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<document id="15B9A37A973DBAE16A6EE2263BCA552E" ID="10.11646/zootaxa.4140.1.1" ID-DOI="10.11646/zootaxa.4140.1.1" ID-GBIF-Dataset="01995ca4-7d04-4402-b21a-926957b1a391" ID-ISSN="1175-5326" ID-Zenodo-Dep="256507" ID-ZooBank="AD1F6B4E-3B0D-422C-B47C-0495BD385857" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1469426945103" checkinUser="plazi" docAuthor="Nguyen, Tung T., Nguyen, Anh D., Tran, Binh T. T. &amp; Blakemore, Robert J." docDate="2016" docId="B5727931FFB57902FF3C23EF67538CBA" docLanguage="en" docName="zootaxa.4140.1.1.pdf" docOrigin="Zootaxa 4140 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Glyphidrilus Horst 1889" docType="treatment" docVersion="21" lastPageNumber="6" masterDocId="494B0149FFB17907FFAB254162078C42" masterDocTitle="A comprehensive checklist of earthworm species and subspecies from Vietnam (Annelida: Clitellata: Oligochaeta: Almidae, Eudrilidae, Glossoscolecidae, Lumbricidae, Megascolecidae, Moniligastridae, Ocnerodrilidae, Octochaetidae)" masterLastPageNumber="92" masterPageNumber="1" pageNumber="5" updateTime="1698365898991" updateUser="plazi">
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<mods:title id="D72808384A7FB950B5F1277376CD4ACF">A comprehensive checklist of earthworm species and subspecies from Vietnam (Annelida: Clitellata: Oligochaeta: Almidae, Eudrilidae, Glossoscolecidae, Lumbricidae, Megascolecidae, Moniligastridae, Ocnerodrilidae, Octochaetidae)</mods:title>
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<mods:namePart id="03742442F67FEA468DA129226E0478D4">Nguyen, Tung T.</mods:namePart>
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<mods:namePart id="674F9D2978E5336511BDC5A3314B855B">Nguyen, Anh D.</mods:namePart>
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<mods:namePart id="BE908B36AB5044F6FF92E9F03DA6EC10">Tran, Binh T. T.</mods:namePart>
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<mods:roleTerm id="321E9C6257C26B80615EE094D52BFBD5">Author</mods:roleTerm>
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<mods:namePart id="56AD435ECB62E227C195C89C1BE54892">Blakemore, Robert J.</mods:namePart>
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<mods:title id="76320232AFCE724F947A01EB9C5479D1">Zootaxa</mods:title>
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<mods:part id="40C5771B21F8E8F59EAEA2BA9EA51DB6">
<mods:date id="A9DC19C361F34F76B9A6702D7D29E9C4">2016</mods:date>
<mods:detail id="791334E270D705B3A895CE64D2A9768C" type="volume">
<mods:number id="093EF7D54A5C5B3141523CE02F5F1E73">4140</mods:number>
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<treatment id="B5727931FFB57902FF3C23EF67538CBA" ID-DOI="http://doi.org/10.5281/zenodo.3513821" ID-GBIF-Taxon="123207203" ID-Zenodo-Dep="3513821" LSID="urn:lsid:plazi:treatment:B5727931FFB57902FF3C23EF67538CBA" httpUri="http://treatment.plazi.org/id/B5727931FFB57902FF3C23EF67538CBA" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
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<heading id="662C7F4BFFB57903FF3C23EF60198A8A" bold="true" box="[151,542,1710,1736]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<emphasis id="0FAF1435FFB57903FF3C23EF60198A8A" bold="true" box="[151,542,1710,1736]" pageId="4" pageNumber="5">
Genus
<taxonomicName id="FADBB3A4FFB57903FF4523EF60198A8A" authority="Horst, 1889" authorityName="Horst" authorityYear="1889" box="[238,542,1710,1736]" class="Clitellata" family="Glossoscolecidae" genus="Glyphidrilus" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="5" phylum="Annelida" rank="genus">
<emphasis id="0FAF1435FFB57903FF4523EF63828A8A" bold="true" box="[238,389,1710,1736]" italics="true" pageId="4" pageNumber="5">Glyphidrilus</emphasis>
<bibRefCitation id="594AB5D6FFB57903FE2723EF60198A8A" author="Horst" box="[396,542,1710,1736]" pageId="4" pageNumber="5" refString="Horst, R. (1889) Over eene nieuwe soort order de Lumbricinen door Prof. Max Weber uit nedenl. Indie medegebracht. Tijdschrift der Nederlandsche Dierkundige Vereeniging, 2, 1 - 77." type="journal article" year="1889">Horst, 1889</bibRefCitation>
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<paragraph id="3D64C827FFB57903FF3C23B460348B68" blockId="4.[151,1436,1781,1895]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="1DCBEF09FFB57903FF3C23B460348B68" pageId="4" pageNumber="5">
<taxonomicName id="FADBB3A4FFB57903FF3C23B463C08B49" authority="Horst, 1889: 76" authorityName="Horst" authorityPageNumber="76" authorityYear="1889" box="[151,455,1781,1803]" class="Clitellata" family="Glossoscolecidae" genus="Glyphidrilus" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="5" phylum="Annelida" rank="genus">
<emphasis id="0FAF1435FFB57903FF3C23B463108B49" box="[151,279,1781,1803]" italics="true" pageId="4" pageNumber="5">Glyphidrilus</emphasis>
<treatmentCitation id="BC7AEE36FFB57903FE8B23B763C08B49" author="Horst" box="[288,455,1781,1803]" page="76" pageId="4" pageNumber="5" year="1889">
<bibRefCitation id="594AB5D6FFB57903FE8B23B763998B49" author="Horst" box="[288,414,1781,1803]" pageId="4" pageNumber="5" refString="Horst, R. (1889) Over eene nieuwe soort order de Lumbricinen door Prof. Max Weber uit nedenl. Indie medegebracht. Tijdschrift der Nederlandsche Dierkundige Vereeniging, 2, 1 - 77." type="journal article" year="1889">Horst, 1889</bibRefCitation>
: 76
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="BC7AEE36FFB57903FE7E23B760858B49" author="Horst" box="[469,642,1781,1803]" page="37" pageId="4" pageNumber="5" year="1893">
<bibRefCitation id="594AB5D6FFB57903FE7E23B7605D8B49" author="Horst" box="[469,602,1781,1803]" pageId="4" pageNumber="5" refString="Horst, R. (1893 b) Earthworms from Malay Archipelago. In Max Weber. Zoologische Ergebnisse einer Reise in Niederlandisch Ost Indien, 3, 28 - 83." type="journal article" year="1893" yearSuffix="b">Horst 1893b</bibRefCitation>
: 37
</treatmentCitation>
;
<treatmentCitation id="BC7AEE36FFB57903FD3B23B461618B49" author="Beddard" box="[656,870,1781,1803]" page="679" pageId="4" pageNumber="5" year="1895">
<bibRefCitation id="594AB5D6FFB57903FD3B23B461368B49" author="Beddard" box="[656,817,1781,1803]" pageId="4" pageNumber="5" refString="Beddard, F. E. (1895 a) A monograph of the Order Oligochaeta. The Clarendon Press, Oxford, 769 pp. http: // dx. doi. org / 10.5962 / bhl. title. 28557" type="book" year="1895" yearSuffix="a">Beddard 1895a</bibRefCitation>
: 679
</treatmentCitation>
;
<treatmentCitation id="BC7AEE36FFB57903FCDF23B466598B49" author="Michaelsen" box="[884,1118,1781,1803]" page="459" pageId="4" pageNumber="5" year="1900">
<bibRefCitation id="594AB5D6FFB57903FCDF23B4662F8B49" author="Michaelsen" box="[884,1064,1781,1803]" pageId="4" pageNumber="5" refString="Michaelsen, W. (1900) Oligochaeta. Das Tierreich, 10, 1 - 575." type="journal article" year="1900">Michaelsen 1900</bibRefCitation>
: 459
</treatmentCitation>
;
<treatmentCitation id="BC7AEE36FFB57903FBC023B467528B49" author="Stephenson" box="[1131,1365,1781,1803]" page="901" pageId="4" pageNumber="5" year="1930">
<bibRefCitation id="594AB5D6FFB57903FBC023B467188B49" author="Stephenson" box="[1131,1311,1781,1803]" pageId="4" pageNumber="5" refString="Stephenson, J. (1930) The Oligochaeta. Clarendon Press, Oxford, 978 pp." type="book" year="1930">Stephenson 1930</bibRefCitation>
: 901
</treatmentCitation>
;
<treatmentCitation id="BC7AEE36FFB57903FAC923B463378B68" author="Gates" page="234" pageId="4" pageNumber="5" year="1972">
<bibRefCitation id="594AB5D6FFB57903FAC923B462FA8B68" author="Gates" pageId="4" pageNumber="5" refString="Gates, G. E. (1972) Burmese Earthworms: An Introduction to the Systematics and Biology of Megadrile Oligochaetes with Special Reference to Southeast Asia. Transactions of the American Philosophical Society, N. S. 62 (7), 1 - 326. http: // dx. doi. org / 10.2307 / 1006214" type="journal article" year="1972">Gates 1972</bibRefCitation>
: 234
</treatmentCitation>
;
<treatmentCitation id="BC7AEE36FFB57903FE91225560288B68" ID-DOI="http://doi.org/10.5281/zenodo.3507051" ID-Zenodo-Dep="3507051" author="Chanabun" box="[314,559,1812,1834]" httpUri="http://treatment.plazi.org/id/ADA54A271EA35569EBDA5B3697DC61D4" page="9" pageId="4" pageNumber="5" year="2013">
<bibRefCitation id="594AB5D6FFB57903FE91225560118B68" author="Chanabun" box="[314,534,1812,1834]" pageId="4" pageNumber="5" refString="Chanabun, R., Sutcharit, C., Tongkerd, P. &amp; Panha, S. (2013) The semi-aquatic freshwater earthworms of the genus Glyphidrilus Horst, 1889 from Thailand (Oligochaeta, Almidae) with re - descriptions of several species. ZooKeys, 265, 1 - 76. http: // dx. doi. org / 10.3897 / zookeys. 265.3911" type="journal article" year="2013">
Chanabun
<emphasis id="0FAF1435FFB57903FE0C225463DD8B68" box="[423,474,1812,1834]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
2013
</bibRefCitation>
: 9
</treatmentCitation>
.
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</paragraph>
<paragraph id="3D64C827FFB57903FF3C227360738B0A" blockId="4.[151,1436,1781,1895]" box="[151,628,1842,1864]" pageId="4" pageNumber="5">
<emphasis id="0FAF1435FFB57903FF3C227362E08B0A" box="[151,231,1842,1864]" italics="true" pageId="4" pageNumber="5">Bilimba</emphasis>
Rosa, 1890: 386;
<bibRefCitation id="594AB5D6FFB57903FE0B2273603A8B0A" author="Beddard" box="[416,573,1842,1864]" pageId="4" pageNumber="5" refString="Beddard, F. E. (1895 a) A monograph of the Order Oligochaeta. The Clarendon Press, Oxford, 769 pp. http: // dx. doi. org / 10.5962 / bhl. title. 28557" type="book" year="1895" yearSuffix="a">Beddard 1895a</bibRefCitation>
: 686.
</paragraph>
</subSubSection>
<subSubSection id="75C19BACFFB57903FF3C2210609D8B25" box="[151,666,1873,1895]" pageId="4" pageNumber="5" type="description">
<paragraph id="3D64C827FFB57903FF3C2210609D8B25" blockId="4.[151,1436,1781,1895]" box="[151,666,1873,1895]" pageId="4" pageNumber="5">
<emphasis id="0FAF1435FFB57903FF3C2210630F8B25" box="[151,264,1873,1895]" italics="true" pageId="4" pageNumber="5">Annadrilus</emphasis>
<bibRefCitation id="594AB5D6FFB57903FEA5221363918B25" author="Horst" box="[270,406,1873,1895]" pageId="4" pageNumber="5" refString="Horst, R. (1893 b) Earthworms from Malay Archipelago. In Max Weber. Zoologische Ergebnisse einer Reise in Niederlandisch Ost Indien, 3, 28 - 83." type="journal article" year="1893" yearSuffix="b">Horst, 1893b</bibRefCitation>
: 44;
<bibRefCitation id="594AB5D6FFB57903FE6E221060648B25" author="Beddard" box="[453,611,1873,1895]" pageId="4" pageNumber="5" refString="Beddard, F. E. (1895 a) A monograph of the Order Oligochaeta. The Clarendon Press, Oxford, 769 pp. http: // dx. doi. org / 10.5962 / bhl. title. 28557" type="book" year="1895" yearSuffix="a">Beddard 1895a</bibRefCitation>
: 680.
</paragraph>
</subSubSection>
<subSubSection id="75C19BACFFB57903FF3C22D560AB8BEE" box="[151,684,1940,1965]" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="3D64C827FFB57903FF3C22D560AB8BEE" blockId="4.[151,1436,1940,2037]" box="[151,684,1940,1965]" pageId="4" pageNumber="5">
<emphasis id="0FAF1435FFB57903FF3C22D563358BEF" bold="true" box="[151,306,1940,1965]" pageId="4" pageNumber="5">Type species.</emphasis>
<taxonomicName id="FADBB3A4FFB57903FE9222D560AF8BEE" authority="Horst, 1889" authorityName="Horst" authorityYear="1889" box="[313,680,1940,1965]" class="Clitellata" family="Glossoscolecidae" genus="Glyphidrilus" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="5" phylum="Annelida" rank="species" species="weberi">
<emphasis id="0FAF1435FFB57903FE9222D5601E8BEF" box="[313,537,1940,1965]" italics="true" pageId="4" pageNumber="5">Glyphidrilus weberi</emphasis>
<bibRefCitation id="594AB5D6FFB57903FD8A22D460AF8BEE" author="Horst" box="[545,680,1940,1964]" pageId="4" pageNumber="5" refString="Horst, R. (1889) Over eene nieuwe soort order de Lumbricinen door Prof. Max Weber uit nedenl. Indie medegebracht. Tijdschrift der Nederlandsche Dierkundige Vereeniging, 2, 1 - 77." type="journal article" year="1889">Horst, 1889</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="75C19BACFFB57903FF6C22F967958BB6" pageId="4" pageNumber="5" type="distribution">
<paragraph id="3D64C827FFB57903FF6C22F967958BB6" blockId="4.[151,1436,1940,2037]" pageId="4" pageNumber="5">
<emphasis id="0FAF1435FFB57903FF6C22F963678B93" bold="true" box="[199,352,1976,2001]" pageId="4" pageNumber="5">Distribution.</emphasis>
<collectingCountry id="45CC88B7FFB57903FEC622F963D28B93" box="[365,469,1976,2001]" name="Tanzania" pageId="4" pageNumber="5">Tanzania</collectingCountry>
,
<collectingCountry id="45CC88B7FFB57903FE4C22F960598B93" box="[487,606,1976,2001]" name="Sri Lanka" pageId="4" pageNumber="5">
<collectingRegion id="FF1F06C5FFB57903FE4C22F9600F8B93" box="[487,520,1976,2001]" country="United Kingdom" name="Surrey" pageId="4" pageNumber="5">Sri</collectingRegion>
Lanka
</collectingCountry>
,
<collectingCountry id="45CC88B7FFB57903FDDB22F960AB8B93" box="[624,684,1976,2001]" name="India" pageId="4" pageNumber="5">India</collectingCountry>
,
<collectingCountry id="45CC88B7FFB57903FD1522F8612A8B93" box="[702,813,1977,2001]" name="Myanmar" pageId="4" pageNumber="5">Myanmar</collectingCountry>
, Malay peninsula,
<collectingCountry id="45CC88B7FFB57903FBB822F966878B93" box="[1043,1152,1976,2001]" name="Indonesia" pageId="4" pageNumber="5">Indonesia</collectingCountry>
(
<collectingRegion id="FF1F06C5FFB57903FB3E22F966F08B92" box="[1173,1271,1976,2000]" country="Indonesia" name="Sumatera Utara" pageId="4" pageNumber="5">Sumatra</collectingRegion>
,
<collectingRegion id="FF1F06C5FFB57903FAA222F8673A8B92" box="[1289,1341,1977,2000]" country="Indonesia" name="Jawa Timur" pageId="4" pageNumber="5">Java</collectingRegion>
,
<collectingCountry id="45CC88B7FFB57903FAE422F9679F8B93" box="[1359,1432,1976,2001]" name="Indonesia" pageId="4" pageNumber="5">Flores</collectingCountry>
, Borneo,
<collectingRegion id="FF1F06C5FFB57903FF5C229D63528BB7" box="[247,341,2012,2037]" country="Indonesia" name="Sulawesi Utara" pageId="4" pageNumber="5">Celebes</collectingRegion>
);
<collectingCountry id="45CC88B7FFB57903FEC3229D63AC8BB7" box="[360,427,2012,2037]" name="China" pageId="4" pageNumber="5">China</collectingCountry>
(
<collectingRegion id="FF1F06C5FFB57903FE11229D60168BB7" box="[442,529,2012,2037]" country="China" name="Hainan" pageId="4" pageNumber="5">Hainan</collectingRegion>
),
<collectingCountry id="45CC88B7FFB57903FD88229D608E8BB7" box="[547,649,2012,2037]" name="Thailand" pageId="4" pageNumber="5">Thailand</collectingCountry>
,
<collectingCountry id="45CC88B7FFB57903FD3E229D610C8BB7" box="[661,779,2012,2037]" name="Cambodia" pageId="4" pageNumber="5">Cambodia</collectingCountry>
,
<collectingCountry id="45CC88B7FFB57903FCBD229C61488BB6" box="[790,847,2013,2036]" name="Laos" pageId="4" pageNumber="5">Laos</collectingCountry>
,
<collectingCountry id="45CC88B7FFB57903FCF1229D61BF8BB7" box="[858,952,2012,2037]" name="Vietnam" pageId="4" pageNumber="5">Vietnam</collectingCountry>
(
<bibRefCitation id="594AB5D6FFB57903FC62229D66818BB6" author="Blakemore" box="[969,1158,2012,2037]" pageId="4" pageNumber="5" refString="Blakemore, R. J. (2002) Cosmopolitan Earthworms - an Eco - Taxonomic Guide to the Peregrine Species of the World. 2 nd Ed. VermEcology, PO BOX 414 Kippax, ACT 2615, Australia. August, 2002." type="journal article" year="2002">Blakemore 2002</bibRefCitation>
;
<bibRefCitation id="594AB5D6FFB57903FB39229D67808BB6" author="Chanabun" box="[1170,1415,2012,2037]" pageId="4" pageNumber="5" refString="Chanabun, R., Sutcharit, C., Tongkerd, P. &amp; Panha, S. (2013) The semi-aquatic freshwater earthworms of the genus Glyphidrilus Horst, 1889 from Thailand (Oligochaeta, Almidae) with re - descriptions of several species. ZooKeys, 265, 1 - 76. http: // dx. doi. org / 10.3897 / zookeys. 265.3911" type="journal article" year="2013">
Chanabun
<emphasis id="0FAF1435FFB57903FAA0229C67448BB7" box="[1291,1347,2012,2037]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
2013
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="75C19BACFFB47902FF6C25D667538CBA" pageId="5" pageNumber="6" type="discussion">
<paragraph id="3D64C827FFB47902FF6C25D667538CBA" blockId="5.[151,1436,151,248]" pageId="5" pageNumber="6">
<materialsCitation id="8DB3C27AFFB47902FF6C25D667538CBA" ID-GBIF-Occurrence="1288043694" country="Vietnam" location="A" pageId="5" pageNumber="6" specimenCount="1" stateProvince="A">
<emphasis id="0FAF1435FFB47902FF6C25D6633C8CF2" bold="true" box="[199,315,151,176]" pageId="5" pageNumber="6">Remarks.</emphasis>
<collectingRegion id="FF1F06C5FFB47902FEE825D963508CF2" box="[323,343,152,176]" country="Nigeria" name="Rivers" pageId="5" pageNumber="6">A</collectingRegion>
total of 3 6 species have been described (
<bibRefCitation id="594AB5D6FFB47902FC8325D661BD8CF2" author="Csuzdi" box="[808,954,151,176]" pageId="5" pageNumber="6" refString="Csuzdi, Cs. (2012) Earthworm species, a searchable database. Opuscula Zoologica, 43 (1), 97 - 99. Avaliable from: http: // earthworm. uw. hu / (Accessed 21 Jul. 2016)" type="journal article" year="2012">Csuzdi 2012</bibRefCitation>
;
<bibRefCitation id="594AB5D6FFB47902FC6C25D666BA8CF2" author="Chanabun" box="[967,1213,151,176]" pageId="5" pageNumber="6" refString="Chanabun, R., Sutcharit, C., Tongkerd, P. &amp; Panha, S. (2013) The semi-aquatic freshwater earthworms of the genus Glyphidrilus Horst, 1889 from Thailand (Oligochaeta, Almidae) with re - descriptions of several species. ZooKeys, 265, 1 - 76. http: // dx. doi. org / 10.3897 / zookeys. 265.3911" type="journal article" year="2013">
Chanabun
<emphasis id="0FAF1435FFB47902FBEA25D8667D8CF2" box="[1089,1146,151,176]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
2013
</bibRefCitation>
); of these, only one species is known from
<collectingCountry id="45CC88B7FFB47902FE3C25FD63FD8C97" box="[407,506,188,213]" name="Vietnam" pageId="5" pageNumber="6">Vietnam</collectingCountry>
. Records of a second species (
<taxonomicName id="FADBB3A4FFB47902FCE725FD66F68C97" authority="Gates, 1945" authorityName="Gates" authorityYear="1945" box="[844,1265,188,213]" class="Clitellata" family="Glossoscolecidae" genus="Glyphidrilus" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="6" phylum="Annelida" rank="species" species="ceylonensis">
<emphasis id="0FAF1435FFB47902FCE725FD66658C97" box="[844,1122,188,213]" italics="true" pageId="5" pageNumber="6">Glyphidrilus ceylonensis</emphasis>
Gates, 1945
</taxonomicName>
), in
<bibRefCitation id="594AB5D6FFB47902FA8A25FD62D88CBA" author="Blakemore" pageId="5" pageNumber="6" refString="Blakemore, R. J. (2008 b) List of earthworms from Cambodia, Laos, and Vietnam (Excel spreadsheet). In: A Series of Searchable Texts on Earthworm Biodiversity, Ecology and Systematics from Various Regions of the World - 2 nd Edition (2008). VermEcology, Yokohama. CD - ROM. Avaliable from: http: // www. annelida. net / earthworm / Vietnam % 20 + % 20 Lao s % 20 + % 20 Cambodia 1. xls (Accessed 21 Jul. 2016)" type="journal article" year="2008" yearSuffix="b">Blakemore 2008b</bibRefCitation>
, based on http://www.mscwbif.org/earthworm/g.php, (currently unaccessible) are not confirmed here.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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