treatments-xml/data/11/6A/87/116A87EAFFCFFFE7448588FDFF7A15DF.xml

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<mods:title id="539D82C2B9163846341CB86EAB631C93">Two new species of Clidicus Laporte, 1832 (Coleoptera: Staphylinidae: Scydmaeninae) from Yunnan, China</mods:title>
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<mods:namePart id="F1A7FA7584DCF3023BD72FD3090FF2A9">Cheng, Zhi-Fei</mods:namePart>
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<mods:namePart id="10506F457F840AD26DE9478F88566139">Yin, Zi-Wei</mods:namePart>
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<mods:date id="4176E7AA8D1C108D3E585CB7198574AD">2019</mods:date>
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<subSubSection id="D1D96577FFCFFFE3448588FDFE2011C8" box="[151,426,225,252]" pageId="1" pageNumber="322" type="nomenclature">
<paragraph id="997C36FCFFCFFFE3448588FDFE2011C8" blockId="1.[151,426,225,288]" box="[151,426,225,252]" pageId="1" pageNumber="322">
<heading id="C2348190FFCFFFE3448588FDFE2011C8" bold="true" box="[151,426,225,252]" fontSize="11" level="1" pageId="1" pageNumber="322" reason="1">
<emphasis id="ABB7EAEEFFCFFFE3448588FDFE2011C8" bold="true" box="[151,426,225,252]" pageId="1" pageNumber="322">
<taxonomicName id="5EC34D7FFFCFFFE3448588FDFECF11CF" ID-CoL="8XXTC" authority="Cheng &amp; Yin, 2019" authorityName="Cheng &amp; Yin" authorityYear="2019" box="[151,325,225,252]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="322" phylum="Arthropoda" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="ABB7EAEEFFCFFFE3448588FDFECF11CF" bold="true" box="[151,325,225,252]" italics="true" pageId="1" pageNumber="322">Clidicus qiuae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B0845795FFCFFFE3455E88FEFE2011C8" box="[332,426,226,252]" pageId="1" pageNumber="322" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="D1D96577FFCFFFE344858919FEAD1014" box="[151,295,261,288]" pageId="1" pageNumber="322" type="description">
<paragraph id="997C36FCFFCFFFE344858919FEAD1014" blockId="1.[151,426,225,288]" box="[151,295,261,288]" pageId="1" pageNumber="322">
(
<figureCitation id="01F82A79FFCFFFE3448D8919FE89102B" box="[159,259,261,287]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,250,1958,1983]" captionStartId-1="2.[151,250,1631,1656]" captionStartId-2="3.[151,250,1924,1949]" captionStartId-3="4.[151,250,1936,1961]" captionTargetBox-0="[151,1435,578,1924]" captionTargetBox-1="[151,1424,676,1608]" captionTargetBox-2="[232,1358,167,1898]" captionTargetBox-3="[151,1437,813,1917]" captionTargetId-0="figure@1.[151,1436,576,1924]" captionTargetId-1="figure@2.[151,1436,668,1611]" captionTargetId-2="figure@3.[227,1361,166,1905]" captionTargetId-3="figure@4.[151,1437,813,1917]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionText-0="FIGURE 1. Dorsal habitus of Clidicus qiuae. A. Male. B. Female. Scale bars: 2 mm." captionText-1="FIGURE 2. Diagnostic features of Clidicus qiuae male. A. Head dorsum. B. Left antenna. C. Labrum. D. Right mandible. E. Left maxilla. F. Labium. Scale bars: 0.5 mm in A–B; 0.2 mm in C–F." captionText-2="FIGURE 3. Diagnostic features of Clidicus qiuae (A–D. Male. E–F. Female). A–C. Aedeagus, dorsal (A), lateral (B), and ventral (C). D. Sperm pump. E. Female genital segment. F. Spermatheca, enlarged. Scale bars: 0.5 mm in A–D; 0.2 mm in E; 0.1 mm in F." captionText-3="FIGURE 4. Collecting environment of Clidicus qiuae. A, B. Collecting site—a slope at the edge of the forest. C, D. Exposed adults of Clidicus qiuae after the leaf litter was stirred (pictures provided by Hao Xu)." httpUri-0="https://zenodo.org/record/3255479/files/figure.png" httpUri-1="https://zenodo.org/record/3255481/files/figure.png" httpUri-2="https://zenodo.org/record/3255483/files/figure.png" httpUri-3="https://zenodo.org/record/3255485/files/figure.png" pageId="1" pageNumber="322">Figs 1–4</figureCitation>
, 
<figureCitation id="01F82A79FFCFFFE34502891AFE951014" box="[272,287,262,288]" captionStart="FIGURE 7" captionStartId="8.[151,250,1375,1400]" captionTargetBox="[167,1421,489,1350]" captionTargetId="figure@8.[167,1421,489,1350]" captionTargetPageId="8" captionText="FIGURE 7. Distribution of extant species of Clidicus." httpUri="https://zenodo.org/record/3255489/files/figure.png" pageId="1" pageNumber="322">7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D1D96577FFCFFFE344858952FE9A132F" pageId="1" pageNumber="322" type="materials_examined">
<paragraph id="997C36FCFFCFFFE344858952FE9A132F" blockId="1.[151,1437,332,539]" pageId="1" pageNumber="322">
<emphasis id="ABB7EAEEFFCFFFE344858952FECF105C" bold="true" box="[151,325,333,360]" pageId="1" pageNumber="322">Type material</emphasis>
(71 exx). 
<materialsCitation id="29AB3CA1FFCFFFE345DA8952FCAF10B8" ID-GBIF-Occurrence="2269395173" collectionCode="RFH" collectorName="IJ Xb" country="China" county="Maguan County" location="Wenshan City" municipality="China" pageId="1" pageNumber="322" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="4678885EFFCFFFE345DA8952FDBF105C" box="[456,565,334,360]" pageId="1" pageNumber="322" type="holotype">
<emphasis id="ABB7EAEEFFCFFFE345DA8952FDBF105C" bold="true" box="[456,565,334,360]" pageId="1" pageNumber="322">Holotype</emphasis>
</typeStatus>
: 
<emphasis id="ABB7EAEEFFCFFFE3465B8951FD25105C" bold="true" box="[585,687,333,360]" pageId="1" pageNumber="322">
<collectingCountry id="E1D4766CFFCFFFE3465B8951FD21105C" box="[585,683,333,360]" name="China" pageId="1" pageNumber="322">CHINA</collectingCountry>
:
</emphasis>
<specimenCount id="8FC5FD75FFCFFFE346AE8952FD59105C" box="[700,723,334,360]" pageId="1" pageNumber="322" type="male">♂</specimenCount>
, ‘ 
<collectingMunicipality id="7918AC86FFCFFFE346FA8951FCBE1053" box="[744,820,333,359]" pageId="1" pageNumber="322">China</collectingMunicipality>
: 
<collectingRegion id="5B07F81EFFCFFFE347568952FC15105C" box="[836,927,334,360]" country="China" name="Yunnan" pageId="1" pageNumber="322">Yunnan</collectingRegion>
, 
<location id="9C1C6027FFCFFFE347A28951FBDD1053" LSID="urn:lsid:plazi:treatment:116A87EAFFCFFFE7448588FDFF7A15DF:9C1C6027FFCFFFE347A28951FBDD1053" box="[944,1111,333,359]" country="China" county="Maguan County" municipality="China" name="Wenshan City" pageId="1" pageNumber="322" stateProvince="Yunnan">Wenshan City</location>
, 
<collectingCounty id="701D4E70FFCFFFE3407A8952FAA81053" box="[1128,1314,333,360]" pageId="1" pageNumber="322">Maguan County</collectingCounty>
(
<collectorName id="3436532AFFCFFFE3412A8950FA011057" box="[1336,1419,332,355]" pageId="1" pageNumber="322">IJẌƃ</collectorName>
), 
<location id="9C1C6027FFCFFFE34485896DFEC610B8" LSID="urn:lsid:plazi:treatment:116A87EAFFCFFFE7448588FDFF7A15DF:9C1C6027FFCFFFE34485896DFEC610B8" box="[151,332,369,396]" country="China" county="Maguan County" municipality="China" name="Gulinjing Town" pageId="1" pageNumber="322" stateProvince="Yunnan">Gulinjing Town</location>
(Ħẇḃş), 
<location id="9C1C6027FFCFFFE345F4896DFD3F10BF" LSID="urn:lsid:plazi:treatment:116A87EAFFCFFFE7448588FDFF7A15DF:9C1C6027FFCFFFE345F4896DFD3F10BF" box="[486,693,369,395]" country="China" county="Maguan County" municipality="China" name="Piaochang Village" pageId="1" pageNumber="322" stateProvince="Yunnan">Piaochang Village</location>
(
<collectionCode id="FFD2AE39FFCFFFE346D4896CFC9010B3" box="[710,794,368,391]" pageId="1" pageNumber="322">ṞFḤ</collectionCode>
)
</materialsCitation>
, Gulinjing N. R. (ĦẇḃssküDṽ), 
<geoCoordinate id="FCF7503BFFCFFFE340FF896DFA1210B8" box="[1261,1432,369,396]" degrees="22" direction="north" minutes="51" orientation="latitude" pageId="1" pageNumber="322" precision="1" seconds="20.03" value="22.855564">22°51′20.03″N</geoCoordinate>
, 
<geoCoordinate id="FCF7503BFFCFFFE344858989FEC71084" box="[151,333,405,432]" degrees="103" direction="east" minutes="59" orientation="longitude" pageId="1" pageNumber="322" precision="1" seconds="10.27" value="103.98618">103°59′10.27″E</geoCoordinate>
, alt. 
<quantity id="5E3B9B19FFCFFFE345918989FE6B1084" box="[387,481,405,432]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="322" unit="m" value="1700.0">1,700 m</quantity>
, 
<date id="ED7D103CFFCFFFE345FF8989FDEB109B" box="[493,609,405,431]" pageId="1" pageNumber="322" value="2018-02-27">27.ii.2018</date>
, Jian-Yue Qiu &amp; Hao Xu leg. / 
<materialsCitation id="29AB3CA1FFCFFFE347D18989FDD910E7" ID-GBIF-Occurrence="2269395171" collectionCode="SNUC" country="China" determinerName="Cheng &amp; Yin" pageId="1" pageNumber="322" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="4678885EFFCFFFE347D18989FBDD109B" box="[963,1111,405,431]" pageId="1" pageNumber="322" type="holotype">HOLOTYPE</typeStatus>
(red) 
<specimenCount id="8FC5FD75FFCFFFE3408E898AFB391084" box="[1180,1203,406,432]" pageId="1" pageNumber="322" type="male">♂</specimenCount>
, 
<taxonomicName id="5EC34D7FFFCFFFE340AD898AFA94109B" authority="Cheng &amp; Yin, 2019" authorityName="Cheng &amp; Yin" authorityYear="2019" box="[1215,1310,406,432]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="322" phylum="Arthropoda" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="ABB7EAEEFFCFFFE340AD898AFA94109B" box="[1215,1310,406,432]" italics="true" pageId="1" pageNumber="322">C. qiuae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B0845795FFCFFFE34137898AFAEF1084" box="[1317,1381,406,432]" pageId="1" pageNumber="322" rank="species">
<emphasis id="ABB7EAEEFFCFFFE34137898AFAEF1084" bold="true" box="[1317,1381,406,432]" pageId="1" pageNumber="322">sp. n.</emphasis>
</taxonomicNameLabel>
, det. 
<determinerName id="6CA0DA38FFCFFFE3448589A5FF6A10E7" box="[151,224,441,467]" pageId="1" pageNumber="322">Cheng</determinerName>
&amp; 
<determinerName id="6CA0DA38FFCFFFE3451389A5FEA610E7" box="[257,300,441,467]" pageId="1" pageNumber="322">Yin</determinerName>
, 2019, SNUC’ (in 
<collectionCode id="FFD2AE39FFCFFFE345E989A5FDC110E7" box="[507,587,441,467]" pageId="1" pageNumber="322">SNUC</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="29AB3CA1FFCFFFE3464F89A6FE4610C3" ID-GBIF-Occurrence="2269395169" country="China" location="CHINA" pageId="1" pageNumber="322" specimenCount="70" specimenCount-female="42" specimenCount-male="28" typeStatus="paratype">
<emphasis id="ABB7EAEEFFCFFFE3464F89A6FCC010E0" bold="true" box="[605,842,441,468]" pageId="1" pageNumber="322">
<typeStatus id="4678885EFFCFFFE3464F89A6FD5310E0" box="[605,729,442,468]" pageId="1" pageNumber="322" type="paratype">Paratypes</typeStatus>
: 
<collectingCountry id="E1D4766CFFCFFFE346F689A5FCCC10E0" box="[740,838,441,468]" name="China" pageId="1" pageNumber="322">CHINA</collectingCountry>
:
</emphasis>
<specimenCount id="8FC5FD75FFCFFFE3474389A5FC1510E0" box="[849,927,441,468]" pageId="1" pageNumber="322" type="male">28 ♂♂</specimenCount>
, 
<specimenCount id="8FC5FD75FFCFFFE347B889A5FC7D10E0" box="[938,1015,441,468]" pageId="1" pageNumber="322" type="female">42 ♀♀</specimenCount>
, same label data as the 
<typeStatus id="4678885EFFCFFFE340E889A5FAD510E7" box="[1274,1375,441,467]" pageId="1" pageNumber="322" type="holotype">holotype</typeStatus>
, each with a following type label
</materialsCitation>
: ‘ 
<materialsCitation id="29AB3CA1FFCFFFE345C989C1FE86132F" ID-GBIF-Occurrence="2269395172" collectionCode="SNUC" country="China" determinerName="Cheng &amp; Yin" pageId="1" pageNumber="322" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="4678885EFFCFFFE345C989C1FDE110C3" box="[475,619,477,503]" pageId="1" pageNumber="322" type="paratype">PARATYPE</typeStatus>
(yellow) 
<specimenCount id="8FC5FD75FFCFFFE346C889C2FD6510CC" box="[730,751,478,504]" pageId="1" pageNumber="322" type="male">♂</specimenCount>
(or 
<specimenCount id="8FC5FD75FFCFFFE3470D89C2FCB810CC" box="[799,818,478,504]" pageId="1" pageNumber="322" type="female">♀</specimenCount>
), 
<taxonomicName id="5EC34D7FFFCFFFE3475E89C2FC2610C3" authority="Cheng &amp; Yin, 2019" authorityName="Cheng &amp; Yin" authorityYear="2019" box="[844,940,478,504]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="322" phylum="Arthropoda" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="ABB7EAEEFFCFFFE3475E89C2FC2610C3" box="[844,940,478,504]" italics="true" pageId="1" pageNumber="322">C. qiuae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B0845795FFCFFFE347A689C2FC7B10CC" box="[948,1009,478,504]" pageId="1" pageNumber="322" rank="species">sp. n.</taxonomicNameLabel>
, det. 
<determinerName id="6CA0DA38FFCFFFE3402289C1FBF310C3" box="[1072,1145,477,503]" pageId="1" pageNumber="322">Cheng</determinerName>
&amp; 
<determinerName id="6CA0DA38FFCFFFE3408F89C1FB4210C3" box="[1181,1224,477,503]" pageId="1" pageNumber="322">Yin</determinerName>
, 2019, SNUC’ (all in 
<collectionCode id="FFD2AE39FFCFFFE344A68A1DFE8E132F" box="[180,260,513,539]" pageId="1" pageNumber="322">SNUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="CDBC6674FFCFFFE344858FBAFC73168A" box="[151,1017,1958,1983]" httpUri="https://zenodo.org/record/3255479/files/figure.png" pageId="1" pageNumber="322" startId="1.[151,250,1958,1983]" targetBox="[151,1435,578,1924]" targetPageId="1">
<paragraph id="997C36FCFFCFFFE344858FBAFC73168A" blockId="1.[151,1017,1958,1983]" box="[151,1017,1958,1983]" pageId="1" pageNumber="322">
<emphasis id="ABB7EAEEFFCFFFE344858FBAFE99168B" bold="true" box="[151,275,1958,1983]" pageId="1" pageNumber="322">FIGURE 1.</emphasis>
Dorsal habitus of 
<taxonomicName id="5EC34D7FFFCFFFE345DC8FBBFDEB168B" authorityName="Cheng &amp; Yin" authorityYear="2019" box="[462,609,1959,1983]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="322" phylum="Arthropoda" rank="species" species="qiuae">
<emphasis id="ABB7EAEEFFCFFFE345DC8FBBFDEB168B" box="[462,609,1959,1983]" italics="true" pageId="1" pageNumber="322">Clidicus qiuae</emphasis>
</taxonomicName>
. 
<emphasis id="ABB7EAEEFFCFFFE3467C8FBAFD0D168B" bold="true" box="[622,647,1958,1983]" pageId="1" pageNumber="322">A.</emphasis>
Male. 
<emphasis id="ABB7EAEEFFCFFFE346DC8FBBFD6F168B" bold="true" box="[718,741,1959,1983]" pageId="1" pageNumber="322">B.</emphasis>
Female. Scale bars: 2 mm.
</paragraph>
</caption>
<subSubSection id="D1D96577FFCCFFE044D58884FE66102A" pageId="2" pageNumber="323" type="diagnosis">
<paragraph id="997C36FCFFCCFFE044D58884FE66102A" blockId="2.[151,1437,152,647]" pageId="2" pageNumber="323">
<emphasis id="ABB7EAEEFFCCFFE044D58884FEC81187" bold="true" box="[199,322,152,179]" pageId="2" pageNumber="323">Diagnosis.</emphasis>
Body often larger than 5.0 mm. Anterior margin of frons broadly emarginate. Anterior margin of labrum with small, triangular emargination at middle. Maxillary palpomeres 2–3 relatively short in relation to palpomere 4. Antennomeres 2–11 each slightly to distinctly elongate. Aedeagus moderately elongate, broadest near middle; parameres symmetric.
</paragraph>
</subSubSection>
<subSubSection id="D1D96577FFCCFFE744D58934FBB31293" lastPageId="5" lastPageNumber="326" pageId="2" pageNumber="323" type="description">
<paragraph id="997C36FCFFCCFFE044D58934FAE31052" blockId="2.[151,1437,152,647]" pageId="2" pageNumber="323">
<emphasis id="ABB7EAEEFFCCFFE044D58934FED31077" bold="true" box="[199,345,296,323]" pageId="2" pageNumber="323">Description.</emphasis>
Male. BL 
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, body (
<figureCitation id="01F82A79FFCCFFE046D18934FC921077" box="[707,792,296,323]" captionStart="FIGURE 1" captionStartId="1.[151,250,1958,1983]" captionTargetBox="[151,1435,578,1924]" captionTargetId="figure@1.[151,1436,576,1924]" captionTargetPageId="1" captionText="FIGURE 1. Dorsal habitus of Clidicus qiuae. A. Male. B. Female. Scale bars: 2 mm." httpUri="https://zenodo.org/record/3255479/files/figure.png" pageId="2" pageNumber="323">Fig. 1A</figureCitation>
) moderately stout, with flattened subrectangular head and strongly convex pronotum and elytra, covered with coarse and dense setae, reddish-brown, tarsi slightly paler.
</paragraph>
<paragraph id="997C36FCFFCCFFE044D5896CFEF613B3" blockId="2.[151,1437,152,647]" pageId="2" pageNumber="323">
Head (
<figureCitation id="01F82A79FFCCFFE04506896CFEE510BF" box="[276,367,368,395]" captionStart="FIGURE 2" captionStartId="2.[151,250,1631,1656]" captionTargetBox="[151,1424,676,1608]" captionTargetId="figure@2.[151,1436,668,1611]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Clidicus qiuae male. A. Head dorsum. B. Left antenna. C. Labrum. D. Right mandible. E. Left maxilla. F. Labium. Scale bars: 0.5 mm in A–B; 0.2 mm in C–F." httpUri="https://zenodo.org/record/3255481/files/figure.png" pageId="2" pageNumber="323">Fig. 2A</figureCitation>
) subrectangular and transverse, broadest at temples, HL 
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, HW 
<quantity id="5E3B9B19FFCCFFE040FD896DFA1210BF" box="[1263,1432,369,395]" metricMagnitude="-3" metricUnit="m" metricValue="1.255" metricValueMax="1.27" metricValueMin="1.24" pageId="2" pageNumber="323" unit="mm" value="1.255" valueMax="1.27" valueMin="1.24">1.24–1.27 mm</quantity>
; tempora about 3.5× length of eye in dorsal view, evenly rounded; vertex with large and deep posteromedian irregular impression; frons broadly emarginate at anterior margin; clypeus transverse and convex, demarcated from frons by deep and distinct fronto-clypeal groove; supraantennal tubercles small but distinctly elevated. Eyes small, nearly round, not projecting laterally from head silhouette, with fine facets. Punctures on vertex and frons small, but distinct. Setae on head moderately long, sparse and suberect to erect. Antennae (
<figureCitation id="01F82A79FFCCFFE040088A38FBFA130B" box="[1050,1136,548,575]" captionStart="FIGURE 2" captionStartId="2.[151,250,1631,1656]" captionTargetBox="[151,1424,676,1608]" captionTargetId="figure@2.[151,1436,668,1611]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Clidicus qiuae male. A. Head dorsum. B. Left antenna. C. Labrum. D. Right mandible. E. Left maxilla. F. Labium. Scale bars: 0.5 mm in A–B; 0.2 mm in C–F." httpUri="https://zenodo.org/record/3255481/files/figure.png" pageId="2" pageNumber="323">Fig. 2B</figureCitation>
) slender, all antennomeres elongate, relative lengths of antennomeres 1–11: 4.58: 1.0: 1.25: 1.33: 1.33: 1.33: 1.42: 1.42: 1.33: 1.33: 1.58; AnL 
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.
</paragraph>
<caption id="CDBC6674FFCCFFE044858E43FCD217A8" httpUri="https://zenodo.org/record/3255481/files/figure.png" pageId="2" pageNumber="323" startId="2.[151,250,1631,1656]" targetBox="[151,1424,676,1608]" targetPageId="2">
<paragraph id="997C36FCFFCCFFE044858E43FCD217A8" blockId="2.[151,1436,1631,1692]" pageId="2" pageNumber="323">
<emphasis id="ABB7EAEEFFCCFFE044858E43FE9E174C" bold="true" box="[151,276,1631,1656]" pageId="2" pageNumber="323">FIGURE 2.</emphasis>
Diagnostic features of 
<taxonomicName id="5EC34D7FFFCCFFE046118E7CFD1C1743" authorityName="Cheng &amp; Yin" authorityYear="2019" box="[515,662,1632,1656]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="323" phylum="Arthropoda" rank="species" species="qiuae">
<emphasis id="ABB7EAEEFFCCFFE046118E7CFD1C1743" box="[515,662,1632,1656]" italics="true" pageId="2" pageNumber="323">Clidicus qiuae</emphasis>
</taxonomicName>
male. 
<emphasis id="ABB7EAEEFFCCFFE046CE8E43FD7F174C" bold="true" box="[732,757,1631,1656]" pageId="2" pageNumber="323">A.</emphasis>
Head dorsum. 
<emphasis id="ABB7EAEEFFCCFFE047808E7CFC23174C" bold="true" box="[914,937,1632,1656]" pageId="2" pageNumber="323">B.</emphasis>
Left antenna. 
<emphasis id="ABB7EAEEFFCCFFE0402F8E43FBDC174C" bold="true" box="[1085,1110,1631,1656]" pageId="2" pageNumber="323">C.</emphasis>
Labrum. 
<emphasis id="ABB7EAEEFFCCFFE040A88E7CFB59174C" bold="true" box="[1210,1235,1632,1656]" pageId="2" pageNumber="323">D.</emphasis>
Right mandible. 
<emphasis id="ABB7EAEEFFCCFFE041978E7CFA16174C" bold="true" box="[1413,1436,1632,1656]" pageId="2" pageNumber="323">E.</emphasis>
Left maxilla. 
<emphasis id="ABB7EAEEFFCCFFE045328E98FEBC17A8" bold="true" box="[288,310,1668,1692]" pageId="2" pageNumber="323">F.</emphasis>
Labium. Scale bars: 0.5 mm in A–B; 0.2 mm in C–F.
</paragraph>
</caption>
<paragraph id="997C36FCFFCCFFE644D58ED6FD921077" blockId="2.[151,1437,1738,2017]" lastBlockId="4.[151,1437,153,791]" lastPageId="4" lastPageNumber="325" pageId="2" pageNumber="323">
Labrum (
<figureCitation id="01F82A79FFCCFFE0453C8ED6FE0817D0" box="[302,386,1738,1764]" captionStart="FIGURE 2" captionStartId="2.[151,250,1631,1656]" captionTargetBox="[151,1424,676,1608]" captionTargetId="figure@2.[151,1436,668,1611]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Clidicus qiuae male. A. Head dorsum. B. Left antenna. C. Labrum. D. Right mandible. E. Left maxilla. F. Labium. Scale bars: 0.5 mm in A–B; 0.2 mm in C–F." httpUri="https://zenodo.org/record/3255481/files/figure.png" pageId="2" pageNumber="323">Fig. 2C</figureCitation>
) subrectangular in dorsal view, with small distinct triangular emargination at middle of anterior margin; anterior margin with single row of five pairs of long and erect setae. Mandible (
<figureCitation id="01F82A79FFCCFFE040778EF2FB36163C" box="[1125,1212,1774,1800]" captionStart="FIGURE 2" captionStartId="2.[151,250,1631,1656]" captionTargetBox="[151,1424,676,1608]" captionTargetId="figure@2.[151,1436,668,1611]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Clidicus qiuae male. A. Head dorsum. B. Left antenna. C. Labrum. D. Right mandible. E. Left maxilla. F. Labium. Scale bars: 0.5 mm in A–B; 0.2 mm in C–F." httpUri="https://zenodo.org/record/3255481/files/figure.png" pageId="2" pageNumber="323">Fig. 2D</figureCitation>
) subtriangular, with broad base and curved distal tooth; teeth robust; prostheca with short and dense setae from base to nearly middle of mandible. Maxilla (
<figureCitation id="01F82A79FFCCFFE045658F2AFE471664" box="[375,461,1846,1872]" captionStart="FIGURE 2" captionStartId="2.[151,250,1631,1656]" captionTargetBox="[151,1424,676,1608]" captionTargetId="figure@2.[151,1436,668,1611]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Clidicus qiuae male. A. Head dorsum. B. Left antenna. C. Labrum. D. Right mandible. E. Left maxilla. F. Labium. Scale bars: 0.5 mm in A–B; 0.2 mm in C–F." httpUri="https://zenodo.org/record/3255481/files/figure.png" pageId="2" pageNumber="323">Fig. 2E</figureCitation>
) with triangular stipes bearing single long seta, subtriangular palpifer with long setae on external margin, elongate galea and broad lacinia with short and dense setae. Maxillary palpus with dense short setae and additionally several long thick setae; palpomere 1 minute and elongate; palpomere 2 moderately long and slender, broadened distally and strongly curved; palpomere 3 large and broadened, broadest near apex; palpomere 4 short, subconical. Labium (
<figureCitation id="01F82A79FFCCFFE045F68FDAFDB216D4" box="[484,568,1990,2016]" captionStart="FIGURE 2" captionStartId="2.[151,250,1631,1656]" captionTargetBox="[151,1424,676,1608]" captionTargetId="figure@2.[151,1436,668,1611]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Clidicus qiuae male. A. Head dorsum. B. Left antenna. C. Labrum. D. Right mandible. E. Left maxilla. F. Labium. Scale bars: 0.5 mm in A–B; 0.2 mm in C–F." httpUri="https://zenodo.org/record/3255481/files/figure.png" pageId="2" pageNumber="323">Fig. 2F</figureCitation>
) elongate, with rectangular mentum about 2.5× as wide as long; prementum moderately large, with anterior surface flattened and divided by longitudinal groove, with several short and long setae. Labial palpus with several long setae; palpomere 1 short and broad, slightly elongate, with one long seta on antero-mesal margin; palpomere 2 strongly elongate, constricted at base and broadened apically, with truncate apex; palpomere 3 subconical with pointed apex, much shorter and narrower than palpomere 2. Lateral lobe of hypopharynx long, with dense microtrichia.
</paragraph>
<caption id="CDBC6674FFCDFFE144858F98FE9016D0" httpUri="https://zenodo.org/record/3255483/files/figure.png" pageId="3" pageNumber="324" startId="3.[151,250,1924,1949]" targetBox="[232,1358,167,1898]" targetPageId="3">
<paragraph id="997C36FCFFCDFFE144858F98FE9016D0" blockId="3.[151,1437,1924,2021]" pageId="3" pageNumber="324">
<emphasis id="ABB7EAEEFFCDFFE144858F98FE9F16A9" bold="true" box="[151,277,1924,1949]" pageId="3" pageNumber="324">FIGURE 3.</emphasis>
Diagnostic features of 
<taxonomicName id="5EC34D7FFFCDFFE1461E8F99FD2B16A9" authorityName="Cheng &amp; Yin" authorityYear="2019" box="[524,673,1925,1949]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="324" phylum="Arthropoda" rank="species" species="qiuae">
<emphasis id="ABB7EAEEFFCDFFE1461E8F99FD2B16A9" box="[524,673,1925,1949]" italics="true" pageId="3" pageNumber="324">Clidicus qiuae</emphasis>
</taxonomicName>
(A–D. Male. E–F. Female). 
<emphasis id="ABB7EAEEFFCDFFE147C08F98FB8016A9" bold="true" box="[978,1034,1924,1949]" pageId="3" pageNumber="324">A–C.</emphasis>
Aedeagus, dorsal (A), lateral (B), and ventral (C). 
<emphasis id="ABB7EAEEFFCDFFE145018FB5FEA616F5" bold="true" box="[275,300,1961,1985]" pageId="3" pageNumber="324">D.</emphasis>
Sperm pump. 
<emphasis id="ABB7EAEEFFCDFFE145D18FB5FE5016F5" bold="true" box="[451,474,1961,1985]" pageId="3" pageNumber="324">E.</emphasis>
Female genital segment. 
<emphasis id="ABB7EAEEFFCDFFE146F28FB5FD7C16F5" bold="true" box="[736,758,1961,1985]" pageId="3" pageNumber="324">F.</emphasis>
Spermatheca, enlarged. Scale bars: 0.5 mm in A–D; 0.2 mm in E; 0.1 mm in F.
</paragraph>
</caption>
<paragraph id="997C36FCFFCAFFE644D58951FAB1109B" blockId="4.[151,1437,153,791]" pageId="4" pageNumber="325">
Pronotum in dorsal view inversely subtrapezoidal, broadest near anterior 1/4, PL 
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, PWb 
<quantity id="5E3B9B19FFCAFFE6413E8951FF4F10BF" metricMagnitude="-4" metricUnit="m" metricValue="9.299999999999999" metricValueMax="9.6" metricValueMin="9.0" pageId="4" pageNumber="325" unit="mm" value="0.9299999999999999" valueMax="0.96" valueMin="0.9">0.90–0.96 mm</quantity>
, PWm 
<quantity id="5E3B9B19FFCAFFE6450E896DFE4E10BF" box="[284,452,369,395]" metricMagnitude="-3" metricUnit="m" metricValue="1.285" metricValueMax="1.33" metricValueMin="1.24" pageId="4" pageNumber="325" unit="mm" value="1.285" valueMax="1.33" valueMin="1.24">1.24–1.33 mm</quantity>
; disc convex, sharply demarcated from narrow posterior ‘collar’ by narrow transverse groove with small, slightly unevenly distributed pits. Punctures and setae on disc similar to those on head.
</paragraph>
<paragraph id="997C36FCFFCAFFE644D589A5FF431357" blockId="4.[151,1437,153,791]" pageId="4" pageNumber="325">
Elytra in dorsal view oval, broadest near anterior 2/5, strongly narrowing toward apices, EL 
<quantity id="5E3B9B19FFCAFFE640D189A5FAED10E7" box="[1219,1383,441,467]" metricMagnitude="-3" metricUnit="m" metricValue="2.99" metricValueMax="3.1" metricValueMin="2.88" pageId="4" pageNumber="325" unit="mm" value="2.99" valueMax="3.1" valueMin="2.88">2.88–3.10 mm</quantity>
, EW 
<quantity id="5E3B9B19FFCAFFE6448589C0FEB610C3" box="[151,316,476,503]" metricMagnitude="-3" metricUnit="m" metricValue="1.92" metricValueMax="1.95" metricValueMin="1.89" pageId="4" pageNumber="325" unit="mm" value="1.92" valueMax="1.95" valueMin="1.89">1.89–1.95 mm</quantity>
, EI 1.50–1.64. Each elytron bears five dorsal and one lateral rows of large and deep pits, second row shortened anteriorly, cuticle between rows with fine and dense setiferous punctures; setae moderately long, denser and thicker than those on head and pronotum, recumbent towards apex. Hind wings absent. Scutellum hardly visible.
</paragraph>
<paragraph id="997C36FCFFCAFFE644D58A71FB3C1223" blockId="4.[151,1437,153,791]" pageId="4" pageNumber="325">
Aedeagus (
<figureCitation id="01F82A79FFCAFFE645548A71FE0D13B3" box="[326,391,621,647]" captionStart="FIGURE 3" captionStartId="3.[151,250,1924,1949]" captionTargetBox="[232,1358,167,1898]" captionTargetId="figure@3.[227,1361,166,1905]" captionTargetPageId="3" captionText="FIGURE 3. Diagnostic features of Clidicus qiuae (A–D. Male. E–F. Female). A–C. Aedeagus, dorsal (A), lateral (B), and ventral (C). D. Sperm pump. E. Female genital segment. F. Spermatheca, enlarged. Scale bars: 0.5 mm in A–D; 0.2 mm in E; 0.1 mm in F." httpUri="https://zenodo.org/record/3255483/files/figure.png" pageId="4" pageNumber="325">Fig. 3</figureCitation>
A–C) stout, AeL 
<quantity id="5E3B9B19FFCAFFE646598A70FD7B13B3" box="[587,753,620,647]" metricMagnitude="-3" metricUnit="m" metricValue="1.325" metricValueMax="1.35" metricValueMin="1.3" pageId="4" pageNumber="325" unit="mm" value="1.325" valueMax="1.35" valueMin="1.3">1.30–1.35 mm</quantity>
, in lateral view strongly curved; parameres symmetric, slen- der, much longer than median lobe, strongly sclerotized, with blunt apices, base and each internal edge fused with median lobe. Sperm pump (
<figureCitation id="01F82A79FFCAFFE645C38AA9FDA313FB" box="[465,553,693,719]" captionStart="FIGURE 3" captionStartId="3.[151,250,1924,1949]" captionTargetBox="[232,1358,167,1898]" captionTargetId="figure@3.[227,1361,166,1905]" captionTargetPageId="3" captionText="FIGURE 3. Diagnostic features of Clidicus qiuae (A–D. Male. E–F. Female). A–C. Aedeagus, dorsal (A), lateral (B), and ventral (C). D. Sperm pump. E. Female genital segment. F. Spermatheca, enlarged. Scale bars: 0.5 mm in A–D; 0.2 mm in E; 0.1 mm in F." httpUri="https://zenodo.org/record/3255483/files/figure.png" pageId="4" pageNumber="325">Fig. 3D</figureCitation>
) with largest diameter at external margins of funnels, 
<quantity id="5E3B9B19FFCAFFE6409C8AA9FAB813FB" box="[1166,1330,693,719]" metricMagnitude="-4" metricUnit="m" metricValue="6.55" metricValueMax="6.8" metricValueMin="6.3" pageId="4" pageNumber="325" unit="mm" value="0.655" valueMax="0.68" valueMin="0.63">0.63–0.68 mm</quantity>
in length (measured between external margins of funnels), short and stout, with short basal funnel and longer distal funnel, distance between funnels moderately long, lumen of internal duct broadened between funnels.
</paragraph>
<caption id="CDBC6674FFCAFFE644858F8CFC7416F9" httpUri="https://zenodo.org/record/3255485/files/figure.png" pageId="4" pageNumber="325" startId="4.[151,250,1936,1961]" targetBox="[151,1437,813,1917]" targetPageId="4">
<paragraph id="997C36FCFFCAFFE644858F8CFC7416F9" blockId="4.[151,1436,1936,1997]" pageId="4" pageNumber="325">
<emphasis id="ABB7EAEEFFCAFFE644858F8CFE99169D" bold="true" box="[151,275,1936,1961]" pageId="4" pageNumber="325">FIGURE 4.</emphasis>
Collecting environment of 
<taxonomicName id="5EC34D7FFFCAFFE6463C8F8DFD4B169C" authorityName="Cheng &amp; Yin" authorityYear="2019" box="[558,705,1937,1961]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="325" phylum="Arthropoda" rank="species" species="qiuae">
<emphasis id="ABB7EAEEFFCAFFE6463C8F8DFD4B169C" box="[558,705,1937,1961]" italics="true" pageId="4" pageNumber="325">Clidicus qiuae</emphasis>
</taxonomicName>
. 
<emphasis id="ABB7EAEEFFCAFFE646DD8F8CFC8F169D" bold="true" box="[719,773,1936,1961]" pageId="4" pageNumber="325">A, B.</emphasis>
Collecting site—a slope at the edge of the forest. 
<emphasis id="ABB7EAEEFFCAFFE641148F8CFAB4169D" bold="true" box="[1286,1342,1936,1961]" pageId="4" pageNumber="325">C, D.</emphasis>
Exposed adults of 
<taxonomicName id="5EC34D7FFFCAFFE644E78FA9FE0216F8" authorityName="Cheng &amp; Yin" authorityYear="2019" box="[245,392,1973,1997]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="325" phylum="Arthropoda" rank="species" species="qiuae">
<emphasis id="ABB7EAEEFFCAFFE644E78FA9FE0216F8" box="[245,392,1973,1997]" italics="true" pageId="4" pageNumber="325">Clidicus qiuae</emphasis>
</taxonomicName>
after the leaf litter was stirred (pictures provided by Hao Xu).
</paragraph>
</caption>
<paragraph id="997C36FCFFCBFFE744D58885FEF8109B" blockId="5.[151,1437,153,575]" pageId="5" pageNumber="326">
Female (
<figureCitation id="01F82A79FFCBFFE745398885FE091187" box="[299,387,153,179]" captionStart="FIGURE 1" captionStartId="1.[151,250,1958,1983]" captionTargetBox="[151,1435,578,1924]" captionTargetId="figure@1.[151,1436,576,1924]" captionTargetPageId="1" captionText="FIGURE 1. Dorsal habitus of Clidicus qiuae. A. Male. B. Female. Scale bars: 2 mm." httpUri="https://zenodo.org/record/3255479/files/figure.png" pageId="5" pageNumber="326">Fig. 1B</figureCitation>
). Externally similar to the male, but differs in much shorter antennae, and relatively stouter elytra. Measurements: BL 
<quantity id="5E3B9B19FFCBFFE745D488A0FDE711E3" box="[454,621,188,215]" metricMagnitude="-3" metricUnit="m" metricValue="5.18" metricValueMax="5.58" metricValueMin="4.78" pageId="5" pageNumber="326" unit="mm" value="5.18" valueMax="5.58" valueMin="4.78">4.78–5.58 mm</quantity>
; HL 
<quantity id="5E3B9B19FFCBFFE746B588A1FCC511E3" box="[679,847,189,215]" metricMagnitude="-4" metricUnit="m" metricValue="9.75" metricValueMax="10.2" metricValueMin="9.3" pageId="5" pageNumber="326" unit="mm" value="0.975" valueMax="1.02" valueMin="0.93">0.93–1.02 mm</quantity>
; HW 
<quantity id="5E3B9B19FFCBFFE7478088A1FBB011E3" box="[914,1082,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="1.29" metricValueMax="1.4" metricValueMin="1.18" pageId="5" pageNumber="326" unit="mm" value="1.29" valueMax="1.4" valueMin="1.18">1.18–1.40 mm</quantity>
; AnL 
<quantity id="5E3B9B19FFCBFFE7409288A1FAA211E3" box="[1152,1320,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="2.9699999999999998" metricValueMax="3.02" metricValueMin="2.92" pageId="5" pageNumber="326" unit="mm" value="2.9699999999999998" valueMax="3.02" valueMin="2.92">2.92–3.02 mm</quantity>
; PL 
<quantity id="5E3B9B19FFCBFFE7414F88A1FE8A11CF" metricMagnitude="-3" metricUnit="m" metricValue="1.27" metricValueMax="1.36" metricValueMin="1.18" pageId="5" pageNumber="326" unit="mm" value="1.27" valueMax="1.36" valueMin="1.18">1.18– 1.36 mm</quantity>
; PWb 
<quantity id="5E3B9B19FFCBFFE7455D88FDFE7D11CF" box="[335,503,225,251]" metricMagnitude="-4" metricUnit="m" metricValue="9.15" metricValueMax="10.2" metricValueMin="8.1" pageId="5" pageNumber="326" unit="mm" value="0.915" valueMax="1.02" valueMin="0.81">0.81–1.02 mm</quantity>
; PWm 
<quantity id="5E3B9B19FFCBFFE7465C88FDFD7C11CF" box="[590,758,225,251]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" metricValueMax="1.36" metricValueMin="1.24" pageId="5" pageNumber="326" unit="mm" value="1.3" valueMax="1.36" valueMin="1.24">1.24–1.36 mm</quantity>
; EL 
<quantity id="5E3B9B19FFCBFFE7473C88FDFC5C11CF" box="[814,982,225,251]" metricMagnitude="-3" metricUnit="m" metricValue="2.945" metricValueMax="3.22" metricValueMin="2.67" pageId="5" pageNumber="326" unit="mm" value="2.945" valueMax="3.22" valueMin="2.67">2.67–3.22 mm</quantity>
; EW 
<quantity id="5E3B9B19FFCBFFE7400588FDFB4A11CF" box="[1047,1216,225,251]" metricMagnitude="-3" metricUnit="m" metricValue="2.015" metricValueMax="2.17" metricValueMin="1.86" pageId="5" pageNumber="326" unit="mm" value="2.015" valueMax="2.17" valueMin="1.86">1.86–2.17 mm</quantity>
; EI 
<quantity id="5E3B9B19FFCBFFE740E388FDFA1311CF" box="[1265,1433,225,251]" metricMagnitude="-3" metricUnit="m" metricValue="1.455" metricValueMax="1.49" metricValueMin="1.42" pageId="5" pageNumber="326" unit="mm" value="1.455" valueMax="1.49" valueMin="1.42">1.42–1.49 mm</quantity>
. Female genitalia (
<figureCitation id="01F82A79FFCBFFE745738919FE3F102B" box="[353,437,261,287]" captionStart="FIGURE 3" captionStartId="3.[151,250,1924,1949]" captionTargetBox="[232,1358,167,1898]" captionTargetId="figure@3.[227,1361,166,1905]" captionTargetPageId="3" captionText="FIGURE 3. Diagnostic features of Clidicus qiuae (A–D. Male. E–F. Female). A–C. Aedeagus, dorsal (A), lateral (B), and ventral (C). D. Sperm pump. E. Female genital segment. F. Spermatheca, enlarged. Scale bars: 0.5 mm in A–D; 0.2 mm in E; 0.1 mm in F." httpUri="https://zenodo.org/record/3255483/files/figure.png" pageId="5" pageNumber="326">Fig. 3E</figureCitation>
) with long and entangled ductus of complicated form around base of spermatheca; bursa copulatrix elongate, finger-shaped; spermatheca (
<figureCitation id="01F82A79FFCBFFE746AD8935FC981077" box="[703,786,297,323]" captionStart="FIGURE 3" captionStartId="3.[151,250,1924,1949]" captionTargetBox="[232,1358,167,1898]" captionTargetId="figure@3.[227,1361,166,1905]" captionTargetPageId="3" captionText="FIGURE 3. Diagnostic features of Clidicus qiuae (A–D. Male. E–F. Female). A–C. Aedeagus, dorsal (A), lateral (B), and ventral (C). D. Sperm pump. E. Female genital segment. F. Spermatheca, enlarged. Scale bars: 0.5 mm in A–D; 0.2 mm in E; 0.1 mm in F." httpUri="https://zenodo.org/record/3255483/files/figure.png" pageId="5" pageNumber="326">Fig. 3F</figureCitation>
) strongly elongate, 
<quantity id="5E3B9B19FFCBFFE747E28935FB191077" box="[1008,1171,297,323]" metricMagnitude="-4" metricUnit="m" metricValue="3.95" metricValueMax="4.1" metricValueMin="3.8" pageId="5" pageNumber="326" unit="mm" value="0.395" valueMax="0.41" valueMin="0.38">0.38–0.41 mm</quantity>
(largest diameter of the spermatheca) long, with long connecting duct of accessory gland; proctiger rhomboidal, slightly wider than long, posterior margin densely fringed with long and dense setae; coxites elongate, 1.5× as long as wide, apex rounded, with setose margin.
</paragraph>
<paragraph id="997C36FCFFCBFFE744D589A4FA96130B" blockId="5.[151,1437,153,575]" pageId="5" pageNumber="326">
<emphasis id="ABB7EAEEFFCBFFE744D589A4FE3810E7" bold="true" box="[199,434,440,467]" pageId="5" pageNumber="326">Comparative notes.</emphasis>
The new species is morphologically close to several congeners from Indochina, 
<emphasis id="ABB7EAEEFFCBFFE7414289A6FAD210E7" box="[1360,1368,442,467]" italics="true" pageId="5" pageNumber="326">i</emphasis>
. 
<emphasis id="ABB7EAEEFFCBFFE7414D89A6FAE110E7" box="[1375,1387,442,467]" italics="true" pageId="5" pageNumber="326">e</emphasis>
., 
<taxonomicName id="5EC34D7FFFCBFFE7419189A5FEC710C3" authority="Orousset" authorityName="Orousset" authorityYear="2014" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="326" phylum="Arthropoda" rank="species" species="beroni">
<emphasis id="ABB7EAEEFFCBFFE7419189A5FA1C10E7" box="[1411,1430,441,467]" italics="true" pageId="5" pageNumber="326">C</emphasis>
. 
<emphasis id="ABB7EAEEFFCBFFE7448589C1FF5510C3" box="[151,223,477,503]" italics="true" pageId="5" pageNumber="326">beroni</emphasis>
Orousset
</taxonomicName>
, 
<taxonomicName id="5EC34D7FFFCBFFE7454889C1FCC810C3" authority="Jaloszynski, Hlavac &amp; Nomura" authorityName="Jaloszynski, Hlavac &amp; Nomura" authorityYear="2003" box="[346,834,477,503]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="326" phylum="Arthropoda" rank="species" species="bellator">
<emphasis id="ABB7EAEEFFCBFFE7454889C1FEE710C3" box="[346,365,477,503]" italics="true" pageId="5" pageNumber="326">C</emphasis>
. 
<emphasis id="ABB7EAEEFFCBFFE7456E89C1FE5F10C3" box="[380,469,477,503]" italics="true" pageId="5" pageNumber="326">bellator</emphasis>
Jałoszyński, Hlaváč &amp; Nomura
</taxonomicName>
, 
<taxonomicName id="5EC34D7FFFCBFFE7475F89C1FACC10C3" authority="Jaloszynski, Hlavac &amp; Nomura" authorityName="Jaloszynski, Hlavac &amp; Nomura" authorityYear="2003" box="[845,1350,477,503]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="326" phylum="Arthropoda" rank="species" species="rufescens">
<emphasis id="ABB7EAEEFFCBFFE7475F89C1FCEA10C3" box="[845,864,477,503]" italics="true" pageId="5" pageNumber="326">C</emphasis>
. 
<emphasis id="ABB7EAEEFFCBFFE7477D89C1FC5310C3" box="[879,985,477,503]" italics="true" pageId="5" pageNumber="326">rufescens</emphasis>
Jałoszyński, Hlaváč &amp; Nomura
</taxonomicName>
, and 
<taxonomicName id="5EC34D7FFFCBFFE7419189C1FDC6132F" authority="Jaloszynski, Hlavac &amp; Nomura" authorityName="Jaloszynski, Hlavac &amp; Nomura" authorityYear="2003" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="326" phylum="Arthropoda" rank="species" species="omoios">
<emphasis id="ABB7EAEEFFCBFFE7419189C1FA1C10C3" box="[1411,1430,477,503]" italics="true" pageId="5" pageNumber="326">C</emphasis>
. 
<emphasis id="ABB7EAEEFFCBFFE744858A1EFF62132F" box="[151,232,514,539]" italics="true" pageId="5" pageNumber="326">omoios</emphasis>
Jałoszyński, Hlaváč &amp; Nomura
</taxonomicName>
, sharing with them a more or less similarly general stout aedeagus with sym- metric parameres. 
<taxonomicName id="5EC34D7FFFCBFFE745758A39FD83130B" authorityName="Cheng &amp; Yin" authorityYear="2019" box="[359,521,549,575]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="326" phylum="Arthropoda" rank="species" species="qiuae">
<emphasis id="ABB7EAEEFFCBFFE745758A39FD83130B" box="[359,521,549,575]" italics="true" pageId="5" pageNumber="326">Clidicus qiuae</emphasis>
</taxonomicName>
can be readily separated from these species by the following features:
</paragraph>
<paragraph id="997C36FCFFCBFFE744858A71FCD7139F" blockId="5.[151,1437,620,935]" pageId="5" pageNumber="326">
- from 
<taxonomicName id="5EC34D7FFFCBFFE744F78A71FEC413B3" authorityName="Orousset" authorityYear="2014" box="[229,334,621,647]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="326" phylum="Arthropoda" rank="species" species="beroni">
<emphasis id="ABB7EAEEFFCBFFE744F78A71FF7213B3" box="[229,248,621,647]" italics="true" pageId="5" pageNumber="326">C</emphasis>
. 
<emphasis id="ABB7EAEEFFCBFFE745148A71FEC413B3" box="[262,334,621,647]" italics="true" pageId="5" pageNumber="326">beroni</emphasis>
</taxonomicName>
by a larger body size (
<quantity id="5E3B9B19FFCBFFE7465E8A70FD7B13B3" box="[588,753,620,647]" metricMagnitude="-3" metricUnit="m" metricValue="5.1899999999999995" metricValueMax="5.3" metricValueMin="5.08" pageId="5" pageNumber="326" unit="mm" value="5.1899999999999995" valueMax="5.3" valueMin="5.08">5.08–5.30 mm</quantity>
vs. 
<quantity id="5E3B9B19FFCBFFE7470D8A71FC0D13B3" box="[799,903,621,647]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="5" pageNumber="326" unit="mm" value="4.2">4.20 mm</quantity>
), laterally expanded parameres, and a relatively larger distal funnel of a much less elongate sperm pump;
</paragraph>
<paragraph id="997C36FCFFCBFFE744858AA9FC6F13C7" blockId="5.[151,1437,620,935]" pageId="5" pageNumber="326">
- from 
<taxonomicName id="5EC34D7FFFCBFFE744F58AA9FEE813FB" authorityName="Jaloszynski, Hlavac &amp; Nomura" authorityYear="2003" box="[231,354,693,719]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="326" phylum="Arthropoda" rank="species" species="bellator">
<emphasis id="ABB7EAEEFFCBFFE744F58AA9FF7013FB" box="[231,250,693,719]" italics="true" pageId="5" pageNumber="326">C</emphasis>
. 
<emphasis id="ABB7EAEEFFCBFFE7451B8AA9FEE813FB" box="[265,354,693,719]" italics="true" pageId="5" pageNumber="326">bellator</emphasis>
</taxonomicName>
by a larger body size (
<quantity id="5E3B9B19FFCBFFE746748AA8FC8613FB" box="[614,780,692,719]" metricMagnitude="-3" metricUnit="m" metricValue="5.1899999999999995" metricValueMax="5.3" metricValueMin="5.08" pageId="5" pageNumber="326" unit="mm" value="5.1899999999999995" valueMax="5.3" valueMin="5.08">5.08–5.30 mm</quantity>
vs. 
<quantity id="5E3B9B19FFCBFFE747298AA8FC6913FB" box="[827,995,692,719]" metricMagnitude="-3" metricUnit="m" metricValue="4.275" metricValueMax="4.4" metricValueMin="4.15" pageId="5" pageNumber="326" unit="mm" value="4.275" valueMax="4.4" valueMin="4.15">4.15–4.40 mm</quantity>
), more elongate antennomeres, a much more elongate sperm pump, and a different form of the spermatheca;
</paragraph>
<paragraph id="997C36FCFFCBFFE744858AE1FDB5126B" blockId="5.[151,1437,620,935]" pageId="5" pageNumber="326">
- from 
<taxonomicName id="5EC34D7FFFCBFFE744F58AE1FEF91223" authorityName="Jaloszynski, Hlavac &amp; Nomura" authorityYear="2003" box="[231,371,765,791]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="326" phylum="Arthropoda" rank="species" species="rufescens">
<emphasis id="ABB7EAEEFFCBFFE744F58AE1FF701223" box="[231,250,765,791]" italics="true" pageId="5" pageNumber="326">C</emphasis>
. 
<emphasis id="ABB7EAEEFFCBFFE7451B8AE1FEF91223" box="[265,371,765,791]" italics="true" pageId="5" pageNumber="326">rufescens</emphasis>
</taxonomicName>
by a much larger body size (
<quantity id="5E3B9B19FFCBFFE746AD8AE0FCEF1223" box="[703,869,764,791]" metricMagnitude="-3" metricUnit="m" metricValue="5.1899999999999995" metricValueMax="5.3" metricValueMin="5.08" pageId="5" pageNumber="326" unit="mm" value="5.1899999999999995" valueMax="5.3" valueMin="5.08">5.08–5.30 mm</quantity>
vs. 
<quantity id="5E3B9B19FFCBFFE747878AE0FBB71223" box="[917,1085,764,791]" metricMagnitude="-3" metricUnit="m" metricValue="3.585" metricValueMax="3.72" metricValueMin="3.45" pageId="5" pageNumber="326" unit="mm" value="3.585" valueMax="3.72" valueMin="3.45">3.45–3.72 mm</quantity>
), an impressed anterior margin of the frons (flat in 
<taxonomicName id="5EC34D7FFFCBFFE745AD8B3DFDC0120F" authorityName="Jaloszynski, Hlavac &amp; Nomura" authorityYear="2003" box="[447,586,801,827]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="326" phylum="Arthropoda" rank="species" species="rufescens">
<emphasis id="ABB7EAEEFFCBFFE745AD8B3DFE58120F" box="[447,466,801,827]" italics="true" pageId="5" pageNumber="326">C</emphasis>
. 
<emphasis id="ABB7EAEEFFCBFFE745F28B3DFDC0120F" box="[480,586,801,827]" italics="true" pageId="5" pageNumber="326">rufescens</emphasis>
</taxonomicName>
), the more elongate antennomeres, and different structures of the aedeagus, sperm pump, and spermatheca;
</paragraph>
<paragraph id="997C36FCFFCBFFE744858B75FBB31293" blockId="5.[151,1437,620,935]" pageId="5" pageNumber="326">
- from 
<taxonomicName id="5EC34D7FFFCBFFE744F68B75FEDF12B7" authorityName="Jaloszynski, Hlavac &amp; Nomura" authorityYear="2003" box="[228,341,873,899]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="326" phylum="Arthropoda" rank="species" species="omoios">
<emphasis id="ABB7EAEEFFCBFFE744F68B75FF7D12B7" box="[228,247,873,899]" italics="true" pageId="5" pageNumber="326">C</emphasis>
. 
<emphasis id="ABB7EAEEFFCBFFE745168B76FEDF12B7" box="[260,341,874,899]" italics="true" pageId="5" pageNumber="326">omoios</emphasis>
</taxonomicName>
by a larger body size (
<quantity id="5E3B9B19FFCBFFE746428B74FD7E12B7" box="[592,756,872,899]" metricMagnitude="-3" metricUnit="m" metricValue="5.1899999999999995" metricValueMax="5.3" metricValueMin="5.08" pageId="5" pageNumber="326" unit="mm" value="5.1899999999999995" valueMax="5.3" valueMin="5.08">5.08–5.30 mm</quantity>
vs. 
<quantity id="5E3B9B19FFCBFFE747328B75FC4D12B7" box="[800,967,873,899]" metricMagnitude="-3" metricUnit="m" metricValue="4.31" metricValueMax="4.4" metricValueMin="4.22" pageId="5" pageNumber="326" unit="mm" value="4.31" valueMax="4.4" valueMin="4.22">4.22–4.40 mm</quantity>
), the more elongate antennomeres, a more elongate aedeagus, and different forms of the sperm pump and spermatheca.
</paragraph>
</subSubSection>
<subSubSection id="D1D96577FFCBFFE744D58BC8FB2E154B" pageId="5" pageNumber="326" type="discussion">
<paragraph id="997C36FCFFCBFFE744D58BC8FB2E154B" blockId="5.[151,1437,980,1259]" pageId="5" pageNumber="326">
<emphasis id="ABB7EAEEFFCBFFE744D58BC8FEC512DB" bold="true" box="[199,335,980,1007]" pageId="5" pageNumber="326">Comments.</emphasis>
All individuals were collected under moist, thick layer of leaf litter within an hour in the morning. The collecting site was on a slope at the edge of the forest, with the searching area of approximately 
<quantity id="5E3B9B19FFCBFFE7410D8BE5FAD31527" box="[1311,1369,1017,1043]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="326" unit="m" value="30.0">30 m</quantity>
<superScript id="6EB69BB4FFCBFFE7414B8BE4FAEB1532" attach="left" box="[1369,1377,1016,1030]" fontSize="6" pageId="5" pageNumber="326">2</superScript>
(
<figureCitation id="01F82A79FFCBFFE7417D8BE5FF2C1503" captionStart="FIGURE 4" captionStartId="4.[151,250,1936,1961]" captionTargetBox="[151,1437,813,1917]" captionTargetId="figure@4.[151,1437,813,1917]" captionTargetPageId="4" captionText="FIGURE 4. Collecting environment of Clidicus qiuae. A, B. Collecting site—a slope at the edge of the forest. C, D. Exposed adults of Clidicus qiuae after the leaf litter was stirred (pictures provided by Hao Xu)." httpUri="https://zenodo.org/record/3255485/files/figure.png" pageId="5" pageNumber="326">Fig. 4</figureCitation>
A–B). Both collectors were looking for larvae of cetoniine beetles beneath the leaf litter layer, and then noticed the red, shining reflection of these large-sized scydmaenines after stirring the litters. After being exposed, the beetles were observed crawling around on the ground and the surface of the litter layer (
<figureCitation id="01F82A79FFCBFFE7400E8C79FBD4154B" box="[1052,1118,1125,1151]" captionStart="FIGURE 4" captionStartId="4.[151,250,1936,1961]" captionTargetBox="[151,1437,813,1917]" captionTargetId="figure@4.[151,1437,813,1917]" captionTargetPageId="4" captionText="FIGURE 4. Collecting environment of Clidicus qiuae. A, B. Collecting site—a slope at the edge of the forest. C, D. Exposed adults of Clidicus qiuae after the leaf litter was stirred (pictures provided by Hao Xu)." httpUri="https://zenodo.org/record/3255485/files/figure.png" pageId="5" pageNumber="326">Fig. 4</figureCitation>
C–D).
</paragraph>
</subSubSection>
<subSubSection id="D1D96577FFCBFFE744D58C94FD6A1597" box="[199,736,1160,1187]" pageId="5" pageNumber="326" type="distribution">
<paragraph id="997C36FCFFCBFFE744D58C94FD6A1597" blockId="5.[151,1437,980,1259]" box="[199,736,1160,1187]" pageId="5" pageNumber="326">
<emphasis id="ABB7EAEEFFCBFFE744D58C94FEEB1597" bold="true" box="[199,353,1160,1187]" pageId="5" pageNumber="326">Distribution.</emphasis>
Southern 
<collectingCountry id="E1D4766CFFCBFFE745C68C95FD911597" box="[468,539,1161,1187]" name="China" pageId="5" pageNumber="326">China</collectingCountry>
: 
<collectingRegion id="5B07F81EFFCBFFE746378C95FDF41597" box="[549,638,1161,1187]" country="China" name="Yunnan" pageId="5" pageNumber="326">Yunnan</collectingRegion>
(
<figureCitation id="01F82A79FFCBFFE7469F8C95FD581597" box="[653,722,1161,1187]" captionStart="FIGURE 7" captionStartId="8.[151,250,1375,1400]" captionTargetBox="[167,1421,489,1350]" captionTargetId="figure@8.[167,1421,489,1350]" captionTargetPageId="8" captionText="FIGURE 7. Distribution of extant species of Clidicus." httpUri="https://zenodo.org/record/3255489/files/figure.png" pageId="5" pageNumber="326">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D1D96577FFCBFFE744D58CB1FF7A15DF" pageId="5" pageNumber="326" type="etymology">
<paragraph id="997C36FCFFCBFFE744D58CB1FF7A15DF" blockId="5.[151,1437,980,1259]" pageId="5" pageNumber="326">
<emphasis id="ABB7EAEEFFCBFFE744D58CB1FEC515F3" bold="true" box="[199,335,1197,1223]" pageId="5" pageNumber="326">Etymology.</emphasis>
The new species is named after our friend, Jian-Yue Qiu, who was one of the collectors of the new species.
</paragraph>
</subSubSection>
</treatment>
</document>