treatments-xml/data/E3/72/22/E37222277574FF8EE3C41BA3FCB5FD76.xml

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<mods:title id="66DFFA6A39E3D83825A379398B89B17F">The new genus Habeastrum, with two new species (Gastropoda, Diplommatinidae) in Mato Grosso do Sul caves, Brazil</mods:title>
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<mods:namePart id="CA9B4E998128CA14C1C7E7A25D08F351">Simone, Luiz Ricardo L.</mods:namePart>
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<treatment id="E37222277574FF8EE3C41BA3FCB5FD76" ID-DOI="http://doi.org/10.5281/zenodo.5941067" ID-GBIF-Taxon="154529781" ID-Zenodo-Dep="5941067" LSID="urn:lsid:zoobank.org:act:6CA1A8C0-4891-4F7C-B1C1-489B9E1B89F7" httpUri="http://treatment.plazi.org/id/E37222277574FF8EE3C41BA3FCB5FD76" lastPageId="1" lastPageNumber="288" pageId="0" pageNumber="287">
<subSubSection id="23C1C0BA7574FF8FE3C41BA3FC35F8FE" pageId="0" pageNumber="287" type="nomenclature">
<paragraph id="6B6493317574FF8FE3C41BA3FC9DF91F" blockId="0.[151,1436,1762,2028]" box="[151,783,1762,1789]" pageId="0" pageNumber="287">
<heading id="302C245D7574FF8FE3C41BA3FC9DF91F" bold="true" box="[151,783,1762,1789]" fontSize="11" level="1" pageId="0" pageNumber="287" reason="1">
<emphasis id="59AF4F237574FF8FE3C41BA3FC9DF91F" bold="true" box="[151,783,1762,1789]" pageId="0" pageNumber="287">
<taxonomicName id="ACDBE8B27574FF8FE3C41BA3FE2BF91E" ID-CoL="3J3L3" authority="Simone, 2019" authorityName="Simone" authorityYear="2019" box="[151,441,1762,1788]" class="Gastropoda" family="Diplommatinidae" genus="Habeastrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="0" pageNumber="287" phylum="Mollusca" rank="species" species="parafusum" status="sp. nov.">
<emphasis id="59AF4F237574FF8FE3C41BA3FE2BF91E" bold="true" box="[151,441,1762,1788]" italics="true" pageId="0" pageNumber="287">Habeastrum parafusum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="429CF2587574FF8FE2EC1BA2FDDDF91F" box="[447,591,1763,1789]" pageId="0" pageNumber="287" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="F3E08FB47574FF8FE10D1BA2FD43F91F" box="[606,721,1763,1789]" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="0" pageNumber="287">Figures 1</figureCitation>
A–J)
</emphasis>
</heading>
</paragraph>
<paragraph id="6B6493317574FF8FE3C41A44FC35F8FE" blockId="0.[151,1436,1762,2028]" box="[151,935,1797,1820]" pageId="0" pageNumber="287">
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urn:lsid:zoobank.org:act: 
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.
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<subSubSection id="23C1C0BA7574FF8FE3C41A69FDEAF883" pageId="0" pageNumber="287" type="description">
<paragraph id="6B6493317574FF8FE3C41A69FDEAF883" blockId="0.[151,1436,1762,2028]" pageId="0" pageNumber="287">
<emphasis id="59AF4F237574FF8FE3C41A69FF4CF8DD" bold="true" box="[151,222,1832,1855]" pageId="0" pageNumber="287">Types.</emphasis>
<materialsCitation id="DBB3996C7574FF8FE3B51A69FD54F8DD" ID-GBIF-Occurrence="2147525214" box="[230,710,1832,1855]" collectionCode="MZSP" pageId="0" pageNumber="287" specimenCode="MZSP 134301" specimenCount="2" typeStatus="holotype">
<typeStatus id="B4602D937574FF8FE3B51A69FED1F8DD" box="[230,323,1832,1855]" pageId="0" pageNumber="287" type="holotype">holotype</typeStatus>
<specimenCode id="3B7D3B4A7574FF8FE21A1A69FE77F8DD" box="[329,485,1832,1855]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="0" pageNumber="287" type="Museum">MZSP 134301</specimenCode>
(
<figureCitation id="F3E08FB47574FF8FE2A01A69FDD5F8DD" box="[499,583,1832,1855]" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="0" pageNumber="287">Fig. 1A</figureCitation>
). 
<typeStatus id="B4602D937574FF8FE1091A69FD54F8DD" box="[602,710,1832,1855]" pageId="0" pageNumber="287" type="paratype">Paratypes</typeStatus>
</materialsCitation>
: 
<materialsCitation id="DBB3996C7574FF8FE19C1A69FB8FF8DD" ID-GBIF-Occurrence="2147524942" box="[719,1053,1832,1855]" collectionCode="MZSP" pageId="0" pageNumber="287" specimenCode="MZSP 135584, 7" specimenCount="1" typeStatus="paratype">
<specimenCode id="3B7D3B4A7574FF8FE19C1A69FCFCF8DD" box="[719,878,1832,1855]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="0" pageNumber="287" type="Museum">MZSP 135584</specimenCode>
, 
<specimenCode id="3B7D3B4A7574FF8FE02B1A69FC17F8DD" box="[888,901,1832,1855]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="0" pageNumber="287" type="Museum">7</specimenCode>
figured shells
</materialsCitation>
, 
<materialsCitation id="DBB3996C7574FF8FE77B1A69FAA2F8DD" ID-GBIF-Occurrence="2147525113" box="[1064,1328,1832,1855]" collectionCode="MZSP" pageId="0" pageNumber="287" specimenCode="MZSP 135585, 15" specimenCount="1" typeStatus="paratype">
<specimenCode id="3B7D3B4A7574FF8FE77B1A69FB54F8DD" box="[1064,1222,1832,1855]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="0" pageNumber="287" type="Museum">MZSP 135585</specimenCode>
, 
<specimenCode id="3B7D3B4A7574FF8FE7831A69FB79F8DD" box="[1232,1259,1832,1855]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="0" pageNumber="287" type="Museum">15</specimenCode>
shells
</materialsCitation>
, MNRJ, 2 shells, USNM, 2 shells all from type locality.
</paragraph>
</subSubSection>
<subSubSection id="23C1C0BA7574FF8FE3941A2CFE2EF845" pageId="0" pageNumber="287" type="materials_examined">
<paragraph id="6B6493317574FF8FE3941A2CFE2EF845" blockId="0.[151,1436,1762,2028]" pageId="0" pageNumber="287">
<materialsCitation id="DBB3996C7574FF8FE3941A2CFE25F845" ID-GBIF-Occurrence="2147525050" collectingDate="1992-07-16" collectorName="Trajano &amp; Gnaspini" country="Brazil" latitude="-21.11028" location="Gruta Pitangueiras" longLatPrecision="21" longitude="-56.581112" municipality="Bonito" pageId="0" pageNumber="287" specimenCount="1" stateProvince="Mato Grosso do Sul" typeStatus="holotype">
<emphasis id="59AF4F237574FF8FE3941A2CFEF2F866" bold="true" box="[199,352,1901,1924]" pageId="0" pageNumber="287">
<typeStatus id="B4602D937574FF8FE3941A2CFF6DF866" box="[199,255,1901,1924]" pageId="0" pageNumber="287">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="13CCD3A17574FF8FE23F1A2CFE42F866" box="[364,464,1901,1924]" name="Brazil" pageId="0" pageNumber="287">BRAZIL</collectingCountry>
. 
<collectingRegion id="A91F5DD37574FF8FE28E1A2CFD53F866" box="[477,705,1901,1924]" country="Brazil" name="Mato Grosso do Sul" pageId="0" pageNumber="287">Mato Grosso do Sul</collectingRegion>
, 
<collectingMunicipality id="8B00094B7574FF8FE1831A2CFC89F866" box="[720,795,1901,1924]" pageId="0" pageNumber="287">Bonito</collectingMunicipality>
, 
<location id="6E04C5EA7574FF8FE0791A2CFC6AF866" LSID="urn:lsid:plazi:treatment:E37222277574FF8EE3C41BA3FCB5FD76:6E04C5EA7574FF8FE0791A2CFC6AF866" box="[810,1016,1901,1924]" country="Brazil" latitude="-21.11028" longLatPrecision="21" longitude="-56.581112" municipality="Bonito" name="Gruta Pitangueiras" pageId="0" pageNumber="287" stateProvince="Mato Grosso do Sul">Gruta Pitangueiras</location>
, 
<geoCoordinate id="0EEFF5F67574FF8FE7551A2CFB11F866" box="[1030,1155,1901,1924]" direction="south" orientation="latitude" pageId="0" pageNumber="287" precision="15" value="-21.11028">21°06’37”S</geoCoordinate>
<geoCoordinate id="0EEFF5F67574FF8FE7DD1A2CFA85F866" box="[1166,1303,1901,1924]" direction="west" orientation="longitude" pageId="0" pageNumber="287" precision="15" value="-56.581112">56°34’52”W</geoCoordinate>
(
<collectorName id="C62EF6E77574FF8FE67B1A2CFAEEF866" box="[1320,1404,1901,1924]" pageId="0" pageNumber="287">Trajano</collectorName>
&amp; 
<collectorName id="C62EF6E77574FF8FE3C41AD1FF65F845" box="[151,247,1936,1959]" pageId="0" pageNumber="287">Gnaspini</collectorName>
col., 
<date id="1F65B5F17574FF8FE2611AD1FE3DF845" box="[306,431,1936,1959]" pageId="0" pageNumber="287" value="1992-07-16">
<collectingDate id="0F214C197574FF8FE2611AD1FE3DF845" box="[306,431,1936,1959]" pageId="0" pageNumber="287" value="1992-07-16">16.vii.1992</collectingDate>
</date>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="23C1C0BA7574FF8FE3941AF3FAABF82B" box="[199,1337,1970,1993]" pageId="0" pageNumber="287" type="diagnosis">
<paragraph id="6B6493317574FF8FE3941AF3FAABF82B" blockId="0.[151,1436,1762,2028]" box="[199,1337,1970,1993]" pageId="0" pageNumber="287">
<emphasis id="59AF4F237574FF8FE3941AF3FEAEF82B" bold="true" box="[199,316,1970,1993]" pageId="0" pageNumber="287">Diagnosis.</emphasis>
Shell with about 7 whorls, 3–4 whorls preceding aperture increasing relatively less in diameter.
</paragraph>
</subSubSection>
<subSubSection id="23C1C0BA7574FF8EE3941A94FC89FE43" lastPageId="1" lastPageNumber="288" pageId="0" pageNumber="287" type="description">
<paragraph id="6B6493317574FF8FE3941A94FA09F80E" blockId="0.[151,1436,1762,2028]" box="[199,1435,2005,2028]" pageId="0" pageNumber="287">
<emphasis id="59AF4F237574FF8FE3941A94FEC0F80E" bold="true" box="[199,338,2005,2028]" pageId="0" pageNumber="287">Description.</emphasis>
Shell turriform, narrowly conical, almost cylindrical. First three whorls slightly sharper pointed,
</paragraph>
<paragraph id="6B6493317575FF8EE3C41DD9FC89FE43" blockId="1.[151,1435,152,660]" pageId="1" pageNumber="288">
profile angle ~40°; gradually becoming more cylindrical in 4–5 remaining whorls, with inflation relatively small, outline angle ~20° (
<figureCitation id="F3E08FB47575FF8EE2491DFBFECBFF33" box="[282,345,186,209]" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="1" pageNumber="288">Figs 1</figureCitation>
A–G). Total of about 7 whorls. Protoconch 
<quantity id="AC233ED47575FF8EE07D1DFBFC1FFF33" box="[814,909,186,209]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="1" pageNumber="288" unit="mm" value="0.39">0.39 mm</quantity>
wide, smooth, except for minute radial wrinkles, translucent (
<figureCitation id="F3E08FB47575FF8EE2481D9CFEE7FF16" box="[283,373,221,244]" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="1" pageNumber="288">Figs 1H</figureCitation>
, 10); occupying about 43% of shell width, about 15% of shell length. Teleoconch up to 6 whorls, sometimes with narrow profile (
<figureCitation id="F3E08FB47575FF8EE2B81C41FDCCFEF5" box="[491,606,256,279]" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="1" pageNumber="288">Figs 1B, F</figureCitation>
), sometimes slightly wider (
<figureCitation id="F3E08FB47575FF8EE0D81C41FB90FEF5" box="[907,1026,256,279]" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="1" pageNumber="288">Figs 1A, C</figureCitation>
), sometimes intermediate (
<figureCitation id="F3E08FB47575FF8EE6721C41FF3CFEDB" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="1" pageNumber="288">Figs 1D, E, G</figureCitation>
). Teleoconch sculpture as for genus (Fig. H), with about 65 axial cords on penultimate whorl. Suture deep, whorl profile rounded. Aperture rounded, not deviating from whorl profile, slightly prosocline (
<figureCitation id="F3E08FB47575FF8EE72F1C04FA87FEBE" box="[1148,1301,325,348]" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="1" pageNumber="288">Figs 1D, G, I</figureCitation>
); minimally deformed by penultimate whorl; occupying about 44% of shell width, about 25% of shell length. Umbilicus deep, narrow, occupying about 12% of inferior shell area (
<figureCitation id="F3E08FB47575FF8EE1971CCBFC9FFE43" box="[708,781,394,417]" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="1" pageNumber="288">
Fig. 
<date id="1F65B5F17575FF8EE1A51CCBFC9FFE43" box="[758,781,394,417]" pageId="1" pageNumber="288">1I</date>
</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="23C1C0BA7575FF8EE3941CECFD69FE05" pageId="1" pageNumber="288" type="distribution">
<paragraph id="6B6493317575FF8EE3941CECFC0FFE26" blockId="1.[151,1435,152,660]" box="[199,925,429,452]" pageId="1" pageNumber="288">
<emphasis id="59AF4F237575FF8EE3941CECFECBFE26" bold="true" box="[199,345,429,452]" pageId="1" pageNumber="288">Distribution.</emphasis>
Known only from the 
<typeStatus id="B4602D937575FF8EE11F1CECFDE8FE26" box="[588,634,429,452]" pageId="1" pageNumber="288">type</typeStatus>
locality, possibly endemic.
</paragraph>
<paragraph id="6B6493317575FF8EE3941C91FD69FE05" blockId="1.[151,1435,152,660]" box="[199,763,464,487]" pageId="1" pageNumber="288">
<emphasis id="59AF4F237575FF8EE3941C91FEB7FE05" bold="true" box="[199,293,464,487]" pageId="1" pageNumber="288">Habitat.</emphasis>
In soil in caves (no living specimen found).
</paragraph>
</subSubSection>
<subSubSection id="23C1C0BA7575FF8EE3941CB3FC5DFDAD" pageId="1" pageNumber="288" type="description">
<paragraph id="6B6493317575FF8EE3941CB3FC5DFDAD" blockId="1.[151,1435,152,660]" pageId="1" pageNumber="288">
<emphasis id="59AF4F237575FF8EE3941CB3FEFCFDEB" bold="true" box="[199,366,498,521]" pageId="1" pageNumber="288">Measurements</emphasis>
(length and width in mm): 
<typeStatus id="B4602D937575FF8EE1C21CB3FD66FDEB" box="[657,756,498,521]" pageId="1" pageNumber="288" type="holotype">Holotype</typeStatus>
<quantity id="AC233ED47575FF8EE1A91CB3FCD9FDEB" box="[762,843,498,521]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="288" unit="mm" value="1.4">1.4 mm</quantity>
by 
<quantity id="AC233ED47575FF8EE0211CB3FC56FDEB" box="[882,964,498,521]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="288" unit="mm" value="0.8">0.8 mm</quantity>
(
<figureCitation id="F3E08FB47575FF8EE0811CB3FBB4FDEB" box="[978,1062,498,521]" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="1" pageNumber="288">Fig. 1A</figureCitation>
); 
<typeStatus id="B4602D937575FF8EE7691CB3FB30FDEB" box="[1082,1186,498,521]" pageId="1" pageNumber="288" type="paratype">Paratypes</typeStatus>
MZSP 135584: #2: 
<quantity id="AC233ED47575FF8EE6291CB3FF53FDCE" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="288" unit="mm" value="1.7">1.7 mm</quantity>
by 
<quantity id="AC233ED47575FF8EE3B81F54FEACFDCE" box="[235,318,533,556]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="288" unit="mm" value="0.8">0.8 mm</quantity>
(
<figureCitation id="F3E08FB47575FF8EE21D1F54FE30FDCE" box="[334,418,533,556]" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="1" pageNumber="288">Fig. 1B</figureCitation>
); #3, 
<quantity id="AC233ED47575FF8EE2B21F54FDA7FDCE" box="[481,565,533,556]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="288" unit="mm" value="1.6">1.6 mm</quantity>
by 
<quantity id="AC233ED47575FF8EE10C1F54FD23FDCE" box="[607,689,533,556]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="288" unit="mm" value="0.9">0.9 mm</quantity>
(
<figureCitation id="F3E08FB47575FF8EE1911F54FC87FDCE" box="[706,789,533,556]" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="1" pageNumber="288">Fig. 1C</figureCitation>
); #4, 
<quantity id="AC233ED47575FF8EE0051F54FC3AFDCE" box="[854,936,533,556]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="288" unit="mm" value="1.6">1.6 mm</quantity>
by 
<quantity id="AC233ED47575FF8EE0801F54FBB4FDCE" box="[979,1062,533,556]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="288" unit="mm" value="0.8">0.8 mm</quantity>
(
<figureCitation id="F3E08FB47575FF8EE7651F54FB19FDCE" box="[1078,1163,533,556]" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="1" pageNumber="288">Fig. 1D</figureCitation>
); #5, 
<quantity id="AC233ED47575FF8EE7981F54FA8CFDCE" box="[1227,1310,533,556]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="288" unit="mm" value="1.4">1.4 mm</quantity>
by 
<quantity id="AC233ED47575FF8EE61A1F54FA09FDCE" box="[1353,1435,533,556]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="288" unit="mm" value="0.7">0.7 mm</quantity>
(
<figureCitation id="F3E08FB47575FF8EE3CC1F79FF62FDAD" box="[159,240,568,591]" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="1" pageNumber="288">Fig. 1E</figureCitation>
); #6, 
<quantity id="AC233ED47575FF8EE27E1F79FEECFDAD" box="[301,382,568,591]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="288" unit="mm" value="1.1">1.1 mm</quantity>
by 
<quantity id="AC233ED47575FF8EE2F51F79FE6AFDAD" box="[422,504,568,591]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="288" unit="mm" value="0.7">0.7 mm</quantity>
(
<figureCitation id="F3E08FB47575FF8EE1541F79FDC5FDAD" box="[519,599,568,591]" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="1" pageNumber="288">Fig. 1F</figureCitation>
); #7, 
<quantity id="AC233ED47575FF8EE1C01F79FD76FDAD" box="[659,740,568,591]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="288" unit="mm" value="1.2">1.2 mm</quantity>
by 
<quantity id="AC233ED47575FF8EE05E1F79FCCDFDAD" box="[781,863,568,591]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="288" unit="mm" value="0.7">0.7 mm</quantity>
(
<figureCitation id="F3E08FB47575FF8EE03E1F79FC50FDAD" box="[877,962,568,591]" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="1" pageNumber="288">Fig. 1G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="23C1C0BA7575FF8EE3941F1BFCB5FD76" pageId="1" pageNumber="288" type="etymology">
<paragraph id="6B6493317575FF8EE3941F1BFCB5FD76" blockId="1.[151,1435,152,660]" pageId="1" pageNumber="288">
<emphasis id="59AF4F237575FF8EE3941F1BFED5FD93" bold="true" box="[199,327,602,625]" pageId="1" pageNumber="288">Etymology.</emphasis>
The specific epithet refers to the general appearance of the shell, resembling a screw, “ 
<emphasis id="59AF4F237575FF8EE7BA1F18FADAFD93" box="[1257,1352,601,625]" italics="true" pageId="1" pageNumber="288">parafuso</emphasis>
” in Portuguese, arbitrarily Latinized with the neutral termination.
</paragraph>
</subSubSection>
</treatment>
</document>