treatments-xml/data/1B/74/87/1B7487E9E419FFEEC5C5FB3D0CA0F86F.xml

<document id="3BB27A4C1CECDD22AEA3EC0AA47224ED" ID-DOI="10.11646/zootaxa.4564.1.12" ID-GBIF-Dataset="586d0df0-d8b0-4479-a391-9ba99a387cdb" ID-ISSN="1175-5326" ID-Zenodo-Dep="2588597" ID-ZooBank="9C5D394E-A86D-40BD-931E-31397EC9BA58" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1552159566916" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2019" docId="1B7487E9E419FFEEC5C5FB3D0CA0F86F" docLanguage="en" docName="zootaxa.4564.1.12.pdf" docOrigin="Zootaxa 4564 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Clavomicrus Jałoszyński 2019, gen. n." docType="treatment" docVersion="11" lastPageNumber="291" masterDocId="E74DFF91E418FFECC552FFD20E6DFFB3" masterDocTitle="' Genus X 2 ' finally named: discovery of males of Clavomicrus gen. n. in the Philippines (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="294" masterPageNumber="289" pageNumber="290" updateTime="1727964259133" updateUser="ExternalLinkService">
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<mods:title id="B71C245AC18720DCA7E59B6E4089EDDF">' Genus X 2 ' finally named: discovery of males of Clavomicrus gen. n. in the Philippines (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:namePart id="3A030847FAA3BAD0B4506D4430174A14">Jałoszyński, Paweł</mods:namePart>
<mods:affiliation id="E044558EABFFB70DB42E23A54445DE24">Museum of Natural History, University of Wrocław, Sienkiewicza 21, 50 - 335 Wrocław, Poland.</mods:affiliation>
<mods:nameIdentifier id="B57749DC56E0BDF4969AC409F065126D" type="email">scydmaenus@yahoo.com</mods:nameIdentifier>
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<mods:date id="E0F50404BF6E68F1E01B785FFBF54BEF">2019</mods:date>
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<treatment id="1B7487E9E419FFEEC5C5FB3D0CA0F86F" ID-DOI="http://doi.org/10.5281/zenodo.5930737" ID-GBIF-Taxon="154366331" ID-Zenodo-Dep="5930737" LSID="urn:lsid:plazi:treatment:1B7487E9E419FFEEC5C5FB3D0CA0F86F" httpUri="http://treatment.plazi.org/id/1B7487E9E419FFEEC5C5FB3D0CA0F86F" lastPageId="2" lastPageNumber="291" pageId="1" pageNumber="290">
<subSubSection id="DBC76574E419FFEDC5C5FB3D0FE4FAB9" box="[151,393,1263,1290]" pageId="1" pageNumber="290" type="nomenclature">
<paragraph id="936236FFE419FFEDC5C5FB3D0FE4FAB9" blockId="1.[151,393,1263,1290]" box="[151,393,1263,1290]" pageId="1" pageNumber="290">
<heading id="C82A8193E419FFEDC5C5FB3D0FE4FAB9" bold="true" box="[151,393,1263,1290]" fontSize="11" level="1" pageId="1" pageNumber="290" reason="1">
<emphasis id="A1A9EAEDE419FFEDC5C5FB3D0FE4FAB9" bold="true" box="[151,393,1263,1290]" pageId="1" pageNumber="290">
<taxonomicName id="54DD4D7CE419FFEDC5C5FB3D0F5DFABA" ID-CoL="8XKG9" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[151,304,1263,1289]" class="Insecta" family="Staphylinidae" genus="Clavomicrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="290" phylum="Arthropoda" rank="genus" status="gen. nov." subFamily="Scydmaeninae">
<emphasis id="A1A9EAEDE419FFEDC5C5FB3D0F5DFABA" bold="true" box="[151,304,1263,1289]" italics="true" pageId="1" pageNumber="290">Clavomicrus</emphasis>
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<taxonomicNameLabel id="BA9A5796E419FFEDC465FB220FE4FAB9" box="[311,393,1264,1290]" pageId="1" pageNumber="290" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="DBC76574E419FFEDC5C5FAE50D68FAFF" box="[151,773,1334,1357]" pageId="1" pageNumber="290" type="type_taxon">
<paragraph id="936236FFE419FFEDC5C5FAE50D68FAFF" blockId="1.[151,773,1334,1357]" box="[151,773,1334,1357]" pageId="1" pageNumber="290">
<typeStatus id="4C66885DE419FFEDC5C5FAE50EA4FAFF" box="[151,201,1335,1356]" pageId="1" pageNumber="290">Type</typeStatus>
species: 
<taxonomicName id="54DD4D7CE419FFEDC474FAE40C64FAFE" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[294,521,1334,1357]" class="Insecta" family="Staphylinidae" genus="Clavomicrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="290" phylum="Arthropoda" rank="species" species="orousseti" status="sp. nov." subFamily="Scydmaeninae">
<emphasis id="A1A9EAEDE419FFEDC474FAE40C64FAFE" box="[294,521,1334,1357]" italics="true" pageId="1" pageNumber="290">Clavomicrus orousseti</emphasis>
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<taxonomicNameLabel id="BA9A5796E419FFEDC742FAE50C24FAFF" box="[528,585,1335,1356]" pageId="1" pageNumber="290" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(here designated).
</paragraph>
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<subSubSection id="DBC76574E419FFEDC5C5FAAB0DAFFA69" pageId="1" pageNumber="290" type="diagnosis">
<paragraph id="936236FFE419FFEDC5C5FAAB0DAFFA69" blockId="1.[151,1436,1401,2038]" pageId="1" pageNumber="290">
<emphasis id="A1A9EAEDE419FFEDC5C5FAAB0F7CFA21" bold="true" box="[151,273,1401,1426]" pageId="1" pageNumber="290">Diagnosis.</emphasis>
Antenna composed of ten antennomeres, with conspicuous oval club composed of completely fused antennomeres X and XI; prothorax lacking dorsal glandular openings and internal 'cavities'; pronotum with two pairs of lateral antebasal pits, inner pair connected by transverse groove.
</paragraph>
</subSubSection>
<subSubSection id="DBC76574E419FFEEC595FA370D94F9BB" lastPageId="2" lastPageNumber="291" pageId="1" pageNumber="290" type="description">
<paragraph id="936236FFE419FFEDC595FA370D9EFA4E" blockId="1.[151,1436,1401,2038]" box="[199,1011,1509,1534]" pageId="1" pageNumber="290">
<emphasis id="A1A9EAEDE419FFEDC595FA370F34FA4D" bold="true" box="[199,345,1509,1534]" pageId="1" pageNumber="290">Description.</emphasis>
Body (
<figureCitation id="0BE62A7AE419FFEDC4F8FA370C60FA4D" box="[426,525,1509,1534]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,899,922]" captionTargetBox="[155,1436,185,881]" captionTargetId="figure@3.[151,1436,185,881]" captionTargetPageId="3" captionText="FIGURES 1–5. Clavomicrus orousseti gen. &amp; sp. n. Dorsal (1) and lateral (2) habitus; head, prothorax and elytral base in dorsal view (3); distal portion of right antenna in dorsal view (4); head and prothorax in ventral view (5). Abbreviations: abp, antebasal pit; a8–11, antennomeres VIII–XI; bef, basal elytral pit; gp, gular plate; gs, gular suture; hy, hypomeron; hyr, hypomeral ridge; md, mandible; mn, mentum; pmn, prementum; psp, prosternal process; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.2588599" httpUri="https://zenodo.org/record/2588599/files/figure.png" pageId="1" pageNumber="290">Figs 1–2</figureCitation>
) suboval, strongly convex, sparsely setose.
</paragraph>
<paragraph id="936236FFE419FFEEC595F9DB0C32FF4B" blockId="1.[151,1436,1401,2038]" lastBlockId="2.[151,1437,151,2013]" lastPageId="2" lastPageNumber="291" pageId="1" pageNumber="290">
Head capsule (
<figureCitation id="0BE62A7AE419FFEDC43CF9DB0C63F991" box="[366,526,1545,1570]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,899,922]" captionTargetBox="[155,1436,185,881]" captionTargetId="figure@3.[151,1436,185,881]" captionTargetPageId="3" captionText="FIGURES 1–5. Clavomicrus orousseti gen. &amp; sp. n. Dorsal (1) and lateral (2) habitus; head, prothorax and elytral base in dorsal view (3); distal portion of right antenna in dorsal view (4); head and prothorax in ventral view (5). Abbreviations: abp, antebasal pit; a8–11, antennomeres VIII–XI; bef, basal elytral pit; gp, gular plate; gs, gular suture; hy, hypomeron; hyr, hypomeral ridge; md, mandible; mn, mentum; pmn, prementum; psp, prosternal process; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.2588599" httpUri="https://zenodo.org/record/2588599/files/figure.png" pageId="1" pageNumber="290">Figs 1–2, 3, 5</figureCitation>
; and Fig. 14B in 
<bibRefCitation id="F74C4B0EE419FFEDC784F9DB0DADF991" author="Jaloszynski, P." box="[726,960,1545,1570]" pageId="1" pageNumber="290" pagination="470 - 496" refId="ref3245" refString="Jaloszynski, P. (2011 b) Cephenniini with prothoracic glands and internal ' cavities': new taxa, enigmatic characters and phylogeny of the Cephennomicrus group of genera (Coleoptera, Staphylinidae, Scydmaeninae). Systematic Entomology, 36, 470 - 496. https: // doi. org / 10.1111 / j. 1365 - 3113.2011.00574. x" type="journal article" year="2011">Jałoszyński (2011b))</bibRefCitation>
short and broad, in intact beetles tilted so strongly that plane of vertex is perpendicular to the long body axis (
<figureCitation id="0BE62A7AE419FFEDC6C0F9FF0DBAF9F5" box="[914,983,1581,1606]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,899,922]" captionTargetBox="[155,1436,185,881]" captionTargetId="figure@3.[151,1436,185,881]" captionTargetPageId="3" captionText="FIGURES 1–5. Clavomicrus orousseti gen. &amp; sp. n. Dorsal (1) and lateral (2) habitus; head, prothorax and elytral base in dorsal view (3); distal portion of right antenna in dorsal view (4); head and prothorax in ventral view (5). Abbreviations: abp, antebasal pit; a8–11, antennomeres VIII–XI; bef, basal elytral pit; gp, gular plate; gs, gular suture; hy, hypomeron; hyr, hypomeral ridge; md, mandible; mn, mentum; pmn, prementum; psp, prosternal process; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.2588599" httpUri="https://zenodo.org/record/2588599/files/figure.png" pageId="1" pageNumber="290">Fig. 2</figureCitation>
); frons and vertex confluent, transverse, convex; supraantennal tubercles indistinct; frons with a pair of tiny diffuse tubercles; eyes large, coarsely faceted, strongly convex; gular plate (
<figureCitation id="0BE62A7AE419FFEDC4A3F9A70C57F93D" box="[497,570,1653,1678]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,899,922]" captionTargetBox="[155,1436,185,881]" captionTargetId="figure@3.[151,1436,185,881]" captionTargetPageId="3" captionText="FIGURES 1–5. Clavomicrus orousseti gen. &amp; sp. n. Dorsal (1) and lateral (2) habitus; head, prothorax and elytral base in dorsal view (3); distal portion of right antenna in dorsal view (4); head and prothorax in ventral view (5). Abbreviations: abp, antebasal pit; a8–11, antennomeres VIII–XI; bef, basal elytral pit; gp, gular plate; gs, gular suture; hy, hypomeron; hyr, hypomeral ridge; md, mandible; mn, mentum; pmn, prementum; psp, prosternal process; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.2588599" httpUri="https://zenodo.org/record/2588599/files/figure.png" pageId="1" pageNumber="290">Fig. 5</figureCitation>
; 
<emphasis id="A1A9EAEDE419FFEDC719F9A40C0AF93E" box="[587,615,1654,1677]" italics="true" pageId="1" pageNumber="290">gp</emphasis>
) large, transverse, with sharply marked gular sutures (
<figureCitation id="0BE62A7AE419FFEDC1BEF9A70B58F93D" box="[1260,1333,1653,1678]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,899,922]" captionTargetBox="[155,1436,185,881]" captionTargetId="figure@3.[151,1436,185,881]" captionTargetPageId="3" captionText="FIGURES 1–5. Clavomicrus orousseti gen. &amp; sp. n. Dorsal (1) and lateral (2) habitus; head, prothorax and elytral base in dorsal view (3); distal portion of right antenna in dorsal view (4); head and prothorax in ventral view (5). Abbreviations: abp, antebasal pit; a8–11, antennomeres VIII–XI; bef, basal elytral pit; gp, gular plate; gs, gular suture; hy, hypomeron; hyr, hypomeral ridge; md, mandible; mn, mentum; pmn, prementum; psp, prosternal process; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.2588599" httpUri="https://zenodo.org/record/2588599/files/figure.png" pageId="1" pageNumber="290">Fig. 5</figureCitation>
; 
<emphasis id="A1A9EAEDE419FFEDC014F9A40B32F93E" box="[1350,1375,1654,1677]" italics="true" pageId="1" pageNumber="290">gs</emphasis>
) and transverse reticulate microsculpture; posterior tentorial pits indiscernible; submentum (
<figureCitation id="0BE62A7AE419FFEDC138F94B0AC2F901" box="[1130,1199,1689,1714]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,899,922]" captionTargetBox="[155,1436,185,881]" captionTargetId="figure@3.[151,1436,185,881]" captionTargetPageId="3" captionText="FIGURES 1–5. Clavomicrus orousseti gen. &amp; sp. n. Dorsal (1) and lateral (2) habitus; head, prothorax and elytral base in dorsal view (3); distal portion of right antenna in dorsal view (4); head and prothorax in ventral view (5). Abbreviations: abp, antebasal pit; a8–11, antennomeres VIII–XI; bef, basal elytral pit; gp, gular plate; gs, gular suture; hy, hypomeron; hyr, hypomeral ridge; md, mandible; mn, mentum; pmn, prementum; psp, prosternal process; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.2588599" httpUri="https://zenodo.org/record/2588599/files/figure.png" pageId="1" pageNumber="290">Fig. 5</figureCitation>
; 
<emphasis id="A1A9EAEDE419FFEDC1EEF9480A84F902" box="[1212,1257,1690,1713]" italics="true" pageId="1" pageNumber="290">smn</emphasis>
) very short and strongly transverse. Mentum (
<figureCitation id="0BE62A7AE419FFEDC752F96F0C20F965" box="[512,589,1725,1750]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,899,922]" captionTargetBox="[155,1436,185,881]" captionTargetId="figure@3.[151,1436,185,881]" captionTargetPageId="3" captionText="FIGURES 1–5. Clavomicrus orousseti gen. &amp; sp. n. Dorsal (1) and lateral (2) habitus; head, prothorax and elytral base in dorsal view (3); distal portion of right antenna in dorsal view (4); head and prothorax in ventral view (5). Abbreviations: abp, antebasal pit; a8–11, antennomeres VIII–XI; bef, basal elytral pit; gp, gular plate; gs, gular suture; hy, hypomeron; hyr, hypomeral ridge; md, mandible; mn, mentum; pmn, prementum; psp, prosternal process; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.2588599" httpUri="https://zenodo.org/record/2588599/files/figure.png" pageId="1" pageNumber="290">Fig. 5</figureCitation>
; 
<emphasis id="A1A9EAEDE419FFEDC730F96C0CE8F966" box="[610,645,1726,1749]" italics="true" pageId="1" pageNumber="290">mn</emphasis>
and Fig. 14F in 
<bibRefCitation id="F74C4B0EE419FFEDC63AF96F0A34F965" author="Jaloszynski, P." box="[872,1113,1725,1750]" pageId="1" pageNumber="290" pagination="470 - 496" refId="ref3245" refString="Jaloszynski, P. (2011 b) Cephenniini with prothoracic glands and internal ' cavities': new taxa, enigmatic characters and phylogeny of the Cephennomicrus group of genera (Coleoptera, Staphylinidae, Scydmaeninae). Systematic Entomology, 36, 470 - 496. https: // doi. org / 10.1111 / j. 1365 - 3113.2011.00574. x" type="journal article" year="2011">Jałoszyński (2011b))</bibRefCitation>
subrectangular, transverse; prementum (
<figureCitation id="0BE62A7AE419FFEDC478F9330F1EF949" box="[298,371,1761,1786]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,899,922]" captionTargetBox="[155,1436,185,881]" captionTargetId="figure@3.[151,1436,185,881]" captionTargetPageId="3" captionText="FIGURES 1–5. Clavomicrus orousseti gen. &amp; sp. n. Dorsal (1) and lateral (2) habitus; head, prothorax and elytral base in dorsal view (3); distal portion of right antenna in dorsal view (4); head and prothorax in ventral view (5). Abbreviations: abp, antebasal pit; a8–11, antennomeres VIII–XI; bef, basal elytral pit; gp, gular plate; gs, gular suture; hy, hypomeron; hyr, hypomeral ridge; md, mandible; mn, mentum; pmn, prementum; psp, prosternal process; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.2588599" httpUri="https://zenodo.org/record/2588599/files/figure.png" pageId="1" pageNumber="290">Fig. 5</figureCitation>
; 
<emphasis id="A1A9EAEDE419FFEDC4D7F9300FD8F94A" box="[389,437,1762,1785]" italics="true" pageId="1" pageNumber="290">pmn</emphasis>
) short, with an unknown number of paired suckers, labial palps minute and broadly separated, inserted at sides of prementum, with palpomere I largest, slightly longer than broad, palpomere II much shorter and slightly narrower than I, shorter than broad, palpomere III much narrower and much longer than II, rodlike with narrowed apex, about 2.5 × as long as broad. Maxilla (Fig. 14E in 
<bibRefCitation id="F74C4B0EE419FFEDC153F89F0A81F8D5" author="Jaloszynski, P." box="[1025,1260,1869,1894]" pageId="1" pageNumber="290" pagination="470 - 496" refId="ref3245" refString="Jaloszynski, P. (2011 b) Cephenniini with prothoracic glands and internal ' cavities': new taxa, enigmatic characters and phylogeny of the Cephennomicrus group of genera (Coleoptera, Staphylinidae, Scydmaeninae). Systematic Entomology, 36, 470 - 496. https: // doi. org / 10.1111 / j. 1365 - 3113.2011.00574. x" type="journal article" year="2011">Jałoszyński (2011b))</bibRefCitation>
with triangular basistipes, elongate mediostipes, elongate lacinia and galea, each with sparse and thick mesal setae; elongate palpifer; maxillary palp (
<figureCitation id="0BE62A7AE419FFEDC4FCF8470F9EF81D" box="[430,499,1941,1966]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,899,922]" captionTargetBox="[155,1436,185,881]" captionTargetId="figure@3.[151,1436,185,881]" captionTargetPageId="3" captionText="FIGURES 1–5. Clavomicrus orousseti gen. &amp; sp. n. Dorsal (1) and lateral (2) habitus; head, prothorax and elytral base in dorsal view (3); distal portion of right antenna in dorsal view (4); head and prothorax in ventral view (5). Abbreviations: abp, antebasal pit; a8–11, antennomeres VIII–XI; bef, basal elytral pit; gp, gular plate; gs, gular suture; hy, hypomeron; hyr, hypomeral ridge; md, mandible; mn, mentum; pmn, prementum; psp, prosternal process; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.2588599" httpUri="https://zenodo.org/record/2588599/files/figure.png" pageId="1" pageNumber="290">Fig. 5</figureCitation>
; 
<emphasis id="A1A9EAEDE419FFEDC752F8440C43F81E" box="[512,558,1942,1965]" italics="true" pageId="1" pageNumber="290">mxp</emphasis>
; Fig. 14E in 
<bibRefCitation id="F74C4B0EE419FFEDC793F8470DC7F81D" author="Jaloszynski, P." box="[705,938,1941,1966]" pageId="1" pageNumber="290" pagination="470 - 496" refId="ref3245" refString="Jaloszynski, P. (2011 b) Cephenniini with prothoracic glands and internal ' cavities': new taxa, enigmatic characters and phylogeny of the Cephennomicrus group of genera (Coleoptera, Staphylinidae, Scydmaeninae). Systematic Entomology, 36, 470 - 496. https: // doi. org / 10.1111 / j. 1365 - 3113.2011.00574. x" type="journal article" year="2011">Jałoszyński (2011b))</bibRefCitation>
with minute palpomere I, strongly elongate, clavate palpomere II, strongly broadened palpomere III which is less than twice as long as broad and has truncate apex, palpomere IV broad and very short, button-like. Mandible (
<figureCitation id="0BE62A7AE419FFEDC63DF80F0DDEF845" box="[879,947,2013,2038]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,899,922]" captionTargetBox="[155,1436,185,881]" captionTargetId="figure@3.[151,1436,185,881]" captionTargetPageId="3" captionText="FIGURES 1–5. Clavomicrus orousseti gen. &amp; sp. n. Dorsal (1) and lateral (2) habitus; head, prothorax and elytral base in dorsal view (3); distal portion of right antenna in dorsal view (4); head and prothorax in ventral view (5). Abbreviations: abp, antebasal pit; a8–11, antennomeres VIII–XI; bef, basal elytral pit; gp, gular plate; gs, gular suture; hy, hypomeron; hyr, hypomeral ridge; md, mandible; mn, mentum; pmn, prementum; psp, prosternal process; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.2588599" httpUri="https://zenodo.org/record/2588599/files/figure.png" pageId="1" pageNumber="290">Fig. 5</figureCitation>
; 
<emphasis id="A1A9EAEDE419FFEDC6EDF80F0D8CF845" box="[959,993,2013,2038]" italics="true" pageId="1" pageNumber="290">md</emphasis>
; Fig. 14D in 
<bibRefCitation id="F74C4B0EE419FFEDC127F80F0B30F845" author="Jaloszynski, P." box="[1141,1373,2013,2038]" pageId="1" pageNumber="290" pagination="470 - 496" refId="ref3245" refString="Jaloszynski, P. (2011 b) Cephenniini with prothoracic glands and internal ' cavities': new taxa, enigmatic characters and phylogeny of the Cephennomicrus group of genera (Coleoptera, Staphylinidae, Scydmaeninae). Systematic Entomology, 36, 470 - 496. https: // doi. org / 10.1111 / j. 1365 - 3113.2011.00574. x" type="journal article" year="2011">Jałoszyński (2011b))</bibRefCitation>
short and robust, with subquadrate basal portion and large subtriangular, relatively blunt apical tooth bent mesad at a nearly right angle, setose prostheca absent. Labrum (Fig. 14C in 
<bibRefCitation id="F74C4B0EE41AFFEEC6C1FF6E0AEDFF66" author="Jaloszynski, P." box="[915,1152,188,213]" pageId="2" pageNumber="291" pagination="470 - 496" refId="ref3245" refString="Jaloszynski, P. (2011 b) Cephenniini with prothoracic glands and internal ' cavities': new taxa, enigmatic characters and phylogeny of the Cephennomicrus group of genera (Coleoptera, Staphylinidae, Scydmaeninae). Systematic Entomology, 36, 470 - 496. https: // doi. org / 10.1111 / j. 1365 - 3113.2011.00574. x" type="journal article" year="2011">Jałoszyński (2011b))</bibRefCitation>
subrectangular, strongly transverse, with anterior margin straight.
</paragraph>
<paragraph id="936236FFE41AFFEEC595FED60A6DFEF3" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="291">
Antennae (
<figureCitation id="0BE62A7AE41AFFEEC413FED60FAFFEAF" box="[321,450,260,285]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,899,922]" captionTargetBox="[155,1436,185,881]" captionTargetId="figure@3.[151,1436,185,881]" captionTargetPageId="3" captionText="FIGURES 1–5. Clavomicrus orousseti gen. &amp; sp. n. Dorsal (1) and lateral (2) habitus; head, prothorax and elytral base in dorsal view (3); distal portion of right antenna in dorsal view (4); head and prothorax in ventral view (5). Abbreviations: abp, antebasal pit; a8–11, antennomeres VIII–XI; bef, basal elytral pit; gp, gular plate; gs, gular suture; hy, hypomeron; hyr, hypomeral ridge; md, mandible; mn, mentum; pmn, prementum; psp, prosternal process; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.2588599" httpUri="https://zenodo.org/record/2588599/files/figure.png" pageId="2" pageNumber="291">Figs 1–2, 4</figureCitation>
; Fig. 14G in 
<bibRefCitation id="F74C4B0EE41AFFEEC706FED60D51FEAE" author="Jaloszynski, P." box="[596,828,260,285]" pageId="2" pageNumber="291" pagination="470 - 496" refId="ref3245" refString="Jaloszynski, P. (2011 b) Cephenniini with prothoracic glands and internal ' cavities': new taxa, enigmatic characters and phylogeny of the Cephennomicrus group of genera (Coleoptera, Staphylinidae, Scydmaeninae). Systematic Entomology, 36, 470 - 496. https: // doi. org / 10.1111 / j. 1365 - 3113.2011.00574. x" type="journal article" year="2011">Jałoszyński (2011b))</bibRefCitation>
long and slender, composed of 10 antennomeres, with a composite oval and abruptly delimited club (fused antennomeres X and XI).
</paragraph>
<paragraph id="936236FFE41AFFEEC595FE9F0C1EFE63" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="291">
Pronotum (
<figureCitation id="0BE62A7AE41AFFEEC417FE9E0FEAFED7" box="[325,391,332,357]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,899,922]" captionTargetBox="[155,1436,185,881]" captionTargetId="figure@3.[151,1436,185,881]" captionTargetPageId="3" captionText="FIGURES 1–5. Clavomicrus orousseti gen. &amp; sp. n. Dorsal (1) and lateral (2) habitus; head, prothorax and elytral base in dorsal view (3); distal portion of right antenna in dorsal view (4); head and prothorax in ventral view (5). Abbreviations: abp, antebasal pit; a8–11, antennomeres VIII–XI; bef, basal elytral pit; gp, gular plate; gs, gular suture; hy, hypomeron; hyr, hypomeral ridge; md, mandible; mn, mentum; pmn, prementum; psp, prosternal process; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.2588599" httpUri="https://zenodo.org/record/2588599/files/figure.png" pageId="2" pageNumber="291">Fig. 3</figureCitation>
) in dorsal view subtrapezoidal with rounded anterior margin, rounded anterior corners which are not visible in dorsal view, shallowly serrate and complete lateral carinae, distinct posterior corners and posterior margin with small median emargination. Pronotal base with two pairs of small lateral antebasal pits (
<figureCitation id="0BE62A7AE41AFFEEC05DFE460B39FE1F" box="[1295,1364,404,429]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,899,922]" captionTargetBox="[155,1436,185,881]" captionTargetId="figure@3.[151,1436,185,881]" captionTargetPageId="3" captionText="FIGURES 1–5. Clavomicrus orousseti gen. &amp; sp. n. Dorsal (1) and lateral (2) habitus; head, prothorax and elytral base in dorsal view (3); distal portion of right antenna in dorsal view (4); head and prothorax in ventral view (5). Abbreviations: abp, antebasal pit; a8–11, antennomeres VIII–XI; bef, basal elytral pit; gp, gular plate; gs, gular suture; hy, hypomeron; hyr, hypomeral ridge; md, mandible; mn, mentum; pmn, prementum; psp, prosternal process; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.2588599" httpUri="https://zenodo.org/record/2588599/files/figure.png" pageId="2" pageNumber="291">Fig. 3</figureCitation>
; 
<emphasis id="A1A9EAEDE41AFFEEC030FE460BE1FE1E" box="[1378,1420,404,429]" italics="true" pageId="2" pageNumber="291">abp</emphasis>
), inner pair connected by transverse groove.
</paragraph>
<paragraph id="936236FFE41AFFEEC595FE0F0D7AFD14" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="291">
Prosternum (
<figureCitation id="0BE62A7AE41AFFEEC408FE0E0FCDFE46" box="[346,416,476,501]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,899,922]" captionTargetBox="[155,1436,185,881]" captionTargetId="figure@3.[151,1436,185,881]" captionTargetPageId="3" captionText="FIGURES 1–5. Clavomicrus orousseti gen. &amp; sp. n. Dorsal (1) and lateral (2) habitus; head, prothorax and elytral base in dorsal view (3); distal portion of right antenna in dorsal view (4); head and prothorax in ventral view (5). Abbreviations: abp, antebasal pit; a8–11, antennomeres VIII–XI; bef, basal elytral pit; gp, gular plate; gs, gular suture; hy, hypomeron; hyr, hypomeral ridge; md, mandible; mn, mentum; pmn, prementum; psp, prosternal process; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.2588599" httpUri="https://zenodo.org/record/2588599/files/figure.png" pageId="2" pageNumber="291">Fig. 5</figureCitation>
; Fig. 14H in 
<bibRefCitation id="F74C4B0EE41AFFEEC76CFE0E0D47FE46" author="Jaloszynski, P." box="[574,810,476,501]" pageId="2" pageNumber="291" pagination="470 - 496" refId="ref3245" refString="Jaloszynski, P. (2011 b) Cephenniini with prothoracic glands and internal ' cavities': new taxa, enigmatic characters and phylogeny of the Cephennomicrus group of genera (Coleoptera, Staphylinidae, Scydmaeninae). Systematic Entomology, 36, 470 - 496. https: // doi. org / 10.1111 / j. 1365 - 3113.2011.00574. x" type="journal article" year="2011">Jałoszyński (2011b))</bibRefCitation>
with basisternal portion much shorter than coxal part; prosternal process (
<figureCitation id="0BE62A7AE41AFFEEC425FE2D0FD2FDAB" box="[375,447,511,536]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,899,922]" captionTargetBox="[155,1436,185,881]" captionTargetId="figure@3.[151,1436,185,881]" captionTargetPageId="3" captionText="FIGURES 1–5. Clavomicrus orousseti gen. &amp; sp. n. Dorsal (1) and lateral (2) habitus; head, prothorax and elytral base in dorsal view (3); distal portion of right antenna in dorsal view (4); head and prothorax in ventral view (5). Abbreviations: abp, antebasal pit; a8–11, antennomeres VIII–XI; bef, basal elytral pit; gp, gular plate; gs, gular suture; hy, hypomeron; hyr, hypomeral ridge; md, mandible; mn, mentum; pmn, prementum; psp, prosternal process; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.2588599" httpUri="https://zenodo.org/record/2588599/files/figure.png" pageId="2" pageNumber="291">Fig. 5</figureCitation>
; 
<emphasis id="A1A9EAEDE41AFFEEC49DFDD30F9BFDAB" box="[463,502,513,536]" italics="true" pageId="2" pageNumber="291">psp</emphasis>
) in ventral view subtriangular, in lateral view (Fig. 14H in 
<bibRefCitation id="F74C4B0EE41AFFEEC1E2FE2D0BF1FDAB" author="Jaloszynski, P." box="[1200,1436,511,536]" pageId="2" pageNumber="291" pagination="470 - 496" refId="ref3245" refString="Jaloszynski, P. (2011 b) Cephenniini with prothoracic glands and internal ' cavities': new taxa, enigmatic characters and phylogeny of the Cephennomicrus group of genera (Coleoptera, Staphylinidae, Scydmaeninae). Systematic Entomology, 36, 470 - 496. https: // doi. org / 10.1111 / j. 1365 - 3113.2011.00574. x" type="journal article" year="2011">Jałoszyński (2011b))</bibRefCitation>
subtriangular with rounded apex, elongate but not expanding ventrad beyond procoxae, slightly bent posterad; notosternal sutures obliterated in front of procoxae; procoxal cavities broadly closed by posterolateral lobes of prosternum that are fused with postcoxal lobes of hypomera; hypomeral ridges (
<figureCitation id="0BE62A7AE41AFFEEC178FDBE0A1DFD36" box="[1066,1136,620,645]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,899,922]" captionTargetBox="[155,1436,185,881]" captionTargetId="figure@3.[151,1436,185,881]" captionTargetPageId="3" captionText="FIGURES 1–5. Clavomicrus orousseti gen. &amp; sp. n. Dorsal (1) and lateral (2) habitus; head, prothorax and elytral base in dorsal view (3); distal portion of right antenna in dorsal view (4); head and prothorax in ventral view (5). Abbreviations: abp, antebasal pit; a8–11, antennomeres VIII–XI; bef, basal elytral pit; gp, gular plate; gs, gular suture; hy, hypomeron; hyr, hypomeral ridge; md, mandible; mn, mentum; pmn, prementum; psp, prosternal process; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.2588599" httpUri="https://zenodo.org/record/2588599/files/figure.png" pageId="2" pageNumber="291">Fig. 5</figureCitation>
; 
<emphasis id="A1A9EAEDE41AFFEEC12CFDBE0ACEFD36" box="[1150,1187,620,645]" italics="true" pageId="2" pageNumber="291">hyr</emphasis>
) complete; hypomera (
<figureCitation id="0BE62A7AE41AFFEEC5CDFD5D0E89FD1B" box="[159,228,655,680]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,899,922]" captionTargetBox="[155,1436,185,881]" captionTargetId="figure@3.[151,1436,185,881]" captionTargetPageId="3" captionText="FIGURES 1–5. Clavomicrus orousseti gen. &amp; sp. n. Dorsal (1) and lateral (2) habitus; head, prothorax and elytral base in dorsal view (3); distal portion of right antenna in dorsal view (4); head and prothorax in ventral view (5). Abbreviations: abp, antebasal pit; a8–11, antennomeres VIII–XI; bef, basal elytral pit; gp, gular plate; gs, gular suture; hy, hypomeron; hyr, hypomeral ridge; md, mandible; mn, mentum; pmn, prementum; psp, prosternal process; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.2588599" httpUri="https://zenodo.org/record/2588599/files/figure.png" pageId="2" pageNumber="291">Fig. 5</figureCitation>
; 
<emphasis id="A1A9EAEDE41AFFEEC5A2FD5D0F67FD1B" box="[240,266,655,680]" italics="true" pageId="2" pageNumber="291">hy</emphasis>
) outside hypomeral ridges broad and concave.
</paragraph>
<paragraph id="936236FFE41AFFEEC595FD670B0CFD5C" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="291">
Mesonotum (Fig. 14J in 
<bibRefCitation id="F74C4B0EE41AFFEEC4B8FD660CBAFD7E" author="Jaloszynski, P." box="[490,727,692,717]" pageId="2" pageNumber="291" pagination="470 - 496" refId="ref3245" refString="Jaloszynski, P. (2011 b) Cephenniini with prothoracic glands and internal ' cavities': new taxa, enigmatic characters and phylogeny of the Cephennomicrus group of genera (Coleoptera, Staphylinidae, Scydmaeninae). Systematic Entomology, 36, 470 - 496. https: // doi. org / 10.1111 / j. 1365 - 3113.2011.00574. x" type="journal article" year="2011">Jałoszyński (2011b))</bibRefCitation>
subtrapezoidal, with subtriangular, transverse mesoscutellum largely exposed between elytral bases (
<figureCitation id="0BE62A7AE41AFFEEC71FFD050CFDFD43" box="[589,656,727,752]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,899,922]" captionTargetBox="[155,1436,185,881]" captionTargetId="figure@3.[151,1436,185,881]" captionTargetPageId="3" captionText="FIGURES 1–5. Clavomicrus orousseti gen. &amp; sp. n. Dorsal (1) and lateral (2) habitus; head, prothorax and elytral base in dorsal view (3); distal portion of right antenna in dorsal view (4); head and prothorax in ventral view (5). Abbreviations: abp, antebasal pit; a8–11, antennomeres VIII–XI; bef, basal elytral pit; gp, gular plate; gs, gular suture; hy, hypomeron; hyr, hypomeral ridge; md, mandible; mn, mentum; pmn, prementum; psp, prosternal process; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.2588599" httpUri="https://zenodo.org/record/2588599/files/figure.png" pageId="2" pageNumber="291">Fig. 3</figureCitation>
), scutoscutellar suture present, mesoscutum strongly transverse.
</paragraph>
<paragraph id="936236FFE41AFFEEC595FD2E0E94FCEE" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="291">
Elytra (
<figureCitation id="0BE62A7AE41AFFEEC448FD2E0FEDFCA7" box="[282,384,764,789]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,899,922]" captionTargetBox="[155,1436,185,881]" captionTargetId="figure@3.[151,1436,185,881]" captionTargetPageId="3" captionText="FIGURES 1–5. Clavomicrus orousseti gen. &amp; sp. n. Dorsal (1) and lateral (2) habitus; head, prothorax and elytral base in dorsal view (3); distal portion of right antenna in dorsal view (4); head and prothorax in ventral view (5). Abbreviations: abp, antebasal pit; a8–11, antennomeres VIII–XI; bef, basal elytral pit; gp, gular plate; gs, gular suture; hy, hypomeron; hyr, hypomeral ridge; md, mandible; mn, mentum; pmn, prementum; psp, prosternal process; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.2588599" httpUri="https://zenodo.org/record/2588599/files/figure.png" pageId="2" pageNumber="291">Figs 1–3</figureCitation>
; Fig. 14M in 
<bibRefCitation id="F74C4B0EE41AFFEEC748FD2E0D6EFCA6" author="Jaloszynski, P." box="[538,771,764,789]" pageId="2" pageNumber="291" pagination="470 - 496" refId="ref3245" refString="Jaloszynski, P. (2011 b) Cephenniini with prothoracic glands and internal ' cavities': new taxa, enigmatic characters and phylogeny of the Cephennomicrus group of genera (Coleoptera, Staphylinidae, Scydmaeninae). Systematic Entomology, 36, 470 - 496. https: // doi. org / 10.1111 / j. 1365 - 3113.2011.00574. x" type="journal article" year="2011">Jałoszyński (2011b))</bibRefCitation>
oval, each with a distinct elongate subhumeral impression, lacking humeral denticle, each elytron with one vestigial asetose basal fovea (
<figureCitation id="0BE62A7AE41AFFEEC150FCCD0A2BFC8B" box="[1026,1094,799,824]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,899,922]" captionTargetBox="[155,1436,185,881]" captionTargetId="figure@3.[151,1436,185,881]" captionTargetPageId="3" captionText="FIGURES 1–5. Clavomicrus orousseti gen. &amp; sp. n. Dorsal (1) and lateral (2) habitus; head, prothorax and elytral base in dorsal view (3); distal portion of right antenna in dorsal view (4); head and prothorax in ventral view (5). Abbreviations: abp, antebasal pit; a8–11, antennomeres VIII–XI; bef, basal elytral pit; gp, gular plate; gs, gular suture; hy, hypomeron; hyr, hypomeral ridge; md, mandible; mn, mentum; pmn, prementum; psp, prosternal process; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.2588599" httpUri="https://zenodo.org/record/2588599/files/figure.png" pageId="2" pageNumber="291">Fig. 3</figureCitation>
; 
<emphasis id="A1A9EAEDE41AFFEEC101FCCD0A18FC8B" box="[1107,1141,799,824]" italics="true" pageId="2" pageNumber="291">bef</emphasis>
), apices of elytra rounded together.
</paragraph>
<paragraph id="936236FFE41AFFEEC595FCB50B45FC5F" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="291">
Mesoventrite (
<figureCitation id="0BE62A7AE41AFFEEC43FFCB50FD9FC33" box="[365,436,871,896]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1713,1736]" captionTargetBox="[161,1422,735,1691]" captionTargetId="figure@4.[161,1426,735,1691]" captionTargetPageId="4" captionText="FIGURES 6–10. Clavomicrus orousseti gen. &amp; sp. n. Pterothorax and part of abdomen in ventral view (6); aedeagus in ventral (7, 9) and lateral (8, 10) views. Abbreviations: ar, anterior ridge; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." figureDoi="http://doi.org/10.5281/zenodo.2588601" httpUri="https://zenodo.org/record/2588601/files/figure.png" pageId="2" pageNumber="291">Fig. 6</figureCitation>
; Fig. 14L in 
<bibRefCitation id="F74C4B0EE41AFFEEC703FCB50D50FC33" author="Jaloszynski, P." box="[593,829,871,896]" pageId="2" pageNumber="291" pagination="470 - 496" refId="ref3245" refString="Jaloszynski, P. (2011 b) Cephenniini with prothoracic glands and internal ' cavities': new taxa, enigmatic characters and phylogeny of the Cephennomicrus group of genera (Coleoptera, Staphylinidae, Scydmaeninae). Systematic Entomology, 36, 470 - 496. https: // doi. org / 10.1111 / j. 1365 - 3113.2011.00574. x" type="journal article" year="2011">Jałoszyński (2011b))</bibRefCitation>
with narrow anterior ridge (
<figureCitation id="0BE62A7AE41AFFEEC1DAFCB50AA2FC33" box="[1160,1231,871,896]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1713,1736]" captionTargetBox="[161,1422,735,1691]" captionTargetId="figure@4.[161,1426,735,1691]" captionTargetPageId="4" captionText="FIGURES 6–10. Clavomicrus orousseti gen. &amp; sp. n. Pterothorax and part of abdomen in ventral view (6); aedeagus in ventral (7, 9) and lateral (8, 10) views. Abbreviations: ar, anterior ridge; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." figureDoi="http://doi.org/10.5281/zenodo.2588601" httpUri="https://zenodo.org/record/2588601/files/figure.png" pageId="2" pageNumber="291">Fig. 6</figureCitation>
; 
<emphasis id="A1A9EAEDE41AFFEEC18DFCBB0A95FC33" box="[1247,1272,873,896]" italics="true" pageId="2" pageNumber="291">ar</emphasis>
); mesoventral intercoxal process (
<figureCitation id="0BE62A7AE41AFFEEC426FC5E0FD7FC16" box="[372,442,908,933]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1713,1736]" captionTargetBox="[161,1422,735,1691]" captionTargetId="figure@4.[161,1426,735,1691]" captionTargetPageId="4" captionText="FIGURES 6–10. Clavomicrus orousseti gen. &amp; sp. n. Pterothorax and part of abdomen in ventral view (6); aedeagus in ventral (7, 9) and lateral (8, 10) views. Abbreviations: ar, anterior ridge; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." figureDoi="http://doi.org/10.5281/zenodo.2588601" httpUri="https://zenodo.org/record/2588601/files/figure.png" pageId="2" pageNumber="291">Fig. 6</figureCitation>
; 
<emphasis id="A1A9EAEDE41AFFEEC49BFC5F0C6EFC17" box="[457,515,909,932]" italics="true" pageId="2" pageNumber="291">msvp</emphasis>
) very broad but with its median region raised and forming diffuse and relatively narrow longitudinal elevation, posterior margin of mesoventral process concave, posterior arms present, long. Lateral impressions functioning as procoxal rests (Fig. 14; 
<emphasis id="A1A9EAEDE41AFFEEC679FC070D3DFC5F" box="[811,848,981,1004]" italics="true" pageId="2" pageNumber="291">pcr</emphasis>
) present, separated at middle and asetose.
</paragraph>
<paragraph id="936236FFE41AFFEEC595FC2A0C6CFB87" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="291">
Metanotum (Fig. 14K in 
<bibRefCitation id="F74C4B0EE41AFFEEC4AAFC250C8AFBA3" author="Jaloszynski, P." box="[504,743,1015,1040]" pageId="2" pageNumber="291" pagination="470 - 496" refId="ref3245" refString="Jaloszynski, P. (2011 b) Cephenniini with prothoracic glands and internal ' cavities': new taxa, enigmatic characters and phylogeny of the Cephennomicrus group of genera (Coleoptera, Staphylinidae, Scydmaeninae). Systematic Entomology, 36, 470 - 496. https: // doi. org / 10.1111 / j. 1365 - 3113.2011.00574. x" type="journal article" year="2011">Jałoszyński (2011b))</bibRefCitation>
largely membranous, with strongly reduced alacristae not reaching middle of metascutum.
</paragraph>
<paragraph id="936236FFE41AFFEEC595FBED0FF2FBEB" blockId="2.[151,1437,151,2013]" box="[199,415,1087,1112]" pageId="2" pageNumber="291">Hind wings absent.</paragraph>
<paragraph id="936236FFE41AFFEEC595FBB60F1AFB5B" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="291">
Metaventrite (
<figureCitation id="0BE62A7AE41AFFEEC43AFBB60FC3FBCE" box="[360,430,1124,1149]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1713,1736]" captionTargetBox="[161,1422,735,1691]" captionTargetId="figure@4.[161,1426,735,1691]" captionTargetPageId="4" captionText="FIGURES 6–10. Clavomicrus orousseti gen. &amp; sp. n. Pterothorax and part of abdomen in ventral view (6); aedeagus in ventral (7, 9) and lateral (8, 10) views. Abbreviations: ar, anterior ridge; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." figureDoi="http://doi.org/10.5281/zenodo.2588601" httpUri="https://zenodo.org/record/2588601/files/figure.png" pageId="2" pageNumber="291">Fig. 6</figureCitation>
; Fig. 14L in 
<bibRefCitation id="F74C4B0EE41AFFEEC71BFBB60D59FBCE" author="Jaloszynski, P." box="[585,820,1124,1149]" pageId="2" pageNumber="291" pagination="470 - 496" refId="ref3245" refString="Jaloszynski, P. (2011 b) Cephenniini with prothoracic glands and internal ' cavities': new taxa, enigmatic characters and phylogeny of the Cephennomicrus group of genera (Coleoptera, Staphylinidae, Scydmaeninae). Systematic Entomology, 36, 470 - 496. https: // doi. org / 10.1111 / j. 1365 - 3113.2011.00574. x" type="journal article" year="2011">Jałoszyński (2011b))</bibRefCitation>
short and transverse; posterior margins of mesocoxal rests not carinate; sides slightly convergent posterad, posterior margin weakly concave at each metacoxa, with broad and short metaventral intercoxal process (
<figureCitation id="0BE62A7AE41AFFEEC792FB7E0D6AFB76" box="[704,775,1196,1221]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1713,1736]" captionTargetBox="[161,1422,735,1691]" captionTargetId="figure@4.[161,1426,735,1691]" captionTargetPageId="4" captionText="FIGURES 6–10. Clavomicrus orousseti gen. &amp; sp. n. Pterothorax and part of abdomen in ventral view (6); aedeagus in ventral (7, 9) and lateral (8, 10) views. Abbreviations: ar, anterior ridge; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." figureDoi="http://doi.org/10.5281/zenodo.2588601" httpUri="https://zenodo.org/record/2588601/files/figure.png" pageId="2" pageNumber="291">Fig. 6</figureCitation>
; 
<emphasis id="A1A9EAEDE41AFFEEC644FB7F0D21FB77" box="[790,844,1197,1220]" italics="true" pageId="2" pageNumber="291">mtvp</emphasis>
) slightly emarginate at middle. Metanepisterna and metepimera narrow.
</paragraph>
<paragraph id="936236FFE41AFFEEC595FB260D77FA83" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="291">
Metendosternite (= metafurca) (Fig. 14 E in 
<bibRefCitation id="F74C4B0EE41AFFEEC7E7FB260DF3FABE" author="Jaloszynski, P." box="[693,926,1268,1293]" pageId="2" pageNumber="291" pagination="470 - 496" refId="ref3245" refString="Jaloszynski, P. (2011 b) Cephenniini with prothoracic glands and internal ' cavities': new taxa, enigmatic characters and phylogeny of the Cephennomicrus group of genera (Coleoptera, Staphylinidae, Scydmaeninae). Systematic Entomology, 36, 470 - 496. https: // doi. org / 10.1111 / j. 1365 - 3113.2011.00574. x" type="journal article" year="2011">Jałoszyński (2011b))</bibRefCitation>
with very short and broad stalk and divergent lateral furcal arms with adjacent bases (nearly V-shaped).
</paragraph>
<paragraph id="936236FFE41AFFEEC595FAEF0E83FA2E" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="291">
Legs (
<figureCitation id="0BE62A7AE41AFFEEC45CFAEE0FC0FAE6" box="[270,429,1340,1365]" captionStart-0="FIGURES 1–5" captionStart-1="FIGURES 6–10" captionStartId-0="3.[151,264,899,922]" captionStartId-1="4.[151,264,1713,1736]" captionTargetBox-0="[155,1436,185,881]" captionTargetBox-1="[161,1422,735,1691]" captionTargetId-0="figure@3.[151,1436,185,881]" captionTargetId-1="figure@4.[161,1426,735,1691]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 1–5. Clavomicrus orousseti gen. &amp; sp. n. Dorsal (1) and lateral (2) habitus; head, prothorax and elytral base in dorsal view (3); distal portion of right antenna in dorsal view (4); head and prothorax in ventral view (5). Abbreviations: abp, antebasal pit; a8–11, antennomeres VIII–XI; bef, basal elytral pit; gp, gular plate; gs, gular suture; hy, hypomeron; hyr, hypomeral ridge; md, mandible; mn, mentum; pmn, prementum; psp, prosternal process; smn, submentum." captionText-1="FIGURES 6–10. Clavomicrus orousseti gen. &amp; sp. n. Pterothorax and part of abdomen in ventral view (6); aedeagus in ventral (7, 9) and lateral (8, 10) views. Abbreviations: ar, anterior ridge; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." figureDoi-0="http://doi.org/10.5281/zenodo.2588599" figureDoi-1="http://doi.org/10.5281/zenodo.2588601" httpUri-0="https://zenodo.org/record/2588599/files/figure.png" httpUri-1="https://zenodo.org/record/2588601/files/figure.png" pageId="2" pageNumber="291">Figs 1–2, 5–6</figureCitation>
) moderately long and slender; pro- and mesocoxae oval, metacoxae strongly transverse; all trochanters short and subtriangular; all femora distinctly clavate; tibiae broadening distally; tarsi moderately slender.
</paragraph>
<paragraph id="936236FFE41AFFEEC595FA750AD7FA73" blockId="2.[151,1437,151,2013]" box="[199,1210,1447,1472]" pageId="2" pageNumber="291">
Abdominal sternites (
<figureCitation id="0BE62A7AE41AFFEEC4EAFA750F91FA73" box="[440,508,1447,1472]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1713,1736]" captionTargetBox="[161,1422,735,1691]" captionTargetId="figure@4.[161,1426,735,1691]" captionTargetPageId="4" captionText="FIGURES 6–10. Clavomicrus orousseti gen. &amp; sp. n. Pterothorax and part of abdomen in ventral view (6); aedeagus in ventral (7, 9) and lateral (8, 10) views. Abbreviations: ar, anterior ridge; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." figureDoi="http://doi.org/10.5281/zenodo.2588601" httpUri="https://zenodo.org/record/2588601/files/figure.png" pageId="2" pageNumber="291">Fig. 6</figureCitation>
) unmodified, sternite III (i.e., first visible) twice as long as IV.
</paragraph>
<paragraph id="936236FFE41AFFEEC595FA1E0D94F9BB" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="291">
Aedeagus (
<figureCitation id="0BE62A7AE41AFFEEC414FA1E0FD4FA57" box="[326,441,1484,1509]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1713,1736]" captionTargetBox="[161,1422,735,1691]" captionTargetId="figure@4.[161,1426,735,1691]" captionTargetPageId="4" captionText="FIGURES 6–10. Clavomicrus orousseti gen. &amp; sp. n. Pterothorax and part of abdomen in ventral view (6); aedeagus in ventral (7, 9) and lateral (8, 10) views. Abbreviations: ar, anterior ridge; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." figureDoi="http://doi.org/10.5281/zenodo.2588601" httpUri="https://zenodo.org/record/2588601/files/figure.png" pageId="2" pageNumber="291">Figs 7–10</figureCitation>
) elongate, with symmetrical median lobe and endophallus, diaphragm present, circular, sub-basal on ventral wall; parameres slender, with apical and subapical setae.
</paragraph>
</subSubSection>
<subSubSection id="DBC76574E41AFFEEC595F9C60F70F9E3" pageId="2" pageNumber="291" type="distribution">
<paragraph id="936236FFE41AFFEEC595F9C60F70F9E3" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="291">
<emphasis id="A1A9EAEDE41AFFEEC595F9C60C58F99E" bold="true" box="[199,565,1556,1581]" pageId="2" pageNumber="291">Distribution and composition.</emphasis>
<taxonomicName id="54DD4D7CE41AFFEEC712F9C60CA2F99E" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[576,719,1556,1581]" class="Insecta" family="Staphylinidae" genus="Clavomicrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="290" phylum="Arthropoda" rank="genus" status="gen. nov." subFamily="Scydmaeninae">
<emphasis id="A1A9EAEDE41AFFEEC712F9C60CA2F99E" box="[576,719,1556,1581]" italics="true" pageId="2" pageNumber="291">Clavomicrus</emphasis>
</taxonomicName>
is represented by one species known to occur in Luzon, the 
<collectingCountry id="EBCA766FE41AFFEEC5C5F9E50F74F9E3" box="[151,281,1591,1616]" name="Philippines" pageId="2" pageNumber="291">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="DBC76574E41AFFEEC595F98E0F7DF90E" pageId="2" pageNumber="291" type="etymology">
<paragraph id="936236FFE41AFFEEC595F98E0F7DF90E" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="291">
<emphasis id="A1A9EAEDE41AFFEEC595F98E0F23F9C6" bold="true" box="[199,334,1628,1653]" pageId="2" pageNumber="291">Etymology.</emphasis>
The name 
<taxonomicName id="54DD4D7CE41AFFEEC484F98E0C08F9C6" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[470,613,1628,1653]" class="Insecta" family="Staphylinidae" genus="Clavomicrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="290" phylum="Arthropoda" rank="genus" status="gen. nov." subFamily="Scydmaeninae">
<emphasis id="A1A9EAEDE41AFFEEC484F98E0C08F9C6" box="[470,613,1628,1653]" italics="true" pageId="2" pageNumber="291">Clavomicrus</emphasis>
</taxonomicName>
combines the prefix 
<emphasis id="A1A9EAEDE41AFFEEC633F98E0DF0F9C6" box="[865,925,1628,1653]" italics="true" pageId="2" pageNumber="291">clavo</emphasis>
- derived from the Latin 
<emphasis id="A1A9EAEDE41AFFEEC197F98E0B6CF9C6" box="[1221,1281,1628,1653]" italics="true" pageId="2" pageNumber="291">clava</emphasis>
(a club) that refers to the antennal structure, with the stem - 
<emphasis id="A1A9EAEDE41AFFEEC7E9F9530D6AF92B" box="[699,775,1665,1688]" italics="true" pageId="2" pageNumber="291">micrus</emphasis>
, to emphasize the similarity to 
<taxonomicName id="54DD4D7CE41AFFEEC129F9AD0B54F92B" authorityName="Reitter" authorityYear="1907" box="[1147,1337,1663,1688]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="A1A9EAEDE41AFFEEC129F9AD0B54F92B" box="[1147,1337,1663,1688]" italics="true" pageId="2" pageNumber="291">Cephennomicrus</emphasis>
</taxonomicName>
. Gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="DBC76574E41AFFEEC595F9150CA0F86F" pageId="2" pageNumber="291" type="discussion">
<paragraph id="936236FFE41AFFEEC595F9150CA0F86F" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="291">
<emphasis id="A1A9EAEDE41AFFEEC595F9150F56F953" bold="true" box="[199,315,1735,1760]" pageId="2" pageNumber="291">Remarks.</emphasis>
Among Cephenniini, decamerous antennae with a large, oval club can be found only in 
<taxonomicName id="54DD4D7CE41AFFEEC5C5F93E0F4BF8B6" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[151,294,1772,1797]" class="Insecta" family="Staphylinidae" genus="Clavomicrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="290" phylum="Arthropoda" rank="genus" status="gen. nov." subFamily="Scydmaeninae">
<emphasis id="A1A9EAEDE41AFFEEC5C5F93E0F4BF8B6" box="[151,294,1772,1797]" italics="true" pageId="2" pageNumber="291">Clavomicrus</emphasis>
</taxonomicName>
and 
<taxonomicName id="54DD4D7CE41AFFEEC430F93E0FDCF8B6" authorityName="Jaloszynski" authorityYear="2009" box="[354,433,1772,1797]" class="Insecta" family="Staphylinidae" genus="Trurlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="A1A9EAEDE41AFFEEC430F93E0FDCF8B6" box="[354,433,1772,1797]" italics="true" pageId="2" pageNumber="291">Trurlia</emphasis>
</taxonomicName>
. The new genus differs from 
<taxonomicName id="54DD4D7CE41AFFEEC65DF93E0D33F8B6" authorityName="Jaloszynski" authorityYear="2009" box="[783,862,1772,1797]" class="Insecta" family="Staphylinidae" genus="Trurlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="A1A9EAEDE41AFFEEC65DF93E0D33F8B6" box="[783,862,1772,1797]" italics="true" pageId="2" pageNumber="291">Trurlia</emphasis>
</taxonomicName>
(among other, minor characters) in the prothorax lacking a pair of large glandular openings on its dorsal surface (present in 
<taxonomicName id="54DD4D7CE41AFFEEC681F8DD0A4CF89B" authorityName="Jaloszynski" authorityYear="2009" box="[979,1057,1807,1832]" class="Insecta" family="Staphylinidae" genus="Trurlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="A1A9EAEDE41AFFEEC681F8DD0A4CF89B" box="[979,1057,1807,1832]" italics="true" pageId="2" pageNumber="291">Trurlia</emphasis>
</taxonomicName>
), the mandibles lacking preapical teeth (two teeth present in 
<taxonomicName id="54DD4D7CE41AFFEEC496F8E60C7EF8FE" authorityName="Jaloszynski" authorityYear="2009" box="[452,531,1844,1869]" class="Insecta" family="Staphylinidae" genus="Trurlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="A1A9EAEDE41AFFEEC496F8E60C7EF8FE" box="[452,531,1844,1869]" italics="true" pageId="2" pageNumber="291">Trurlia</emphasis>
</taxonomicName>
), the prosternal process subtriangular in ventral view and lacking an anterior tip (suboval and with its ventral surface flattened in 
<taxonomicName id="54DD4D7CE41AFFEEC79FF8850D76F8C3" authorityName="Jaloszynski" authorityYear="2009" box="[717,795,1879,1904]" class="Insecta" family="Staphylinidae" genus="Trurlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="A1A9EAEDE41AFFEEC79FF8850D76F8C3" box="[717,795,1879,1904]" italics="true" pageId="2" pageNumber="291">Trurlia</emphasis>
</taxonomicName>
, and with a small anterior projection that in the ventral view is visible as an anterior tip projecting in front of the anterior prosternal margin), a much broader mesoventral intercoxal process, smooth posterior margins of mesocoxal rests (sharply carinate in 
<taxonomicName id="54DD4D7CE41AFFEEC13CF8720AD1F80A" authorityName="Jaloszynski" authorityYear="2009" box="[1134,1212,1952,1977]" class="Insecta" family="Staphylinidae" genus="Trurlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="A1A9EAEDE41AFFEEC13CF8720AD1F80A" box="[1134,1212,1952,1977]" italics="true" pageId="2" pageNumber="291">Trurlia</emphasis>
</taxonomicName>
), and one vestigial basal elytral fovea (two distinct foveae in 
<taxonomicName id="54DD4D7CE41AFFEEC73FF8160CD1F86E" authorityName="Jaloszynski" authorityYear="2009" box="[621,700,1988,2013]" class="Insecta" family="Staphylinidae" genus="Trurlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="A1A9EAEDE41AFFEEC73FF8160CD1F86E" box="[621,700,1988,2013]" italics="true" pageId="2" pageNumber="291">Trurlia</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>