128 lines
15 KiB
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128 lines
15 KiB
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<document id="B6748C590388D1DC7B769F2686CD56BD" ID="10.11646/zootaxa.4157.1.1" ID-DOI="10.11646/zootaxa.4157.1.1" ID-GBIF-Dataset="2389f078-1812-4b4e-826a-561cd7e46654" ID-ISSN="1175-5326" ID-Zenodo-Dep="262413" ID-ZooBank="6BF4514A-892F-499F-BC1E-B7920C7A00B0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1472198526050" checkinUser="plazi" docAuthor="Jałoszyński, Paweł & Perkovsky, Evgeny" docDate="2016" docId="03A187A9292DFFE5FF03C5B59E59FD76" docLanguage="en" docName="zootaxa.4157.1.1.pdf" docOrigin="Zootaxa 4157 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Vertheia" docType="treatment" docVersion="9" lastPageNumber="21" masterDocId="FF98FFD12939FFF1FF94C52D9D54FFD1" masterDocTitle="Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber" masterLastPageNumber="85" masterPageNumber="1" pageNumber="21" updateTime="1698382676775" updateUser="plazi">
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<mods:mods id="883C21EAE9785526B14B8A404D4990CA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="58086ABA910A1A27E42160C48E44B1B1">
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<mods:title id="23C3EC5B74907597AD6DFEC4B2D444AF">Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber</mods:title>
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</mods:titleInfo>
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<mods:name id="1FFC2B8B1360559344BABD7255FBB72C" type="personal">
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<mods:role id="E9647B7B9250BDDE1E2C2953207EE0BA">
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<mods:roleTerm id="0B4C06193C70BAA00BC5058D563CE689">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3CDAC58B3D524A1564B0659278E940EF">Jałoszyński, Paweł</mods:namePart>
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</mods:name>
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<mods:name id="73F83B550D6B8C01073ABEC9375F1BE1" type="personal">
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<mods:role id="122E0DD5D04ADBDF434FB3F5DBE8C58A">
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<mods:roleTerm id="FCE5590653C15913DC9104868F2F6058">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="67C96BC07636C69A5C697A0B10BF9823">Perkovsky, Evgeny</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="7550DEF079BE9A05DDA6E669F93DAC56">text</mods:typeOfResource>
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<mods:relatedItem id="968C705C30B287B3548F459F35CBBB30" type="host">
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<mods:titleInfo id="C0D7CF0EFFE0D89C176DDE342F2BDC95">
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<mods:title id="25049377698A5C1E1504CB93FAAB44EF">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="1114F1C933DCD69CB779B5EE05E75B25">
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<mods:date id="0AAF513D447A63EFEB909B83E9D866B2">2016</mods:date>
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<mods:detail id="FDCCD731FA5E62D92559774174DE3D47" type="volume">
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<mods:number id="9056214F2625251E84A3413086A92194">4157</mods:number>
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</mods:detail>
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<mods:detail id="48B4DE3BC078CDE46CCD3D270C23D72D" type="issue">
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<mods:number id="3A7DB06725407E6CFCD6CCA344DBC98A">1</mods:number>
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</mods:detail>
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<mods:extent id="59C9CB0CE9C52F68D5C400D27E8406A8" unit="page">
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<mods:start id="6AE1057D5AEF5B6586B4FA219E80B7DB">1</mods:start>
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<mods:end id="C13CB5385044ED1CC73ABAE3249BB5AE">85</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="A6117B74475ABA9361E072DECD65CE04">journal article</mods:classification>
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<mods:identifier id="4BF05714AAE1346A51E471D89DCD55CB" type="DOI">10.11646/zootaxa.4157.1.1</mods:identifier>
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<mods:identifier id="AA17B840BF70D6C7208DDF6EECF1A9F1" type="GBIF-Dataset">2389f078-1812-4b4e-826a-561cd7e46654</mods:identifier>
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<mods:identifier id="778E0700EFB957A8FFB00EC9F4425572" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="C3320AA018E284967844C5068971ACA7" type="Zenodo-Dep">262413</mods:identifier>
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<mods:identifier id="5330E3C1070C4006C5181162E8C9D038" type="ZooBank">6BF4514A-892F-499F-BC1E-B7920C7A00B0</mods:identifier>
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</mods:mods>
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<treatment id="03A187A9292DFFE5FF03C5B59E59FD76" ID-DOI="http://doi.org/10.5281/zenodo.5681739" ID-GBIF-Taxon="124407824" ID-Zenodo-Dep="5681739" LSID="urn:lsid:plazi:treatment:03A187A9292DFFE5FF03C5B59E59FD76" httpUri="http://treatment.plazi.org/id/03A187A9292DFFE5FF03C5B59E59FD76" lastPageNumber="21" pageId="20" pageNumber="21">
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<subSubSection id="C3126534292DFFE5FF03C5B59C17FF02" pageId="20" pageNumber="21" type="nomenclature">
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<paragraph id="8BB736BF292DFFE5FF03C5B59C1FFF63" blockId="20.[151,331,151,211]" box="[151,331,151,178]" pageId="20" pageNumber="21">
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<heading id="D0FF81D3292DFFE5FF03C5B59C1FFF63" bold="true" box="[151,331,151,178]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
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<emphasis id="B97CEAAD292DFFE5FF03C5B59C1FFF63" bold="true" box="[151,331,151,178]" pageId="20" pageNumber="21">
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†
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<taxonomicName id="4C084D3C292DFFE5FF31C5BA9C53FF60" box="[165,263,151,177]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B97CEAAD292DFFE5FF31C5BA9C53FF60" bold="true" box="[165,263,151,177]" italics="true" pageId="20" pageNumber="21">Vertheia</emphasis>
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</taxonomicName>
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sp. 1
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB736BF292DFFE5FF03C5979C17FF02" blockId="20.[151,331,151,211]" box="[151,323,186,211]" pageId="20" pageNumber="21">
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(
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<figureCitation id="13332A3A292DFFE5FF0BC5979DB3FF02" box="[159,231,186,211]" captionStart="FIGURES 1 – 20" captionStartId="7.[151,264,1794,1817]" captionTargetBox="[157,1423,512,1763]" captionTargetId="figure@7.[151,1436,503,1772]" captionTargetPageId="7" captionText="FIGURES 1 – 20. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 27418, Rovnoleptochromus ableptonoides sp. n. (1); K- 243, Vertheia quadrisetosa sp. n. (2); UA- 442, Vertheia sp. 1 (3); K- 800, Cephennodes sp. 1 (4); K- 6011, Cephennomicrus giganteus sp. n. (5); K- 8289, Cephennomicrus sp. 1 (6); K- 7010, Cephennomicrus sp. 2 (7); K- 27565, Cephennomicrus sp. 3 (8); K- 8912, Cephennomicrus sp. 4 (9); DU- 32, Cephennomicrus sp. 5 (10); K- 5770, Glaesoconnus unicus sp. n. (11); K- 9579, Rovnoscydmus frontalis sp. n. (12); K- 347, Rovnoscydmus microscopicus sp. n. (13); K- 25702, Rovnoscydmus sp. specimen 1 (14); K- 26348, Rovnoscydmus sp. specimen 2 (15); UA- 1556, Rovnoscydmus sp. specimen 3 (16); K- 8970, Rovnoscydmus sp. specimen 4 (17); K- 8968, Rovnoscydmus sp. specimen 5 (18); K- 7350, Rovnoscydmus sp. specimen 6 (19); K- 3037, Rovnoscydmus sp. specimen 7 (20)." httpUri="https://zenodo.org/record/262414/files/figure.png" pageId="20" pageNumber="21">Figs 3</figureCitation>
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,
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<figureCitation id="13332A3A292DFFE5FF67C5979C6FFF02" box="[243,315,186,211]" captionStart-0="FIGURES 56 – 60" captionStart-1="FIGURES 61 – 64" captionStartId-0="19.[151,264,1962,1985]" captionStartId-1="21.[151,264,1846,1869]" captionTargetBox-0="[151,1436,628,1941]" captionTargetBox-1="[160,1393,591,1818]" captionTargetId-0="figure@19.[151,1436,628,1941]" captionTargetId-1="figure@21.[151,1436,591,1824]" captionTargetPageId-0="19" captionTargetPageId-1="21" captionText-0="FIGURES 56 – 60. Vertheia sp. 1 (UA- 442). Approximately anterodorsal view (56, 57), lateral view (58), ventral view of head and posteroventral view of thorax and abdomen (59, 60)." captionText-1="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri-0="https://zenodo.org/record/262421/files/figure.png" httpUri-1="https://zenodo.org/record/262422/files/figure.png" pageId="20" pageNumber="21">56–64</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126534292DFFE5FF03C42F9E40FEEE" pageId="20" pageNumber="21" type="materials_examined">
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<paragraph id="8BB736BF292DFFE5FF03C42F9E40FEEE" blockId="20.[151,1436,258,679]" pageId="20" pageNumber="21">
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<materialsCitation id="3B603CE2292DFFE5FF03C42F9E40FEEE" ID-GBIF-Occurrence="1300093512" collectionCode="SIZK" county="Material" location="Europe" municipality="Late Eocene" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Rovno">
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<emphasis id="B97CEAAD292DFFE5FF03C42F9E61FECA" bold="true" box="[151,821,258,283]" pageId="20" pageNumber="21">
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<collectingCounty id="62D64E33292DFFE5FF03C42F9DABFECA" box="[151,255,258,283]" pageId="20" pageNumber="21">Material</collectingCounty>
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studied.
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<collectingMunicipality id="6BD3ACC5292DFFE5FEFFC42F9CAAFECA" box="[363,510,258,283]" pageId="20" pageNumber="21">Late Eocene</collectingMunicipality>
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of
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<location id="8ED76064292DFFE5FDB7C42F9F29FECA" LSID="urn:lsid:plazi:treatment:03A187A9292DFFE5FF03C5B59E59FD76:8ED76064292DFFE5FDB7C42F9F29FECA" box="[547,637,258,283]" county="Material" municipality="Late Eocene" name="Europe" pageId="20" pageNumber="21" stateProvince="Rovno">Europe</location>
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,
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<collectingRegion id="49CCF85D292DFFE5FD1DC42F9F83FECA" box="[649,727,258,283]" country="Ukraine" name="Rivne" pageId="20" pageNumber="21">Rovno</collectingRegion>
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amber:
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</emphasis>
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sex unknown; inclusion in an elongate prism of amber
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<quantity id="4CF09B5A292DFFE5FF03C40A9DB2FEEF" box="[151,230,295,319]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="21" unit="mm" value="20.0">20 mm</quantity>
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long (
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<figureCitation id="13332A3A292DFFE5FEBAC40B9C25FEEE" box="[302,369,294,319]" captionStart="FIGURES 1 – 20" captionStartId="7.[151,264,1794,1817]" captionTargetBox="[157,1423,512,1763]" captionTargetId="figure@7.[151,1436,503,1772]" captionTargetPageId="7" captionText="FIGURES 1 – 20. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 27418, Rovnoleptochromus ableptonoides sp. n. (1); K- 243, Vertheia quadrisetosa sp. n. (2); UA- 442, Vertheia sp. 1 (3); K- 800, Cephennodes sp. 1 (4); K- 6011, Cephennomicrus giganteus sp. n. (5); K- 8289, Cephennomicrus sp. 1 (6); K- 7010, Cephennomicrus sp. 2 (7); K- 27565, Cephennomicrus sp. 3 (8); K- 8912, Cephennomicrus sp. 4 (9); DU- 32, Cephennomicrus sp. 5 (10); K- 5770, Glaesoconnus unicus sp. n. (11); K- 9579, Rovnoscydmus frontalis sp. n. (12); K- 347, Rovnoscydmus microscopicus sp. n. (13); K- 25702, Rovnoscydmus sp. specimen 1 (14); K- 26348, Rovnoscydmus sp. specimen 2 (15); UA- 1556, Rovnoscydmus sp. specimen 3 (16); K- 8970, Rovnoscydmus sp. specimen 4 (17); K- 8968, Rovnoscydmus sp. specimen 5 (18); K- 7350, Rovnoscydmus sp. specimen 6 (19); K- 3037, Rovnoscydmus sp. specimen 7 (20)." httpUri="https://zenodo.org/record/262414/files/figure.png" pageId="20" pageNumber="21">Fig. 3</figureCitation>
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), collection number UA-442 (
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<collectionCode id="ED19AE7A292DFFE5FD50C40A9E5CFEEE" box="[708,776,295,319]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="20" pageNumber="21">SIZK</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126534292DFFE5FF53C4679E59FD76" pageId="20" pageNumber="21" type="discussion">
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<paragraph id="8BB736BF292DFFE5FF53C4679EFFFD8E" blockId="20.[151,1436,258,679]" pageId="20" pageNumber="21">
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<emphasis id="B97CEAAD292DFFE5FF53C4679C6FFEB2" bold="true" box="[199,315,330,355]" pageId="20" pageNumber="21">Remarks.</emphasis>
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This specimen is very similar to
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<taxonomicName id="4C084D3C292DFFE5FD77C4619EC2FEB2" box="[739,918,330,355]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="quadrisetosa">
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<emphasis id="B97CEAAD292DFFE5FD77C4619EC2FEB2" box="[739,918,330,355]" italics="true" pageId="20" pageNumber="21">V. quadrisetosa</emphasis>
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</taxonomicName>
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, but it lacks any remarkable characters to unambiguously define a new species. The differences include significantly lower EI (i.e., stouter elytra), slightly smaller BL, shorter antennae, lack of the long setae near posteromesal margins of eyes and lateral pronotal setae, and the presence of complete transverse antebasal groove, i.e., connecting lateral antebasal impressions. However, the setae may be broken off, the pronotal groove may show some variation, and proportions of the body parts and the body length may be different in males and females of the same species. For this reason, the specimen of
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<taxonomicName id="4C084D3C292DFFE5FF03C70F9DA6FDEA" box="[151,242,546,571]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B97CEAAD292DFFE5FF03C70F9DA6FDEA" box="[151,242,546,571]" italics="true" pageId="20" pageNumber="21">Vertheia</emphasis>
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</taxonomicName>
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that shows more unique, remarkable characters was selected as a
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<typeStatus id="54B3881D292DFFE5FC76C70F9910FDEA" box="[994,1092,546,571]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
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of a new species, whereas the other one, slightly different and less remarkable, remains undescribed.
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</paragraph>
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<paragraph id="8BB736BF292DFFE5FF53C7469E59FD76" blockId="20.[151,1436,258,679]" pageId="20" pageNumber="21">
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Measurements of
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<taxonomicName id="4C084D3C292DFFE5FE07C7479CBAFD52" box="[403,494,618,643]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B97CEAAD292DFFE5FE07C7479CBAFD52" box="[403,494,618,643]" italics="true" pageId="20" pageNumber="21">Vertheia</emphasis>
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</taxonomicName>
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sp. (UA-442): BL
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<quantity id="4CF09B5A292DFFE5FD58C7469E60FD53" box="[716,820,619,643]" metricMagnitude="-4" metricUnit="m" metricValue="9.3" pageId="20" pageNumber="21" unit="mm" value="0.93">0.93 mm</quantity>
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; HL
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<quantity id="4CF09B5A292DFFE5FCFBC7479E83FD53" box="[879,983,618,643]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="21" unit="mm" value="0.15">0.15 mm</quantity>
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, HW
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<quantity id="4CF09B5A292DFFE5FB8FC74699D7FD53" box="[1051,1155,619,643]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="20" pageNumber="21" unit="mm" value="0.24">0.24 mm</quantity>
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, AnL
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<quantity id="4CF09B5A292DFFE5FB58C7469861FD53" box="[1228,1333,619,643]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="20" pageNumber="21" unit="mm" value="0.33">0.33 mm</quantity>
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; PL
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<quantity id="4CF09B5A292DFFE5FAFFC7469D91FD77" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="20" pageNumber="21" unit="mm" value="0.28">0.28 mm</quantity>
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, PW
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<quantity id="4CF09B5A292DFFE5FE95C7A29C3CFD77" box="[257,360,655,679]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="20" pageNumber="21" unit="mm" value="0.3">0.30 mm</quantity>
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; EL
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<quantity id="4CF09B5A292DFFE5FE09C7A39F57FD77" box="[413,515,654,679]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="20" pageNumber="21" unit="mm" value="0.5">0.50 mm</quantity>
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, EW
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<quantity id="4CF09B5A292DFFE5FDD5C7A29FF3FD77" box="[577,679,655,679]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="20" pageNumber="21" unit="mm" value="0.48">0.48 mm</quantity>
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, EI 1.05.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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