2</td>
<td id="76EB103BFFC9003AFC6F90EBFBE5E29B" box="[974,1099,1428,1450]" gridcol="3" gridrow="15" pageId="14" pageNumber="345">9.6</td>
<td id="76EB103BFFC9003AFB3B90EBFAB9E29B" box="[1178,1303,1428,1450]" gridcol="4" gridrow="15" pageId="14" pageNumber="345">11.0</td>
</tr>
<tr id="353A7947FFC9003AFF3E90C2FAB9E2E2" box="[159,1303,1469,1491]" gridrow="16" pageId="14" pageNumber="345">
<th id="76EB103BFFC9003AFF3E90C2FE29E2E2" box="[159,391,1469,1491]" gridcol="0" gridrow="16" pageId="14" pageNumber="345">EN</th>
<td id="76EB103BFFC9003AFE4B90C2FDC6E2E2" box="[490,616,1469,1491]" gridcol="1" gridrow="16" pageId="14" pageNumber="345">8.5</td>
<td id="76EB103BFFC9003AFD4A90C2FCC6E2E2" box="[747,872,1469,1491]" gridcol="2" gridrow="16" pageId="14" pageNumber="345">9.0</td>
<td id="76EB103BFFC9003AFC6F90C2FBE5E2E2" box="[974,1099,1469,1491]" gridcol="3" gridrow="16" pageId="14" pageNumber="345">8.1</td>
<td id="76EB103BFFC9003AFB3B90C2FAB9E2E2" box="[1178,1303,1469,1491]" gridcol="4" gridrow="16" pageId="14" pageNumber="345">8.4</td>
</tr>
<tr id="353A7947FFC9003AFF3E9099FAB9E2CD" box="[159,1303,1510,1532]" gridrow="17" pageId="14" pageNumber="345">
<th id="76EB103BFFC9003AFF3E9099FE29E2CD" box="[159,391,1510,1532]" gridcol="0" gridrow="17" pageId="14" pageNumber="345">EL</th>
<td id="76EB103BFFC9003AFE4B9099FDC6E2CD" box="[490,616,1510,1532]" gridcol="1" gridrow="17" pageId="14" pageNumber="345">2.6</td>
<td id="76EB103BFFC9003AFD4A9099FCC6E2CD" box="[747,872,1510,1532]" gridcol="2" gridrow="17" pageId="14" pageNumber="345">2.4</td>
<td id="76EB103BFFC9003AFC6F9099FBE5E2CD" box="[974,1099,1510,1532]" gridcol="3" gridrow="17" pageId="14" pageNumber="345">2.1</td>
<td id="76EB103BFFC9003AFB3B9099FAB9E2CD" box="[1178,1303,1510,1532]" gridcol="4" gridrow="17" pageId="14" pageNumber="345">1.8</td>
</tr>
<tr id="353A7947FFC9003AFF3E936FFAB9E117" box="[159,1303,1552,1574]" gridrow="18" pageId="14" pageNumber="345">
<th id="76EB103BFFC9003AFF3E936FFE29E117" box="[159,391,1552,1574]" gridcol="0" gridrow="18" pageId="14" pageNumber="345">IO</th>
<td id="76EB103BFFC9003AFE4B936FFDC6E117" box="[490,616,1552,1574]" gridcol="1" gridrow="18" pageId="14" pageNumber="345">2.9</td>
<td id="76EB103BFFC9003AFD4A936FFCC6E117" box="[747,872,1552,1574]" gridcol="2" gridrow="18" pageId="14" pageNumber="345">3.2</td>
<td id="76EB103BFFC9003AFC6F936FFBE5E117" box="[974,1099,1552,1574]" gridcol="3" gridrow="18" pageId="14" pageNumber="345">2.9</td>
<td id="76EB103BFFC9003AFB3B936FFAB9E117" box="[1178,1303,1552,1574]" gridcol="4" gridrow="18" pageId="14" pageNumber="345">3.0</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB57B05FFC9FFCAFF669315FC48E7C6" blockId="14.[151,1436,1642,2027]" lastBlockId="15.[151,1436,151,248]" lastPageId="15" lastPageNumber="346" pageId="14" pageNumber="345">
<emphasis id="B97EA717FFC9FFCBFF669315FEF4E1B2" bold="true" box="[199,346,1642,1667]" pageId="14" pageNumber="345">Comparison</emphasis>
. 
<taxonomicName id="4C0A0086FFC9FFCBFED19314FD2DE1B2" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[368,643,1643,1667]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="345" phylum="Chordata" rank="species" species="limajalur" status="sp. nov.">
<emphasis id="B97EA717FFC9FFCBFED19314FD2DE1B2" box="[368,643,1643,1667]" italics="true" pageId="14" pageNumber="345">Cyrtodactylus limajalur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFC9FFCBFD339315FD5CE1B2" box="[658,754,1642,1667]" pageId="14" pageNumber="345" rank="species">
<emphasis id="B97EA717FFC9FFCBFD339315FD5CE1B2" bold="true" box="[658,754,1642,1667]" pageId="14" pageNumber="345">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from all its Bornean congeners by one or more morphological characteristics. The new species is distinguished from 
<taxonomicName id="4C0A0086FFC9FFCBFC1A93F0FBE7E196" baseAuthorityName="Mocquard" baseAuthorityYear="1890" box="[955,1097,1679,1703]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="345" phylum="Chordata" rank="species" species="baluensis">
<emphasis id="B97EA717FFC9FFCBFC1A93F0FBE7E196" box="[955,1097,1679,1703]" italics="true" pageId="14" pageNumber="345">C. baluensis</emphasis>
</taxonomicName>
by having a distinct banding pattern as opposed to bands and blotches; it is distinguished from 
<taxonomicName id="4C0A0086FFC9FFCBFC3093CCFBEDE1FA" box="[913,1091,1715,1739]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="345" phylum="Chordata" rank="species" species="cavernicolus">
<emphasis id="B97EA717FFC9FFCBFC3093CCFBEDE1FA" box="[913,1091,1715,1739]" italics="true" pageId="14" pageNumber="345">C. cavernicolus</emphasis>
</taxonomicName>
by having a lower number of ventral scales (33–42 
<emphasis id="B97EA717FFC9FFCBFE2A93A7FE7DE1DE" box="[395,467,1752,1775]" italics="true" pageId="14" pageNumber="345">versus</emphasis>
51–58), enlarged subcaudals, and a precloacal pit as opposed to a precloacal groove; it is distinguished from 
<taxonomicName id="4C0A0086FFC9FFCBFE2E9384FD94E022" baseAuthorityName="Peters" baseAuthorityYear="1871" box="[399,570,1787,1811]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="345" phylum="Chordata" rank="species" species="consobrinus">
<emphasis id="B97EA717FFC9FFCBFE2E9384FD94E022" box="[399,570,1787,1811]" italics="true" pageId="14" pageNumber="345">C. consobrinus</emphasis>
</taxonomicName>
by having a smaller adult maximum SVL (
<quantity id="4CF2D6E0FFC9FFCBFB8C9384FBD1E023" box="[1069,1151,1787,1811]" metricMagnitude="-2" metricUnit="m" metricValue="9.4" pageId="14" pageNumber="345" unit="mm" value="94.0">94 mm</quantity>
<emphasis id="B97EA717FFC9FFCBFB269384FB61E023" box="[1159,1231,1787,1810]" italics="true" pageId="14" pageNumber="345">versus</emphasis>
<quantity id="4CF2D6E0FFC9FFCBFB769385FA94E022" box="[1239,1338,1786,1811]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" pageId="14" pageNumber="345" unit="mm" value="125.0">125 mm</quantity>
), and no white reticulated pattern on occiput; it is distinguished from 
<taxonomicName id="4C0A0086FFC9FFCBFCE59260FC07E006" authorityName="Hikida" authorityYear="1990" box="[836,937,1823,1847]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="345" phylum="Chordata" rank="species" species="ingeri">
<emphasis id="B97EA717FFC9FFCBFCE59260FC07E006" box="[836,937,1823,1847]" italics="true" pageId="14" pageNumber="345">C. ingeri</emphasis>
</taxonomicName>
by having a precloacal pit as opposed to no depression, 5–6 enlarged femoral scales as opposed to none, and a distinct banding pattern as opposed to bands and blotches; it is distinguished from 
<taxonomicName id="4C0A0086FFC9FFCBFDB49218FD1EE04E" box="[533,688,1895,1919]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="345" phylum="Chordata" rank="species" species="malayanus">
<emphasis id="B97EA717FFC9FFCBFDB49218FD1EE04E" box="[533,688,1895,1919]" italics="true" pageId="14" pageNumber="345">C. malayanus</emphasis>
</taxonomicName>
by having no tubercles on the upper arm, and a lower number of ventral scales (33–42 
<emphasis id="B97EA717FFC9FFCBFE3D92F4FE4AE093" box="[412,484,1931,1954]" italics="true" pageId="14" pageNumber="345">versus</emphasis>
58–62); it is distinguished from 
<taxonomicName id="4C0A0086FFC9FFCBFCD392F4FC45E093" authorityName="Hikida" authorityYear="1990" box="[882,1003,1931,1954]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="345" phylum="Chordata" rank="species" species="matsuii">
<emphasis id="B97EA717FFC9FFCBFCD392F4FC45E093" box="[882,1003,1931,1954]" italics="true" pageId="14" pageNumber="345">C. matsuii</emphasis>
</taxonomicName>
by having enlarged subcaudals, 5–6 enlarged femoral scales as opposed to none, and a smaller maximum SVL (
<quantity id="4CF2D6E0FFC9FFCBFB9092D0FB26E0F6" box="[1073,1160,1967,1991]" metricMagnitude="-2" metricUnit="m" metricValue="9.4" pageId="14" pageNumber="345" unit="mm" value="94.0">94 mm</quantity>
<emphasis id="B97EA717FFC9FFCBFB3492CFFB73E0F6" box="[1173,1245,1968,1991]" italics="true" pageId="14" pageNumber="345">versus</emphasis>
<quantity id="4CF2D6E0FFC9FFCBFB4B92D1FAFCE0F6" box="[1258,1362,1966,1991]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" pageId="14" pageNumber="345" unit="mm" value="105.0">105 mm</quantity>
); it is distinguished from 
<taxonomicName id="4C0A0086FFC9FFCBFED392ACFDA5E0DA" authorityName="Inger" authorityYear="1958" box="[370,523,2003,2027]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="345" phylum="Chordata" rank="species" species="pubisulcus">
<emphasis id="B97EA717FFC9FFCBFED392ACFDA5E0DA" box="[370,523,2003,2027]" italics="true" pageId="14" pageNumber="345">C. pubisulcus</emphasis>
</taxonomicName>
by having enlarged subcaudals, 5–6 enlarged femoral scales as opposed to none, and a precloacal pit as opposed to a precloacal groove; and it is distinguished from 
<taxonomicName id="4C0A0086FFC8FFCAFB9495E7FB36E781" authorityName="Hikida" authorityYear="1990" box="[1077,1176,152,176]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="346" phylum="Chordata" rank="species" species="yoshii">
<emphasis id="B97EA717FFC8FFCAFB9495E7FB36E781" box="[1077,1176,152,176]" italics="true" pageId="15" pageNumber="346">C. yoshii</emphasis>
</taxonomicName>
by having no tubercles on the upper arm, a lower number of ventral scales (33–42 
<emphasis id="B97EA717FFC8FFCAFC9C95C2FC2AE7E5" box="[829,900,189,212]" italics="true" pageId="15" pageNumber="346">versus</emphasis>
50–58), a lower number of subdigital lamellae (19–22 
<emphasis id="B97EA717FFC8FFCAFF4C959EFE9AE7C9" box="[237,308,225,248]" italics="true" pageId="15" pageNumber="346">versus</emphasis>
25–30), and 5–6 enlarged femoral scales as opposed to none.
</paragraph>
</subSubSection>
<caption id="DF752B8DFFC8FFCAFF369205FD01E0FC" httpUri="https://zenodo.org/record/3242446/files/figure.png" pageId="15" pageNumber="346" startId="15.[151,250,1914,1936]" targetBox="[161,1425,438,1779]" targetPageId="15">
<paragraph id="8BB57B05FFC8FFCAFF369205FD01E0FC" blockId="15.[151,1436,1914,1997]" pageId="15" pageNumber="346">
<emphasis id="B97EA717FFC8FFCAFF369205FEA3E0BE" bold="true" box="[151,269,1914,1936]" pageId="15" pageNumber="346">FIGURE 5</emphasis>
. Preserved type series of 
<taxonomicName id="4C0A0086FFC8FFCAFDB89205FCA4E0A1" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[537,778,1914,1936]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="346" phylum="Chordata" rank="species" species="limajalur" status="sp. nov.">Cyrtodactylus limajalur</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFC8FFCAFCB19204FCCCE0A1" box="[784,866,1915,1936]" pageId="15" pageNumber="346" rank="species">sp. nov.</taxonomicNameLabel>
; A) type series displaying dorsal side of body (from left to right: CAS 262849; CAS 262848; CAS 262847; and CAS 262846); and B) type series displaying ventral side of body (specimens arranged in the same order as figure 5A).
</paragraph>
</caption>
<caption id="DF752B8DFFD7FFD5FF36963CFBEEE468" box="[151,1088,835,857]" httpUri="https://zenodo.org/record/3242448/files/figure.png" pageId="16" pageNumber="347" startId="16.[151,250,835,857]" targetBox="[151,1436,179,814]" targetPageId="16">
<paragraph id="8BB57B05FFD7FFD5FF36963CFBEEE468" blockId="16.[151,1088,835,857]" box="[151,1088,835,857]" pageId="16" pageNumber="347">
<emphasis id="B97EA717FFD7FFD5FF36963CFEA3E469" bold="true" box="[151,269,835,857]" pageId="16" pageNumber="347">FIGURE 6</emphasis>
. Photograph of live paratype of 
<taxonomicName id="4C0A0086FFD7FFD5FDFA963CFCE5E468" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[603,843,835,857]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" species="limajalur" status="sp. nov.">Cyrtodactylus limajalur</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFD7FFD5FCF0963BFC0EE468" box="[849,928,836,857]" pageId="16" pageNumber="347" rank="species">sp. nov.</taxonomicNameLabel>
(CAS 262846).
</paragraph>
</caption>
<subSubSection id="C310288EFFD7FFD5FF6696FAFF41E15F" pageId="16" pageNumber="347" type="description">
<paragraph id="8BB57B05FFD7FFD5FF6696FAFD86E28B" blockId="16.[151,1436,901,1646]" pageId="16" pageNumber="347">
<emphasis id="B97EA717FFD7FFD5FF6696FAFE7FE4AF" bold="true" box="[199,465,901,926]" pageId="16" pageNumber="347">Phylogenetic analyses.</emphasis>
The ML tree shown herein corroborates the morphological data distinguishing the two lineages as new species (
<figureCitation id="13316780FFD7FFD5FE1296D6FE57E4F3" box="[435,505,937,962]" captionStart="FIGURE 7" captionStartId="17.[151,250,1900,1922]" captionTargetBox="[289,1242,439,1872]" captionTargetId="figure@17.[284,1302,439,1875]" captionTargetPageId="17" captionText="FIGURE 7. ML tree of Bornean Cyrtodactylus and their closely related congeners. Phylogenetic position of C. limajalur sp. nov. and C. muluensis sp. nov. is indicated by the purple and green boxes, respectively. Black circles at the node denote UF values of 95 or greater and posterior probabilities of 0.95 or greater. White circles at the node denote nodes at which either the UF or posterior probability demonstrated support, but not both." httpUri="https://zenodo.org/record/3242450/files/figure.png" pageId="16" pageNumber="347">Fig. 7</figureCitation>
). The ML analysis places both 
<taxonomicName id="4C0A0086FFD7FFD5FCC096D5FC42E4F3" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[865,1004,938,962]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" species="limajalur" status="sp. nov.">
<emphasis id="B97EA717FFD7FFD5FCC096D5FC42E4F3" box="[865,1004,938,962]" italics="true" pageId="16" pageNumber="347">C. limajalur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFD7FFD5FC5596D6FBE1E4F3" box="[1012,1103,937,962]" pageId="16" pageNumber="347" rank="species">
<emphasis id="B97EA717FFD7FFD5FC5596D6FBE1E4F3" bold="true" box="[1012,1103,937,962]" pageId="16" pageNumber="347">sp. nov.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="4C0A0086FFD7FFD5FB2996D5FAB4E4F3" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[1160,1306,938,962]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" species="muluensis" status="sp. nov.">
<emphasis id="B97EA717FFD7FFD5FB2996D5FAB4E4F3" box="[1160,1306,938,962]" italics="true" pageId="16" pageNumber="347">C. muluensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFD7FFD5FA8296D6FAD0E4F3" box="[1315,1406,937,962]" pageId="16" pageNumber="347" rank="species">
<emphasis id="B97EA717FFD7FFD5FA8296D6FAD0E4F3" bold="true" box="[1315,1406,937,962]" pageId="16" pageNumber="347">sp. nov.</emphasis>
</taxonomicNameLabel>
in separate deeply divergent clades with both being distinct from 
<emphasis id="B97EA717FFD7FFD5FC3596B1FB45E4D7" box="[916,1259,974,998]" italics="true" pageId="16" pageNumber="347">
<taxonomicName id="4C0A0086FFD7FFD5FC3596B1FBE4E4D7" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[916,1098,974,998]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" sensu="stricto" species="cavernicolus">C. cavernicolus</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFD7FFD5FBF996B0FB45E4D7" box="[1112,1259,975,998]" pageId="16" pageNumber="347" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
, as previously suggested. 
<taxonomicName id="4C0A0086FFD7FFD5FEB5968DFD84E33B" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[276,554,1010,1034]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" species="muluensis" status="sp. nov.">
<emphasis id="B97EA717FFD7FFD5FEB5968DFD84E33B" box="[276,554,1010,1034]" italics="true" pageId="16" pageNumber="347">Cyrtodactylus muluensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFD7FFD5FD93968EFD23E33B" box="[562,653,1009,1034]" pageId="16" pageNumber="347" rank="species">
<emphasis id="B97EA717FFD7FFD5FD93968EFD23E33B" bold="true" box="[562,653,1009,1034]" pageId="16" pageNumber="347">sp. nov.</emphasis>
</taxonomicNameLabel>
is well supported as the sister species to 
<emphasis id="B97EA717FFD7FFD5FBC5968DFA3AE338" box="[1124,1428,1010,1034]" italics="true" pageId="16" pageNumber="347">
<taxonomicName id="4C0A0086FFD7FFD5FBC5968DFB50E33B" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[1124,1278,1010,1034]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" sensu="stricto" species="pubisulcus">C. pubisulcus</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFD7FFD5FAA7968DFA3AE338" box="[1286,1428,1010,1033]" pageId="16" pageNumber="347" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
, another Bornean endemic. Furthermore, 
<taxonomicName id="4C0A0086FFD7FFD5FDCE9169FCAAE31F" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[623,772,1046,1070]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" species="muluensis" status="sp. nov.">
<emphasis id="B97EA717FFD7FFD5FDCE9169FCAAE31F" box="[623,772,1046,1070]" italics="true" pageId="16" pageNumber="347">C. muluensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFD7FFD5FCAF916AFCC2E31F" box="[782,876,1045,1070]" pageId="16" pageNumber="347" rank="species">
<emphasis id="B97EA717FFD7FFD5FCAF916AFCC2E31F" bold="true" box="[782,876,1045,1070]" pageId="16" pageNumber="347">sp. nov.</emphasis>
</taxonomicNameLabel>
is embedded within the monophyletic group of primarily Philippine endemics, including 
<emphasis id="B97EA717FFD7FFD5FDC99145FAB5E363" box="[616,1307,1082,1106]" italics="true" pageId="16" pageNumber="347">
<taxonomicName id="4C0A0086FFD7FFD5FDC99145FCB0E363" baseAuthorityName="Steindachner" baseAuthorityYear="1867" box="[616,798,1082,1106]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" species="philippinicus">C. philippinicus</taxonomicName>
, 
<taxonomicName id="4C0A0086FFD7FFD5FC889145FC61E363" box="[809,975,1082,1106]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" species="redimiculus">C. redimiculus</taxonomicName>
, 
<taxonomicName id="4C0A0086FFD7FFD5FC7B9145FBD2E363" box="[986,1148,1082,1106]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" species="jambangan">C. jambangan</taxonomicName>
, 
<taxonomicName id="4C0A0086FFD7FFD5FB269145FAB5E363" box="[1159,1307,1082,1106]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" species="tautborum">C. tautborum</taxonomicName>
</emphasis>
, and the 
<taxonomicName id="4C0A0086FFD7FFD5FA239145FE8EE347" baseAuthorityName="Taylor" baseAuthorityYear="1915" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" species="agusanensis">
<emphasis id="B97EA717FFD7FFD5FA239145FE8EE347" italics="true" pageId="16" pageNumber="347">C. agusanensis</emphasis>
</taxonomicName>
species complex. Nodal support is lacking for some of the noted relationships: 
<taxonomicName id="4C0A0086FFD7FFD5FB0E9121FAE7E347" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[1199,1353,1118,1142]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" species="pubisulcus">
<emphasis id="B97EA717FFD7FFD5FB0E9121FAE7E347" box="[1199,1353,1118,1142]" italics="true" pageId="16" pageNumber="347">C. pubisulcus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0A0086FFD7FFD5FA239121FEAFE3AB" baseAuthorityName="Mocquard" baseAuthorityYear="1890" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" species="baluensis">
<emphasis id="B97EA717FFD7FFD5FA239121FEAFE3AB" italics="true" pageId="16" pageNumber="347">C. baluensis</emphasis>
</taxonomicName>
as outgroups to the monophyly containing 
<taxonomicName id="4C0A0086FFD7FFD5FD4B91FDFCD5E3AB" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[746,891,1154,1178]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" species="muluensis" status="sp. nov.">
<emphasis id="B97EA717FFD7FFD5FD4B91FDFCD5E3AB" box="[746,891,1154,1178]" italics="true" pageId="16" pageNumber="347">C. muluensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFD7FFD5FC2291FEFC73E3AB" box="[899,989,1153,1178]" pageId="16" pageNumber="347" rank="species">
<emphasis id="B97EA717FFD7FFD5FC2291FEFC73E3AB" bold="true" box="[899,989,1153,1178]" pageId="16" pageNumber="347">sp. nov.</emphasis>
</taxonomicNameLabel>
; 
<taxonomicName id="4C0A0086FFD7FFD5FC4D91FDFB3EE3AB" box="[1004,1168,1154,1178]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" species="redimiculus">
<emphasis id="B97EA717FFD7FFD5FC4D91FDFB3EE3AB" box="[1004,1168,1154,1178]" italics="true" pageId="16" pageNumber="347">C. redimiculus</emphasis>
</taxonomicName>
as the nearest outgroup to the 
<taxonomicName id="4C0A0086FFD7FFD5FF4291D9FED8E38F" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[227,374,1190,1214]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" species="muluensis" status="sp. nov.">
<emphasis id="B97EA717FFD7FFD5FF4291D9FED8E38F" box="[227,374,1190,1214]" italics="true" pageId="16" pageNumber="347">C. muluensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFD7FFD5FE2191DAFE73E38F" box="[384,477,1189,1214]" pageId="16" pageNumber="347" rank="species">
<emphasis id="B97EA717FFD7FFD5FE2191DAFE73E38F" bold="true" box="[384,477,1189,1214]" pageId="16" pageNumber="347">sp. nov.</emphasis>
</taxonomicNameLabel>
, 
<emphasis id="B97EA717FFD7FFD5FE4F91D9FC12E38F" box="[494,956,1190,1214]" italics="true" pageId="16" pageNumber="347">
<taxonomicName id="4C0A0086FFD7FFD5FE4F91D9FD27E38F" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[494,649,1190,1214]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" sensu="stricto" species="pubisulcus">C. pubisulcus</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFD7FFD5FD3291D8FC8BE38F" box="[659,805,1191,1214]" pageId="16" pageNumber="347" sensu="stricto">sensu stricto</taxonomicNameLabel>
, 
<taxonomicName id="4C0A0086FFD7FFD5FC9591D9FC17E38F" authorityName="Grismer" authorityYear="2005" box="[820,953,1190,1214]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" species="aurensis">C. aurensis</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C0A0086FFD7FFD5FC5891D9FB02E38F" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[1017,1196,1190,1214]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" species="cavernicolus">
<emphasis id="B97EA717FFD7FFD5FC5891D9FB02E38F" box="[1017,1196,1190,1214]" italics="true" pageId="16" pageNumber="347">C. cavernicolus</emphasis>
</taxonomicName>
monophyly; and the sister clade relationship between the 
<taxonomicName id="4C0A0086FFD7FFD5FD9691B5FD66E3D3" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[567,712,1226,1250]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" species="muluensis" status="sp. nov.">
<emphasis id="B97EA717FFD7FFD5FD9691B5FD66E3D3" box="[567,712,1226,1250]" italics="true" pageId="16" pageNumber="347">C. muluensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFD7FFD5FD7191B6FC84E3D3" box="[720,810,1225,1250]" pageId="16" pageNumber="347" rank="species">
<emphasis id="B97EA717FFD7FFD5FD7191B6FC84E3D3" bold="true" box="[720,810,1225,1250]" pageId="16" pageNumber="347">sp. nov.</emphasis>
</taxonomicNameLabel>
group and the 
<taxonomicName id="4C0A0086FFD7FFD5FC7591B5FBD0E3D0" baseAuthorityName="Taylor" baseAuthorityYear="1915" box="[980,1150,1226,1249]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" species="agusanensis">
<emphasis id="B97EA717FFD7FFD5FC7591B5FBD0E3D0" box="[980,1150,1226,1249]" italics="true" pageId="16" pageNumber="347">C. agusanensis</emphasis>
</taxonomicName>
clade; but there is strong support for the deepest node forming the Philippine clade. The phylogeny has strong support (92/1.00) for designating 
<taxonomicName id="4C0A0086FFD7FFD5FEBF906DFE00E21B" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[286,430,1298,1322]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" species="muluensis" status="sp. nov.">
<emphasis id="B97EA717FFD7FFD5FEBF906DFE00E21B" box="[286,430,1298,1322]" italics="true" pageId="16" pageNumber="347">C. muluensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFD7FFD5FE15906EFDA0E21B" box="[436,526,1297,1322]" pageId="16" pageNumber="347" rank="species">
<emphasis id="B97EA717FFD7FFD5FE15906EFDA0E21B" bold="true" box="[436,526,1297,1322]" pageId="16" pageNumber="347">sp. nov.</emphasis>
</taxonomicNameLabel>
as a distinct lineage. The two subclades demonstrated within the 
<taxonomicName id="4C0A0086FFD7FFD5FB45906DFADAE21B" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[1252,1396,1298,1322]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" species="muluensis" status="sp. nov.">
<emphasis id="B97EA717FFD7FFD5FB45906DFADAE21B" box="[1252,1396,1298,1322]" italics="true" pageId="16" pageNumber="347">C. muluensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFD7FFD5FADB906EFF66E27F" pageId="16" pageNumber="347" rank="species">
<emphasis id="B97EA717FFD7FFD5FADB906EFF66E27F" bold="true" pageId="16" pageNumber="347">sp. nov.</emphasis>
</taxonomicNameLabel>
clade correlate with the karst formation in which they were found in. The clade comprised of specimens CAS 262995–262997 originated from the karst formation around the Clearwater Cave, and the specimens in the other subclade were from the karst formation around the Lang Cave. No morphological differences were found between the individuals in the two subclades
</paragraph>
<paragraph id="8BB57B05FFD7FFD5FF6690B9FF41E15F" blockId="16.[151,1436,901,1646]" pageId="16" pageNumber="347">
<taxonomicName id="4C0A0086FFD7FFD5FF6690B9FE7BE2EF" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[199,469,1478,1502]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" species="limajalur" status="sp. nov.">
<emphasis id="B97EA717FFD7FFD5FF6690B9FE7BE2EF" box="[199,469,1478,1502]" italics="true" pageId="16" pageNumber="347">Cyrtodactylus limajalur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFD7FFD5FE7C90BAFD96E2EF" box="[477,568,1477,1502]" pageId="16" pageNumber="347" rank="species">
<emphasis id="B97EA717FFD7FFD5FE7C90BAFD96E2EF" bold="true" box="[477,568,1477,1502]" pageId="16" pageNumber="347">sp. nov.</emphasis>
</taxonomicNameLabel>
forms a clade that is well supported (99/0.99) as the sister species to the 
<emphasis id="B97EA717FFD7FFD5FA2390B9FE0FE133" italics="true" pageId="16" pageNumber="347">
<taxonomicName id="4C0A0086FFD7FFD5FA2390B9FEB4E133" baseAuthorityName="Peters" baseAuthorityYear="1871" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" species="consobrinus">C. consobrinus</taxonomicName>
/ 
<taxonomicName id="4C0A0086FFD7FFD5FE869095FE0FE133" box="[295,417,1514,1538]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" species="malayanus">malayanus</taxonomicName>
</emphasis>
group. While these latter two species are difficult to distinguish from one another morphologically (Hikida 1991; references therein), they are well-supported as phylogenetically distinct lineages based on the concatenated dataset. Regardless, 
<taxonomicName id="4C0A0086FFD7FFD5FD06934DFC81E17B" authority="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das, 2019" authorityName="Davis &amp; Bauer &amp; Jackman &amp; Nashriq &amp; Das" authorityYear="2019" box="[679,815,1586,1610]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="347" phylum="Chordata" rank="species" species="limajalur" status="sp. nov.">
<emphasis id="B97EA717FFD7FFD5FD06934DFC81E17B" box="[679,815,1586,1610]" italics="true" pageId="16" pageNumber="347">C. limajalur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D1A6CFFD7FFD5FC97934EFC3EE17B" box="[822,912,1585,1610]" pageId="16" pageNumber="347" rank="species">
<emphasis id="B97EA717FFD7FFD5FC97934EFC3EE17B" bold="true" box="[822,912,1585,1610]" pageId="16" pageNumber="347">sp. nov.</emphasis>
</taxonomicNameLabel>
has strong support for its designation as a new species.
</paragraph>
</subSubSection>
</treatment>
</document>