treatments-xml/data/F7/1D/87/F71D87D5FFBB856C62C2C71EFA989354.xml

<document id="C292E606AF3FB8D4DC857B9922D10120" ID-DOI="10.11646/zootaxa.4006.3.5" ID-GBIF-Dataset="cf6a819b-c985-4328-b16b-ce638a3c5dd6" ID-ISSN="1175-5326" ID-Zenodo-Dep="231910" ID-ZooBank="8F133516-4CB5-4BBD-9DBF-2EE7C2EA2F55" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461214103462" checkinUser="plazi" docAuthor="Podenas, Sigitas, Podeniene, Virginija &amp; Byun, Hye-Woo" docDate="2015" docId="F71D87D5FFBB856C62C2C71EFA989354" docLanguage="en" docName="zt04006p520.pdf" docOrigin="Zootaxa 4006 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heterangaeus koreanus Podenas, Podeniene &amp; Byun, 2015, n. sp." docType="treatment" docVersion="9" lastPageNumber="518" masterDocId="0B24FFADFFBE85606255C53DFFA6962E" masterDocTitle="Heterangaeus Alexander, 1925 crane flies (Diptera: Pediciidae) of Korea" masterLastPageNumber="520" masterPageNumber="506" pageNumber="511" updateTime="1698602364234" updateUser="plazi">
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<mods:title id="067C0BD676EED05DA174B86F6594E1D7">Heterangaeus Alexander, 1925 crane flies (Diptera: Pediciidae) of Korea</mods:title>
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<mods:namePart id="75CF9BC3FCCEE549B9F4788B8015D2F8">Podenas, Sigitas</mods:namePart>
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<mods:namePart id="F51CA46BF9BFE3A43DC1F97CF002A4D4">Podeniene, Virginija</mods:namePart>
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<mods:namePart id="CA0D15A52400D245E0D9F2ADD19A1C31">Byun, Hye-Woo</mods:namePart>
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<mods:date id="88BEFCE2625862A9AE0FC43969346627">2015</mods:date>
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<treatment id="F71D87D5FFBB856C62C2C71EFA989354" ID-DOI="http://doi.org/10.5281/zenodo.6100844" ID-GBIF-Taxon="119630752" ID-Zenodo-Dep="6100844" LSID="urn:lsid:plazi:treatment:F71D87D5FFBB856C62C2C71EFA989354" httpUri="http://treatment.plazi.org/id/F71D87D5FFBB856C62C2C71EFA989354" lastPageId="12" lastPageNumber="518" pageId="5" pageNumber="511">
<subSubSection id="37AE6548FFBB856562C2C71EFDA79410" box="[151,513,547,574]" pageId="5" pageNumber="511" type="nomenclature">
<paragraph id="7F0B36C3FFBB856562C2C71EFDA79410" blockId="5.[151,513,547,574]" box="[151,513,547,574]" pageId="5" pageNumber="511">
<heading id="244381AFFFBB856562C2C71EFDA79410" bold="true" box="[151,513,547,574]" fontSize="11" level="1" pageId="5" pageNumber="511" reason="1">
<emphasis id="4DC0EAD1FFBB856562C2C71EFDA79410" bold="true" box="[151,513,547,574]" pageId="5" pageNumber="511">
<taxonomicName id="B8B44D40FFBB856562C2C71EFE119413" ID-CoL="8GT3R" box="[151,439,547,573]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="511" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFBB856562C2C71EFE119413" bold="true" box="[151,439,547,573]" italics="true" pageId="5" pageNumber="511">Heterangaeus koreanus</emphasis>
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<taxonomicNameLabel id="56F357AAFFBB856563EAC719FDA79410" box="[447,513,548,574]" pageId="5" pageNumber="511" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="37AE6548FFBB856562C2C757FC01923B" pageId="5" pageNumber="511" type="diagnosis">
<paragraph id="7F0B36C3FFBB856562C2C757FC01923B" blockId="5.[151,1437,618,2029]" pageId="5" pageNumber="511">
<emphasis id="4DC0EAD1FFBB856562C2C757FEAC94AD" bold="true" box="[151,266,618,643]" pageId="5" pageNumber="511">Diagnosis</emphasis>
. Medium-size yellowish species. Antenna very short, if bent backwards, reaching pronotum at most; dark brown at base, turning whitish at apex. Distinguished from other species in genus by shape of wing, details of venation, wing pattern, and structure of male terminalia. Wing (
<figureCitation id="E78F2A46FFBB856561CCC78FFC4794E5" box="[921,993,690,715]" captionStart="FIGURES 5 – 11" captionStartId="7.[151,264,1874,1897]" captionTargetBox="[171,1425,213,1834]" captionTargetId="figure@7.[151,1436,193,1853]" captionTargetPageId="7" captionText="FIGURES 5 – 11. Heterangaeus koreanus n. sp., paratypes. 5, wing; 6, gonostylus; 7, male genitalia, dorsal view; 8, aedeagus; 9, interbase; 10, ninth sternite; 11, ovipositor, lateral view." httpUri="https://zenodo.org/record/231912/files/figure.png" pageId="5" pageNumber="511">Fig. 5</figureCitation>
) paddle-shaped, with clearly sinuous posterior margin, distinct emarginations not only at tips of both anal veins, but also at posterior margin of second anal cell. Anal angle large, second anal cell wide. Vein 
<emphasis id="4DC0EAD1FFBB85656147C7C6FC929539" box="[786,820,763,791]" italics="true" pageId="5" pageNumber="511">Sc2</emphasis>
well before base of 
<emphasis id="4DC0EAD1FFBB85656675C7C1FB9A953D" box="[1056,1084,764,787]" italics="true" pageId="5" pageNumber="511">Rs</emphasis>
; cell 
<emphasis id="4DC0EAD1FFBB85656628C7C1FB3C9539" box="[1149,1178,764,791]" italics="true" pageId="5" pageNumber="511">m1</emphasis>
with short stem; basal deflection of 
<emphasis id="4DC0EAD1FFBB8565636FC61FFED29510" box="[314,372,802,830]" italics="true" pageId="5" pageNumber="511">CuA1</emphasis>
distinctly beyond branching point of 
<emphasis id="4DC0EAD1FFBB85656111C61FFCFD9517" box="[836,859,802,825]" italics="true" pageId="5" pageNumber="511">M</emphasis>
. Wing with distinct and abundant dark pattern. Pigmented areas surrounded by darker margins. The most distinctive are two cross-bands, connected along vein 
<emphasis id="4DC0EAD1FFBB8565672CC674FA33954E" box="[1401,1429,841,864]" italics="true" pageId="5" pageNumber="511">Rs</emphasis>
, thus forming “8”-shaped light “window” in the middle; Costal cell mostly clear, basally with weak brownish tinge. Male gonostylus (
<figureCitation id="E78F2A46FFBB85656332C6ADFE0B9587" box="[359,429,912,937]" captionStart="FIGURES 5 – 11" captionStartId="7.[151,264,1874,1897]" captionTargetBox="[171,1425,213,1834]" captionTargetId="figure@7.[151,1436,193,1853]" captionTargetPageId="7" captionText="FIGURES 5 – 11. Heterangaeus koreanus n. sp., paratypes. 5, wing; 6, gonostylus; 7, male genitalia, dorsal view; 8, aedeagus; 9, interbase; 10, ninth sternite; 11, ovipositor, lateral view." httpUri="https://zenodo.org/record/231912/files/figure.png" pageId="5" pageNumber="511">Fig. 6</figureCitation>
) club-shaped, with single strong subapical spine; interbase (
<figureCitation id="E78F2A46FFBB85656639C6ADFB159586" box="[1132,1203,912,937]" captionStart="FIGURES 5 – 11" captionStartId="7.[151,264,1874,1897]" captionTargetBox="[171,1425,213,1834]" captionTargetId="figure@7.[151,1436,193,1853]" captionTargetPageId="7" captionText="FIGURES 5 – 11. Heterangaeus koreanus n. sp., paratypes. 5, wing; 6, gonostylus; 7, male genitalia, dorsal view; 8, aedeagus; 9, interbase; 10, ninth sternite; 11, ovipositor, lateral view." httpUri="https://zenodo.org/record/231912/files/figure.png" pageId="5" pageNumber="511">Fig. 9</figureCitation>
) horn-shaped; ninth tergite (
<figureCitation id="E78F2A46FFBB856562B8C689FE9795E3" box="[237,305,948,973]" captionStart="FIGURES 5 – 11" captionStartId="7.[151,264,1874,1897]" captionTargetBox="[171,1425,213,1834]" captionTargetId="figure@7.[151,1436,193,1853]" captionTargetPageId="7" captionText="FIGURES 5 – 11. Heterangaeus koreanus n. sp., paratypes. 5, wing; 6, gonostylus; 7, male genitalia, dorsal view; 8, aedeagus; 9, interbase; 10, ninth sternite; 11, ovipositor, lateral view." httpUri="https://zenodo.org/record/231912/files/figure.png" pageId="5" pageNumber="511">Fig. 7</figureCitation>
) with smooth posterior margin and narrow lateral incisions; ninth sternite (
<figureCitation id="E78F2A46FFBB8565662BC689FB6995E3" box="[1150,1231,948,973]" captionStart="FIGURES 5 – 11" captionStartId="7.[151,264,1874,1897]" captionTargetBox="[171,1425,213,1834]" captionTargetId="figure@7.[151,1436,193,1853]" captionTargetPageId="7" captionText="FIGURES 5 – 11. Heterangaeus koreanus n. sp., paratypes. 5, wing; 6, gonostylus; 7, male genitalia, dorsal view; 8, aedeagus; 9, interbase; 10, ninth sternite; 11, ovipositor, lateral view." httpUri="https://zenodo.org/record/231912/files/figure.png" pageId="5" pageNumber="511">Fig. 10</figureCitation>
) with “U“-shaped incision in the middle. Ovipositor (
<figureCitation id="E78F2A46FFBB85656018C6E5FD0395DE" box="[589,677,984,1009]" captionStart="FIGURES 5 – 11" captionStartId="7.[151,264,1874,1897]" captionTargetBox="[171,1425,213,1834]" captionTargetId="figure@7.[151,1436,193,1853]" captionTargetPageId="7" captionText="FIGURES 5 – 11. Heterangaeus koreanus n. sp., paratypes. 5, wing; 6, gonostylus; 7, male genitalia, dorsal view; 8, aedeagus; 9, interbase; 10, ninth sternite; 11, ovipositor, lateral view." httpUri="https://zenodo.org/record/231912/files/figure.png" pageId="5" pageNumber="511">Fig. 11</figureCitation>
) with wide arched cercus and straight, comparatively narrow hypovalvae. Dorsal margin of hypovalvae bearing seven long bristles.
</paragraph>
</subSubSection>
<subSubSection id="37AE6548FFBB856A6292C11DFC0E914B" lastPageId="10" lastPageNumber="516" pageId="5" pageNumber="511" type="description">
<paragraph id="7F0B36C3FFBB85686292C11DFB5D9232" blockId="5.[151,1437,618,2029]" lastBlockId="8.[151,1437,151,2036]" lastPageId="8" lastPageNumber="514" pageId="5" pageNumber="511">
<emphasis id="4DC0EAD1FFBB85656292C11DFEFF9217" bold="true" box="[199,345,1056,1081]" pageId="5" pageNumber="511">Description.</emphasis>
Adult, male (
<figureCitation id="E78F2A46FFBB856563AFC11DFDDE9217" box="[506,632,1056,1081]" captionStart="FIGURES 5 – 11" captionStartId="7.[151,264,1874,1897]" captionTargetBox="[171,1425,213,1834]" captionTargetId="figure@7.[151,1436,193,1853]" captionTargetPageId="7" captionText="FIGURES 5 – 11. Heterangaeus koreanus n. sp., paratypes. 5, wing; 6, gonostylus; 7, male genitalia, dorsal view; 8, aedeagus; 9, interbase; 10, ninth sternite; 11, ovipositor, lateral view." httpUri="https://zenodo.org/record/231912/files/figure.png" pageId="5" pageNumber="511">Figs. 5–10</figureCitation>
). Body length 
<quantity id="B84C9B26FFBB85656171C11DFC169216" box="[804,944,1056,1080]" metricMagnitude="-3" metricUnit="m" metricValue="8.65" metricValueMax="8.9" metricValueMin="8.4" pageId="5" pageNumber="511" unit="mm" value="8.65" valueMax="8.9" valueMin="8.4">8.4–8.9 mm</quantity>
; wing length 
<quantity id="B84C9B26FFBB8565661AC11DFB7A9216" box="[1103,1244,1056,1080]" metricMagnitude="-3" metricUnit="m" metricValue="9.85" metricValueMax="9.9" metricValueMin="9.8" pageId="5" pageNumber="511" unit="mm" value="9.85" valueMax="9.9" valueMin="9.8">9.8–9.9 mm</quantity>
. Body yellowish brown. Head dark brown, somewhat lighter ventrally, covered with brownish gray pruinosity, vertex covered with sparse erect whitish setae. Rostrum black. Mouth parts brown. Antenna 
<quantity id="B84C9B26FFBB856561EEC155FBF892AE" box="[955,1118,1128,1153]" metricMagnitude="-4" metricUnit="m" metricValue="7.6" metricValueMax="8.7" metricValueMin="6.5" pageId="5" pageNumber="511" unit="mm" value="0.76" valueMax="0.87" valueMin="0.65">0.65–0.87 mm</quantity>
long, if bent backwards, not reaching beyond pronotum. Scape cylindrical, elongate, black ventrally, brown dorsally, covered with sparse grayish pruinosity. Pedicel less than half the length of scape, getting slightly wider toward distal end, reddish brown with somewhat lighter distal end. Both basal segments bearing strong sparse brown setae. Flagellum 11- segmented, basal segments brownish yellow, middle segments yellow, distal segments whitish. Basal flagellomeres oval, remaining segments subglobular, distal flagellomeres elongate, nearly cylindrical. Verticils white and long, approximately three to four times as long as respective segments. Flagellomeres ventrally covered with very short white pubescence. Palpus very long, distinctly longer than antenna, blackish, length of distal segment distinctly exceeds that of preceding segment. Palpus covered with dark erect setae. Thorax generally brownish yellow. Pronotum brown dorsally, with yellowish lateral margins. Prescutum uniformly colored, reddish yellow in lighter specimens, light brown in darker specimens. Pseudosutural fovea yellowish. Prescutal stripes not expressed. Scutal lobes slightly darker than prescutum, especially frontally. Scutellum yellow to fulvous. Mediotergite frontally yellow, turning fulvous posteriorly in lighter specimens, in darker specimens brownish yellow. Pleuron generally yellow, area surrounding wing base and ventral part darkened. Darker specimens with rather distinct oblique dark line extending from head through katepisternum to coxae of middle and posterior legs. Katepisternum without setae. Wing (
<figureCitation id="E78F2A46FFBB85656378C3B9FED790B3" box="[301,369,1668,1693]" captionStart="FIGURES 5 – 11" captionStartId="7.[151,264,1874,1897]" captionTargetBox="[171,1425,213,1834]" captionTargetId="figure@7.[151,1436,193,1853]" captionTargetPageId="7" captionText="FIGURES 5 – 11. Heterangaeus koreanus n. sp., paratypes. 5, wing; 6, gonostylus; 7, male genitalia, dorsal view; 8, aedeagus; 9, interbase; 10, ninth sternite; 11, ovipositor, lateral view." httpUri="https://zenodo.org/record/231912/files/figure.png" pageId="5" pageNumber="511">Fig. 5</figureCitation>
) iridescent with heavy dark pattern. Dark spots surrounding all cross-veins, branching points and tips of all longitudinal veins. Pigmented areas surrounded by darker margins. The costal cell darkened just at wing base before humeral vein, beyond that it is light with just very tip darkened. Wing base darkened frontally to first anal vein. Separate spot surrounds 
<emphasis id="4DC0EAD1FFBB85656006C3CCFDD09123" box="[595,630,1777,1805]" italics="true" pageId="5" pageNumber="511">Sc2</emphasis>
. Cross-band starting at base of 
<emphasis id="4DC0EAD1FFBB8565618CC3CCFC539126" box="[985,1013,1777,1800]" italics="true" pageId="5" pageNumber="511">Rs</emphasis>
and connected with spot surrounding additional crossvein in the middle of cell 
<emphasis id="4DC0EAD1FFBB85656025C225FD229101" box="[624,644,1816,1839]" italics="true" pageId="5" pageNumber="511">m</emphasis>
. Another cross-band extends along cord. Both cross-bands connected along vein 
<emphasis id="4DC0EAD1FFBB85656347C201FE88917D" box="[274,302,1852,1875]" italics="true" pageId="5" pageNumber="511">Rs</emphasis>
, forming characteristic “8“-shaped figure. Another oblique band extends from tip of 
<emphasis id="4DC0EAD1FFBB856566B7C201FB5D9176" box="[1250,1275,1852,1880]" italics="true" pageId="5" pageNumber="511">R3</emphasis>
through distal cross-veins and branching points and reaching basal deflection of 
<emphasis id="4DC0EAD1FFBB8565612AC25FFC1E9150" box="[895,952,1890,1918]" italics="true" pageId="5" pageNumber="511">CuA1</emphasis>
. Shape of spots at distal wing end variable even in same specimen. Cell 
<emphasis id="4DC0EAD1FFBB85656389C2B7FE51918F" box="[476,503,1930,1953]" italics="true" pageId="5" pageNumber="511">cu</emphasis>
mostly darkened with some light spots in the dark background, but these spots also variable in different specimens. Large spot at tip of first anal vein is connected with dark area in cell 
<emphasis id="4DC0EAD1FFBB85656742C293FA9491EB" box="[1303,1330,1966,1989]" italics="true" pageId="5" pageNumber="511">cu</emphasis>
and with dark spots at tips of 
<emphasis id="4DC0EAD1FFBB8565632AC2ECFE1C91C3" box="[383,442,2001,2029]" italics="true" pageId="5" pageNumber="511">CuA1</emphasis>
and 
<emphasis id="4DC0EAD1FFBB856563A4C2ECFD8D91C3" box="[497,555,2001,2029]" italics="true" pageId="5" pageNumber="511">CuA2</emphasis>
. Large spot with light nucleus is at base of first anal cell. It is connected with large dark spot at tip of second anal vein. That spot covers whole distal part of second anal cell. Venation: vein 
<emphasis id="4DC0EAD1FFB68568672FC5A5FA3A969A" box="[1402,1436,152,180]" italics="true" pageId="8" pageNumber="514">Sc1</emphasis>
very long, reaching beyond branching point of 
<emphasis id="4DC0EAD1FFB6856860FFC5FDFD7396F5" box="[682,725,192,219]" italics="true" pageId="8" pageNumber="514">R2+3</emphasis>
and 
<emphasis id="4DC0EAD1FFB685686159C5FDFC8396F5" box="[780,805,192,219]" italics="true" pageId="8" pageNumber="514">R4</emphasis>
. Vein 
<emphasis id="4DC0EAD1FFB68568613BC582FC3796F5" box="[878,913,191,219]" italics="true" pageId="8" pageNumber="514">Sc2</emphasis>
situated long distance before base of 
<emphasis id="4DC0EAD1FFB68568676EC5FDFAF196F9" box="[1339,1367,192,215]" italics="true" pageId="8" pageNumber="514">Rs</emphasis>
. Base of 
<emphasis id="4DC0EAD1FFB6856862E0C5DBFF7796D3" box="[181,209,230,253]" italics="true" pageId="8" pageNumber="514">Rs</emphasis>
distinctly angulated and long-spurred in some specimens, in other spur is missing, or 
<emphasis id="4DC0EAD1FFB6856866C6C5DBFB0996D3" box="[1171,1199,230,253]" italics="true" pageId="8" pageNumber="514">Rs</emphasis>
base arched. Base of 
<emphasis id="4DC0EAD1FFB6856862C2C437FF679708" box="[151,193,266,294]" italics="true" pageId="8" pageNumber="514">R2+3</emphasis>
angulated and short-spurred. Additional cross-veins in cells 
<emphasis id="4DC0EAD1FFB685686138C437FC269708" box="[877,896,266,294]" italics="true" pageId="8" pageNumber="514">r1</emphasis>
, 
<emphasis id="4DC0EAD1FFB6856861DBC437FC079708" box="[910,929,266,294]" italics="true" pageId="8" pageNumber="514">r3</emphasis>
, 
<emphasis id="4DC0EAD1FFB6856861FAC437FC649708" box="[943,962,266,294]" italics="true" pageId="8" pageNumber="514">r4</emphasis>
, 
<emphasis id="4DC0EAD1FFB685686185C437FC4A9708" box="[976,1004,266,294]" italics="true" pageId="8" pageNumber="514">m1</emphasis>
and 
<emphasis id="4DC0EAD1FFB685686676C437FB91970F" box="[1059,1079,266,289]" italics="true" pageId="8" pageNumber="514">m</emphasis>
. Cell 
<emphasis id="4DC0EAD1FFB685686629C437FB3E9708" box="[1148,1176,266,294]" italics="true" pageId="8" pageNumber="514">m1</emphasis>
with short stem, discal cell comparatively short, less than half as long as wide. Additional cross-vein in cell 
<emphasis id="4DC0EAD1FFB685686612C40CFBFD9766" box="[1095,1115,305,328]" italics="true" pageId="8" pageNumber="514">m</emphasis>
approximately at same level as base of 
<emphasis id="4DC0EAD1FFB685686359C468FE8E9742" box="[268,296,341,364]" italics="true" pageId="8" pageNumber="514">Rs</emphasis>
. Basal deflection of 
<emphasis id="4DC0EAD1FFB685686045C468FDEC975F" box="[528,586,341,369]" italics="true" pageId="8" pageNumber="514">CuA1</emphasis>
distinctly beyond branching point of 
<emphasis id="4DC0EAD1FFB6856861BAC468FBA09742" box="[1007,1030,341,364]" italics="true" pageId="8" pageNumber="514">M</emphasis>
. Both anal veins strongly divergent. Posterior wing margin distinctly wavy, distinct emarginations not only at tips of both anal veins, but also at posterior margin of second anal cell. Anal angle comparatively big, second anal cell widest in the central part. Halter 
<quantity id="B84C9B26FFB6856862BCC4FEFE3497F5" box="[233,402,451,476]" metricMagnitude="-3" metricUnit="m" metricValue="1.39" metricValueMax="1.45" metricValueMin="1.33" pageId="8" pageNumber="514" unit="mm" value="1.39" valueMax="1.45" valueMin="1.33">1.33–1.45 mm</quantity>
long, stem whitish, knob yellowish. Legs: frontal coxa yellow, middle coxa brownish, posterior coxa yellowish with brownish infuscation. Trochanters yellow, ventral margin narrowly darkened. Femorae of all legs generally yellow, with whitish bases and distinctly darkened distal ends. Tibiae white with darkened apices. Basal tarsomere yellow with whitish base and narrowly darkened distal end. Second tarsomere light brown basally, turning dark brown distally. Remaining tarsomeres dark brown. Short, semi-erect setae, covering legs, same color as area on which they are distributed, yellow on femur, white on tibiae and dark brown on darkened apices of segments. Femur I: 
<quantity id="B84C9B26FFB68568603BC7A6FCB6949D" box="[622,784,667,692]" metricMagnitude="-3" metricUnit="m" metricValue="5.025" metricValueMax="5.3" metricValueMin="4.75" pageId="8" pageNumber="514" unit="mm" value="5.025" valueMax="5.3" valueMin="4.75">4.75–5.30 mm</quantity>
long, II: 4.20–6.00 mm, III: 
<quantity id="B84C9B26FFB685686605C7A6FB50949D" box="[1104,1270,667,692]" metricMagnitude="-3" metricUnit="m" metricValue="5.625" metricValueMax="5.75" metricValueMin="5.5" pageId="8" pageNumber="514" unit="mm" value="5.625" valueMax="5.75" valueMin="5.5">5.50–5.75 mm</quantity>
; tibiae I: 5.10– 6.00 mm, II: 
<quantity id="B84C9B26FFB68568637DC782FE6B94F9" box="[296,461,703,728]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" metricValueMax="5.5" metricValueMin="4.9" pageId="8" pageNumber="514" unit="mm" value="5.2" valueMax="5.5" valueMin="4.9">4.90–5.50 mm</quantity>
, III: 
<quantity id="B84C9B26FFB685686057C782FD0194F9" box="[514,679,703,728]" metricMagnitude="-3" metricUnit="m" metricValue="5.85" metricValueMax="6.3" metricValueMin="5.4" pageId="8" pageNumber="514" unit="mm" value="5.85" valueMax="6.3" valueMin="5.4">5.40–6.30 mm</quantity>
; tarsus I: 
<quantity id="B84C9B26FFB685686144C782FC1394F9" box="[785,949,703,728]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" metricValueMax="7.6" metricValueMin="7.2" pageId="8" pageNumber="514" unit="mm" value="7.4" valueMax="7.6" valueMin="7.2">7.20–7.60 mm</quantity>
, II: 
<quantity id="B84C9B26FFB6856861B4C782FB2094F9" box="[993,1158,703,728]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" metricValueMax="6.2" metricValueMin="5.6" pageId="8" pageNumber="514" unit="mm" value="5.9" valueMax="6.2" valueMin="5.6">5.60–6.20 mm</quantity>
, III: 
<quantity id="B84C9B26FFB6856866EEC782FAFB94F9" box="[1211,1373,703,728]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="6.2" metricValueMin="5.8" pageId="8" pageNumber="514" unit="mm" value="6.0" valueMax="6.2" valueMin="5.8">5.80–6.20 mm</quantity>
long. Abdominal tergites yellowish brown. Basal tergite with whitish lateral margins. Posterior tergites dark brown. First abdominal sternite brownish with whitish lateral margins. Succeeding sternites generally brownish yellow, with brown posterior margin and fuzzy brownish lateral stripes. Posterior sternites dark brown. Abdomen covered with scarce semi-erect, whitish setae. Male terminalia (
<figureCitation id="E78F2A46FFB6856860B3C672FC889549" box="[742,814,847,872]" captionStart="FIGURES 5 – 11" captionStartId="7.[151,264,1874,1897]" captionTargetBox="[171,1425,213,1834]" captionTargetId="figure@7.[151,1436,193,1853]" captionTargetPageId="7" captionText="FIGURES 5 – 11. Heterangaeus koreanus n. sp., paratypes. 5, wing; 6, gonostylus; 7, male genitalia, dorsal view; 8, aedeagus; 9, interbase; 10, ninth sternite; 11, ovipositor, lateral view." httpUri="https://zenodo.org/record/231912/files/figure.png" pageId="8" pageNumber="514">Fig. 7</figureCitation>
) reddish brown. Ninth tergite with straight posterior margin, which is covered by pubescence, lateral margins of ninth tergite with narrow incisions. Gonocoxite with widened distal end covered in short black spines and separated from the rest of gonocoxite by somewhat narrower “neck“. One pair of gonostyli. Gonostylus (
<figureCitation id="E78F2A46FFB6856860C9C686FD4295FA" box="[668,740,955,980]" captionStart="FIGURES 5 – 11" captionStartId="7.[151,264,1874,1897]" captionTargetBox="[171,1425,213,1834]" captionTargetId="figure@7.[151,1436,193,1853]" captionTargetPageId="7" captionText="FIGURES 5 – 11. Heterangaeus koreanus n. sp., paratypes. 5, wing; 6, gonostylus; 7, male genitalia, dorsal view; 8, aedeagus; 9, interbase; 10, ninth sternite; 11, ovipositor, lateral view." httpUri="https://zenodo.org/record/231912/files/figure.png" pageId="8" pageNumber="514">Fig. 6</figureCitation>
) elongated, club-shaped with strong subapical black spine, margin on the opposite side covered with a few fine setae. Interbase (
<figureCitation id="E78F2A46FFB6856861BEC6E2FB9395D9" box="[1003,1077,991,1016]" captionStart="FIGURES 5 – 11" captionStartId="7.[151,264,1874,1897]" captionTargetBox="[171,1425,213,1834]" captionTargetId="figure@7.[151,1436,193,1853]" captionTargetPageId="7" captionText="FIGURES 5 – 11. Heterangaeus koreanus n. sp., paratypes. 5, wing; 6, gonostylus; 7, male genitalia, dorsal view; 8, aedeagus; 9, interbase; 10, ninth sternite; 11, ovipositor, lateral view." httpUri="https://zenodo.org/record/231912/files/figure.png" pageId="8" pageNumber="514">Fig. 9</figureCitation>
) horn-shaped, slightly arched. Aedeagus (
<figureCitation id="E78F2A46FFB685686340C13EFEF19235" box="[277,343,1027,1052]" captionStart="FIGURES 5 – 11" captionStartId="7.[151,264,1874,1897]" captionTargetBox="[171,1425,213,1834]" captionTargetId="figure@7.[151,1436,193,1853]" captionTargetPageId="7" captionText="FIGURES 5 – 11. Heterangaeus koreanus n. sp., paratypes. 5, wing; 6, gonostylus; 7, male genitalia, dorsal view; 8, aedeagus; 9, interbase; 10, ninth sternite; 11, ovipositor, lateral view." httpUri="https://zenodo.org/record/231912/files/figure.png" pageId="8" pageNumber="514">Fig. 8</figureCitation>
) short and wide with darkened distal part. Ninth sternite with “U“-shaped incision.
</paragraph>
<caption id="2BCB664BFFB8856662C2C203FF7E91BC" httpUri="https://zenodo.org/record/231911/files/figure.png" pageId="6" pageNumber="512" targetBox="[162,1422,204,1810]" targetPageId="6">
<paragraph id="7F0B36C3FFB8856662C2C203FF7E91BC" blockId="6.[151,1436,1854,1938]" pageId="6" pageNumber="512">
<emphasis id="4DC0EAD1FFB8856662C2C203FE9D917A" bold="true" box="[151,315,1854,1877]" pageId="6" pageNumber="512">FIGURES 1–4.</emphasis>
<taxonomicName id="B8B44D40FFB885666314C27DFD3C917B" box="[321,666,1855,1877]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="512" phylum="Arthropoda" rank="subSpecies" species="gloriosus" subSpecies="gloriosus">
<emphasis id="4DC0EAD1FFB885666314C27DFD3C917B" box="[321,666,1855,1877]" italics="true" pageId="6" pageNumber="512">Heterangaeus gloriosus gloriosus</emphasis>
</taxonomicName>
. 1, wing of female from Paiktusan, N. Korea; 2–4, male genitalia, paratype from Saghalien Island: 2, gonostylus; 3, gonocoxite with gonostylus and interbase; 4, interbase; 5, ninth sternite of male from Japan.
</paragraph>
</caption>
<caption id="2BCB664BFFB9856762C2C26FFD4391A9" httpUri="https://zenodo.org/record/231912/files/figure.png" pageId="7" pageNumber="513" targetBox="[171,1425,213,1834]" targetPageId="7">
<paragraph id="7F0B36C3FFB9856762C2C26FFD4391A9" blockId="7.[151,1436,1874,1927]" pageId="7" pageNumber="513">
<emphasis id="4DC0EAD1FFB9856762C2C26FFEE09149" bold="true" box="[151,326,1874,1897]" pageId="7" pageNumber="513">FIGURES 5–11.</emphasis>
<taxonomicName id="B8B44D40FFB985676319C26EFD9A9146" box="[332,572,1874,1896]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="513" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB985676319C26EFD9A9146" box="[332,572,1874,1896]" italics="true" pageId="7" pageNumber="513">Heterangaeus koreanus</emphasis>
</taxonomicName>
<emphasis id="4DC0EAD1FFB985676016C26EFDDA9146" bold="true" box="[579,636,1875,1896]" pageId="7" pageNumber="513">
<taxonomicNameLabel id="56F357AAFFB985676016C26EFDDA9146" box="[579,636,1875,1896]" pageId="7" pageNumber="513" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, paratypes. 5, wing; 6, gonostylus; 7, male genitalia, dorsal view; 8, aedeagus; 9, interbase; 10, ninth sternite; 11, ovipositor, lateral view.
</paragraph>
</caption>
<paragraph id="7F0B36C3FFB685686292C11AFCF29336" blockId="8.[151,1437,151,2036]" pageId="8" pageNumber="514">
Female (
<figureCitation id="E78F2A46FFB68568637DC11AFEDC9211" box="[296,378,1063,1088]" captionStart="FIGURES 5 – 11" captionStartId="7.[151,264,1874,1897]" captionTargetBox="[171,1425,213,1834]" captionTargetId="figure@7.[151,1436,193,1853]" captionTargetPageId="7" captionText="FIGURES 5 – 11. Heterangaeus koreanus n. sp., paratypes. 5, wing; 6, gonostylus; 7, male genitalia, dorsal view; 8, aedeagus; 9, interbase; 10, ninth sternite; 11, ovipositor, lateral view." httpUri="https://zenodo.org/record/231912/files/figure.png" pageId="8" pageNumber="514">Fig. 11</figureCitation>
). Body length 
<quantity id="B84C9B26FFB68568604BC11AFD129211" box="[542,692,1063,1088]" metricMagnitude="-2" metricUnit="m" metricValue="1.065" metricValueMax="1.15" metricValueMin="0.98" pageId="8" pageNumber="514" unit="mm" value="10.65" valueMax="11.5" valueMin="9.8">9.8–11.5 mm</quantity>
; wing length 
<quantity id="B84C9B26FFB68568611FC11AFC169211" box="[842,944,1063,1087]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="8" pageNumber="514" unit="mm" value="11.4">11.4 mm</quantity>
. Body coloration generally similar to that of male, but more whitish, than yellowish. Antenna 
<quantity id="B84C9B26FFB685686095C176FCC5924D" box="[704,867,1099,1124]" metricMagnitude="-3" metricUnit="m" metricValue="1.39" metricValueMax="1.85" metricValueMin="0.93" pageId="8" pageNumber="514" unit="mm" value="1.39" valueMax="1.85" valueMin="0.93">0.93–1.85 mm</quantity>
long. Halter 
<quantity id="B84C9B26FFB6856861A3C176FB3F924D" box="[1014,1177,1099,1124]" metricMagnitude="-3" metricUnit="m" metricValue="1.525" metricValueMax="1.65" metricValueMin="1.4" pageId="8" pageNumber="514" unit="mm" value="1.525" valueMax="1.65" valueMin="1.4">1.40–1.65 mm</quantity>
long. Femur I: 
<quantity id="B84C9B26FFB68568671DC176FF6592A9" metricMagnitude="-3" metricUnit="m" metricValue="5.449999999999999" metricValueMax="5.6" metricValueMin="5.3" pageId="8" pageNumber="514" unit="mm" value="5.449999999999999" valueMax="5.6" valueMin="5.3">5.3–5.6 mm</quantity>
long, II: 
<quantity id="B84C9B26FFB68568637CC152FE1492A9" box="[297,434,1135,1160]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" metricValueMax="5.7" metricValueMin="5.5" pageId="8" pageNumber="514" unit="mm" value="5.6" valueMax="5.7" valueMin="5.5">5.5–5.7 mm</quantity>
, III: 
<quantity id="B84C9B26FFB6856863B2C152FDD792A9" box="[487,625,1135,1160]" metricMagnitude="-3" metricUnit="m" metricValue="6.199999999999999" metricValueMax="6.3" metricValueMin="6.1" pageId="8" pageNumber="514" unit="mm" value="6.199999999999999" valueMax="6.3" valueMin="6.1">6.1–6.3 mm</quantity>
; tibiae I: 
<quantity id="B84C9B26FFB68568608CC152FCC792A9" box="[729,865,1135,1160]" metricMagnitude="-3" metricUnit="m" metricValue="5.55" metricValueMax="5.6" metricValueMin="5.5" pageId="8" pageNumber="514" unit="mm" value="5.55" valueMax="5.6" valueMin="5.5">5.5–5.6 mm</quantity>
, II: 
<quantity id="B84C9B26FFB6856861D8C152FBB092A9" box="[909,1046,1135,1160]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="5.7" metricValueMin="5.3" pageId="8" pageNumber="514" unit="mm" value="5.5" valueMax="5.7" valueMin="5.3">5.3–5.7 mm</quantity>
, III: 
<quantity id="B84C9B26FFB685686619C152FB7092A9" box="[1100,1238,1135,1160]" metricMagnitude="-3" metricUnit="m" metricValue="6.199999999999999" metricValueMax="6.3" metricValueMin="6.1" pageId="8" pageNumber="514" unit="mm" value="6.199999999999999" valueMax="6.3" valueMin="6.1">6.1–6.3 mm</quantity>
: tarsus I: 
<quantity id="B84C9B26FFB685686715C152FA3192A9" box="[1344,1431,1135,1160]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="8" pageNumber="514" unit="mm" value="7.6">7.6 mm</quantity>
, II: 5.0– 
<quantity id="B84C9B26FFB6856862BEC1AEFEE29285" box="[235,324,1171,1196]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="8" pageNumber="514" unit="mm" value="6.2">6.2 mm</quantity>
, III: 
<quantity id="B84C9B26FFB685686328C1AEFDA39285" box="[381,517,1171,1196]" metricMagnitude="-3" metricUnit="m" metricValue="6.35" metricValueMax="6.9" metricValueMin="5.8" pageId="8" pageNumber="514" unit="mm" value="6.35" valueMax="6.9" valueMin="5.8">5.8–6.9 mm</quantity>
long. Posterior end of abdomen less darkened than in male. Ovipositor (
<figureCitation id="E78F2A46FFB685686717C1AEFA359285" box="[1346,1427,1171,1196]" captionStart="FIGURES 5 – 11" captionStartId="7.[151,264,1874,1897]" captionTargetBox="[171,1425,213,1834]" captionTargetId="figure@7.[151,1436,193,1853]" captionTargetPageId="7" captionText="FIGURES 5 – 11. Heterangaeus koreanus n. sp., paratypes. 5, wing; 6, gonostylus; 7, male genitalia, dorsal view; 8, aedeagus; 9, interbase; 10, ninth sternite; 11, ovipositor, lateral view." httpUri="https://zenodo.org/record/231912/files/figure.png" pageId="8" pageNumber="514">Fig. 11</figureCitation>
) grayish brown. Cercus wide, tip raised upwards. Hypovalvae straight and comparatively narrow, dorsal margin bearing seven pairs of strong setae. Distal-most pair separated from the rest, six frontal pairs follow each other in comb-like fashion, frontal pair the shortest, caudal the longest.
</paragraph>
<paragraph id="7F0B36C3FFB685686292C019FD689068" blockId="8.[151,1437,151,2036]" pageId="8" pageNumber="514">
Larva (
<figureCitation id="E78F2A46FFB685686374C01EFE129315" box="[289,436,1315,1340]" captionStart="FIGURES 12 – 18" captionStartId="9.[151,264,1681,1704]" captionTargetBox="[162,1414,213,1635]" captionTargetId="figure@9.[151,1436,193,1660]" captionTargetPageId="9" captionText="FIGURES 12 – 18. Heterangaeus koreanus n. sp., larva. 12, tenth and eleventh body segments, dorsal view; 13, tenth and eleventh body segments, ventral view; 14, labrum, clypeus, left antenna and uppermost part of frons, dorsal view; 15, left maxilla and hypostoma, ventral view; 16, labrum and clypeus, dorsal view; 17, left mandible, dorsal view; 18, prementum, ventral view." httpUri="https://zenodo.org/record/231913/files/figure.png" pageId="8" pageNumber="514">Figs. 12–18</figureCitation>
). Last instar larvae. Body length 13.0– 
<quantity id="B84C9B26FFB6856861CAC01EFBA99315" box="[927,1039,1315,1339]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="8" pageNumber="514" unit="mm" value="14.9">14.9 mm</quantity>
; width 
<quantity id="B84C9B26FFB685686621C01EFAAE9315" box="[1140,1288,1315,1339]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.4" metricValueMin="1.7" pageId="8" pageNumber="514" unit="mm" value="2.05" valueMax="2.4" valueMin="1.7">1.7–2.4 mm</quantity>
. Larvae are hemicephalic, metapneunistic. Body covered with long dark brown setae, which gives body brown-greenish color. Tergum and sternum are covered with longer setae forming longitudinal lines. Line of longer setae on tergum is interrupted by area of very short pubescence on the seventh abdominal segment (
<figureCitation id="E78F2A46FFB685686678C0BAFB2793B1" box="[1069,1153,1415,1440]" captionStart="FIGURES 12 – 18" captionStartId="9.[151,264,1681,1704]" captionTargetBox="[162,1414,213,1635]" captionTargetId="figure@9.[151,1436,193,1660]" captionTargetPageId="9" captionText="FIGURES 12 – 18. Heterangaeus koreanus n. sp., larva. 12, tenth and eleventh body segments, dorsal view; 13, tenth and eleventh body segments, ventral view; 14, labrum, clypeus, left antenna and uppermost part of frons, dorsal view; 15, left maxilla and hypostoma, ventral view; 16, labrum and clypeus, dorsal view; 17, left mandible, dorsal view; 18, prementum, ventral view." httpUri="https://zenodo.org/record/231913/files/figure.png" pageId="8" pageNumber="514">Fig. 12</figureCitation>
). Line of longer setae on sternum is continuous and reach anal field (
<figureCitation id="E78F2A46FFB6856860D6C095FD7193EF" box="[643,727,1448,1473]" captionStart="FIGURES 12 – 18" captionStartId="9.[151,264,1681,1704]" captionTargetBox="[162,1414,213,1635]" captionTargetId="figure@9.[151,1436,193,1660]" captionTargetPageId="9" captionText="FIGURES 12 – 18. Heterangaeus koreanus n. sp., larva. 12, tenth and eleventh body segments, dorsal view; 13, tenth and eleventh body segments, ventral view; 14, labrum, clypeus, left antenna and uppermost part of frons, dorsal view; 15, left maxilla and hypostoma, ventral view; 16, labrum and clypeus, dorsal view; 17, left mandible, dorsal view; 18, prementum, ventral view." httpUri="https://zenodo.org/record/231913/files/figure.png" pageId="8" pageNumber="514">Fig. 13</figureCitation>
). The first thoracic segment and abdominal segments 3 – 7 are longer than wider. Two other thoracic and first two abdominal segments are longer than wider. Abdominal segments 4 – 7 with well developed ventral creeping welts, laterally elevated into humps. Creeping welts on 5 – 7 segments are slightly longer than on 4th segment and their length increases gradually towards posterior end. Apical part of each welt bears short, dark brown spinulae.
</paragraph>
<paragraph id="7F0B36C3FFB6856A6292C372FBF0900A" blockId="8.[151,1437,151,2036]" lastBlockId="10.[151,1437,1114,1930]" lastPageId="10" lastPageNumber="516" pageId="8" pageNumber="514">
<emphasis id="4DC0EAD1FFB685686292C372FEC59046" box="[199,355,1615,1640]" italics="true" pageId="8" pageNumber="514">Head capsule</emphasis>
. Length 
<quantity id="B84C9B26FFB685686385C372FDDF9049" box="[464,633,1615,1639]" metricMagnitude="-3" metricUnit="m" metricValue="1.18" metricValueMax="1.22" metricValueMin="1.14" pageId="8" pageNumber="514" unit="mm" value="1.18" valueMax="1.22" valueMin="1.14">1.14–1.22 mm</quantity>
, width 
<quantity id="B84C9B26FFB685686081C372FCD89049" box="[724,894,1615,1640]" metricMagnitude="-4" metricUnit="m" metricValue="5.15" metricValueMax="5.3" metricValueMin="5.0" pageId="8" pageNumber="514" unit="mm" value="0.515" valueMax="0.53" valueMin="0.5">0.50–0.53 mm</quantity>
. Well-sclerotized, massive, dorsally depressed, narrow and elongate in shape, partially retracted within thorax. Lateral margins of head capsule parallel, posterior parts of genae are heavily sclerotised. Dorsal sutures short, arched and directed outward. Frontoclypeal suture straight with side parts raised up. Mandible with long setae at the base. Eight sensory pits located around the setae and one pit near the middle line (
<figureCitation id="E78F2A46FFB68568604DC3E2FDC890D9" box="[536,622,1759,1784]" captionStart="FIGURES 12 – 18" captionStartId="9.[151,264,1681,1704]" captionTargetBox="[162,1414,213,1635]" captionTargetId="figure@9.[151,1436,193,1660]" captionTargetPageId="9" captionText="FIGURES 12 – 18. Heterangaeus koreanus n. sp., larva. 12, tenth and eleventh body segments, dorsal view; 13, tenth and eleventh body segments, ventral view; 14, labrum, clypeus, left antenna and uppermost part of frons, dorsal view; 15, left maxilla and hypostoma, ventral view; 16, labrum and clypeus, dorsal view; 17, left mandible, dorsal view; 18, prementum, ventral view." httpUri="https://zenodo.org/record/231913/files/figure.png" pageId="8" pageNumber="514">Fig. 14</figureCitation>
). A sensory pit is near the middle of frons. One long and one medium length setae and three sensory pits are at the base of maxilla (
<figureCitation id="E78F2A46FFB685686119C23EFC069132" box="[844,928,1795,1820]" captionStart="FIGURES 12 – 18" captionStartId="9.[151,264,1681,1704]" captionTargetBox="[162,1414,213,1635]" captionTargetId="figure@9.[151,1436,193,1660]" captionTargetPageId="9" captionText="FIGURES 12 – 18. Heterangaeus koreanus n. sp., larva. 12, tenth and eleventh body segments, dorsal view; 13, tenth and eleventh body segments, ventral view; 14, labrum, clypeus, left antenna and uppermost part of frons, dorsal view; 15, left maxilla and hypostoma, ventral view; 16, labrum and clypeus, dorsal view; 17, left mandible, dorsal view; 18, prementum, ventral view." httpUri="https://zenodo.org/record/231913/files/figure.png" pageId="8" pageNumber="514">Fig. 15</figureCitation>
). One long seta is at the base of maxilla near hypostoma. Length of groove, formed by frontal suture, is 2/ 
<date id="0B0A1003FFB68568611BC215FCCE916E" box="[846,872,1832,1856]" pageId="8" pageNumber="514">3x</date>
the head capsule. Posterior part of frons pointed. Clypeus wide and trapezoidal, with rounded margins. Labrum saddle-shaped with several sensory structures: one seta of medium length and two sensory papillae near the anterior margin, four short sensory papillae and a tubercle with sensory pegs on anteriolateral part, a long seta and four short papillae are in the middle (
<figureCitation id="E78F2A46FFB6856866E4C2AEFAA39182" box="[1201,1285,1939,1964]" captionStart="FIGURES 12 – 18" captionStartId="9.[151,264,1681,1704]" captionTargetBox="[162,1414,213,1635]" captionTargetId="figure@9.[151,1436,193,1660]" captionTargetPageId="9" captionText="FIGURES 12 – 18. Heterangaeus koreanus n. sp., larva. 12, tenth and eleventh body segments, dorsal view; 13, tenth and eleventh body segments, ventral view; 14, labrum, clypeus, left antenna and uppermost part of frons, dorsal view; 15, left maxilla and hypostoma, ventral view; 16, labrum and clypeus, dorsal view; 17, left mandible, dorsal view; 18, prementum, ventral view." httpUri="https://zenodo.org/record/231913/files/figure.png" pageId="8" pageNumber="514">Fig. 16</figureCitation>
). Epipharynx covered with short dense bristles. Antenna short, reaching apical part of clypeus (
<figureCitation id="E78F2A46FFB685686629C28AFB7191FE" box="[1148,1239,1975,2000]" captionStart="FIGURES 12 – 18" captionStartId="9.[151,264,1681,1704]" captionTargetBox="[162,1414,213,1635]" captionTargetId="figure@9.[151,1436,193,1660]" captionTargetPageId="9" captionText="FIGURES 12 – 18. Heterangaeus koreanus n. sp., larva. 12, tenth and eleventh body segments, dorsal view; 13, tenth and eleventh body segments, ventral view; 14, labrum, clypeus, left antenna and uppermost part of frons, dorsal view; 15, left maxilla and hypostoma, ventral view; 16, labrum and clypeus, dorsal view; 17, left mandible, dorsal view; 18, prementum, ventral view." httpUri="https://zenodo.org/record/231913/files/figure.png" pageId="8" pageNumber="514">Fig. 14</figureCitation>
). Basal segment cylindrical, with sensory pit near the base, twice as long as wide, with two very long but equal in length apical 
<emphasis id="4DC0EAD1FFB4856A62C2C6C3FEF5923A" bold="true" box="[151,339,1022,1045]" pageId="10" pageNumber="516">
<figureCitation id="E78F2A46FFB4856A62C2C6C3FEE8923A" box="[151,334,1022,1045]" captionStart="FIGURES 12 – 18" captionStartId="9.[151,264,1681,1704]" captionTargetBox="[162,1414,213,1635]" captionTargetId="figure@9.[151,1436,193,1660]" captionTargetPageId="9" captionText="FIGURES 12 – 18. Heterangaeus koreanus n. sp., larva. 12, tenth and eleventh body segments, dorsal view; 13, tenth and eleventh body segments, ventral view; 14, labrum, clypeus, left antenna and uppermost part of frons, dorsal view; 15, left maxilla and hypostoma, ventral view; 16, labrum and clypeus, dorsal view; 17, left mandible, dorsal view; 18, prementum, ventral view." httpUri="https://zenodo.org/record/231913/files/figure.png" pageId="10" pageNumber="516">FIGURES 12–18</figureCitation>
.
</emphasis>
(Continued) papillae (1.3x the length of basal segment itself) and four short sensory papillae on the apex of basal segment of antenna (
<figureCitation id="E78F2A46FFB4856A62ABC141FEF492BA" box="[254,338,1148,1173]" captionStart="FIGURES 12 – 18" captionStartId="9.[151,264,1681,1704]" captionTargetBox="[162,1414,213,1635]" captionTargetId="figure@9.[151,1436,193,1660]" captionTargetPageId="9" captionText="FIGURES 12 – 18. Heterangaeus koreanus n. sp., larva. 12, tenth and eleventh body segments, dorsal view; 13, tenth and eleventh body segments, ventral view; 14, labrum, clypeus, left antenna and uppermost part of frons, dorsal view; 15, left maxilla and hypostoma, ventral view; 16, labrum and clypeus, dorsal view; 17, left mandible, dorsal view; 18, prementum, ventral view." httpUri="https://zenodo.org/record/231913/files/figure.png" pageId="10" pageNumber="516">Fig. 14</figureCitation>
). Mandible large, heavily sclerotised and sickle-shaped. Apical tooth is the largest (
<figureCitation id="E78F2A46FFB4856A6755C141FAF292BA" box="[1280,1364,1148,1173]" captionStart="FIGURES 12 – 18" captionStartId="9.[151,264,1681,1704]" captionTargetBox="[162,1414,213,1635]" captionTargetId="figure@9.[151,1436,193,1660]" captionTargetPageId="9" captionText="FIGURES 12 – 18. Heterangaeus koreanus n. sp., larva. 12, tenth and eleventh body segments, dorsal view; 13, tenth and eleventh body segments, ventral view; 14, labrum, clypeus, left antenna and uppermost part of frons, dorsal view; 15, left maxilla and hypostoma, ventral view; 16, labrum and clypeus, dorsal view; 17, left mandible, dorsal view; 18, prementum, ventral view." httpUri="https://zenodo.org/record/231913/files/figure.png" pageId="10" pageNumber="516">Fig. 17</figureCitation>
). First and second dorsal teeth are small, similar in size, triangular, with blunt apex. Third tooth is more than twice as long as other dorsal teeth, horn-shaped. First and second ventral teeth small and blunt. Third and fourth teeth large, triangular. Fourth tooth curved and sharply pointed. Two long setae at base of mandible. A sensory pit located on dorsal surface of mandible, near the dorsal teeth. Maxillae well developed, consists of outer lobe (palpiger) and inner lobe (galea) (
<figureCitation id="E78F2A46FFB4856A632FC01FFE749315" box="[378,466,1314,1339]" captionStart="FIGURES 12 – 18" captionStartId="9.[151,264,1681,1704]" captionTargetBox="[162,1414,213,1635]" captionTargetId="figure@9.[151,1436,193,1660]" captionTargetPageId="9" captionText="FIGURES 12 – 18. Heterangaeus koreanus n. sp., larva. 12, tenth and eleventh body segments, dorsal view; 13, tenth and eleventh body segments, ventral view; 14, labrum, clypeus, left antenna and uppermost part of frons, dorsal view; 15, left maxilla and hypostoma, ventral view; 16, labrum and clypeus, dorsal view; 17, left mandible, dorsal view; 18, prementum, ventral view." httpUri="https://zenodo.org/record/231913/files/figure.png" pageId="10" pageNumber="516">Fig. 15</figureCitation>
). Palpiger cylindrical, more prominent, with large sclerite at the base. Apical part surrounded by sclerotised ring. Palpus button-shaped with few sensory structures (exact number was difficult to establish). A sensory pit is on the outer margin and another sensory pit is on the inner margin of apical part of palpiger. Galea finger-shaped with large sclerite at the base. Apical part of galea sclerotised with two long setae and one papilla. Cardo arched, well developed with two very long setae and a sensory pit. Hypostoma heavily sclerotised, divided into two plates each of which bears four sharp teeth (
<figureCitation id="E78F2A46FFB4856A6194C0F4FBB293CC" box="[961,1044,1481,1506]" captionStart="FIGURES 12 – 18" captionStartId="9.[151,264,1681,1704]" captionTargetBox="[162,1414,213,1635]" captionTargetId="figure@9.[151,1436,193,1660]" captionTargetPageId="9" captionText="FIGURES 12 – 18. Heterangaeus koreanus n. sp., larva. 12, tenth and eleventh body segments, dorsal view; 13, tenth and eleventh body segments, ventral view; 14, labrum, clypeus, left antenna and uppermost part of frons, dorsal view; 15, left maxilla and hypostoma, ventral view; 16, labrum and clypeus, dorsal view; 17, left mandible, dorsal view; 18, prementum, ventral view." httpUri="https://zenodo.org/record/231913/files/figure.png" pageId="10" pageNumber="516">Fig. 15</figureCitation>
). The outermost tooth significantly shorter; the other teeth are similar. Plates connected with well developed push-button connection. Prementum as a flattened plate with protruding lateral lobes and set apically with short teeth (
<figureCitation id="E78F2A46FFB4856A61A0C331FBEE900A" box="[1013,1096,1548,1573]" captionStart="FIGURES 12 – 18" captionStartId="9.[151,264,1681,1704]" captionTargetBox="[162,1414,213,1635]" captionTargetId="figure@9.[151,1436,193,1660]" captionTargetPageId="9" captionText="FIGURES 12 – 18. Heterangaeus koreanus n. sp., larva. 12, tenth and eleventh body segments, dorsal view; 13, tenth and eleventh body segments, ventral view; 14, labrum, clypeus, left antenna and uppermost part of frons, dorsal view; 15, left maxilla and hypostoma, ventral view; 16, labrum and clypeus, dorsal view; 17, left mandible, dorsal view; 18, prementum, ventral view." httpUri="https://zenodo.org/record/231913/files/figure.png" pageId="10" pageNumber="516">Fig. 18</figureCitation>
).
</paragraph>
<caption id="2BCB664BFFB7856962C2C3ACFEBC912D" httpUri="https://zenodo.org/record/231913/files/figure.png" pageId="9" pageNumber="515" targetBox="[162,1414,213,1635]" targetPageId="9">
<paragraph id="7F0B36C3FFB7856962C2C3ACFEBC912D" blockId="9.[151,1436,1681,1795]" pageId="9" pageNumber="515">
<emphasis id="4DC0EAD1FFB7856962C2C3ACFEF19089" bold="true" box="[151,343,1681,1704]" pageId="9" pageNumber="515">FIGURES 12–18.</emphasis>
<taxonomicName id="B8B44D40FFB785696335C3AEFDF29086" box="[352,596,1682,1704]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="515" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB785696335C3AEFDF29086" box="[352,596,1682,1704]" italics="true" pageId="9" pageNumber="515">Heterangaeus koreanus</emphasis>
</taxonomicName>
<emphasis id="4DC0EAD1FFB785696008C3AFFD3F9089" bold="true" box="[605,665,1682,1703]" pageId="9" pageNumber="515">
<taxonomicNameLabel id="56F357AAFFB785696008C3AFFD3F9089" box="[605,665,1682,1703]" pageId="9" pageNumber="515" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, larva. 12, tenth and eleventh body segments, dorsal view; 13, tenth and eleventh body segments, ventral view; 14, labrum, clypeus, left antenna and uppermost part of frons, dorsal view; 15, left maxilla and hypostoma, ventral view; 16, labrum and clypeus, dorsal view; 17, left mandible, dorsal view; 18, prementum, ventral view.
</paragraph>
</caption>
<paragraph id="7F0B36C3FFB4856A6292C310FC93909F" blockId="10.[151,1437,1114,1930]" pageId="10" pageNumber="516">
<emphasis id="4DC0EAD1FFB4856A6292C310FED59068" box="[199,371,1581,1606]" italics="true" pageId="10" pageNumber="516">Spiracular disc</emphasis>
. Spiracular disc bears two conical elongated ventral lobes. Each of them more than three times as long as its width at base, pointed apically, with three long setae at the apex (
<figureCitation id="E78F2A46FFB4856A664EC36CFBD69044" box="[1051,1136,1617,1642]" captionStart="FIGURES 12 – 18" captionStartId="9.[151,264,1681,1704]" captionTargetBox="[162,1414,213,1635]" captionTargetId="figure@9.[151,1436,193,1660]" captionTargetPageId="9" captionText="FIGURES 12 – 18. Heterangaeus koreanus n. sp., larva. 12, tenth and eleventh body segments, dorsal view; 13, tenth and eleventh body segments, ventral view; 14, labrum, clypeus, left antenna and uppermost part of frons, dorsal view; 15, left maxilla and hypostoma, ventral view; 16, labrum and clypeus, dorsal view; 17, left mandible, dorsal view; 18, prementum, ventral view." httpUri="https://zenodo.org/record/231913/files/figure.png" pageId="10" pageNumber="516">Fig. 12</figureCitation>
). Spiracles are very small, elongated in shape. The distance between spiracles almost equal to the width of a spiracle. Spiracular disc (except small area around the spiracles) entirely covered with setae.
</paragraph>
<paragraph id="7F0B36C3FFB4856A6292C380FCC990D4" blockId="10.[151,1437,1114,1930]" pageId="10" pageNumber="516">
<emphasis id="4DC0EAD1FFB4856A6292C380FE9090F8" box="[199,310,1725,1750]" italics="true" pageId="10" pageNumber="516">Anal field</emphasis>
. Anus surrounded by four very short, segmented, white and fleshy anal papillae, which are fingershaped and equal in size, one-third the length of spiracular lobes.
</paragraph>
<paragraph id="7F0B36C3FFB4856A6292C238FBBF9130" blockId="10.[151,1437,1114,1930]" box="[199,1049,1797,1822]" pageId="10" pageNumber="516">
<emphasis id="4DC0EAD1FFB4856A6292C238FE3F9130" bold="true" box="[199,409,1797,1822]" pageId="10" pageNumber="516">Period of activity.</emphasis>
Adults are flying from late April through end of August.
</paragraph>
<paragraph id="7F0B36C3FFB4856A6292C214FC0E914B" blockId="10.[151,1437,1114,1930]" pageId="10" pageNumber="516">
<emphasis id="4DC0EAD1FFB4856A6292C214FE85916C" bold="true" box="[199,291,1833,1858]" pageId="10" pageNumber="516">Habitat</emphasis>
(
<figureCitation id="E78F2A46FFB4856A6364C214FE22916C" box="[305,388,1833,1858]" captionStart="FIGURE 19" captionStartId="11.[151,250,1178,1200]" captionTargetBox="[151,1435,193,1156]" captionTargetId="figure@11.[151,1436,193,1156]" captionTargetPageId="11" captionText="FIGURE 19. Habitat of Heterangaeus koreanus n. sp. (Piakol valley)." httpUri="https://zenodo.org/record/231914/files/figure.png" pageId="10" pageNumber="516">Fig. 19</figureCitation>
). Wet muddy mountain slopes covered by deciduous trees well above river level. Larvae were collected from mud accumulated on stones under leaf litter or mosses.
</paragraph>
</subSubSection>
<subSubSection id="37AE6548FFB4856A6292C24CFE5A91A7" box="[199,508,1905,1930]" pageId="10" pageNumber="516" type="distribution">
<paragraph id="7F0B36C3FFB4856A6292C24CFE5A91A7" blockId="10.[151,1437,1114,1930]" box="[199,508,1905,1930]" pageId="10" pageNumber="516">
<emphasis id="4DC0EAD1FFB4856A6292C24CFEC691A4" bold="true" box="[199,352,1905,1930]" pageId="10" pageNumber="516">Distribution.</emphasis>
<collectingCountry id="07A37653FFB4856A6332C24CFE5191A7" box="[359,503,1905,1930]" name="South Korea" pageId="10" pageNumber="516">South Korea</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="2BCB664BFFB5856B62C2C1A7FCC3929E" box="[151,869,1178,1200]" httpUri="https://zenodo.org/record/231914/files/figure.png" pageId="11" pageNumber="517" targetBox="[151,1435,193,1156]" targetPageId="11">
<paragraph id="7F0B36C3FFB5856B62C2C1A7FCC3929E" blockId="11.[151,869,1178,1200]" box="[151,869,1178,1200]" pageId="11" pageNumber="517">
<emphasis id="4DC0EAD1FFB5856B62C2C1A7FEB99281" bold="true" box="[151,287,1178,1200]" pageId="11" pageNumber="517">FIGURE 19.</emphasis>
Habitat of 
<taxonomicName id="B8B44D40FFB5856B63C7C1A6FD24929E" box="[402,642,1178,1200]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="517" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB5856B63C7C1A6FD24929E" box="[402,642,1178,1200]" italics="true" pageId="11" pageNumber="517">Heterangaeus koreanus</emphasis>
</taxonomicName>
<emphasis id="4DC0EAD1FFB5856B60DDC1A7FD679281" bold="true" box="[648,705,1178,1199]" pageId="11" pageNumber="517">
<taxonomicNameLabel id="56F357AAFFB5856B60DDC1A7FD679281" box="[648,705,1178,1199]" pageId="11" pageNumber="517" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(Piakol valley).
</paragraph>
</caption>
<subSubSection id="37AE6548FFB5856B6292C03DFC35908B" pageId="11" pageNumber="517" type="materials_examined">
<paragraph id="7F0B36C3FFB5856B6292C03DFA949313" blockId="11.[151,1436,1280,2033]" pageId="11" pageNumber="517">
<emphasis id="4DC0EAD1FFB5856B6292C03DFD949337" bold="true" box="[199,562,1280,1305]" pageId="11" pageNumber="517">
Examined material. 
<typeStatus id="A00F8861FFB5856B6390C03DFD949337" box="[453,562,1280,1305]" pageId="11" pageNumber="517" type="holotype">Holotype</typeStatus>
</emphasis>
, male (pinned). 
<collectingCountry id="07A37653FFB5856B60AEC03DFC369336" box="[763,912,1280,1305]" name="South Korea" pageId="11" pageNumber="517">South Korea</collectingCountry>
, Jeollanam-do province, Gurve, Toji-myeon, Naedong-ri, Piakol valley, 
<geoCoordinate id="1A805004FFB5856B6391C019FDE29313" box="[452,580,1316,1341]" direction="north" orientation="latitude" pageId="11" pageNumber="517" precision="1" value="35.26586">N35.26586</geoCoordinate>
, 
<geoCoordinate id="1A805004FFB5856B601AC019FD7C9313" box="[591,730,1316,1341]" direction="east" orientation="longitude" pageId="11" pageNumber="517" precision="1" value="127.5809">E127.58090</geoCoordinate>
, altitude 
<quantity id="B84C9B26FFB5856B6115C018FC2F9312" box="[832,905,1317,1341]" metricMagnitude="2" metricUnit="m" metricValue="4.4799999999999995" pageId="11" pageNumber="517" unit="m" value="448.0">448 m</quantity>
, 2012.04.27, collected by S. Podenas.
</paragraph>
<paragraph id="7F0B36C3FFB5856B6292C075FBA0903B" blockId="11.[151,1436,1280,2033]" pageId="11" pageNumber="517">
<emphasis id="4DC0EAD1FFB5856B6292C075FE99934F" bold="true" box="[199,319,1352,1377]" pageId="11" pageNumber="517">
<typeStatus id="A00F8861FFB5856B6292C075FE99934F" box="[199,319,1352,1377]" pageId="11" pageNumber="517" type="paratype">Paratypes</typeStatus>
</emphasis>
. 
<specimenCount id="69B2FD4AFFB5856B6306C074FED3934F" box="[339,373,1353,1377]" pageId="11" pageNumber="517" type="generic">1♂</specimenCount>
(pinned), 
<collectingCountry id="07A37653FFB5856B63A0C075FD2C934E" box="[501,650,1352,1377]" name="South Korea" pageId="11" pageNumber="517">South Korea</collectingCountry>
, Jeollabuk-do, Namwon, Sannae-myeon, Deokdong-ri, 
<geoCoordinate id="1A805004FFB5856B674DC075FA3E934F" box="[1304,1432,1352,1377]" direction="north" orientation="latitude" pageId="11" pageNumber="517" precision="1" value="35.33629">N35.33629</geoCoordinate>
, 
<geoCoordinate id="1A805004FFB5856B62C2C051FE8493AA" box="[151,290,1388,1412]" direction="east" orientation="longitude" pageId="11" pageNumber="517" precision="1" value="127.5323">E127.53230</geoCoordinate>
, altitude 
<quantity id="B84C9B26FFB5856B63DDC050FE7493AA" box="[392,466,1389,1413]" metricMagnitude="2" metricUnit="m" metricValue="7.27" pageId="11" pageNumber="517" unit="m" value="727.0">727 m</quantity>
, 2013.05.0 7, collected by S. Podenas; 
<specimenCount id="69B2FD4AFFB5856B61D1C050FC0F93AB" box="[900,937,1389,1413]" pageId="11" pageNumber="517" type="generic">1♂</specimenCount>
, 2♀♀, (
<specimenCount id="69B2FD4AFFB5856B6653C050FB8C93AB" box="[1030,1066,1389,1413]" pageId="11" pageNumber="517" type="generic">1♂</specimenCount>
, 1♀ pinned; 1♀ body in ethanol, wings slide mounted, ovipositor in microvial on pin), 
<collectingCountry id="07A37653FFB5856B615DC0ADFC3D9386" box="[776,923,1424,1449]" name="South Korea" pageId="11" pageNumber="517">South Korea</collectingCountry>
, Jeollanam-do province, Gurve, Toji-myeon, Naedong-ri, Piakol valley, 
<geoCoordinate id="1A805004FFB5856B6399C089FDEA93E2" box="[460,588,1460,1484]" direction="north" orientation="latitude" pageId="11" pageNumber="517" precision="1" value="35.2658">N35.26580</geoCoordinate>
, 
<geoCoordinate id="1A805004FFB5856B600FC089FD4393E2" box="[602,741,1460,1484]" direction="east" orientation="longitude" pageId="11" pageNumber="517" precision="1" value="127.58128">E127.58128</geoCoordinate>
, altitude 
<quantity id="B84C9B26FFB5856B6104C088FC3A93E2" box="[849,924,1461,1485]" metricMagnitude="2" metricUnit="m" metricValue="3.78" pageId="11" pageNumber="517" unit="m" value="378.0">378 m</quantity>
, 2013.05.10, collected by S. Podenas; 3♂♂, 1♀, (2♂♂, 1♀ pinned, 
<specimenCount id="69B2FD4AFFB5856B63EFC0E4FE7B93DF" box="[442,477,1497,1521]" pageId="11" pageNumber="517" type="generic">1♂</specimenCount>
in ethanol), 
<collectingCountry id="07A37653FFB5856B60D7C0E5FCBF93DE" box="[642,793,1496,1521]" name="South Korea" pageId="11" pageNumber="517">South Korea</collectingCountry>
, Gyeonggi-do, Gapyeong-gun, Buk-myeon, Hwaak-ri, 
<geoCoordinate id="1A805004FFB5856B62C2C0C1FEB1903B" box="[151,279,1532,1557]" direction="north" orientation="latitude" pageId="11" pageNumber="517" precision="1" value="37.99363">N37.99363</geoCoordinate>
, 
<geoCoordinate id="1A805004FFB5856B6377C0C1FE0B903B" box="[290,429,1532,1557]" direction="east" orientation="longitude" pageId="11" pageNumber="517" precision="1" value="127.52102">E127.52102</geoCoordinate>
, altitude 
<quantity id="B84C9B26FFB5856B6046C0C0FDFA903A" box="[531,604,1533,1557]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="11" pageNumber="517" unit="m" value="810.0">810 m</quantity>
, 2014.08.20, collected by S. Podenas.
</paragraph>
<paragraph id="7F0B36C3FFB5856B6292C31DFC35908B" blockId="11.[151,1436,1280,2033]" pageId="11" pageNumber="517">
<emphasis id="4DC0EAD1FFB5856B6292C31DFEBA9017" bold="true" box="[199,284,1568,1593]" pageId="11" pageNumber="517">Larvae</emphasis>
. 4 presumable last instar larvae from S. 
<collectingCountry id="07A37653FFB5856B615EC31CFCF49016" box="[779,850,1569,1592]" name="South Korea" pageId="11" pageNumber="517">Korea</collectingCountry>
, Gyeongsangnam-do, Hamyang, Macheon-myeon, Samjeong-ri, 
<geoCoordinate id="1A805004FFB5856B6364C379FE149073" box="[305,434,1604,1629]" direction="north" orientation="latitude" pageId="11" pageNumber="517" precision="1" value="35.3588">N35.35880</geoCoordinate>
, 
<geoCoordinate id="1A805004FFB5856B63EAC379FDEC9073" box="[447,586,1604,1629]" direction="east" orientation="longitude" pageId="11" pageNumber="517" precision="1" value="127.63672">E127.63672</geoCoordinate>
, altitude 
<quantity id="B84C9B26FFB5856B60E1C379FD599072" box="[692,767,1604,1629]" metricMagnitude="2" metricUnit="m" metricValue="6.92" pageId="11" pageNumber="517" unit="m" value="692.0">692 m</quantity>
, 2013.05.11, collected by V. Podeniene; 7 presumable last instar larvae from 
<collectingCountry id="07A37653FFB5856B6336C355FE5490AE" box="[355,498,1640,1665]" name="South Korea" pageId="11" pageNumber="517">South Korea</collectingCountry>
, Jeollanam-do province, Gurve, Toji-myeon, Naedong-ri, Piakol valley, 
<geoCoordinate id="1A805004FFB5856B674DC355FA3E90AF" box="[1304,1432,1640,1665]" direction="north" orientation="latitude" pageId="11" pageNumber="517" precision="1" value="35.2658">N35.26580</geoCoordinate>
, 
<geoCoordinate id="1A805004FFB5856B62C2C3B1FE84908B" box="[151,290,1676,1701]" direction="east" orientation="longitude" pageId="11" pageNumber="517" precision="1" value="127.58128">E127.58128</geoCoordinate>
, altitude 
<quantity id="B84C9B26FFB5856B63DDC3B0FE77908A" box="[392,465,1677,1701]" metricMagnitude="2" metricUnit="m" metricValue="3.78" pageId="11" pageNumber="517" unit="m" value="378.0">378 m</quantity>
, 2013.05.12, collected by V. Podeniene.
</paragraph>
</subSubSection>
<subSubSection id="37AE6548FFB5856B6292C38DFC6990E6" box="[199,975,1712,1737]" pageId="11" pageNumber="517" type="etymology">
<paragraph id="7F0B36C3FFB5856B6292C38DFC6990E6" blockId="11.[151,1436,1280,2033]" box="[199,975,1712,1737]" pageId="11" pageNumber="517">
<emphasis id="4DC0EAD1FFB5856B6292C38DFEE190E7" bold="true" box="[199,327,1712,1737]" pageId="11" pageNumber="517">Etymology</emphasis>
. Species is named for the country of its discovery, 
<collectingCountry id="07A37653FFB5856B61D7C38CFC6C90E6" box="[898,970,1713,1736]" name="South Korea" pageId="11" pageNumber="517">Korea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="37AE6548FFB5856C6292C3E9FA989354" lastPageId="12" lastPageNumber="518" pageId="11" pageNumber="517" type="discussion">
<paragraph id="7F0B36C3FFB5856C6292C3E9FC8C944D" blockId="11.[151,1436,1280,2033]" lastBlockId="12.[151,1436,151,1403]" lastPageId="12" lastPageNumber="518" pageId="11" pageNumber="517">
<emphasis id="4DC0EAD1FFB5856B6292C3E9FEEA90C3" bold="true" box="[199,332,1748,1773]" pageId="11" pageNumber="517">Discussion.</emphasis>
<taxonomicName id="B8B44D40FFB5856B6300C3EBFE4690C3" box="[341,480,1748,1773]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="517" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB5856B6300C3EBFE4690C3" box="[341,480,1748,1773]" italics="true" pageId="11" pageNumber="517">H. koreanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="56F357AAFFB5856B63BFC3E9FD8D90C3" box="[490,555,1748,1773]" pageId="11" pageNumber="517" rank="species">
<emphasis id="4DC0EAD1FFB5856B63BFC3E9FD8D90C3" bold="true" box="[490,555,1748,1773]" pageId="11" pageNumber="517">n. sp.</emphasis>
</taxonomicNameLabel>
is related to 
<taxonomicName id="B8B44D40FFB5856B6092C3EBFB0490C3" authority="Alexander, 1924" authorityName="Alexander" authorityYear="1924" box="[711,1186,1748,1773]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="517" phylum="Arthropoda" rank="subSpecies" species="gloriosus" subSpecies="gloriosus">
<emphasis id="4DC0EAD1FFB5856B6092C3EBFC6090C3" box="[711,966,1748,1773]" italics="true" pageId="11" pageNumber="517">H. gloriosus gloriosus</emphasis>
(
<bibRefCitation id="1B254B32FFB5856B6182C3E9FB3F90C3" author="Alexander" box="[983,1177,1748,1773]" pageId="11" pageNumber="517" refString="Alexander, C. P. (1924) New or little-known crane flies from northern Japan (Tipulidae, Diptera). Philippine Journal of Science, 24, 531 - 611." type="journal article" year="1924">Alexander, 1924</bibRefCitation>
)
</taxonomicName>
. Main differences are observed in wing pattern and wing venation (
<figureCitation id="E78F2A46FFB5856B60CFC3C5FD4E913F" box="[666,744,1784,1809]" captionStart="FIGURES 1 – 4" captionStartId="6.[151,264,1854,1877]" captionTargetBox="[162,1422,204,1810]" captionTargetId="figure@6.[151,1436,193,1833]" captionTargetPageId="6" captionText="FIGURES 1 – 4. Heterangaeus gloriosus gloriosus. 1, wing of female from Paiktusan, N. Korea; 2 – 4, male genitalia, paratype from Saghalien Island: 2, gonostylus; 3, gonocoxite with gonostylus and interbase; 4, interbase; 5, ninth sternite of male from Japan." httpUri="https://zenodo.org/record/231911/files/figure.png" pageId="11" pageNumber="517">Figs. 1</figureCitation>
, 
<figureCitation id="E78F2A46FFB5856B60A2C3C5FCA0913F" box="[759,774,1784,1809]" captionStart="FIGURES 5 – 11" captionStartId="7.[151,264,1874,1897]" captionTargetBox="[171,1425,213,1834]" captionTargetId="figure@7.[151,1436,193,1853]" captionTargetPageId="7" captionText="FIGURES 5 – 11. Heterangaeus koreanus n. sp., paratypes. 5, wing; 6, gonostylus; 7, male genitalia, dorsal view; 8, aedeagus; 9, interbase; 10, ninth sternite; 11, ovipositor, lateral view." httpUri="https://zenodo.org/record/231912/files/figure.png" pageId="11" pageNumber="517">5</figureCitation>
), as well as in male and female genitalia (
<figureCitation id="E78F2A46FFB5856B66B2C3C5FAF3913F" box="[1255,1365,1784,1809]" captionStart="FIGURES 1 – 4" captionStartId="6.[151,264,1854,1877]" captionTargetBox="[162,1422,204,1810]" captionTargetId="figure@6.[151,1436,193,1833]" captionTargetPageId="6" captionText="FIGURES 1 – 4. Heterangaeus gloriosus gloriosus. 1, wing of female from Paiktusan, N. Korea; 2 – 4, male genitalia, paratype from Saghalien Island: 2, gonostylus; 3, gonocoxite with gonostylus and interbase; 4, interbase; 5, ninth sternite of male from Japan." httpUri="https://zenodo.org/record/231911/files/figure.png" pageId="11" pageNumber="517">Figs. 2–4</figureCitation>
, 
<figureCitation id="E78F2A46FFB5856B6734C3C5FA28913F" box="[1377,1422,1784,1809]" captionStart="FIGURES 5 – 11" captionStartId="7.[151,264,1874,1897]" captionTargetBox="[171,1425,213,1834]" captionTargetId="figure@7.[151,1436,193,1853]" captionTargetPageId="7" captionText="FIGURES 5 – 11. Heterangaeus koreanus n. sp., paratypes. 5, wing; 6, gonostylus; 7, male genitalia, dorsal view; 8, aedeagus; 9, interbase; 10, ninth sternite; 11, ovipositor, lateral view." httpUri="https://zenodo.org/record/231912/files/figure.png" pageId="11" pageNumber="517">6, 9</figureCitation>
). Wing cell 
<emphasis id="4DC0EAD1FFB5856B6359C223FEBE911B" box="[268,280,1822,1845]" italics="true" pageId="11" pageNumber="517">c</emphasis>
beyond humeral vein is clear in 
<taxonomicName id="B8B44D40FFB5856B60D8C223FCB0911B" box="[653,790,1820,1845]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="517" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB5856B60D8C223FCB0911B" box="[653,790,1820,1845]" italics="true" pageId="11" pageNumber="517">H. koreanus</emphasis>
</taxonomicName>
<emphasis id="4DC0EAD1FFB5856B614BC221FCF8911B" bold="true" box="[798,862,1820,1845]" pageId="11" pageNumber="517">
<taxonomicNameLabel id="56F357AAFFB5856B614BC221FCF8911B" box="[798,862,1820,1845]" pageId="11" pageNumber="517" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, when that of 
<taxonomicName id="B8B44D40FFB5856B6656C223FB58911B" box="[1027,1278,1820,1845]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="517" phylum="Arthropoda" rank="subSpecies" species="gloriosus" subSpecies="gloriosus">
<emphasis id="4DC0EAD1FFB5856B6656C223FB58911B" box="[1027,1278,1820,1845]" italics="true" pageId="11" pageNumber="517">H. gloriosus gloriosus</emphasis>
</taxonomicName>
has dark spot above the origin of 
<emphasis id="4DC0EAD1FFB5856B632DC27CFE329176" box="[376,404,1857,1880]" italics="true" pageId="11" pageNumber="517">Rs</emphasis>
. Vein 
<emphasis id="4DC0EAD1FFB5856B638AC27CFE5F9176" box="[479,505,1857,1880]" italics="true" pageId="11" pageNumber="517">Sc</emphasis>
<quantity id="B84C9B26FFB5856B63ACC272FDB99177" box="[505,543,1856,1885]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="11" pageNumber="517" unit="in" value="2.0">
<emphasis id="4DC0EAD1FFB5856B63ACC272FDA79173" box="[505,513,1871,1885]" italics="true" pageId="11" pageNumber="517">2</emphasis>
in
</quantity>
<taxonomicName id="B8B44D40FFB5856B607DC27CFD179177" box="[552,689,1856,1881]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="517" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB5856B607DC27CFD179177" box="[552,689,1856,1881]" italics="true" pageId="11" pageNumber="517">H. koreanus</emphasis>
</taxonomicName>
<emphasis id="4DC0EAD1FFB5856B60ECC27DFD5C9177" bold="true" box="[697,762,1856,1881]" pageId="11" pageNumber="517">
<taxonomicNameLabel id="56F357AAFFB5856B60ECC27DFD5C9177" box="[697,762,1856,1881]" pageId="11" pageNumber="517" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is well before base of 
<emphasis id="4DC0EAD1FFB5856B6655C27CFBBA9176" box="[1024,1052,1857,1880]" italics="true" pageId="11" pageNumber="517">Rs</emphasis>
, thus dark spot, surrounding it, is separate from spot, surrounding base of 
<emphasis id="4DC0EAD1FFB5856B603CC255FD239151" box="[617,645,1896,1919]" italics="true" pageId="11" pageNumber="517">Rs</emphasis>
. Because 
<emphasis id="4DC0EAD1FFB5856B60A9C255FCB19151" box="[764,791,1896,1919]" italics="true" pageId="11" pageNumber="517">Sc</emphasis>
<quantity id="B84C9B26FFB5856B6142C24BFC9891AE" box="[791,830,1895,1924]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="11" pageNumber="517" unit="in" value="2.0">
<emphasis id="4DC0EAD1FFB5856B6142C24BFCB991AA" box="[791,799,1910,1924]" italics="true" pageId="11" pageNumber="517">2</emphasis>
in
</quantity>
<taxonomicName id="B8B44D40FFB5856B611DC255FBE091AE" box="[840,1094,1895,1920]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="517" phylum="Arthropoda" rank="subSpecies" species="gloriosus" subSpecies="gloriosus">
<emphasis id="4DC0EAD1FFB5856B611DC255FBE091AE" box="[840,1094,1895,1920]" italics="true" pageId="11" pageNumber="517">H. gloriosus gloriosus</emphasis>
</taxonomicName>
is close to 
<emphasis id="4DC0EAD1FFB5856B669AC255FB4D9151" box="[1231,1259,1896,1919]" italics="true" pageId="11" pageNumber="517">Rs</emphasis>
base, thus it is surrounded by same dark spot, which surrounds base of 
<emphasis id="4DC0EAD1FFB5856B615FC2B2FC809188" box="[778,806,1935,1958]" italics="true" pageId="11" pageNumber="517">Rs</emphasis>
. 
<taxonomicName id="B8B44D40FFB5856B6161C2B2FC1C9189" box="[820,954,1934,1959]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="517" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB5856B6161C2B2FC1C9189" box="[820,954,1934,1959]" italics="true" pageId="11" pageNumber="517">H. koreanus</emphasis>
</taxonomicName>
<emphasis id="4DC0EAD1FFB5856B6197C2B3FBA79189" bold="true" box="[962,1025,1934,1959]" pageId="11" pageNumber="517">
<taxonomicNameLabel id="56F357AAFFB5856B6197C2B3FBA79189" box="[962,1025,1934,1959]" pageId="11" pageNumber="517" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has extra cross-vein in cell 
<emphasis id="4DC0EAD1FFB5856B6762C2B2FAEC9185" box="[1335,1354,1935,1963]" italics="true" pageId="11" pageNumber="517">r1</emphasis>
, which is missing in 
<taxonomicName id="B8B44D40FFB5856B637CC28BFD8491E3" box="[297,546,1972,1997]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="517" phylum="Arthropoda" rank="subSpecies" species="gloriosus" subSpecies="gloriosus">
<emphasis id="4DC0EAD1FFB5856B637CC28BFD8491E3" box="[297,546,1972,1997]" italics="true" pageId="11" pageNumber="517">H. gloriosus gloriosus</emphasis>
</taxonomicName>
. The most distinct feature of 
<taxonomicName id="B8B44D40FFB5856B613DC28BFC4991E3" box="[872,1007,1972,1997]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="517" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB5856B613DC28BFCDA91E3" box="[872,892,1974,1997]" italics="true" pageId="11" pageNumber="517">H</emphasis>
. 
<emphasis id="4DC0EAD1FFB5856B61DFC289FC4991E3" box="[906,1007,1972,1997]" italics="true" pageId="11" pageNumber="517">koreanus</emphasis>
</taxonomicName>
<emphasis id="4DC0EAD1FFB5856B61A3C289FB9291E3" bold="true" box="[1014,1076,1972,1997]" pageId="11" pageNumber="517">
<taxonomicNameLabel id="56F357AAFFB5856B61A3C289FB9291E3" box="[1014,1076,1972,1997]" pageId="11" pageNumber="517" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
wing is distinctly darkened area along whole 
<emphasis id="4DC0EAD1FFB5856B6372C2E7FEE591DF" box="[295,323,2010,2033]" italics="true" pageId="11" pageNumber="517">Rs</emphasis>
. It makes characteristic “8“-shaped figure in central part of wing. Central part of 
<emphasis id="4DC0EAD1FFB5856B6684C2E7FB4B91DF" box="[1233,1261,2010,2033]" italics="true" pageId="11" pageNumber="517">Rs</emphasis>
not surrounded by dark area in 
<taxonomicName id="B8B44D40FFB2856C631EC5A4FDE1969E" box="[331,583,151,176]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="subSpecies" species="gloriosus" subSpecies="gloriosus">
<emphasis id="4DC0EAD1FFB2856C631EC5A4FEF9969E" box="[331,351,153,176]" italics="true" pageId="12" pageNumber="518">H</emphasis>
. 
<emphasis id="4DC0EAD1FFB2856C633AC5AAFDE1969E" box="[367,583,151,176]" italics="true" pageId="12" pageNumber="518">gloriosus gloriosus</emphasis>
</taxonomicName>
, thus central transparent area of wing is not separated and more resembles transverse light band. Pigmentation of cell 
<emphasis id="4DC0EAD1FFB2856C60D4C580FD3A96FA" box="[641,668,189,212]" italics="true" pageId="12" pageNumber="518">cu</emphasis>
in both species is variable, but that of 
<taxonomicName id="B8B44D40FFB2856C660FC580FB4496FB" box="[1114,1250,188,213]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB2856C660FC580FB4496FB" box="[1114,1250,188,213]" italics="true" pageId="12" pageNumber="518">H. koreanus</emphasis>
</taxonomicName>
<emphasis id="4DC0EAD1FFB2856C66B9C581FA8A96FB" bold="true" box="[1260,1324,188,213]" pageId="12" pageNumber="518">
<taxonomicNameLabel id="56F357AAFFB2856C66B9C581FA8A96FB" box="[1260,1324,188,213]" pageId="12" pageNumber="518" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is mostly darkened with separate small light spots, when that of 
<taxonomicName id="B8B44D40FFB2856C6151C5DCFBA796D6" box="[772,1025,223,248]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="subSpecies" species="gloriosus" subSpecies="gloriosus">
<emphasis id="4DC0EAD1FFB2856C6151C5DCFBA796D6" box="[772,1025,223,248]" italics="true" pageId="12" pageNumber="518">H. gloriosus gloriosus</emphasis>
</taxonomicName>
is mostly light with 6–7 dark spots. Posterior margin of wing distinctly emarginate in 
<taxonomicName id="B8B44D40FFB2856C6091C438FCED9733" box="[708,843,260,285]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB2856C6091C438FCED9733" box="[708,843,260,285]" italics="true" pageId="12" pageNumber="518">H. koreanus</emphasis>
</taxonomicName>
<emphasis id="4DC0EAD1FFB2856C6107C439FC379733" bold="true" box="[850,913,260,285]" pageId="12" pageNumber="518">
<taxonomicNameLabel id="56F357AAFFB2856C6107C439FC379733" box="[850,913,260,285]" pageId="12" pageNumber="518" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, deep emarginations present not only at apices of both anal veins, but also at posterior margin of cell 
<emphasis id="4DC0EAD1FFB2856C6147C414FC8E976A" box="[786,808,297,324]" italics="true" pageId="12" pageNumber="518">a2</emphasis>
. Posterior wing margin of 
<taxonomicName id="B8B44D40FFB2856C6637C414FAC6976E" box="[1122,1376,295,320]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="subSpecies" species="gloriosus" subSpecies="gloriosus">
<emphasis id="4DC0EAD1FFB2856C6637C414FAC6976E" box="[1122,1376,295,320]" italics="true" pageId="12" pageNumber="518">H. gloriosus gloriosus</emphasis>
</taxonomicName>
only shallowly emarginated. Gonostylus of male genitalia is wider, and subapical spine longer in 
<taxonomicName id="B8B44D40FFB2856C6695C46DFAEC9749" box="[1216,1354,334,359]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB2856C6695C46DFAEC9749" box="[1216,1354,334,359]" italics="true" pageId="12" pageNumber="518">H. koreanus</emphasis>
</taxonomicName>
<emphasis id="4DC0EAD1FFB2856C6701C473FA329749" bold="true" box="[1364,1428,334,359]" pageId="12" pageNumber="518">
<taxonomicNameLabel id="56F357AAFFB2856C6701C473FA329749" box="[1364,1428,334,359]" pageId="12" pageNumber="518" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, than in 
<taxonomicName id="B8B44D40FFB2856C62A7C449FE5497A5" box="[242,498,370,395]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="subSpecies" species="gloriosus" subSpecies="gloriosus">
<emphasis id="4DC0EAD1FFB2856C62A7C449FEA097A5" box="[242,262,372,395]" italics="true" pageId="12" pageNumber="518">H</emphasis>
. 
<emphasis id="4DC0EAD1FFB2856C6342C44FFE5497A5" box="[279,498,370,395]" italics="true" pageId="12" pageNumber="518">gloriosus gloriosus</emphasis>
</taxonomicName>
. Gonocoxite of 
<taxonomicName id="B8B44D40FFB2856C60E6C449FC9997A5" box="[691,831,370,395]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB2856C60E6C449FC9997A5" box="[691,831,370,395]" italics="true" pageId="12" pageNumber="518">H. koreanus</emphasis>
</taxonomicName>
<emphasis id="4DC0EAD1FFB2856C611CC44FFC2D97A5" bold="true" box="[841,907,370,395]" pageId="12" pageNumber="518">
<taxonomicNameLabel id="56F357AAFFB2856C611CC44FFC2D97A5" box="[841,907,370,395]" pageId="12" pageNumber="518" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has distinctly narrowed “neck“, which is not visible in 
<typeStatus id="A00F8861FFB2856C635DC4A5FECE979E" box="[264,360,408,432]" pageId="12" pageNumber="518" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="B8B44D40FFB2856C63DAC4A5FD2D9781" box="[399,651,406,431]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="subSpecies" species="gloriosus" subSpecies="gloriosus">
<emphasis id="4DC0EAD1FFB2856C63DAC4A5FD2D9781" box="[399,651,406,431]" italics="true" pageId="12" pageNumber="518">H. gloriosus gloriosus</emphasis>
</taxonomicName>
. Distal part of interbase of 
<taxonomicName id="B8B44D40FFB2856C6193C4A5FBE99781" box="[966,1103,406,431]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB2856C6193C4A5FBE99781" box="[966,1103,406,431]" italics="true" pageId="12" pageNumber="518">H. koreanus</emphasis>
</taxonomicName>
<emphasis id="4DC0EAD1FFB2856C6602C4ABFB3E9781" bold="true" box="[1111,1176,406,431]" pageId="12" pageNumber="518">
<taxonomicNameLabel id="56F357AAFFB2856C6602C4ABFB3E9781" box="[1111,1176,406,431]" pageId="12" pageNumber="518" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is straight and narrow, when that of 
<taxonomicName id="B8B44D40FFB2856C6379C481FD8197FD" box="[300,551,442,467]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="subSpecies" species="gloriosus" subSpecies="gloriosus">
<emphasis id="4DC0EAD1FFB2856C6379C481FD8197FD" box="[300,551,442,467]" italics="true" pageId="12" pageNumber="518">H. gloriosus gloriosus</emphasis>
</taxonomicName>
is wider and slightly arched. Female of 
<taxonomicName id="B8B44D40FFB2856C61A5C481FB4D97FD" box="[1008,1259,442,467]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="subSpecies" species="gloriosus" subSpecies="gloriosus">
<emphasis id="4DC0EAD1FFB2856C61A5C481FB4D97FD" box="[1008,1259,442,467]" italics="true" pageId="12" pageNumber="518">H. gloriosus gloriosus</emphasis>
</taxonomicName>
is undescribed, but we compared 
<taxonomicName id="B8B44D40FFB2856C6326C4DDFDA797D9" box="[371,513,478,503]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB2856C6326C4DDFDA797D9" box="[371,513,478,503]" italics="true" pageId="12" pageNumber="518">H. koreanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="56F357AAFFB2856C605BC4E3FDF297D9" box="[526,596,478,503]" pageId="12" pageNumber="518" rank="species">
<emphasis id="4DC0EAD1FFB2856C605BC4E3FDF297D9" bold="true" box="[526,596,478,503]" pageId="12" pageNumber="518">n. sp.</emphasis>
</taxonomicNameLabel>
females with a female from 
<collectingCountry id="07A37653FFB2856C61EBC4DDFBA797D9" box="[958,1025,480,503]" name="Japan" pageId="12" pageNumber="518">Japan</collectingCountry>
. Ovipositor of Korean female has comparatively shorter and wider cercus and narrower hypovalvae. Dorsal margin of hypovalvae of Korean females has seven pairs of strong setae, Japanese female just six pairs. Caudal pair of setae, separated from the frontal six pairs by wide gap, is totally missing in Japanese specimen.
</paragraph>
<paragraph id="7F0B36C3FFB2856C6292C753FD7095C1" blockId="12.[151,1436,151,1403]" pageId="12" pageNumber="518">
Overall appearance of larva of 
<taxonomicName id="B8B44D40FFB2856C6065C74DFD1D94A9" box="[560,699,622,647]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB2856C6065C74DFD1D94A9" box="[560,699,622,647]" italics="true" pageId="12" pageNumber="518">H. koreanus</emphasis>
</taxonomicName>
<emphasis id="4DC0EAD1FFB2856C6091C753FCA094A9" bold="true" box="[708,774,622,647]" pageId="12" pageNumber="518">
<taxonomicNameLabel id="56F357AAFFB2856C6091C753FCA094A9" box="[708,774,622,647]" pageId="12" pageNumber="518" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is the same as 
<taxonomicName id="B8B44D40FFB2856C61E8C74DFBC594A9" box="[957,1123,622,647]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="mongolicus">
<emphasis id="4DC0EAD1FFB2856C61E8C74DFBC594A9" box="[957,1123,622,647]" italics="true" pageId="12" pageNumber="518">H. mongolicus</emphasis>
</taxonomicName>
. Length and color of setae covering body are different in these two species. 
<taxonomicName id="B8B44D40FFB2856C60EEC7A9FCF89485" box="[699,862,658,683]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="mongolicus">
<emphasis id="4DC0EAD1FFB2856C60EEC7A9FCF89485" box="[699,862,658,683]" italics="true" pageId="12" pageNumber="518">H. mongolicus</emphasis>
</taxonomicName>
is covered with short white setae, when body of 
<taxonomicName id="B8B44D40FFB2856C67D4C7A9FF5B94E1" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB2856C67D4C7A9FF5B94E1" italics="true" pageId="12" pageNumber="518">H. koreanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="56F357AAFFB2856C635DC78BFEED94E1" box="[264,331,694,719]" pageId="12" pageNumber="518" rank="species">
<emphasis id="4DC0EAD1FFB2856C635DC78BFEED94E1" bold="true" box="[264,331,694,719]" pageId="12" pageNumber="518">n. sp.</emphasis>
</taxonomicNameLabel>
is covered with long and dark brown setae. Both species have long setae on tergum forming longitudinal line which is interrupted by shorter setae on last abdominal segment and this once again confirm, that this character could be one of the distinguishing features for larvae of this genus. Line of longer setae on sternum is visible just in 
<taxonomicName id="B8B44D40FFB2856C636DC619FE679515" box="[312,449,802,827]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB2856C636DC619FE679515" box="[312,449,802,827]" italics="true" pageId="12" pageNumber="518">H. koreanus</emphasis>
</taxonomicName>
<emphasis id="4DC0EAD1FFB2856C639CC61FFDAF9515" bold="true" box="[457,521,802,827]" pageId="12" pageNumber="518">
<taxonomicNameLabel id="56F357AAFFB2856C639CC61FFDAF9515" box="[457,521,802,827]" pageId="12" pageNumber="518" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Both species have creeping welts on 7–10 abdominal segments, which look like humps. Differences were noticed in coloration of spinulae covering humps: in 
<taxonomicName id="B8B44D40FFB2856C6659C675FB169571" box="[1036,1200,838,863]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="mongolicus">
<emphasis id="4DC0EAD1FFB2856C6659C675FB169571" box="[1036,1200,838,863]" italics="true" pageId="12" pageNumber="518">H. mongolicus</emphasis>
</taxonomicName>
they are light brown in 
<taxonomicName id="B8B44D40FFB2856C62E2C651FEE495AD" box="[183,322,874,899]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB2856C62E2C651FEE495AD" box="[183,322,874,899]" italics="true" pageId="12" pageNumber="518">H. koreanus</emphasis>
</taxonomicName>
<emphasis id="4DC0EAD1FFB2856C6319C657FE2895AD" bold="true" box="[332,398,874,899]" pageId="12" pageNumber="518">
<taxonomicNameLabel id="56F357AAFFB2856C6319C657FE2895AD" box="[332,398,874,899]" pageId="12" pageNumber="518" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
dark brown. Number and shape of anal papillae are the same in both species. Shape and length of spiracular lobes of 
<taxonomicName id="B8B44D40FFB2856C638EC6ADFDD99589" box="[475,639,910,935]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="mongolicus">
<emphasis id="4DC0EAD1FFB2856C638EC6ADFDD99589" box="[475,639,910,935]" italics="true" pageId="12" pageNumber="518">H. mongolicus</emphasis>
</taxonomicName>
and 
<taxonomicName id="B8B44D40FFB2856C60E3C6ADFC989589" box="[694,830,910,935]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB2856C60E3C6ADFC989589" box="[694,830,910,935]" italics="true" pageId="12" pageNumber="518">H. koreanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="56F357AAFFB2856C6113C6B3FC209589" box="[838,902,910,935]" pageId="12" pageNumber="518" rank="species">
<emphasis id="4DC0EAD1FFB2856C6113C6B3FC209589" bold="true" box="[838,902,910,935]" pageId="12" pageNumber="518">n. sp.</emphasis>
</taxonomicNameLabel>
is very similar. It was noticed that 
<taxonomicName id="B8B44D40FFB2856C6741C6ADFA3D9589" box="[1300,1435,910,935]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB2856C6741C6ADFA3D9589" box="[1300,1435,910,935]" italics="true" pageId="12" pageNumber="518">H. koreanus</emphasis>
</taxonomicName>
<emphasis id="4DC0EAD1FFB2856C62C2C68FFF7395E5" bold="true" box="[151,213,946,971]" pageId="12" pageNumber="518">
<taxonomicNameLabel id="56F357AAFFB2856C62C2C68FFF7395E5" box="[151,213,946,971]" pageId="12" pageNumber="518" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has three setae at the apex of spiracular lobe, but it is hard to say if these setae really absent in 
<taxonomicName id="B8B44D40FFB2856C66ACC689FA3A95E5" box="[1273,1436,946,971]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="mongolicus">
<emphasis id="4DC0EAD1FFB2856C66ACC689FA3A95E5" box="[1273,1436,946,971]" italics="true" pageId="12" pageNumber="518">H. mongolicus</emphasis>
</taxonomicName>
or they are just broken on the specimens examined.
</paragraph>
<paragraph id="7F0B36C3FFB2856C6292C6C7FE9E9379" blockId="12.[151,1436,151,1403]" pageId="12" pageNumber="518">
The general appearance of head capsule of 
<taxonomicName id="B8B44D40FFB2856C60E6C6C1FCF1923D" box="[691,855,1018,1043]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="mongolicus">
<emphasis id="4DC0EAD1FFB2856C60E6C6C1FCF1923D" box="[691,855,1018,1043]" italics="true" pageId="12" pageNumber="518">H. mongolicus</emphasis>
</taxonomicName>
and 
<taxonomicName id="B8B44D40FFB2856C61C5C6C1FBBC923D" box="[912,1050,1018,1043]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB2856C61C5C6C1FBBC923D" box="[912,1050,1018,1043]" italics="true" pageId="12" pageNumber="518">H. koreanus</emphasis>
</taxonomicName>
<emphasis id="4DC0EAD1FFB2856C6674C6C7FBC7923D" bold="true" box="[1057,1121,1018,1043]" pageId="12" pageNumber="518">
<taxonomicNameLabel id="56F357AAFFB2856C6674C6C7FBC7923D" box="[1057,1121,1018,1043]" pageId="12" pageNumber="518" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is very similar. Differences were noticed in number and arrangement of sensory structures on labrum and frons. Antennae of 
<taxonomicName id="B8B44D40FFB2856C66A3C11DFA3A9219" box="[1270,1436,1054,1079]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="mongolicus">
<emphasis id="4DC0EAD1FFB2856C66A3C11DFA3A9219" box="[1270,1436,1054,1079]" italics="true" pageId="12" pageNumber="518">H. mongolicus</emphasis>
</taxonomicName>
and 
<taxonomicName id="B8B44D40FFB2856C6292C179FEF69275" box="[199,336,1090,1115]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB2856C6292C179FEF69275" box="[199,336,1090,1115]" italics="true" pageId="12" pageNumber="518">H. koreanus</emphasis>
</taxonomicName>
<emphasis id="4DC0EAD1FFB2856C630CC17FFE3F9275" bold="true" box="[345,409,1090,1115]" pageId="12" pageNumber="518">
<taxonomicNameLabel id="56F357AAFFB2856C630CC17FFE3F9275" box="[345,409,1090,1115]" pageId="12" pageNumber="518" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
are similar, but 
<taxonomicName id="B8B44D40FFB2856C6006C179FD7A9275" box="[595,732,1090,1115]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB2856C6006C179FD7A9275" box="[595,732,1090,1115]" italics="true" pageId="12" pageNumber="518">H. koreanus</emphasis>
</taxonomicName>
<emphasis id="4DC0EAD1FFB2856C60B1C17FFC839275" bold="true" box="[740,805,1090,1115]" pageId="12" pageNumber="518">
<taxonomicNameLabel id="56F357AAFFB2856C60B1C17FFC839275" box="[740,805,1090,1115]" pageId="12" pageNumber="518" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has four papillae on the apex, when 
<taxonomicName id="B8B44D40FFB2856C669FC179FAC89275" box="[1226,1390,1090,1115]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="mongolicus">
<emphasis id="4DC0EAD1FFB2856C669FC179FAC89275" box="[1226,1390,1090,1115]" italics="true" pageId="12" pageNumber="518">H. mongolicus</emphasis>
</taxonomicName>
has only three. Mandibles of both 
<taxonomicName id="B8B44D40FFB2856C63BEC155FD2E9251" box="[491,648,1128,1151]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="genus">
<emphasis id="4DC0EAD1FFB2856C63BEC155FD2E9251" box="[491,648,1128,1151]" italics="true" pageId="12" pageNumber="518">Heterangaeus</emphasis>
</taxonomicName>
species are similar in shape. Differences were noticed just in shape of fourth ventral teeth. The most significant differences were noticed in hypostoma. Each part of hypostoma of 
<taxonomicName id="B8B44D40FFB2856C67D4C1B1FF5B92E9" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB2856C67D4C1B1FF5B92E9" italics="true" pageId="12" pageNumber="518">H. koreanus</emphasis>
</taxonomicName>
<emphasis id="4DC0EAD1FFB2856C6350C193FEE092E9" bold="true" box="[261,326,1198,1223]" pageId="12" pageNumber="518">
<taxonomicNameLabel id="56F357AAFFB2856C6350C193FEE092E9" box="[261,326,1198,1223]" pageId="12" pageNumber="518" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
bears four teeth, when 
<taxonomicName id="B8B44D40FFB2856C6000C18DFD5C92E9" box="[597,762,1198,1223]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="mongolicus">
<emphasis id="4DC0EAD1FFB2856C6000C18DFD5C92E9" box="[597,762,1198,1223]" italics="true" pageId="12" pageNumber="518">H. mongolicus</emphasis>
</taxonomicName>
bears just three. Two parts of hypostoma are held together with push-button connection (which connects two plates of hypostoma) in 
<taxonomicName id="B8B44D40FFB2856C6648C1E9FB0A92C5" box="[1053,1196,1234,1259]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB2856C6648C1E9FB0A92C5" box="[1053,1196,1234,1259]" italics="true" pageId="12" pageNumber="518">H. koreanus</emphasis>
</taxonomicName>
<emphasis id="4DC0EAD1FFB2856C66ECC1EFFB5892C5" bold="true" box="[1209,1278,1234,1259]" pageId="12" pageNumber="518">
<taxonomicNameLabel id="56F357AAFFB2856C66ECC1EFFB5892C5" box="[1209,1278,1234,1259]" pageId="12" pageNumber="518" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, when in 
<taxonomicName id="B8B44D40FFB2856C67D4C1E9FEBE9321" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="mongolicus">
<emphasis id="4DC0EAD1FFB2856C67D4C1E9FEBE9321" italics="true" pageId="12" pageNumber="518">H. mongolicus</emphasis>
</taxonomicName>
push-button connection is not well developed. Larvae of both species of 
<taxonomicName id="B8B44D40FFB2856C662EC1C5FABE9321" box="[1147,1304,1272,1295]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="genus">
<emphasis id="4DC0EAD1FFB2856C662EC1C5FABE9321" box="[1147,1304,1272,1295]" italics="true" pageId="12" pageNumber="518">Heterangaeus</emphasis>
</taxonomicName>
develop in organic-rich mud; 
<taxonomicName id="B8B44D40FFB2856C633DC021FE57931D" box="[360,497,1306,1331]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB2856C633DC021FE57931D" box="[360,497,1306,1331]" italics="true" pageId="12" pageNumber="518">H. koreanus</emphasis>
</taxonomicName>
<emphasis id="4DC0EAD1FFB2856C63ACC027FD9F931D" bold="true" box="[505,569,1306,1331]" pageId="12" pageNumber="518">
<taxonomicNameLabel id="56F357AAFFB2856C63ACC027FD9F931D" box="[505,569,1306,1331]" pageId="12" pageNumber="518" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
was collected from mud accumulated on the stones and 
<taxonomicName id="B8B44D40FFB2856C66EFC021FAF8931D" box="[1210,1374,1306,1331]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="mongolicus">
<emphasis id="4DC0EAD1FFB2856C66EFC021FAF8931D" box="[1210,1374,1306,1331]" italics="true" pageId="12" pageNumber="518">H. mongolicus</emphasis>
</taxonomicName>
from mud in marsh.
</paragraph>
<paragraph id="7F0B36C3FFB2856C6292C059FA989354" blockId="12.[151,1436,151,1403]" box="[199,1342,1378,1403]" pageId="12" pageNumber="518">
<taxonomicName id="B8B44D40FFB2856C6292C059FEE99355" box="[199,335,1378,1403]" class="Insecta" family="Pediciidae" genus="Heterangaeus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="4DC0EAD1FFB2856C6292C059FEE99355" box="[199,335,1378,1403]" italics="true" pageId="12" pageNumber="518">H. koreanus</emphasis>
</taxonomicName>
<emphasis id="4DC0EAD1FFB2856C6300C05FFE339355" bold="true" box="[341,405,1378,1403]" pageId="12" pageNumber="518">
<taxonomicNameLabel id="56F357AAFFB2856C6300C05FFE339355" box="[341,405,1378,1403]" pageId="12" pageNumber="518" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is the first species of the family 
<taxonomicName id="B8B44D40FFB2856C6157C05FFCD19355" box="[770,887,1378,1403]" class="Insecta" family="Pediciidae" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="518" phylum="Arthropoda" rank="family">Pediciidae</taxonomicName>
listed from the territory of 
<collectingCountry id="07A37653FFB2856C66FCC05FFA9F9354" box="[1193,1337,1378,1403]" name="South Korea" pageId="12" pageNumber="518">South Korea</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>