treatments-xml/data/F7/A0/D5/F7A0D5A99DD1186964EF11836DB83DB6.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.743.23029" ID-GBIF-Dataset="1e5fbbb7-a293-466f-b354-9eb70c8befe7" ID-PMC="PMC5904386" ID-Pensoft-Pub="1313-2970-743-67" ID-PubMed="29670437" ID-ZBK="4D31DBBF07B1479C90489C9207B5103D" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-743-67" ModsDocOrigin="ZooKeys 743" ModsDocTitle="Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae)" checkinTime="1521107529766" checkinUser="pensoft" docAuthor="Kohlmann, Bert, Arriaga-Jimenez, Alfonsina &amp; Roes, Matthias" docDate="2018" docId="F7A0D5A99DD1186964EF11836DB83DB6" docLanguage="en" docName="ZooKeys 743: 67-93" docOrigin="ZooKeys 743" docSource="http://dx.doi.org/10.3897/zookeys.743.23029" docTitle="Geotrupes (Onthotrupes) pecki Howden 1974" docType="treatment" docVersion="4" lastPageNumber="79" masterDocId="FF813C19FFB8FFF8FFB7FFACFF851225" masterDocTitle="Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae)" masterLastPageNumber="93" masterPageNumber="67" pageNumber="79" updateTime="1668165570053" updateUser="ExternalLinkService">
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<mods:title>Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae)</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kohlmann, Bert</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Arriaga-Jimenez, Alfonsina</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Roes, Matthias</mods:namePart>
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<mods:date>2018</mods:date>
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<mods:number>743</mods:number>
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<mods:start>67</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.743.23029</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.743.23029</mods:identifier>
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<treatment ID-GBIF-Taxon="142283983" LSID="urn:lsid:plazi:treatment:F7A0D5A99DD1186964EF11836DB83DB6" httpUri="http://treatment.plazi.org/id/F7A0D5A99DD1186964EF11836DB83DB6" lastPageId="13" lastPageNumber="79" pageId="12" pageNumber="79">
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<paragraph pageId="12" pageNumber="79">
<taxonomicName authority="Howden, 1974" authorityName="Howden" authorityYear="1974" class="Insecta" family="Scarabaeidae" genus="Geotrupes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geotrupes (Onthotrupes) pecki" order="Coleoptera" pageId="12" pageNumber="79" phylum="Arthropoda" rank="species" species="pecki" subGenus="Onthotrupes">Geotrupes (Onthotrupes) pecki Howden, 1974</taxonomicName>
Figs 6, 9 
<normalizedToken originalValue="a–b">a-b</normalizedToken>
, 10b
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="79" type="reference_group">
<paragraph pageId="12" pageNumber="79">
<taxonomicName class="Insecta" family="Scarabaeidae" genus="Geotrupes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geotrupes pecki" order="Coleoptera" pageId="12" pageNumber="79" phylum="Arthropoda" rank="species" species="pecki">Geotrupes pecki</taxonomicName>
: 
<bibRefCitation author="Howden, HF" journalOrPublisher="Canadian Journal of Zoology" pageId="20" pageNumber="87" pagination="567 - 573" title="Additional records and descriptions of North and Central American Geotrupinae (Coleoptera, Scarabaeidae)." url="https://doi.org/10.1139/z74-073" volume="52" year="1974">Howden (1974</bibRefCitation>
: 572, Plate II), 
<bibRefCitation author="Trotta-Moreu, N" journalOrPublisher="Acta Zoologica Mexicana NS" pageId="21" pageNumber="88" pagination="39 - 65" title="Distribucion conocida y potencial de las especies de Geotrupinae (Coleoptera: Scarabaeoidea) en Mexico." volume="24" year="2008">Trotta-Moreu et al. (2008</bibRefCitation>
: 43, 47, 50, 51), 
<bibRefCitation author="Trotta-Moreu, N" journalOrPublisher="Community and Ecosystem Ecology" pageId="21" pageNumber="88" pagination="42 - 49" title="Deriving the species richness distribution of Geotrupinae (Coleoptera: Scarabaeoidea) in Mexico from the overlap of individual model predictions." url="https://doi.org/10.1603/EN08179" volume="39" year="2010">Trotta-Moreu and Lobo (2010</bibRefCitation>
: 46).
</paragraph>
</subSubSection>
<subSubSection lastPageId="13" lastPageNumber="80" pageId="12" pageNumber="79" type="remarks">
<paragraph pageId="12" pageNumber="79">Remarks.</paragraph>
<paragraph pageId="12" pageNumber="79">
This species has been recorded only once in the literature, more than 40 years ago, when Howden described it in 1974 in the Sierra Sur (Sierra de 
<normalizedToken originalValue="Miahuatlán">Miahuatlan</normalizedToken>
). Using the only two known distribution records reported by 
<bibRefCitation author="Howden, HF" journalOrPublisher="Canadian Journal of Zoology" pageId="20" pageNumber="87" pagination="567 - 573" title="Additional records and descriptions of North and Central American Geotrupinae (Coleoptera, Scarabaeidae)." url="https://doi.org/10.1139/z74-073" volume="52" year="1974">Howden (1974)</bibRefCitation>
, 
<bibRefCitation author="Trotta-Moreu, N" journalOrPublisher="Acta Zoologica Mexicana NS" pageId="21" pageNumber="88" pagination="39 - 65" title="Distribucion conocida y potencial de las especies de Geotrupinae (Coleoptera: Scarabaeoidea) en Mexico." volume="24" year="2008">Trotta-Moreu et al. (2008)</bibRefCitation>
performed an analysis of the known and potential distribution of 
<taxonomicName lsidName="G. pecki" pageId="12" pageNumber="79" rank="species" species="pecki">G. pecki</taxonomicName>
in the state of Oaxaca using the MaxEnt prediction model (
<bibRefCitation author="Phillips, SJ" journalOrPublisher="Ecological Modelling" pageId="21" pageNumber="88" pagination="231 - 259" title="Maximum entropy modelling of species geographic distributions." url="https://doi.org/10.1016/j.ecolmodel.2005.03.026" volume="190" year="2006">Phillips et al. 2006</bibRefCitation>
). For 
<taxonomicName lsidName="G. pecki" pageId="12" pageNumber="79" rank="species" species="pecki">G. pecki</taxonomicName>
, 
<bibRefCitation author="Trotta-Moreu, N" journalOrPublisher="Acta Zoologica Mexicana NS" pageId="21" pageNumber="88" pagination="39 - 65" title="Distribucion conocida y potencial de las especies de Geotrupinae (Coleoptera: Scarabaeoidea) en Mexico." volume="24" year="2008">Trotta-Moreu et al. (2008)</bibRefCitation>
estimated a potential distribution area of 5254 km2 and mean values for altitude (1862 m), rainfall (1524 
<normalizedToken originalValue="±">+/-</normalizedToken>
287 mm), and temperature (18.5 
<normalizedToken originalValue="±">+/-</normalizedToken>
5 °C). With the present collecting localities, we can recalculate and correct the 
<bibRefCitation author="Trotta-Moreu, N" journalOrPublisher="Acta Zoologica Mexicana NS" pageId="21" pageNumber="88" pagination="39 - 65" title="Distribucion conocida y potencial de las especies de Geotrupinae (Coleoptera: Scarabaeoidea) en Mexico." volume="24" year="2008">Trotta-Moreu et al. (2008)</bibRefCitation>
too low mean value for altitude for this species.
</paragraph>
<paragraph pageId="12" pageNumber="79">
Below, an updated key for the genus 
<taxonomicName class="Insecta" family="Geotrupidae" genus="Geotrupes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geotrupes" order="Coleoptera" pageId="12" pageNumber="79" phylum="Arthropoda" rank="genus">Geotrupes</taxonomicName>
in Oaxaca is presented. 
<taxonomicName lsidName="G. truncaticornis" pageId="12" pageNumber="79" rank="species" species="truncaticornis">G. truncaticornis</taxonomicName>
Howden is not included in the key; because it is known only from Guerrero. 
<bibRefCitation author="Trotta-Moreu, N" journalOrPublisher="Acta Zoologica Mexicana NS" pageId="21" pageNumber="88" pagination="39 - 65" title="Distribucion conocida y potencial de las especies de Geotrupinae (Coleoptera: Scarabaeoidea) en Mexico." volume="24" year="2008">Trotta-Moreu et al. (2008)</bibRefCitation>
erroneously reported it from Oaxaca (Kohlmann in press).
</paragraph>
<paragraph lastPageId="13" lastPageNumber="80" pageId="12" pageNumber="79">
Material (6 males, 8 females). Holotype: 
<normalizedToken originalValue="México">Mexico</normalizedToken>
.10 km E Sn 
<normalizedToken originalValue="Sebastián">Sebastian</normalizedToken>
<normalizedToken originalValue="Río">Rio</normalizedToken>
Hondo, 
<normalizedToken originalValue="Miahuatlán">Miahuatlan</normalizedToken>
, Oaxaca. 31-VIII-17, coprotrampa, 
<geoCoordinate direction="north" orientation="latitude" precision="15" value="16.198889">16°11'56&quot;N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-96.365">96°21'54&quot;W</geoCoordinate>
, bosque pino/ 
<normalizedToken originalValue="pingüica">pingueica</normalizedToken>
, 2930 m, Arriaga J. A. Col. Allotype: 
<normalizedToken originalValue="México">Mexico</normalizedToken>
. 4 km NO Sto. Domingo Ozolotepec, 
<normalizedToken originalValue="Miahuatlán">Miahuatlan</normalizedToken>
, Oaxaca. 31-VIII-17, coprotrampa, 
<geoCoordinate direction="north" orientation="latitude" precision="15" value="16.199165">16°11'57&quot;N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-96.35">96°21&quot;W</geoCoordinate>
, bosque pino/ 
<normalizedToken originalValue="pingüica">pingueica</normalizedToken>
, 2920 m, Arriaga J. A. Col. Paratypes: 
<normalizedToken originalValue="México">Mexico</normalizedToken>
. 4 
<pageBreakToken pageId="13" pageNumber="80" start="start">km</pageBreakToken>
NW Sto. Domingo Ozolotepec, 
<normalizedToken originalValue="Miahuatlán">Miahuatlan</normalizedToken>
, Oaxaca. 31-VIII-17, coprotrampa, 
<geoCoordinate direction="north" orientation="latitude" precision="15" value="16.198889">16°11'56&quot;N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-96.365555">96°21'56&quot;W</geoCoordinate>
, bosque pino/ 
<normalizedToken originalValue="pingüica">pingueica</normalizedToken>
, 2930 m, Arriaga J. A. Col. (1 male, 1 female), 
<geoCoordinate direction="north" orientation="latitude" precision="15" value="16.199165">16°11'57&quot;N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-96.35">96°21&quot;W</geoCoordinate>
, 2920 m, (one male, three females); 3 km NW Sto. Domingo Ozolotepec, 
<geoCoordinate direction="north" orientation="latitude" precision="15" value="16.198055">16°11'53&quot;N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-96.333336">96°20&quot;W</geoCoordinate>
, 2740 m, (2 males); 8 km E San 
<normalizedToken originalValue="Sebastián">Sebastian</normalizedToken>
<normalizedToken originalValue="Río">Rio</normalizedToken>
Hondo, 30-VIII-17, C. D. exc. Vaca, 
<geoCoordinate direction="north" orientation="latitude" precision="15" value="16.1875">16°11'15&quot;N</geoCoordinate>
, 
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-96.3875">96°23'15&quot;W</geoCoordinate>
, 2900 m, (1 male, 3 females).
</paragraph>
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