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<document id="A17613383E03D261FC1637EE4501393E" ID-DOI="10.5281/zenodo.193717" ID-GBIF-Dataset="ecbfc9d0-d04b-4599-b972-cf6898ce8521" ID-ISSN="1175-5326" ID-Zenodo-Dep="193717" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460168505526" checkinUser="plazi" docAuthor="Wowor, Daisy &amp; Ng, Peter K. L." docDate="2010" docId="03ACA30FE510FFFAFF17FDB66F74FAE4" docLanguage="en" docName="zt02372p052.pdf" docOrigin="Zootaxa 2372" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Arachnochium Wowor &amp; Ng, 2010, gen. nov." docType="treatment" docVersion="10" lastPageNumber="39" masterDocId="FF95DB77E511FFF8FF80FFC46E5BFFD3" masterDocTitle="On two new genera of Asian prawns previously assigned to Macrobrachium (Crustacea: Decapoda: Caridea: Palaemonidae) *" masterLastPageNumber="52" masterPageNumber="37" pageNumber="38" updateTime="1698235898136" updateUser="plazi">
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<mods:title id="8C8486DD8BE9EB819DD984DD313467B1">On two new genera of Asian prawns previously assigned to Macrobrachium (Crustacea: Decapoda: Caridea: Palaemonidae) *</mods:title>
</mods:titleInfo>
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<mods:namePart id="72C880710436ACF7096C3036F7CBC6A2">Wowor, Daisy</mods:namePart>
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<mods:namePart id="0EBD0AD64562A34966EFD02CF3BAC675">Ng, Peter K. L.</mods:namePart>
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<mods:typeOfResource id="2E68E10F7BB95A81133BC672AC7F7E34">text</mods:typeOfResource>
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<mods:title id="DFC282B8A7A39620C19951C79E6A9C7A">Zootaxa</mods:title>
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<mods:date id="A1107C36CDD7D6B839F45178A8EAAACB">2010</mods:date>
<mods:detail id="8FD2DAD994C8F19DF1D5A5AB9F2EE711" type="volume">
<mods:number id="629ACF11BACB8E156E7D452FECB8387F">2372</mods:number>
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<mods:classification id="DB80CA3A2EEEA1181E4985DFD8463557">journal article</mods:classification>
<mods:identifier id="EC4057F6C23DC49E5CFBCA0CCACFAC19" type="DOI">10.5281/zenodo.193717</mods:identifier>
<mods:identifier id="7B00778E4A536F2EC81CE471D2E45787" type="GBIF-Dataset">ecbfc9d0-d04b-4599-b972-cf6898ce8521</mods:identifier>
<mods:identifier id="5B69BA2B10AD2AD9EF418650530F2AC0" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="35469BFE5A0D97CF53FAE3DB55C1325A" type="Zenodo-Dep">193717</mods:identifier>
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<treatment id="03ACA30FE510FFFAFF17FDB66F74FAE4" ID-DOI="http://doi.org/10.5281/zenodo.5259969" ID-GBIF-Taxon="119383406" ID-Zenodo-Dep="5259969" LSID="urn:lsid:plazi:treatment:03ACA30FE510FFFAFF17FDB66F74FAE4" httpUri="http://treatment.plazi.org/id/03ACA30FE510FFFAFF17FDB66F74FAE4" lastPageId="2" lastPageNumber="39" pageId="1" pageNumber="38">
<subSubSection id="C31F4192E510FFF9FF17FDB66F99FD5F" box="[151,450,626,652]" pageId="1" pageNumber="38" type="nomenclature">
<paragraph id="8BBA1219E510FFF9FF17FDB66F99FD5F" blockId="1.[151,450,626,652]" box="[151,450,626,652]" pageId="1" pageNumber="38">
<heading id="D0F2A575E510FFF9FF17FDB66F99FD5F" bold="true" box="[151,450,626,652]" fontSize="11" level="1" pageId="1" pageNumber="38" reason="1">
<emphasis id="B971CE0BE510FFF9FF17FDB66F99FD5F" bold="true" box="[151,450,626,652]" pageId="1" pageNumber="38">
<taxonomicName id="4C05699AE510FFF9FF17FDB66F15FD5F" ID-CoL="fc25e5c8-caec-415b-93f5-17d4f6fa4bd2" box="[151,334,626,652]" class="Malacostraca" family="Palaemonidae" genus="Arachnochium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="38" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B971CE0BE510FFF9FF17FDB66F15FD5F" bold="true" box="[151,334,626,652]" italics="true" pageId="1" pageNumber="38">Arachnochium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2427370E510FFF9FED4FDB66F99FD5F" box="[340,450,626,652]" pageId="1" pageNumber="38" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31F4192E510FFF9FF17FD7E6D84FD07" box="[151,991,698,724]" pageId="1" pageNumber="38" type="materials_examined">
<paragraph id="8BBA1219E510FFF9FF17FD7E6D84FD07" blockId="1.[151,1437,698,2031]" box="[151,991,698,724]" pageId="1" pageNumber="38">
<emphasis id="B971CE0BE510FFF9FF17FD7E6F62FD07" bold="true" box="[151,313,698,724]" pageId="1" pageNumber="38">
<typeStatus id="54BEACBBE510FFF9FF17FD7E6E8FFD07" box="[151,212,698,724]" pageId="1" pageNumber="38">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C05699AE510FFF9FEC0FD7E6C98FD07" authority="Kemp, 1917" authorityName="Kemp" authorityYear="1917" box="[320,707,698,724]" class="Malacostraca" family="Palaemonidae" genus="Palaemon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="B971CE0BE510FFF9FEC0FD7E6C73FD00" box="[320,552,698,723]" italics="true" pageId="1" pageNumber="38">Palaemon mirabilis</emphasis>
<bibRefCitation id="EF946FE8E510FFF9FDB0FD7E6C98FD07" author="Kemp" box="[560,707,698,724]" pageId="1" pageNumber="51" refString="Kemp, S. (1917) Notes on Crustacea Decapoda in the Indian Museum, IX: Leander styliferus, Milne-Edwards, and related forms. Records of the Indian Museum, 13 (4), 203 - 231, pls. 8 - 10." type="journal article" year="1917">Kemp, 1917</bibRefCitation>
</taxonomicName>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="C31F4192E510FFF9FF46FD1B6CEAF9A9" pageId="1" pageNumber="38" type="diagnosis">
<paragraph id="8BBA1219E510FFF9FF46FD1B6CEAF9A9" blockId="1.[151,1437,698,2031]" pageId="1" pageNumber="38">
<emphasis id="B971CE0BE510FFF9FF46FD1B6F1DFD2A" bold="true" box="[198,326,735,761]" pageId="1" pageNumber="38">Diagnosis.</emphasis>
Small-sized prawn of subcylindrical body form. Rostrum well developed, compressed; dorsal carina armed with 14–17 teeth, 4 or 5 teeth completely postorbital; ventral carina armed with at least 1 tooth; plumose setae present between teeth; lateral carina well developed. Carapace with antennal and hepatic spines; branchiostegal groove present, running from base of hepatic spine to antennal carapace margin below antennal spine. Antennule with 3-segmented peduncle, ventromedial tooth on basal segment small; distolateral angle of basal segment well developed; 2 flagella present, upper flagellum biramous with rami fused proximally. Antenna with basicerite unarmed, 1 flagellum present; scaphocerite well developed, distal lamina clearly extending beyond distolateral tooth. Ocular beak well developed. Epistome bilobed. Mouthparts similar to those of
<taxonomicName id="4C05699AE510FFF9FD92FBCE6C8EFBF0" box="[530,725,1034,1059]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE510FFF9FD92FBCE6C8EFBF0" box="[530,725,1034,1059]" italics="true" pageId="1" pageNumber="38">Macrobrachium</emphasis>
</taxonomicName>
; mandible with 3-segmented palp. Second pereiopod very slender, about 1.6 times as wide as first pereiopod, similar in shape, equal in size, maximum length twice as long as carapace. Carpus slightly tapered. All segments glabrous. Pereiopods equally developed in both sexes. Third, fourth and fifth pereiopods glabrous; third pereiopod shorter than second pereipod, fourth pereiopod longer than second pereiopod, fifth pereiopod distinctly longer than fourth pereiopod; third and fourth pereiopods with 5 or 6 ventral spines distributed along length of propodus but without distal paired spines; fifth pereiopod with 8 ventral spines along length of propodus, transverse rows of setae present at outer margin of distoventral propodus. T5 without transverse plate; male T8 without anterior lobes, with median process posteriorly. Fifth segment of abdominal pleura with bilobed posterolateral margin. First pleopod without appendix interna and masculina, endopod smaller than exopod, kidney-shaped, exopod long, slender; second pleopod with appendix interna and masculina in males, appendix masculina slightly longer than appendix interna, exopod and endopod long and slender; third, fourth and fifth pleopods with appendix interna only, exopod and endopod long, slender. Inter-uropodal sclerite not developed, preanal carina absent. Telson elongated, slender, glabrous, 2 pairs of dorsal spines present; median projection of posterior margin shorter than inner pair of posterior spines, inner pair of spines longer than outer pair, with most 2 pairs of long plumose subventral setae, longer than inner pair of spines. Uropods glabrous; exopod with mobile mesial spine, distinctly longer than distolateral tooth.
</paragraph>
</subSubSection>
<subSubSection id="C31F4192E510FFF9FF46F9416BC7F917" pageId="1" pageNumber="38" type="etymology">
<paragraph id="8BBA1219E510FFF9FF46F9416BC7F917" blockId="1.[151,1437,698,2031]" pageId="1" pageNumber="38">
<emphasis id="B971CE0BE510FFF9FF46F9416F08F94C" bold="true" box="[198,339,1669,1695]" pageId="1" pageNumber="38">Etymology.</emphasis>
The genus name
<taxonomicName id="4C05699AE510FFF9FDA6F9416C89F94D" box="[550,722,1669,1694]" class="Malacostraca" family="Palaemonidae" genus="Arachnochium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE510FFF9FDA6F9416C89F94D" box="[550,722,1669,1694]" italics="true" pageId="1" pageNumber="38">Arachnochium</emphasis>
</taxonomicName>
is an arbitrary combination of the Greek words
<emphasis id="B971CE0BE510FFF9FA91F9416B2AF94D" box="[1297,1393,1669,1694]" italics="true" pageId="1" pageNumber="38">arachne</emphasis>
for spider and
<emphasis id="B971CE0BE510FFF9FE97F96F6FDAF917" box="[279,385,1707,1732]" italics="true" pageId="1" pageNumber="38">brachion</emphasis>
for leg, alluding to the unusually long pereiopods, like those of a spider. Gender is neuter.
</paragraph>
</subSubSection>
<subSubSection id="C31F4192E510FFFAFF46F90B6F74FAE4" lastPageId="2" lastPageNumber="39" pageId="1" pageNumber="38" type="discussion">
<paragraph id="8BBA1219E510FFF9FF46F90B6F5AF8E7" blockId="1.[151,1437,698,2031]" pageId="1" pageNumber="38">
<emphasis id="B971CE0BE510FFF9FF46F90B6F1BF93A" bold="true" box="[198,320,1743,1769]" pageId="1" pageNumber="38">Remarks.</emphasis>
<taxonomicName id="4C05699AE510FFF9FEC7F9146C6BF93A" box="[327,560,1744,1769]" class="Malacostraca" family="Palaemonidae" genus="Palaemon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="B971CE0BE510FFF9FEC7F9146C6BF93A" box="[327,560,1744,1769]" italics="true" pageId="1" pageNumber="38">Palaemon mirabilis</emphasis>
</taxonomicName>
was first placed in
<taxonomicName id="4C05699AE510FFF9FC98F9146A04F939" authority="Bate, 1868" authorityName="Bate" authorityYear="1868" box="[792,1119,1744,1770]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE510FFF9FC98F9146D8EF93A" box="[792,981,1744,1769]" italics="true" pageId="1" pageNumber="38">Macrobrachium</emphasis>
<bibRefCitation id="EF946FE8E510FFF9FC5DF9146A04F939" author="Bate" box="[989,1119,1744,1770]" pageId="1" pageNumber="51" refString="Bate, C. S. (1868) On a new genus, with four new species, of freshwater prawns. Proceedings of the Zoological Society of London, 1868, 363 - 368, pls. 30 - 31." type="proceedings paper" year="1868">Bate, 1868</bibRefCitation>
</taxonomicName>
, by
<bibRefCitation id="EF946FE8E510FFF9FB11F9146B60F939" author="Suvatti" box="[1169,1339,1744,1770]" pageId="1" pageNumber="52" refString="Suvatti, C. (1937) A check-list of aquatic fauna in Siam (excluding Fishes), Department of Fisheries, Bangkok, Thailand, 116 pp." type="book" year="1937">Suvatti (1937)</bibRefCitation>
and has been retained there despite doubts expressed (
<bibRefCitation id="EF946FE8E510FFF9FD32F9316DEBF8DC" author="Chace" box="[690,944,1781,1807]" pageId="1" pageNumber="51" refString="Chace, F. A. Jr. &amp; Bruce, A. J. (1993) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 6: Superfamily Palaemonoidea. Smithsonian Contribution to Zoology, 543, 1 - 152." type="journal article" year="1993">Chace &amp; Bruce 1993</bibRefCitation>
). It is herein transferred to
<taxonomicName id="4C05699AE510FFF9FB6FF9316BC7F8DD" box="[1263,1436,1781,1806]" class="Malacostraca" family="Palaemonidae" genus="Arachnochium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="38" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B971CE0BE510FFF9FB6FF9316BC7F8DD" box="[1263,1436,1781,1806]" italics="true" pageId="1" pageNumber="38">Arachnochium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2427370E510FFF9FF17F8DE6F5AF8E7" box="[151,257,1818,1844]" pageId="1" pageNumber="38" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BBA1219E510FFF9FF46F8846BC7F83C" blockId="1.[151,1437,698,2031]" pageId="1" pageNumber="38">
<bibRefCitation id="EF946FE8E510FFF9FF46F8846F32F889" author="Kemp" box="[198,361,1856,1882]" pageId="1" pageNumber="51" refString="Kemp, S. (1917) Notes on Crustacea Decapoda in the Indian Museum, IX: Leander styliferus, Milne-Edwards, and related forms. Records of the Indian Museum, 13 (4), 203 - 231, pls. 8 - 10." type="journal article" year="1917">Kemp (1917)</bibRefCitation>
believed that his
<taxonomicName id="4C05699AE510FFF9FDCAF8846D6CF88A" box="[586,823,1856,1881]" class="Malacostraca" family="Palaemonidae" genus="Palaemon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="B971CE0BE510FFF9FDCAF8846D6CF88A" box="[586,823,1856,1881]" italics="true" pageId="1" pageNumber="38">Palaemon mirabilis</emphasis>
</taxonomicName>
was more closely related to species in the genus
<taxonomicName id="4C05699AE510FFF9FF17F8A16EA5F8AD" box="[151,254,1893,1918]" class="Malacostraca" family="Palaemonidae" genus="Leander" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE510FFF9FF17F8A16EA5F8AD" box="[151,254,1893,1918]" italics="true" pageId="1" pageNumber="38">Leander</emphasis>
</taxonomicName>
(= currently
<taxonomicName id="4C05699AE510FFF9FE25F8A16C25F8AC" authority="Weber" authorityName="Weber" box="[421,638,1893,1919]" class="Malacostraca" family="Palaemonidae" genus="Palaemon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE510FFF9FE25F8A16C79F8AD" box="[421,546,1893,1918]" italics="true" pageId="1" pageNumber="38">Palaemon</emphasis>
Weber
</taxonomicName>
) than to those of
<taxonomicName id="4C05699AE510FFF9FCDCF8A16D82F8AD" box="[860,985,1893,1918]" class="Malacostraca" family="Palaemonidae" genus="Palaemon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE510FFF9FCDCF8A16D82F8AD" box="[860,985,1893,1918]" italics="true" pageId="1" pageNumber="38">Palaemon</emphasis>
</taxonomicName>
(= currently
<taxonomicName id="4C05699AE510FFF9FB00F8A16BD4F8AC" authority="Bate" authorityName="Bate" box="[1152,1423,1893,1919]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE510FFF9FB00F8A16B1CF8AD" box="[1152,1351,1893,1918]" italics="true" pageId="1" pageNumber="38">Macrobrachium</emphasis>
Bate
</taxonomicName>
). However, the presence of a prominent hepatic spine led him to retain this species in
<taxonomicName id="4C05699AE510FFF9FB4BF84F6B12F877" box="[1227,1353,1931,1956]" class="Malacostraca" family="Palaemonidae" genus="Palaemon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE510FFF9FB4BF84F6B12F877" box="[1227,1353,1931,1956]" italics="true" pageId="1" pageNumber="38">Palaemon</emphasis>
</taxonomicName>
. After examining specimens from
<collectingCountry id="F3125289E510FFF9FE68F8746C0CF819" box="[488,599,1968,1994]" name="Thailand" pageId="1" pageNumber="38">Thailand</collectingCountry>
,
<bibRefCitation id="EF946FE8E510FFF9FDE4F8746D22F819" author="Chace" box="[612,889,1968,1994]" pageId="1" pageNumber="51" refString="Chace, F. A. Jr. &amp; Bruce, A. J. (1993) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 6: Superfamily Palaemonoidea. Smithsonian Contribution to Zoology, 543, 1 - 152." type="journal article" year="1993">Chace &amp; Bruce (1993)</bibRefCitation>
concluded that
<taxonomicName id="4C05699AE510FFF9FBBCF8746B76F81A" box="[1084,1325,1968,1993]" class="Malacostraca" family="Palaemonidae" genus="Palaemon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="B971CE0BE510FFF9FBBCF8746B76F81A" box="[1084,1325,1968,1993]" italics="true" pageId="1" pageNumber="38">Palaemon mirabilis</emphasis>
</taxonomicName>
does not belong to
<taxonomicName id="4C05699AE510FFF9FE8DF8116F91F83D" box="[269,458,2005,2030]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE510FFF9FE8DF8116F91F83D" box="[269,458,2005,2030]" italics="true" pageId="1" pageNumber="38">Macrobrachium</emphasis>
</taxonomicName>
, but did not belong to
<taxonomicName id="4C05699AE510FFF9FD57F8116D15F83D" box="[727,846,2005,2030]" class="Malacostraca" family="Palaemonidae" genus="Palaemon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE510FFF9FD57F8116D15F83D" box="[727,846,2005,2030]" italics="true" pageId="1" pageNumber="38">Palaemon</emphasis>
</taxonomicName>
either, because of the presence of a hepatic spine.
</paragraph>
<paragraph id="8BBA1219E513FFFAFF17FF5C6C59FE94" blockId="2.[151,1437,152,1335]" pageId="2" pageNumber="39">
They commented that this species should be assigned to a new genus, but deferred from doing so because they believed they did not have male specimens. In the present study, numerous male specimens were available, and all agreed very well with the diagnostic characters of the females. Interestingly, among the USNM material examined by
<bibRefCitation id="EF946FE8E513FFFAFE20FECC6CEAFEF1" author="Chace" box="[416,689,264,290]" pageId="2" pageNumber="51" refString="Chace, F. A. Jr. &amp; Bruce, A. J. (1993) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 6: Superfamily Palaemonoidea. Smithsonian Contribution to Zoology, 543, 1 - 152." type="journal article" year="1993">Chace &amp; Bruce (1993)</bibRefCitation>
, there was in fact a male specimen which had been incorrectly sexed (see material examined).
</paragraph>
<paragraph id="8BBA1219E513FFFAFF46FE966B1CFC11" blockId="2.[151,1437,152,1335]" pageId="2" pageNumber="39">
<taxonomicName id="4C05699AE513FFFAFF46FE966F28FEB8" box="[198,371,338,363]" class="Malacostraca" family="Palaemonidae" genus="Arachnochium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE513FFFAFF46FE966F28FEB8" box="[198,371,338,363]" italics="true" pageId="2" pageNumber="39">Arachnochium</emphasis>
</taxonomicName>
can be distinguished from
<emphasis id="B971CE0BE513FFFAFD37FE966DE7FEB8" box="[695,956,338,363]" italics="true" pageId="2" pageNumber="39">
<taxonomicName id="4C05699AE513FFFAFD37FE966D2FFEB8" box="[695,884,338,363]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus" sensu="stricto">Macrobrachium</taxonomicName>
<taxonomicNameLabel id="A2427370E513FFFAFCFDFE966DE7FEB8" box="[893,956,338,363]" pageId="2" pageNumber="39" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
by: the absence of a T5 transverse plate (present in
<taxonomicName id="4C05699AE513FFFAFEA4FEBC6FB1FE42" box="[292,490,376,401]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE513FFFAFEA4FEBC6FB1FE42" box="[292,490,376,401]" italics="true" pageId="2" pageNumber="39">Macrobrachium</emphasis>
</taxonomicName>
); absence of anterior lobes on the male T8 (present in
<taxonomicName id="4C05699AE513FFFAFB15FEBC6B01FE42" box="[1173,1370,376,401]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE513FFFAFB15FEBC6B01FE42" box="[1173,1370,376,401]" italics="true" pageId="2" pageNumber="39">Macrobrachium</emphasis>
</taxonomicName>
); the second pereiopod being very slender and glabrous (moderately slender to robust and covered with spines, spinules and scales in
<taxonomicName id="4C05699AE513FFFAFE1EFE066C00FE08" box="[414,603,450,475]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE513FFFAFE1EFE066C00FE08" box="[414,603,450,475]" italics="true" pageId="2" pageNumber="39">Macrobrachium</emphasis>
</taxonomicName>
); the second pereiopod being at most twice as long as the carapace (at least 2.5 times in
<taxonomicName id="4C05699AE513FFFAFEEBFE2C6C71FDD2" box="[363,554,488,513]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE513FFFAFEEBFE2C6C71FDD2" box="[363,554,488,513]" italics="true" pageId="2" pageNumber="39">Macrobrachium</emphasis>
</taxonomicName>
); the second pereiopod being equally developed in both sexes (larger and more developed in fully developed males in
<taxonomicName id="4C05699AE513FFFAFD1DFDC96D01FDF5" box="[669,858,525,550]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE513FFFAFD1DFDC96D01FDF5" box="[669,858,525,550]" italics="true" pageId="2" pageNumber="39">Macrobrachium</emphasis>
</taxonomicName>
); the fourth and the fifth pereiopods being longer than the second pereiopod (shorter in
<taxonomicName id="4C05699AE513FFFAFDE0FDF66D7AFD98" box="[608,801,562,587]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE513FFFAFDE0FDF66D7AFD98" box="[608,801,562,587]" italics="true" pageId="2" pageNumber="39">Macrobrachium</emphasis>
</taxonomicName>
); the fifth pereiopod being distinctly longer than the fourth pereiopod (same length in
<taxonomicName id="4C05699AE513FFFAFDABFD9C6CB7FDA2" box="[555,748,600,625]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE513FFFAFDABFD9C6CB7FDA2" box="[555,748,600,625]" italics="true" pageId="2" pageNumber="39">Macrobrachium</emphasis>
</taxonomicName>
); the propodus of the third and the fourth pereiopods are without distal paired spines (with distal paired spines in
<taxonomicName id="4C05699AE513FFFAFCA7FDB96DBFFD45" box="[807,996,637,662]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE513FFFAFCA7FDB96DBFFD45" box="[807,996,637,662]" italics="true" pageId="2" pageNumber="39">Macrobrachium</emphasis>
</taxonomicName>
); the fifth abdominal pleurite possess a bilobed posterolateral margin (unilobed in
<taxonomicName id="4C05699AE513FFFAFD25FD666D39FD68" box="[677,866,674,699]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE513FFFAFD25FD666D39FD68" box="[677,866,674,699]" italics="true" pageId="2" pageNumber="39">Macrobrachium</emphasis>
</taxonomicName>
); the ventroposterior margin of the telson has at most two pairs of plumose setae (at least eight pairs in
<taxonomicName id="4C05699AE513FFFAFCB7FD0C6DA2FD32" box="[823,1017,712,737]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE513FFFAFCB7FD0C6DA2FD32" box="[823,1017,712,737]" italics="true" pageId="2" pageNumber="39">Macrobrachium</emphasis>
</taxonomicName>
), and the adult females reaching a larger size than adult males (adult females usually distinctly smaller than adult males in
<taxonomicName id="4C05699AE513FFFAFB2CFD296B31FCD5" box="[1196,1386,749,774]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE513FFFAFB2CFD296B31FCD5" box="[1196,1386,749,774]" italics="true" pageId="2" pageNumber="39">Macrobrachium</emphasis>
</taxonomicName>
). In addition to the absence of T8 anterior lobes, the bases of the fifth pereiopods of male
<taxonomicName id="4C05699AE513FFFAFB0CFCD66B62FCF8" box="[1164,1337,786,811]" class="Malacostraca" family="Palaemonidae" genus="Arachnochium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE513FFFAFB0CFCD66B62FCF8" box="[1164,1337,786,811]" italics="true" pageId="2" pageNumber="39">Arachnochium</emphasis>
</taxonomicName>
are also widely separated compared to the condition in
<taxonomicName id="4C05699AE513FFFAFD5BFCFC6DFBFC82" box="[731,928,824,849]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE513FFFAFD5BFCFC6DFBFC82" box="[731,928,824,849]" italics="true" pageId="2" pageNumber="39">Macrobrachium</emphasis>
</taxonomicName>
species. The form of the T8, however, is typical of female
<taxonomicName id="4C05699AE513FFFAFEEAFC996C72FCA5" box="[362,553,861,886]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE513FFFAFEEAFC996C72FCA5" box="[362,553,861,886]" italics="true" pageId="2" pageNumber="39">Macrobrachium</emphasis>
</taxonomicName>
. Therefore, one can easily confuse male specimens of
<taxonomicName id="4C05699AE513FFFAFB3BFC996B32FCA5" box="[1211,1385,861,886]" class="Malacostraca" family="Palaemonidae" genus="Arachnochium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE513FFFAFB3BFC996B32FCA5" box="[1211,1385,861,886]" italics="true" pageId="2" pageNumber="39">Arachnochium</emphasis>
</taxonomicName>
and females of
<taxonomicName id="4C05699AE513FFFAFEA1FC466FBDFC48" box="[289,486,898,923]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE513FFFAFEA1FC466FBDFC48" box="[289,486,898,923]" italics="true" pageId="2" pageNumber="39">Macrobrachium</emphasis>
</taxonomicName>
. Only the presence of a genital flap at the base of the fifth pereiopod and an appendix masculina on the endopod of the second pleopod can ascertain the male sex of the specimen.
</paragraph>
<paragraph id="8BBA1219E513FFFAFF46FC096F89FB3F" blockId="2.[151,1437,152,1335]" pageId="2" pageNumber="39">
An allied species,
<taxonomicName id="4C05699AE513FFFAFE21FC096C89FC35" box="[417,722,973,998]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="kulsiense">
<emphasis id="B971CE0BE513FFFAFE21FC096C89FC35" box="[417,722,973,998]" italics="true" pageId="2" pageNumber="39">Macrobrachium kulsiense</emphasis>
</taxonomicName>
from the Kulsi River of Brahmaputra drainage,
<collectingCountry id="F3125289E513FFFAFA93FC096B09FC34" box="[1299,1362,973,999]" name="India" pageId="2" pageNumber="39">India</collectingCountry>
, has a very high rostral basal crest which a maximum depth distinctly more than the maximum dorsoventral diameter of the cornea and the adult females are also larger than the adult males. Although specimens were not examined for the present study, the description by
<bibRefCitation id="EF946FE8E513FFFAFC8EFBF96A1CFB84" box="[782,1095,1085,1111]" pageId="2" pageNumber="51" refString="Jayachandran, K. V., Lal Mohan, R. S. &amp; Raji, A. V. (2007) A new species of Macrobrachium Bate, 1868 (Decapoda, Palaemonidae) from the dolphin trenches of Kulsi River, N. India, possibly under threat. Zoologischer Anzeiger, 246, 43 - 48." type="journal article">
Jayachandran
<emphasis id="B971CE0BE513FFFAFC34FBF96DB5FB85" box="[948,1006,1085,1110]" italics="true" pageId="2" pageNumber="39">et al.</emphasis>
(2007)
</bibRefCitation>
, especially with regard to the two key generic characters mentioned above, clearly indicates that this species also belongs to
<taxonomicName id="4C05699AE513FFFAFB6FFBA76BC7FBAF" box="[1263,1436,1123,1148]" class="Malacostraca" family="Palaemonidae" genus="Arachnochium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B971CE0BE513FFFAFB6FFBA76BC7FBAF" box="[1263,1436,1123,1148]" italics="true" pageId="2" pageNumber="39">Arachnochium</emphasis>
</taxonomicName>
as presently defined. Unlike
<taxonomicName id="4C05699AE513FFFAFE68FB4C6C37FB72" box="[488,620,1160,1185]" class="Malacostraca" family="Palaemonidae" genus="Arachnochium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="mirabile">
<emphasis id="B971CE0BE513FFFAFE68FB4C6C37FB72" box="[488,620,1160,1185]" italics="true" pageId="2" pageNumber="39">A. mirabile</emphasis>
</taxonomicName>
,
<taxonomicName id="4C05699AE513FFFAFDFCFB4C6D5CFB72" box="[636,775,1160,1185]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="kulsiense" status="comb. nov.">
<emphasis id="B971CE0BE513FFFAFDFCFB4C6D5CFB72" box="[636,775,1160,1185]" italics="true" pageId="2" pageNumber="39">A. kulsiense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2427370E513FFFAFC90FB4C6DCFFB71" box="[784,916,1160,1186]" pageId="2" pageNumber="39">comb. nov.</taxonomicNameLabel>
, has only a few large eggs, a longer rostrum with the tip reaching the end of the scaphocerite and fewer dorsal rostral teeth (see also
<bibRefCitation id="EF946FE8E513FFFAFB36FB696E8EFB3F" author="Jayachandran" pageId="2" pageNumber="51" refString="Jayachandran, K. V., Lal Mohan, R. S. &amp; Raji, A. V. (2007) A new species of Macrobrachium Bate, 1868 (Decapoda, Palaemonidae) from the dolphin trenches of Kulsi River, N. India, possibly under threat. Zoologischer Anzeiger, 246, 43 - 48." type="journal article" year="2007">
Jayachandran
<emphasis id="B971CE0BE513FFFAFAE0FB696BCFFB15" box="[1376,1428,1197,1222]" italics="true" pageId="2" pageNumber="39">et al</emphasis>
. 2007
</bibRefCitation>
; Klotz, pers. comm.).
</paragraph>
<paragraph id="8BBA1219E513FFFAFF46FB336F74FAE4" blockId="2.[151,1437,152,1335]" pageId="2" pageNumber="39">
<emphasis id="B971CE0BE513FFFAFF46FB336FC0FAC2" bold="true" box="[198,411,1271,1297]" pageId="2" pageNumber="39">Species included.</emphasis>
<taxonomicName id="4C05699AE513FFFAFE22FB3C6AF2FAC1" authority="Jayachandran, Lal Mohan &amp; Raji, 2007" authorityName="Jayachandran, Lal Mohan &amp; Raji" authorityYear="2007" box="[418,1193,1272,1298]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="kulsiense">
<emphasis id="B971CE0BE513FFFAFE22FB3C6C8AFAC2" box="[418,721,1272,1297]" italics="true" pageId="2" pageNumber="39">Macrobrachium kulsiense</emphasis>
<bibRefCitation id="EF946FE8E513FFFAFD58FB3C6AF2FAC1" author="Jayachandran" box="[728,1193,1272,1298]" pageId="2" pageNumber="51" refString="Jayachandran, K. V., Lal Mohan, R. S. &amp; Raji, A. V. (2007) A new species of Macrobrachium Bate, 1868 (Decapoda, Palaemonidae) from the dolphin trenches of Kulsi River, N. India, possibly under threat. Zoologischer Anzeiger, 246, 43 - 48." type="journal article" year="2007">Jayachandran, Lal Mohan &amp; Raji, 2007</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C05699AE513FFFAFB34FB3C6F70FAE4" authority="Kemp, 1917" authorityName="Kemp" authorityYear="1917" class="Malacostraca" family="Palaemonidae" genus="Palaemon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="B971CE0BE513FFFAFB34FB3C6BC7FAC2" box="[1204,1436,1272,1297]" italics="true" pageId="2" pageNumber="39">Palaemon mirabilis</emphasis>
<bibRefCitation id="EF946FE8E513FFFAFF17FAD96F70FAE4" author="Kemp" box="[151,299,1309,1335]" pageId="2" pageNumber="51" refString="Kemp, S. (1917) Notes on Crustacea Decapoda in the Indian Museum, IX: Leander styliferus, Milne-Edwards, and related forms. Records of the Indian Museum, 13 (4), 203 - 231, pls. 8 - 10." type="journal article" year="1917">Kemp, 1917</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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