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<mods:title id="01DF230F0FD52954332B731BE921A224">The two southernmost species of the Andean genus Acrographinotus Holmgren (Arachnida, Opiliones, Gonyleptidae) described from Bolivia and Argentina</mods:title>
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<mods:namePart id="D94D1A675B3E676D772A5860100B9C57">Acosta, Luis E.</mods:namePart>
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<mods:title id="7D2DD8DF39E7A8E5C52F5F4AD455EE37">Zootaxa</mods:title>
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<mods:part id="E87FDDDC841080B39511F30A6BE3D53E">
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<mods:date id="D3EC8AE2351114BC34DC00D8D98D6CE6">2024</mods:date>
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<mods:number id="4BCDB20CBC32310550ADF9745AC4CC63">2024-07-19</mods:number>
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<mods:number id="230EED929F4C71737DE3BE2042A2EE35">5</mods:number>
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<taxonomicName id="92F69E849F60FF8570AFFC80FDB4FCF4" ID-CoL="7NMYV" authority="Holmgren, 1916" authorityName="Holmgren" authorityYear="1917" box="[151,564,831,858]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="genus">
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<emphasis id="678239159F60FF8570AFFC80FDB4FCF4" bold="true" box="[151,564,831,858]" pageId="3" pageNumber="550">
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<emphasis id="678239159F60FF8570AFFC80FEE4FCF4" bold="true" box="[151,356,831,858]" italics="true" pageId="3" pageNumber="550">Acrographinotus</emphasis>
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<bibRefCitation id="316798F69F60FF857154FCFFFDB4FCF4" author="Holmgren, N." box="[364,564,832,858]" pageId="3" pageNumber="550" pagination="1 - 303" refId="ref11267" refString="Holmgren, N. (1916) Zur vergleichenden Anatomie des Gehirns von Polychaeten, Onychophoren, Xiphosuren, Arachniden, Crustaceen, Myriopoden und Inseckten. Vorstudien zu einer Phylogenie der Arthropoden. Kungliga Svenska Vetenskapsakademiens handlingar, 56, 1 - 303, 12 pls." type="journal article" year="1916">Holmgren, 1916</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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urn:lsid:zoobank.org:act:
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</uri>
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<paragraph id="5549E5079F60FF8570AFFC17FC2CFC6F" blockId="3.[151,940,936,961]" box="[151,940,936,961]" pageId="3" pageNumber="550">
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<treatmentCitationGroup id="75E6C2299F60FF8570AFFC17FC2CFC6F" box="[151,940,936,961]" pageId="3" pageNumber="550">
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<treatmentCitation id="D457C3169F60FF8570AFFC17FD91FC6F" author="Holmgren, N." box="[151,529,936,961]" page="89" pageId="3" pageNumber="550" year="1916">
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<taxonomicName id="92F69E849F60FF8570AFFC17FD91FC6F" ID-CoL="7NMYV" authority="Holmgren 1916: 89" authorityName="Holmgren" authorityPageNumber="89" authorityYear="1917" box="[151,529,936,961]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="genus">
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<emphasis id="678239159F60FF8570AFFC17FEC2FC6E" box="[151,322,936,960]" italics="true" pageId="3" pageNumber="550">Acrographinotus</emphasis>
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<bibRefCitation id="316798F69F60FF857170FC17FD91FC6F" author="Holmgren, N." box="[328,529,936,961]" pageId="3" pageNumber="550" pagination="1 - 303" refId="ref11267" refString="Holmgren, N. (1916) Zur vergleichenden Anatomie des Gehirns von Polychaeten, Onychophoren, Xiphosuren, Arachniden, Crustaceen, Myriopoden und Inseckten. Vorstudien zu einer Phylogenie der Arthropoden. Kungliga Svenska Vetenskapsakademiens handlingar, 56, 1 - 303, 12 pls." type="journal article" year="1916">Holmgren 1916: 89</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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;
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<treatmentCitation id="D457C3169F60FF857223FC16FD40FC6E" author="Acosta, L. E." box="[539,704,936,961]" page="58" pageId="3" pageNumber="550" year="2001">
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<bibRefCitation id="316798F69F60FF857223FC16FD40FC6E" author="Acosta, L. E." box="[539,704,936,961]" pageId="3" pageNumber="550" pagination="58 - 66" refId="ref10602" refString="Acosta, L. E. (2001) The identity of Acrographinotus erectispina, with a review of the generic diagnosis, and the description of a new species (Opiliones, Gonyleptidae, Pachylinae). Bulletin of the British Arachnological Society, 12 (2), 58 - 66." type="journal article" year="2001">Acosta 2001: 58</bibRefCitation>
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</treatmentCitation>
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[complete synonymy].
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="1DECB68C9F60FF8570AFFC50FDCCFB83" pageId="3" pageNumber="550" type="type_taxon">
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<paragraph id="5549E5079F60FF8570AFFC50FDCCFB83" blockId="3.[151,1437,1006,2041]" pageId="3" pageNumber="550">
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<emphasis id="678239159F60FF8570AFFC50FEB8FBA6" bold="true" box="[151,312,1006,1033]" pageId="3" pageNumber="550">
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<typeStatus id="8A4D5BA59F60FF8570AFFC50FF52FBA7" box="[151,210,1007,1033]" pageId="3" pageNumber="550">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="92F69E849F60FF85717DFC50FCBDFBA7" ID-CoL="7QH8R" authority="Roewer, 1929" authorityName="Roewer" authorityYear="1929" box="[325,829,1007,1033]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="erectispina">
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<emphasis id="678239159F60FF85717DFC50FD0AFBA6" box="[325,650,1007,1033]" italics="true" pageId="3" pageNumber="550">Acrographinotus erectispina</emphasis>
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<bibRefCitation id="316798F69F60FF8572A0FC50FCBDFBA7" author="Roewer, C. F." box="[664,829,1007,1033]" pageId="3" pageNumber="550" pagination="179 - 290" refId="ref11972" refString="Roewer, C. F. (1929) Weitere Weberknechte III. III Erganzung der: " Weberknechte der Erde ", 1923. Abhandlungen vom naturwissenschaftlichen Verein zu Bremen, 27 (2), 179 - 290, pl. 1." type="journal article" year="1929">Roewer, 1929</bibRefCitation>
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</taxonomicName>
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, by subsequent designation of
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<bibRefCitation id="316798F69F60FF85748BFC50FAE2FBA7" author="Roewer, C. F." box="[1203,1378,1007,1033]" pageId="3" pageNumber="550" pagination="179 - 290" refId="ref11972" refString="Roewer, C. F. (1929) Weitere Weberknechte III. III Erganzung der: " Weberknechte der Erde ", 1923. Abhandlungen vom naturwissenschaftlichen Verein zu Bremen, 27 (2), 179 - 290, pl. 1." type="journal article" year="1929">Roewer (1929)</bibRefCitation>
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(see nomenclatural history in
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<bibRefCitation id="316798F69F60FF857196FBACFDBFFB83" author="Acosta, L. E." box="[430,575,1043,1069]" pageId="3" pageNumber="550" pagination="58 - 66" refId="ref10602" refString="Acosta, L. E. (2001) The identity of Acrographinotus erectispina, with a review of the generic diagnosis, and the description of a new species (Opiliones, Gonyleptidae, Pachylinae). Bulletin of the British Arachnological Society, 12 (2), 58 - 66." type="journal article" year="2001">Acosta 2001</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="1DECB68C9F60FF8570FFFB88FBC7FB37" pageId="3" pageNumber="550" type="taxon_list">
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<paragraph id="5549E5079F60FF8570FFFB88FBC7FB37" blockId="3.[151,1437,1006,2041]" pageId="3" pageNumber="550">
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<emphasis id="678239159F60FF8570FFFB88FE12FBFE" bold="true" box="[199,402,1078,1105]" pageId="3" pageNumber="550">Included species.</emphasis>
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<taxonomicName id="92F69E849F60FF8571A2FB88FC00FBFF" ID-CoL="7QH8R" authority="Roewer, 1929" authorityName="Roewer" authorityYear="1929" box="[410,896,1079,1105]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="erectispina">
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<emphasis id="678239159F60FF8571A2FB88FD59FBFE" box="[410,729,1079,1105]" italics="true" pageId="3" pageNumber="550">Acrographinotus erectispina</emphasis>
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<bibRefCitation id="316798F69F60FF8572D9FB88FC00FBFF" author="Roewer, C. F." box="[737,896,1079,1105]" pageId="3" pageNumber="550" pagination="179 - 290" refId="ref11972" refString="Roewer, C. F. (1929) Weitere Weberknechte III. III Erganzung der: " Weberknechte der Erde ", 1923. Abhandlungen vom naturwissenschaftlichen Verein zu Bremen, 27 (2), 179 - 290, pl. 1." type="journal article" year="1929">Roewer, 1929</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="92F69E849F60FF8573B3FB88FB48FBFF" ID-CoL="7QH8Q" authority="Roewer, 1929" authorityName="Roewer" authorityYear="1929" box="[907,1224,1079,1105]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="curvispina">
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<emphasis id="678239159F60FF8573B3FB88FBA1FBFE" box="[907,1057,1079,1105]" italics="true" pageId="3" pageNumber="550">A. curvispina</emphasis>
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<bibRefCitation id="316798F69F60FF857411FB88FB48FBFF" author="Roewer, C. F." box="[1065,1224,1079,1105]" pageId="3" pageNumber="550" pagination="179 - 290" refId="ref11972" refString="Roewer, C. F. (1929) Weitere Weberknechte III. III Erganzung der: " Weberknechte der Erde ", 1923. Abhandlungen vom naturwissenschaftlichen Verein zu Bremen, 27 (2), 179 - 290, pl. 1." type="journal article" year="1929">Roewer, 1929</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="92F69E849F60FF8574ECFB88FF5BFBDB" ID-CoL="7QH8Z" authority="(Roewer, 1957)" baseAuthorityName="Roewer" baseAuthorityYear="1957" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="ortizi">
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<emphasis id="678239159F60FF8574ECFB88FAAEFBFE" box="[1236,1326,1079,1105]" italics="true" pageId="3" pageNumber="550">A. ortizi</emphasis>
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(
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<bibRefCitation id="316798F69F60FF857506FB88FF53FBDB" author="Roewer, C. F." pageId="3" pageNumber="550" pagination="67 - 94" refId="ref12086" refString="Roewer, C. F. (1957) Arachnida Arthrogastra aus Peru, III. Senckenbergiana biologica, 38 (1 / 2), 67 - 94, pls. 3 - 6." type="journal article" year="1957">Roewer, 1957</bibRefCitation>
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)
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</taxonomicName>
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,
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<taxonomicName id="92F69E849F60FF8570D0FBE4FD97FBDB" ID-CoL="7QH8X" authority="(Roewer, 1929)" baseAuthorityName="Roewer" baseAuthorityYear="1929" box="[232,535,1115,1141]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="ornatus">
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<emphasis id="678239159F60FF8570D0FBE4FEDEFBDA" box="[232,350,1115,1141]" italics="true" pageId="3" pageNumber="550">A. ornatus</emphasis>
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(
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<bibRefCitation id="316798F69F60FF857155FBE4FD90FBDB" author="Roewer, C. F." box="[365,528,1115,1141]" pageId="3" pageNumber="550" pagination="179 - 290" refId="ref11972" refString="Roewer, C. F. (1929) Weitere Weberknechte III. III Erganzung der: " Weberknechte der Erde ", 1923. Abhandlungen vom naturwissenschaftlichen Verein zu Bremen, 27 (2), 179 - 290, pl. 1." type="journal article" year="1929">Roewer, 1929</bibRefCitation>
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)
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</taxonomicName>
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,
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<taxonomicName id="92F69E849F60FF85721CFBE4FC69FBDB" ID-CoL="7QH8N" authority="(Soares & Bauab, 1972)" baseAuthorityName="Soares & Bauab" baseAuthorityYear="1972" box="[548,1001,1115,1141]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="ceratopygus">
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|
|
<emphasis id="678239159F60FF85721CFBE4FD4DFBDA" box="[548,717,1115,1141]" italics="true" pageId="3" pageNumber="550">A. ceratopygus</emphasis>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="316798F69F60FF8572E5FBE4FC61FBDB" author="Soares, H. E. M. & Bauab, M. J." box="[733,993,1115,1141]" pageId="3" pageNumber="550" pagination="317 - 342" refId="ref12122" refString="Soares, H. E. M. & Bauab, M. J. (1972) Contribuicao ao estudo dos Opilioes do Peru (Opiliones: Gonyleptidae). Acta Zoologica Lilloana, 29, 317 - 342." type="journal article" year="1972">Soares & Bauab, 1972</bibRefCitation>
|
|||
|
|
)
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="92F69E849F60FF8573CEFBE4FA8FFBDB" ID-CoL="7QH8T" authority="Acosta, 2002 a" authorityName="Acosta" authorityYear="2002" box="[1014,1295,1115,1141]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="mitmaj">
|
|||
|
|
<emphasis id="678239159F60FF8573CEFBE4FBE4FBDA" box="[1014,1124,1115,1141]" italics="true" pageId="3" pageNumber="550">A. mitmaj</emphasis>
|
|||
|
|
<bibRefCitation id="316798F69F60FF857453FBE4FA8FFBDB" author="Acosta, L. E." box="[1131,1295,1115,1141]" pageId="3" pageNumber="550" pagination="78 - 83" refId="ref10656" refString="Acosta, L. E. (2002 a) Acrographinotus mitmaj, a new harvestman species from central Peru (Opiliones, Gonyleptidae, Pachylinae). The Journal of Arachnology, 30 (1), 78 - 83. https: // doi. org / 10.1636 / 0161 - 8202 (2002) 030 [0078: AMANHS] 2.0. CO; 2" type="journal article" year="2002">Acosta, 2002a</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="92F69E849F60FF857524FBE4FEAEFB37" ID-CoL="7QH8V" authority="Acosta, 2001" authorityName="Acosta" authorityYear="2001" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="niawpaq">
|
|||
|
|
<emphasis id="678239159F60FF857524FBE4FA1CFBDA" box="[1308,1436,1115,1141]" italics="true" pageId="3" pageNumber="550">A. niawpaq</emphasis>
|
|||
|
|
<bibRefCitation id="316798F69F60FF8570AFFBC0FEAEFB37" author="Acosta, L. E." box="[151,302,1151,1177]" pageId="3" pageNumber="550" pagination="58 - 66" refId="ref10602" refString="Acosta, L. E. (2001) The identity of Acrographinotus erectispina, with a review of the generic diagnosis, and the description of a new species (Opiliones, Gonyleptidae, Pachylinae). Bulletin of the British Arachnological Society, 12 (2), 58 - 66." type="journal article" year="2001">Acosta, 2001</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="92F69E849F60FF857101FBC0FDDFFB37" ID-CoL="7QH8W" authority="(Roewer, 1952)" baseAuthorityName="Roewer" baseAuthorityYear="1952" box="[313,607,1150,1177]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="opacus">
|
|||
|
|
<emphasis id="678239159F60FF857101FBC0FE27FB36" box="[313,423,1151,1177]" italics="true" pageId="3" pageNumber="550">A. opacus</emphasis>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="316798F69F60FF85718EFBC0FDD8FB37" author="Roewer, C. F." box="[438,600,1150,1177]" pageId="3" pageNumber="550" pagination="37 - 58" refId="ref12017" refString="Roewer, C. F. (1952) Neotropische Arachnida Arthrogastra, zumeist aus Peru [I]. Senckenbergiana, 33 (1 / 3), 37 - 58." type="journal article" year="1952">Roewer, 1952</bibRefCitation>
|
|||
|
|
)
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="92F69E849F60FF857253FBC0FD71FB36" authorityName="Acosta" authorityYear="2024" box="[619,753,1151,1177]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="tariquiae" status="sp. nov.">
|
|||
|
|
<emphasis id="678239159F60FF857253FBC0FD71FB36" box="[619,753,1151,1177]" italics="true" pageId="3" pageNumber="550">A. tariquiae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="678239159F60FF8572C0FBC0FCD1FB37" bold="true" box="[760,849,1151,1177]" pageId="3" pageNumber="550">
|
|||
|
|
<taxonomicNameLabel id="7CB1846E9F60FF8572C0FBC0FCD1FB37" box="[760,849,1151,1177]" pageId="3" pageNumber="550" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="92F69E849F60FF857367FBC0FC66FB37" authorityName="Acosta" authorityYear="2024" box="[863,998,1151,1177]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="calilegua" status="sp. nov.">
|
|||
|
|
<emphasis id="678239159F60FF857367FBC0FC66FB37" box="[863,998,1151,1177]" italics="true" pageId="3" pageNumber="550">A. calilegua</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="678239159F60FF8573D6FBC0FBC7FB37" bold="true" box="[1006,1095,1151,1177]" pageId="3" pageNumber="550">
|
|||
|
|
<taxonomicNameLabel id="7CB1846E9F60FF8573D6FBC0FBC7FB37" box="[1006,1095,1151,1177]" pageId="3" pageNumber="550" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="1DECB68C9F60FF8570FFFB1CFD02FA73" pageId="3" pageNumber="550" type="discussion">
|
|||
|
|
<paragraph id="5549E5079F60FF8570FFFB1CFD02FA73" blockId="3.[151,1437,1006,2041]" pageId="3" pageNumber="550">
|
|||
|
|
<emphasis id="678239159F60FF8570FFFB1CFEBBFB13" bold="true" box="[199,315,1187,1213]" pageId="3" pageNumber="550">Remarks.</emphasis>
|
|||
|
|
The most recent incorporation into the genus is
|
|||
|
|
<taxonomicName id="92F69E849F60FF857340FB1CFC6AFB12" baseAuthorityName="Roewer" baseAuthorityYear="1952" box="[888,1002,1187,1213]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="opacus">
|
|||
|
|
<emphasis id="678239159F60FF857340FB1CFC6AFB12" box="[888,1002,1187,1213]" italics="true" pageId="3" pageNumber="550">A. opacus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, originally described in the Brazilian genus
|
|||
|
|
<taxonomicName id="92F69E849F60FF8570DAFB78FDD5FB4F" authority="Mello-Leitao, 1932" authorityName="Mello-Leitao" authorityYear="1932" box="[226,597,1223,1249]" class="Arachnida" family="Gonyleptidae" genus="Liogyndulus" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="678239159F60FF8570DAFB78FEECFB4F" box="[226,364,1223,1249]" italics="true" pageId="3" pageNumber="550">Liogyndulus</emphasis>
|
|||
|
|
<bibRefCitation id="316798F69F60FF85714EFB78FDD5FB4F" author="Mello-Leitao, C. F. de" box="[374,597,1223,1249]" pageId="3" pageNumber="550" pagination="1 - 505" refId="ref11729" refString="Mello-Leitao, C. F. de (1932) Opilioes do Brasil. Revista do Museu Paulista, 17 (2), 1 - 505, 61 pls." type="journal article" year="1932">Mello-Leitão, 1932</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. As
|
|||
|
|
<bibRefCitation id="316798F69F60FF8572B1FB78FC56FB4F" author="Soares, H. E. M. & Soares, B. A. M." box="[649,982,1223,1249]" pageId="3" pageNumber="550" pagination="301 - 314" refId="ref12163" refString="Soares, H. E. M. & Soares, B. A. M. (1984) Opera opiliologica varia. XXV. (Opiliones, Gonyleptidae). Revista Brasileira de Entomologia, 28 (3), 301 - 314." type="journal article" year="1984">Soares & Soares (1984: 309)</bibRefCitation>
|
|||
|
|
synonymyzed the latter under
|
|||
|
|
<taxonomicName id="92F69E849F60FF857504FB78FEB7FAAB" authority="Roewer, 1913" authorityName="Roewer" authorityYear="1913" class="Arachnida" family="Gonyleptidae" genus="Cobania" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="678239159F60FF857504FB78FA1CFB4F" box="[1340,1436,1223,1249]" italics="true" pageId="3" pageNumber="550">Cobania</emphasis>
|
|||
|
|
<bibRefCitation id="316798F69F60FF8570AFFB54FEB7FAAB" author="Roewer, C. F." box="[151,311,1259,1285]" pageId="3" pageNumber="550" pagination="1 - 256" refId="ref11860" refString="Roewer, C. F. (1913) Die Familie der Gonyleptiden der Opiliones-Laniatores. Archiv fur Naturgeschichte, 79 A (4), 1 - 256, pl. 1 a." type="journal article" year="1913">Roewer, 1913</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, the combination
|
|||
|
|
<taxonomicName id="92F69E849F60FF857238FB54FD2DFAAA" box="[512,685,1259,1285]" class="Arachnida" family="Gonyleptidae" genus="Cobania" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="opaca">
|
|||
|
|
<emphasis id="678239159F60FF857238FB54FD2DFAAA" box="[512,685,1259,1285]" italics="true" pageId="3" pageNumber="550">Cobania opaca</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
was implied (
|
|||
|
|
<bibRefCitation id="316798F69F60FF857375FB54FC60FAAB" author="Acosta, L. E." box="[845,992,1259,1285]" pageId="3" pageNumber="550" pagination="209 - 225" refId="ref10563" refString="Acosta, L. E. (1996) Die Typus-Exemplare der von Carl-Friedrich Roewer beschriebenen Pachylinae (Arachnida: Opiliones: Gonyleptidae). Senckenbergiana Biologica, 76 (1 / 2), 209 - 225." type="journal article" year="1996">Acosta 1996</bibRefCitation>
|
|||
|
|
), though never explicit.
|
|||
|
|
<emphasis id="678239159F60FF8574CBFB54FF66FA86" italics="true" pageId="3" pageNumber="550">
|
|||
|
|
“
|
|||
|
|
<taxonomicName id="92F69E849F60FF8574C5FB54FF66FA86" authorityName="Roewer" authorityYear="1952" class="Arachnida" family="Gonyleptidae" genus="Liogyndulus" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="opacus">Liogyndulus” opacus</taxonomicName>
|
|||
|
|
</emphasis>
|
|||
|
|
was listed by
|
|||
|
|
<bibRefCitation id="316798F69F60FF8571B3FAB0FDD1FA87" author="Kury, A. B." box="[395,593,1294,1321]" pageId="3" pageNumber="550" pagination="5 - 337" refId="ref11368" refString="Kury, A. B. (2003) Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, Volumen Especial Monografico, 1, 5 - 337." type="journal article" year="2003">Kury (2003: 153)</bibRefCitation>
|
|||
|
|
as
|
|||
|
|
<taxonomicName id="92F69E849F60FF857241FAB0FD74FA87" authorityName="Sorensen" authorityYear="1884" box="[633,756,1295,1321]" class="Arachnida" family="Gonyleptidae" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="subFamily" subFamily="Pachylinae">Pachylinae</taxonomicName>
|
|||
|
|
<emphasis id="678239159F60FF8572C4FAB0FC1EFA87" box="[764,926,1295,1321]" italics="true" pageId="3" pageNumber="550">incertae sedis,</emphasis>
|
|||
|
|
because “this species has nothing to do with
|
|||
|
|
<emphasis id="678239159F60FF8570AFFA8CFE87FAE3" box="[151,263,1331,1357]" italics="true" pageId="3" pageNumber="550">
|
|||
|
|
<taxonomicName id="92F69E849F60FF8570AFFA8CFF7DFAE3" authorityName="Roewer" authorityYear="1913" box="[151,253,1331,1357]" class="Arachnida" family="Gonyleptidae" genus="Cobania" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="genus">Cobania</taxonomicName>
|
|||
|
|
”
|
|||
|
|
</emphasis>
|
|||
|
|
. Finally,
|
|||
|
|
<bibRefCitation id="316798F69F60FF857148FA8CFD4FFAE3" author="Kury, A. B. & Pinto-da-Rocha, R." box="[368,719,1331,1357]" pageId="3" pageNumber="550" refId="ref11461" refString="Kury, A. B. & Pinto-da-Rocha, R. (2020) On two false South American Liogyndulus. In: Kury, A. B., Mendes, A. C., Cardoso, L., Kury, M. S. & Granado, A. de A. (Eds.), WCO-Lite: Online World Catalogue of Harvestmen (Arachnida, Opiliones). Version 1.0 - Checklist of all valid nomina in Opiliones with authors and dates of publication up to 2018. Self published, Rio de Janeiro, p. 57." type="journal volume" year="2020">Kury & Pinto-da-Rocha (2020)</bibRefCitation>
|
|||
|
|
placed it in
|
|||
|
|
<taxonomicName id="92F69E849F60FF857360FA8CFB95FAE3" authorityName="Holmgren" authorityYear="1917" box="[856,1045,1331,1357]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="678239159F60FF857360FA8CFB95FAE3" box="[856,1045,1331,1357]" italics="true" pageId="3" pageNumber="550">Acrographinotus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, an action upheld by the diagnoses match (details not given, however) and biogeographic considerations (an Andean species is unlikely to belong to a Brazilian genus). Since the
|
|||
|
|
<typeStatus id="8A4D5BA59F60FF8571F3FAC4FD86FA3B" box="[459,518,1403,1429]" pageId="3" pageNumber="550">types</typeStatus>
|
|||
|
|
and only known specimens of
|
|||
|
|
<taxonomicName id="92F69E849F60FF857359FAC4FBC1FA3A" authorityName="Roewer" authorityYear="1952" box="[865,1089,1403,1429]" class="Arachnida" family="Gonyleptidae" genus="Liogyndulus" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="opacus">
|
|||
|
|
<emphasis id="678239159F60FF857359FAC4FBC1FA3A" box="[865,1089,1403,1429]" italics="true" pageId="3" pageNumber="550">Liogyndulus opacus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
are
|
|||
|
|
<specimenCount id="43F02E8E9F60FF857449FAC4FB79FA3B" box="[1137,1273,1403,1429]" count="2" pageId="3" pageNumber="550" type="female">two females</specimenCount>
|
|||
|
|
and a juvenile (
|
|||
|
|
<bibRefCitation id="316798F69F60FF8570A7FA20FEB1FA17" author="Acosta, L. E." box="[159,305,1439,1465]" pageId="3" pageNumber="550" pagination="209 - 225" refId="ref10563" refString="Acosta, L. E. (1996) Die Typus-Exemplare der von Carl-Friedrich Roewer beschriebenen Pachylinae (Arachnida: Opiliones: Gonyleptidae). Senckenbergiana Biologica, 76 (1 / 2), 209 - 225." type="journal article" year="1996">Acosta 1996</bibRefCitation>
|
|||
|
|
), while sound diagnostic features involve males (
|
|||
|
|
<bibRefCitation id="316798F69F60FF857361FA20FC6BFA17" author="Acosta, L. E." box="[857,1003,1439,1465]" pageId="3" pageNumber="550" pagination="58 - 66" refId="ref10602" refString="Acosta, L. E. (2001) The identity of Acrographinotus erectispina, with a review of the generic diagnosis, and the description of a new species (Opiliones, Gonyleptidae, Pachylinae). Bulletin of the British Arachnological Society, 12 (2), 58 - 66." type="journal article" year="2001">Acosta 2001</bibRefCitation>
|
|||
|
|
), this assignment should be taken with caution and would require further evidence.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="1DECB68C9F60FF8570FFFA59FEC8F9E7" pageId="3" pageNumber="550" type="distribution">
|
|||
|
|
<paragraph id="5549E5079F60FF8570FFFA59FEC8F9E7" blockId="3.[151,1437,1006,2041]" pageId="3" pageNumber="550">
|
|||
|
|
<emphasis id="678239159F60FF8570FFFA59FEE1F9AE" bold="true" box="[199,353,1510,1536]" pageId="3" pageNumber="550">Distribution.</emphasis>
|
|||
|
|
Higher altitudinal belts on Andean and sub-Andean ranges of
|
|||
|
|
<collectingCountry id="2DE1A5979F60FF85743FFA58FBBDF9AF" box="[1031,1085,1511,1537]" name="Peru" pageId="3" pageNumber="550">Perú</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="2DE1A5979F60FF85747EFA58FB18F9AF" box="[1094,1176,1511,1537]" name="Bolivia" pageId="3" pageNumber="550">Bolivia</collectingCountry>
|
|||
|
|
and northern
|
|||
|
|
<collectingCountry id="2DE1A5979F60FF857515FA58FA1CF9AF" box="[1325,1436,1511,1537]" name="Argentina" pageId="3" pageNumber="550">Argentina</collectingCountry>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="CDCDF9829F60FF8570A7F9B4FF62F98B" box="[159,226,1547,1573]" captionStart="FIGURE 1" captionStartId="1.[151,250,1772,1796]" captionTargetBox="[151,1432,181,1744]" captionTargetId="figure-15@1.[151,1435,181,1749]" captionTargetPageId="1" captionText="FIGURE 1. An overview of the known records of genus Acrographinotus (red dots), spread between 5°55’S and 23°41’S along Andean and sub-Andean ranges of Perú, Bolivia and northwestern Argentina (ARG.). The blue arrow points the ‘Andean elbow’ at around 18°S. References for localities cited in the text: 1: Route Olmos-Jaén (presumed doubtful record); 2: Yánac; 3: Unduavi; 4: records in Tarija Department. Inset (left): location of the represented area in South America. Yellow square: area enlarged on Fig. 7A." figureDoi="http://doi.org/10.5281/zenodo.12783060" httpUri="https://zenodo.org/record/12783060/files/figure.png" pageId="3" pageNumber="550">Fig. 1</figureCitation>
|
|||
|
|
); elevation range
|
|||
|
|
<quantity id="920E48E29F60FF85718AF9B5FDCEF98B" box="[434,590,1546,1573]" metricMagnitude="3" metricUnit="m" metricValue="3.2049999999999996" metricValueMax="4.75" metricValueMin="1.66" pageId="3" pageNumber="550" unit="m" value="3205.0" valueMax="4750.0" valueMin="1660.0">1660–4750 m</quantity>
|
|||
|
|
a.s.l. (~29% of records from above
|
|||
|
|
<quantity id="920E48E29F60FF8573D2F9B4FBC4F98B" box="[1002,1092,1547,1573]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="550" unit="m" value="4000.0">4000 m</quantity>
|
|||
|
|
; 77% above
|
|||
|
|
<quantity id="920E48E29F60FF8574EFF9B4FAB0F98B" box="[1239,1328,1547,1573]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="550" unit="m" value="3000.0">3000 m</quantity>
|
|||
|
|
; only 5% below
|
|||
|
|
<quantity id="920E48E29F60FF8570DBF990FEBCF9E7" box="[227,316,1583,1609]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="550" unit="m" value="2000.0">2000 m</quantity>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="1DECB68C9F60FF8270FFF9EDFF60FE27" lastPageId="4" lastPageNumber="551" pageId="3" pageNumber="550" type="diagnosis">
|
|||
|
|
<paragraph id="5549E5079F60FF8270FFF9EDFF60FE27" blockId="3.[151,1437,1006,2041]" lastBlockId="4.[151,1437,151,393]" lastPageId="4" lastPageNumber="551" pageId="3" pageNumber="550">
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<emphasis id="678239159F60FF8570FFF9EDFE1AF9C3" bold="true" box="[199,410,1618,1645]" pageId="3" pageNumber="550">Diagnosis update.</emphasis>
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Ocular mound with a median apophysis, either tall or low, acute or blunt; in species with low armature (
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<emphasis id="678239159F60FF857132F9C8FEB2F93E" box="[266,306,1655,1680]" italics="true" pageId="3" pageNumber="550">e.g.</emphasis>
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,
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<taxonomicName id="92F69E849F60FF857178F9C8FE2CF93E" authorityName="Acosta" authorityYear="2002" box="[320,428,1655,1681]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="mitmaj">
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<emphasis id="678239159F60FF857178F9C8FE2CF93E" box="[320,428,1655,1681]" italics="true" pageId="3" pageNumber="550">A. mitmaj</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="92F69E849F60FF857181F9C8FD92F93E" baseAuthorityName="Roewer" baseAuthorityYear="1957" box="[441,530,1655,1681]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="ortizi">
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<emphasis id="678239159F60FF857181F9C8FD92F93E" box="[441,530,1655,1681]" italics="true" pageId="3" pageNumber="550">A. ortizi</emphasis>
|
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</taxonomicName>
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), intraspecific variation may span from a short apophysis to completely unarmed, even with a median depression or notch instead, giving the false appearance of a paired condition. DS in most cases unarmed, be it smooth or with barely scattered granulation or very small round tubercles; only exceptionally (
|
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<taxonomicName id="92F69E849F60FF8575BCF900FF6DF952" baseAuthorityName="Roewer" baseAuthorityYear="1929" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="ornatus">
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<emphasis id="678239159F60FF8575BCF900FF6DF952" italics="true" pageId="3" pageNumber="550">A. ornatus</emphasis>
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</taxonomicName>
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) tubercles are large in all areas, pustule-like (males) or round-flat (females). Pp femur without a medial subapical spine. Tarsal formula 6:8–10:7:7. Males with FT-III armed by a single median apophysis, sometimes with a row of granules or (seldom) small apophyses on each side. Free tergites I–II and dorsal anal plate unarmed. VAP either completely unarmed, just granulous, or armed with a pair of horn-like apophyses, be it independent or fused to a shelf-like tegumentary projection in between (in
|
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<taxonomicName id="92F69E849F60FF8572CEF8CCFCFEF823" authorityName="Acosta" authorityYear="2024" box="[758,894,1907,1933]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="calilegua" status="sp. nov.">
|
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<emphasis id="678239159F60FF8572CEF8CCFCFEF823" box="[758,894,1907,1933]" italics="true" pageId="3" pageNumber="550">A. calilegua</emphasis>
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</taxonomicName>
|
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<emphasis id="678239159F60FF8573BFF8CCFC61F823" bold="true" box="[903,993,1907,1933]" pageId="3" pageNumber="550">
|
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<taxonomicNameLabel id="7CB1846E9F60FF8573BFF8CCFC61F823" box="[903,993,1907,1933]" pageId="3" pageNumber="550" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
|
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only the latter remains). Coxa IV with a short prolateral apophysis, blunt or truncate, mostly transverse or slightly diagonal; it is rarely large and sigmoid (
|
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<taxonomicName id="92F69E849F60FF857099F804FE98F87A" baseAuthorityName="Roewer" baseAuthorityYear="1929" box="[161,280,1979,2005]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="ornatus">
|
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<emphasis id="678239159F60FF857099F804FE98F87A" box="[161,280,1979,2005]" italics="true" pageId="3" pageNumber="550">A. ornatus</emphasis>
|
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</taxonomicName>
|
|||
|
|
). Trochanter IV sub-trapezoid, articulated obliquely outwards; it is most frequently asymmetric (the prolateral side shorter than the retrolateral one); exceptions are
|
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<taxonomicName id="92F69E849F60FF85735AF860FC58F856" baseAuthorityName="Roewer" baseAuthorityYear="1929" box="[866,984,2015,2041]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="ornatus">
|
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<emphasis id="678239159F60FF85735AF860FC58F856" box="[866,984,2015,2041]" italics="true" pageId="3" pageNumber="550">A. ornatus</emphasis>
|
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</taxonomicName>
|
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and
|
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<taxonomicName id="92F69E849F60FF857437F860FBFDF856" authorityName="Acosta" authorityYear="2002" box="[1039,1149,2015,2041]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="mitmaj">
|
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<emphasis id="678239159F60FF857437F860FBFDF856" box="[1039,1149,2015,2041]" italics="true" pageId="3" pageNumber="550">A. mitmaj</emphasis>
|
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</taxonomicName>
|
|||
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, with trochanter IV more symmetric in shape and articulation fairly straight. FeIV straight or arched, in most species with longitudinal rows of granules or blunt tubercles; in some cases (
|
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<emphasis id="678239159F67FF8272A2FF04FB93FF7A" box="[666,1043,187,213]" italics="true" pageId="4" pageNumber="551">
|
|||
|
|
<taxonomicName id="92F69E849F67FF8272A2FF04FCB2FF7A" authorityName="Roewer" authorityYear="1929" box="[666,818,187,213]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="551" phylum="Arthropoda" rank="species" species="curvispina">A. curvispina</taxonomicName>
|
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,
|
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|
|
<taxonomicName id="92F69E849F67FF827305FF04FC1BFF7A" baseAuthorityName="Roewer" baseAuthorityYear="1957" box="[829,923,187,213]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="551" phylum="Arthropoda" rank="species" species="ortizi">A. ortizi</taxonomicName>
|
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,
|
|||
|
|
<taxonomicName id="92F69E849F67FF82739DFF04FB93FF7A" authorityName="Acosta" authorityYear="2002" box="[933,1043,187,213]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="551" phylum="Arthropoda" rank="species" species="mitmaj">A. mitmaj</taxonomicName>
|
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|
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</emphasis>
|
|||
|
|
) granules and apophysis are acute; only exceptionally spine-like apophyses are very large (
|
|||
|
|
<taxonomicName id="92F69E849F67FF82733FFF60FCFCFF56" baseAuthorityName="Roewer" baseAuthorityYear="1929" box="[775,892,223,249]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="551" phylum="Arthropoda" rank="species" species="ornatus">
|
|||
|
|
<emphasis id="678239159F67FF82733FFF60FCFCFF56" box="[775,892,223,249]" italics="true" pageId="4" pageNumber="551">A. ornatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
). Penis: distal end of VPS dilated and armed with a downward-pointing process; on top of the dilated portion there is, with a few exceptions, a membranous projection (the VPS has been described to resemble a ‘round-crested ibis head’). Besides the armature of ocular mound, DS and FT-III, the feature, VPS shape is the best defining character of
|
|||
|
|
<taxonomicName id="92F69E849F67FF82736EFEF4FB93FECB" authorityName="Holmgren" authorityYear="1917" box="[854,1043,331,357]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="551" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="678239159F67FF82736EFEF4FB93FECB" box="[854,1043,331,357]" italics="true" pageId="4" pageNumber="551">Acrographinotus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
among
|
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|
|
<taxonomicName id="92F69E849F67FF827457FEF4FB6AFECB" authorityName="Sorensen" authorityYear="1884" box="[1135,1258,331,357]" class="Arachnida" family="Gonyleptidae" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="551" phylum="Arthropoda" rank="subFamily" subFamily="Pachylinae">Pachylinae</taxonomicName>
|
|||
|
|
genera (
|
|||
|
|
<bibRefCitation id="316798F69F67FF827575FEF4FF53FE27" author="Acosta, L. E." pageId="4" pageNumber="551" pagination="58 - 66" refId="ref10602" refString="Acosta, L. E. (2001) The identity of Acrographinotus erectispina, with a review of the generic diagnosis, and the description of a new species (Opiliones, Gonyleptidae, Pachylinae). Bulletin of the British Arachnological Society, 12 (2), 58 - 66." type="journal article" year="2001">Acosta 2001</bibRefCitation>
|
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|
|
).
|
|||
|
|
</paragraph>
|
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|
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</subSubSection>
|
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</treatment>
|
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</document>
|