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<document id="8BF997226E64AAB7080B793E6542D49A" ID-CLB-Dataset="24370" ID-DOI="10.11646/zootaxa.4722.2.6" ID-GBIF-Dataset="ae7c5192-65dd-4ee9-839f-96b7021b42a0" ID-ISSN="1175-5326" ID-Zenodo-Dep="3605861" ID-ZooBank="A2AAACA1-41AB-4DF6-9049-6A5D3B9D45C3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1578905277601" checkinUser="plazi" docAuthor="Dantas, Galileu P. S., Huamantinco Araujo, Ana A. &amp; Hamada, Neusa" docDate="2020" docId="03A087B4263B9F46FF07E34D5340107E" docLanguage="en" docName="zootaxa.4722.2.6.pdf" docOrigin="Zootaxa 4722 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rheotanytarsus amaru Dantas &amp; Huamantinco Araujo &amp; Hamada 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="198" masterDocId="FF99FFCC263A9F45FF90E1055063121D" masterDocTitle="A new species of Rheotanytarsus Thienemann &amp; Bause (Diptera: Chironomidae) from Peruvian Andes, with updated key to South American species" masterLastPageNumber="200" masterPageNumber="195" pageNumber="196" updateTime="1698777364426" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="F1A3E98D2B79F7BE6BAFAC8FA17B28F7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="55A0E0704156A18C88678F52B95C2719">
<mods:title id="C1DAC0A867FF8EE142B2D471608AFABA">A new species of Rheotanytarsus Thienemann &amp; Bause (Diptera: Chironomidae) from Peruvian Andes, with updated key to South American species</mods:title>
</mods:titleInfo>
<mods:name id="C8F6BE16E21B0C897D755651F59FAAE8" type="personal">
<mods:role id="405C9C4ADF610F67F450CA0ECCBB9C13">
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<mods:namePart id="DB5CE23DD61469ECB5893184F4200558">Dantas, Galileu P. S.</mods:namePart>
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<mods:namePart id="FEC854E0B10ED7E57658E981C658F148">Huamantinco Araujo, Ana A.</mods:namePart>
</mods:name>
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<mods:role id="02FF90F7C6034EF44B3518354EA749EF">
<mods:roleTerm id="857C325DB2A9C86E9BA7971C5320786C">Author</mods:roleTerm>
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<mods:namePart id="32B74D9B9FDC263E96E2E23B3A230F80">Hamada, Neusa</mods:namePart>
</mods:name>
<mods:typeOfResource id="A817673B23F6236752E230390C6C5C6A">text</mods:typeOfResource>
<mods:relatedItem id="5BD10CB0E1744FC749A0DAFEC9366CAE" type="host">
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<mods:title id="935BB8659AC8809BA6FC89DE7065AF0E">Zootaxa</mods:title>
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<mods:part id="98B19BA2188065B1E89FAAB2B865339E">
<mods:date id="0C2DA6D76F6A1CE3776D798B327B3C1D">2020</mods:date>
<mods:detail id="CACA287ACDA4C4D6D846FB43F1F595CE" type="pubDate">
<mods:number id="0E590FFA7BE4A9F213FF19E63500065A">2020-01-13</mods:number>
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<mods:number id="EC710EC59F120724F9181D6883CBD398">4722</mods:number>
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<mods:number id="13A086A50C71EEAFCD493B0B38DFE565">2</mods:number>
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<mods:start id="9396862E4C5E7F4D093C7DB97D68D499">195</mods:start>
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<mods:classification id="BB4900180F4CCFD3ADB18A3918FEB10D">journal article</mods:classification>
<mods:identifier id="36F4AACC49CA3D76CE39987699458ADF" type="CLB-Dataset">24370</mods:identifier>
<mods:identifier id="A78AF37778EFCDA7B48E84AC840BB673" type="DOI">10.11646/zootaxa.4722.2.6</mods:identifier>
<mods:identifier id="FF619EB8E4220E786B80DF1F7BFBAC88" type="GBIF-Dataset">ae7c5192-65dd-4ee9-839f-96b7021b42a0</mods:identifier>
<mods:identifier id="E06C5C3DC857E7192A067E7FB2752750" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
<treatment id="03A087B4263B9F46FF07E34D5340107E" ID-GBIF-Taxon="161672188" LSID="urn:lsid:plazi:treatment:03A087B4263B9F46FF07E34D5340107E" httpUri="http://treatment.plazi.org/id/03A087B4263B9F46FF07E34D5340107E" lastPageId="3" lastPageNumber="198" pageId="1" pageNumber="196">
<subSubSection id="C3136529263B9F44FF07E34D519C107E" box="[151,511,584,611]" pageId="1" pageNumber="196" type="nomenclature">
<paragraph id="8BB636A2263B9F44FF07E34D519C107E" blockId="1.[151,982,584,683]" box="[151,511,584,611]" pageId="1" pageNumber="196">
<heading id="D0FE81CE263B9F44FF07E34D519C107E" bold="true" box="[151,511,584,611]" fontSize="11" level="1" pageId="1" pageNumber="196" reason="1">
<emphasis id="B97DEAB0263B9F44FF07E34D519C107E" bold="true" box="[151,511,584,611]" pageId="1" pageNumber="196">
<taxonomicName id="4C094D21263B9F44FF07E34D51FC107E" ID-CoL="9HW4D" authority="Dantas &amp; Huamantinco Araujo &amp; Hamada, 2020" authorityName="Dantas &amp; Huamantinco Araujo &amp; Hamada" authorityYear="2020" box="[151,415,584,611]" class="Insecta" family="Chironomidae" genus="Rheotanytarsus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="species" species="amaru" status="sp. nov.">
<emphasis id="B97DEAB0263B9F44FF07E34D51FC107E" bold="true" box="[151,415,584,611]" italics="true" pageId="1" pageNumber="196">Rheotanytarsus amaru</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CB263B9F44FE36E34C519C107E" box="[422,511,585,611]" pageId="1" pageNumber="196" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3136529263B9F44FF07E36953B510B6" pageId="1" pageNumber="196" type="description">
<paragraph id="8BB636A2263B9F44FF07E369517D109A" blockId="1.[151,982,584,683]" box="[151,286,620,647]" pageId="1" pageNumber="196">
(
<figureCitation id="13322A27263B9F44FF30E36950BE109A" box="[160,221,620,647]" captionStart="FIGURE 1" captionStartId="2.[151,250,1977,2002]" captionTargetBox="[186,1401,166,1958]" captionTargetId="figure@2.[185,1403,166,1958]" captionTargetPageId="2" captionText="FIGURE 1. Rheotanytarsus amaru sp. n., Adult male:A. Head, in frontal view. B. Thorax, in lateral view. C. Wing. D. Hypopygium, in dorsal view. E. Hypopygium in dorsal view, with tergite IX removed." figureDoi="http://doi.org/10.5281/zenodo.3605867" httpUri="https://zenodo.org/record/3605867/files/figure.png" pageId="1" pageNumber="196">Fig. 1</figureCitation>
A–E)
</paragraph>
<paragraph id="8BB636A2263B9F44FF07E39553B510B6" blockId="1.[151,982,584,683]" box="[151,982,656,683]" pageId="1" pageNumber="196">
<uri id="FF983AA0263B9F44FF07E39553B510B6" box="[151,982,656,683]" pageId="1" pageNumber="196">
urn:lsid:zoobank.org:act:
<uuid id="FFAF0C77263B9F44FE3BE39553B510B6" box="[427,982,656,683]" pageId="1" pageNumber="196">C0635FB5-CCC0-4D08-8ED7-26A7D67DE29E</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3136529263B9F44FF07E3DC55501126" pageId="1" pageNumber="196" type="materials_examined">
<paragraph id="8BB636A2263B9F44FF07E3DC55501126" blockId="1.[151,1437,728,1763]" pageId="1" pageNumber="196">
<emphasis id="B97DEAB0263B9F44FF07E3DC512410EE" bold="true" box="[151,327,728,755]" pageId="1" pageNumber="196">Type material:</emphasis>
<materialsCitation id="3B613CFF263B9F44FEDEE3DD51CC1126" ID-GBIF-Occurrence="2549107397" collectingDate="2014-08-17" collectionCode="R, MHN-UNMSM" collectorName="N. Hamada &amp; Hamada, M &amp; L. Ferreira-Keppler &amp; J. M. C. Nascimento" country="Peru" county="Calca province" latitude="-13.355556" location="Urubamba river" longLatPrecision="21" longitude="-71.935555" municipality="Lamay district" pageId="1" pageNumber="196" specimenCount="1" stateProvince="Cusco" typeStatus="holotype">
<typeStatus id="54B28800263B9F44FEDEE3DD51DA10EF" box="[334,441,728,754]" pageId="1" pageNumber="196" type="holotype">Holotype</typeStatus>
, adult male:
<emphasis id="B97DEAB0263B9F44FDD9E3DC52F610EE" bold="true" box="[585,661,729,755]" pageId="1" pageNumber="196">
<collectingCountry id="F31E7632263B9F44FDD9E3DC52F610EE" box="[585,661,729,755]" name="Peru" pageId="1" pageNumber="196">PERU</collectingCountry>
</emphasis>
,
<collectingRegion id="49CDF840263B9F44FD33E3DC528A10EE" box="[675,745,729,755]" country="Peru" name="Cusco" pageId="1" pageNumber="196">Cusco</collectingRegion>
department,
<collectingCounty id="62D74E2E263B9F44FCEDE3DD544B10EF" box="[893,1064,728,754]" pageId="1" pageNumber="196">Calca province</collectingCounty>
,
<collectingMunicipality id="6BD2ACD8263B9F44FBA3E3DC54BA10EF" box="[1075,1241,728,755]" pageId="1" pageNumber="196">Lamay district</collectingMunicipality>
,
<location id="8ED66079263B9F44FB74E3DD55FB10EF" LSID="urn:lsid:plazi:treatment:03A087B4263B9F46FF07E34D5340107E:8ED66079263B9F44FB74E3DD55FB10EF" box="[1252,1432,728,754]" country="Peru" county="Calca province" latitude="-13.355556" longLatPrecision="21" longitude="-71.935555" municipality="Lamay district" name="Urubamba river" pageId="1" pageNumber="196" stateProvince="Cusco">Urubamba river</location>
,
<geoCoordinate id="EE3D5065263B9F44FF07E3F85145110A" box="[151,294,765,791]" degrees="13" direction="south" minutes="21" orientation="latitude" pageId="1" pageNumber="196" precision="15" seconds="20" value="-13.355556">13°21’20” S</geoCoordinate>
,
<geoCoordinate id="EE3D5065263B9F44FEA5E3F951AD110A" box="[309,462,764,791]" degrees="71" direction="west" minutes="56" orientation="longitude" pageId="1" pageNumber="196" precision="15" seconds="08" value="-71.935555">71°56’08” W</geoCoordinate>
,
<date id="FFB71062263B9F44FE4BE3F95205110B" box="[475,614,764,790]" pageId="1" pageNumber="196" value="2014-08-17">
<collectingDate id="EFF3E98A263B9F44FE4BE3F95205110B" box="[475,614,764,790]" pageId="1" pageNumber="196" value="2014-08-17">17.viii.2014</collectingDate>
</date>
, leg.
<collectorName id="26FC5374263B9F44FD36E3F85349110B" box="[678,810,764,791]" pageId="1" pageNumber="196">N. Hamada</collectorName>
, M. Bobadilla,
<collectionCode id="ED18AE67263B9F44FC4FE3F85396110A" box="[991,1013,765,791]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="196">R</collectionCode>
.
<collectorName id="26FC5374263B9F44FC68E3F854B5110B" box="[1016,1238,764,791]" pageId="1" pageNumber="196">L. Ferreira-Keppler</collectorName>
&amp;
<collectorName id="26FC5374263B9F44FB6EE3F850BE1126" pageId="1" pageNumber="196">J.M.C. Nascimento</collectorName>
(
<collectionCode id="ED18AE67263B9F44FF7CE22451CB1126" box="[236,424,801,827]" pageId="1" pageNumber="196">MHN-UNMSM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B613CFF263B9F44FE29E22453881126" ID-GBIF-Occurrence="2549107398" box="[441,1003,800,827]" collectionCode="MHN-UNMSM" country="Peru" pageId="1" pageNumber="196" specimenCount="6" specimenCount-male="6" stateProvince="Cusco" typeStatus="paratype">
<typeStatus id="54B28800263B9F44FE29E224524A1126" box="[441,553,801,827]" pageId="1" pageNumber="196" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="9D0FFD2B263B9F44FDA4E22552EE1127" box="[564,653,800,826]" pageId="1" pageNumber="196" type="male">6 males</specimenCount>
, as
<typeStatus id="54B28800263B9F44FD27E225537A1127" box="[695,793,800,826]" pageId="1" pageNumber="196" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED18AE67263B9F44FCB7E22453801126" box="[807,995,801,827]" pageId="1" pageNumber="196">MHN-UNMSM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B613CFF263B9F44FC66E22455531126" ID-GBIF-Occurrence="2549107399" box="[1014,1328,800,827]" collectionCode="INPA" country="Peru" pageId="1" pageNumber="196" specimenCount="3" specimenCount-male="3" stateProvince="Cusco" typeStatus="holotype">
<specimenCount id="9D0FFD2B263B9F44FC66E224542F1127" box="[1014,1100,800,827]" pageId="1" pageNumber="196" type="male">3 males</specimenCount>
as
<typeStatus id="54B28800263B9F44FBE1E22554B01127" box="[1137,1235,800,826]" pageId="1" pageNumber="196" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED18AE67263B9F44FB72E224554B1126" box="[1250,1320,801,827]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="1" pageNumber="196">INPA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3136529263B9F44FF57E24052F311D7" pageId="1" pageNumber="196" type="etymology">
<paragraph id="8BB636A2263B9F44FF57E24052F311D7" blockId="1.[151,1437,728,1763]" pageId="1" pageNumber="196">
<emphasis id="B97DEAB0263B9F44FF57E240512D1142" bold="true" box="[199,334,837,863]" pageId="1" pageNumber="196">Etymology.</emphasis>
In Inca mythology, “Amaru” (quechua) is a huge double-headed serpent that dwells underground, at the bottom of lakes and rivers. The Inca Empire was the largest in pre-Columbian America, the administrative, political and military center of the empire was in the city of
<collectingRegion id="49CDF840263B9F44FCD5E28853ED11BA" box="[837,910,909,935]" country="Peru" name="Cusco" pageId="1" pageNumber="196">Cusco</collectingRegion>
, near the
<typeStatus id="54B28800263B9F44FC6DE288544E11BA" box="[1021,1069,909,935]" pageId="1" pageNumber="196">type</typeStatus>
locality of the new species. The name is to be treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C3136529263B9F44FF57E2D153261646" pageId="1" pageNumber="196" type="diagnosis">
<paragraph id="8BB636A2263B9F44FF57E2D153261646" blockId="1.[151,1437,728,1763]" pageId="1" pageNumber="196">
<emphasis id="B97DEAB0263B9F44FF57E2D1512111F2" bold="true" box="[199,322,980,1007]" pageId="1" pageNumber="196">Diagnosis.</emphasis>
Wing length&gt;
<quantity id="4CF19B47263B9F44FE61E2D0525611F2" box="[497,565,981,1007]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="196" unit="mm" value="2.0">2 mm</quantity>
; spurs on mid and hind tibiae present; anal tergite bands V-shaped; anal point parallel-sided; superior volsella oval, apically tapered; stem of median volsella curved, swollen and rounded at apex, bearing a brush of setiform lamellae not fused into a plate; median and superior volsella about the same size; gonostylus broadest subbasally, evenly tapering toward apex.
</paragraph>
</subSubSection>
<subSubSection id="C3136529263B9F46FF57E561523E13EA" lastPageId="3" lastPageNumber="198" pageId="1" pageNumber="196" type="description">
<paragraph id="8BB636A2263B9F44FF57E56151381662" blockId="1.[151,1437,728,1763]" box="[199,347,1124,1151]" pageId="1" pageNumber="196">
<emphasis id="B97DEAB0263B9F44FF57E56151381662" bold="true" box="[199,347,1124,1151]" pageId="1" pageNumber="196">Description:</emphasis>
</paragraph>
<paragraph id="8BB636A2263B9F44FF57E58D53B116DA" blockId="1.[151,1437,728,1763]" pageId="1" pageNumber="196">
<emphasis id="B97DEAB0263B9F44FF57E58D512F16BE" bold="true" box="[199,332,1160,1187]" pageId="1" pageNumber="196">Adult male</emphasis>
(n = 10). Body size and proportions. Total length
<quantity id="4CF19B47263B9F44FCE6E58D547F16BE" box="[886,1052,1160,1187]" metricMagnitude="-3" metricUnit="m" metricValue="3.485" metricValueMax="3.52" metricValueMin="3.45" pageId="1" pageNumber="196" unit="mm" value="3.485" valueMax="3.52" valueMin="3.45">3.45–3.52 mm</quantity>
. Wing length
<quantity id="4CF19B47263B9F44FB25E58C553916BE" box="[1205,1370,1161,1187]" metricMagnitude="-3" metricUnit="m" metricValue="2.075" metricValueMax="2.11" metricValueMin="2.04" pageId="1" pageNumber="196" unit="mm" value="2.075" valueMax="2.11" valueMin="2.04">2.04–2.11 mm</quantity>
. Total length/wing length 1.67–1.70. Wing length/length of profemur 2.33–2.38.
</paragraph>
<paragraph id="8BB636A2263B9F44FF57E5D554DA1713" blockId="1.[151,1437,728,1763]" pageId="1" pageNumber="196">Coloration. Eyes black. Head capsule and pedicel dark brown; antennal flagellum and palp brown. Thorax dark brown. All legs brown. Wing membrane with pale brownish undertone. Abdomen light brown.</paragraph>
<paragraph id="8BB636A2263B9F44FF57E41D535317DE" blockId="1.[151,1437,728,1763]" pageId="1" pageNumber="196">
Head (
<figureCitation id="13322A27263B9F44FE83E41D510E172E" box="[275,365,1304,1331]" captionStart="FIGURE 1" captionStartId="2.[151,250,1977,2002]" captionTargetBox="[186,1401,166,1958]" captionTargetId="figure@2.[185,1403,166,1958]" captionTargetPageId="2" captionText="FIGURE 1. Rheotanytarsus amaru sp. n., Adult male:A. Head, in frontal view. B. Thorax, in lateral view. C. Wing. D. Hypopygium, in dorsal view. E. Hypopygium in dorsal view, with tergite IX removed." figureDoi="http://doi.org/10.5281/zenodo.3605867" httpUri="https://zenodo.org/record/3605867/files/figure.png" pageId="1" pageNumber="196">Fig. 1A</figureCitation>
). Eyes bare, with well-developed dorsomedian extensions. Antenna with 13 flagellomeres; ultimate flagellomere 485–492 μm long; AR 0.97–0.0.99. Frontal tubercles absent. Tentorium 150–160 μm long, heavily sclerotized. Temporal setae 8–11 on each side. Clypeus with 24–28 setae. Lengths of palpomeres 1–5 (in μm): 43–55, 42–52, 130–141, 130–132, 215–232; third palpomere with 1 sensillum clavatum subapically, 14–17 μm long. Fifth palpomere/ third palpomere ratio 1.65–1.70.
</paragraph>
<paragraph id="8BB636A2263B9F44FF57E4C954081416" blockId="1.[151,1437,728,1763]" pageId="1" pageNumber="196">
Thorax (
<figureCitation id="13322A27263B9F44FEB8E4C9511C17FA" box="[296,383,1484,1511]" captionStart="FIGURE 1" captionStartId="2.[151,250,1977,2002]" captionTargetBox="[186,1401,166,1958]" captionTargetId="figure@2.[185,1403,166,1958]" captionTargetPageId="2" captionText="FIGURE 1. Rheotanytarsus amaru sp. n., Adult male:A. Head, in frontal view. B. Thorax, in lateral view. C. Wing. D. Hypopygium, in dorsal view. E. Hypopygium in dorsal view, with tergite IX removed." figureDoi="http://doi.org/10.5281/zenodo.3605867" httpUri="https://zenodo.org/record/3605867/files/figure.png" pageId="1" pageNumber="196">Fig. 1B</figureCitation>
). Ac 22–24, restricted to anterior region of scutum; Dc 11–13 on each side, uniserial; Pa 1 on each side; Scts 7–9. Scutum projected anteriorly, distinctly overreaching antepronotum.
</paragraph>
<paragraph id="8BB636A2263B9F44FF57E711546B146B" blockId="1.[151,1437,728,1763]" pageId="1" pageNumber="196">
Wing (
<figureCitation id="13322A27263B9F44FE83E71151091432" box="[275,362,1556,1583]" captionStart="FIGURE 1" captionStartId="2.[151,250,1977,2002]" captionTargetBox="[186,1401,166,1958]" captionTargetId="figure@2.[185,1403,166,1958]" captionTargetPageId="2" captionText="FIGURE 1. Rheotanytarsus amaru sp. n., Adult male:A. Head, in frontal view. B. Thorax, in lateral view. C. Wing. D. Hypopygium, in dorsal view. E. Hypopygium in dorsal view, with tergite IX removed." figureDoi="http://doi.org/10.5281/zenodo.3605867" httpUri="https://zenodo.org/record/3605867/files/figure.png" pageId="1" pageNumber="196">Fig. 1C</figureCitation>
). VRCu 1.35–1.43. Membrane covered with macrotrichia. Brachiolum with 1 seta, C with about 360 setae, R with about 20, R
<subScript id="178D34E7263B9F44FE76E74F518D1445" attach="left" box="[486,494,1610,1624]" fontSize="6" pageId="1" pageNumber="196">1</subScript>
with about 30, R
<subScript id="178D34E7263B9F44FD23E74F52AF1445" attach="left" box="[691,716,1610,1624]" fontSize="6" pageId="1" pageNumber="196">4+5</subScript>
with about 60, M
<subScript id="178D34E7263B9F44FC03E74F53CE1445" attach="left" box="[915,941,1610,1624]" fontSize="6" pageId="1" pageNumber="196">1+2</subScript>
with about 65, M
<subScript id="178D34E7263B9F44FBE7E74F54F21445" attach="left" box="[1143,1169,1610,1624]" fontSize="6" pageId="1" pageNumber="196">3+4</subScript>
with about 45, Cu with about 25, Cu
<subScript id="178D34E7263B9F44FEB7E76B514C1461" attach="left" box="[295,303,1646,1660]" fontSize="6" pageId="1" pageNumber="196">1</subScript>
with about 25 and An with about 40 setae, remaining veins bare.
</paragraph>
<paragraph id="8BB636A2263B9F44FF57E78452B414FE" blockId="1.[151,1437,728,1763]" pageId="1" pageNumber="196">
Legs. Fore leg tibia with a sinuous spur 25–27 μm long. Tibial combs of mid and hind legs separated; spurs of mid leg equal, markedly bent at apex, about 22–23 μm long; spurs of hind leg equal: slightly bent at apex, 25–26 μm long. Lengths and proportions of legs as in
<tableCitation id="C68B0319263B9F44FDEDE7CD52B214FE" box="[637,721,1736,1763]" captionStart="TABLE 1" captionStartId="1.[151,239,1810,1834]" captionTargetPageId="1" captionText="TABLE 1. Lengths (in μm) and proportions of legs of Rheotanytarsus amaru sp. n., male (n = 10)." httpUri="http://table.plazi.org/id/DF76662A263B9F44FF07E61754DB1537" pageId="1" pageNumber="196" tableUuid="DF76662A263B9F44FF07E61754DB1537">Table 1</tableCitation>
.
</paragraph>
<caption id="DF76662A263B9F44FF07E61754DB1537" ID-Table-UUID="DF76662A263B9F44FF07E61754DB1537" box="[151,1208,1810,1834]" httpUri="http://table.plazi.org/id/DF76662A263B9F44FF07E61754DB1537" pageId="1" pageNumber="196" targetBox="[166,1421,1846,1989]" targetIsTable="true" targetPageId="1">
<paragraph id="8BB636A2263B9F44FF07E61754DB1537" blockId="1.[151,1208,1810,1834]" box="[151,1208,1810,1834]" pageId="1" pageNumber="196">
<emphasis id="B97DEAB0263B9F44FF07E61751691537" bold="true" box="[151,266,1810,1834]" pageId="1" pageNumber="196">TABLE 1.</emphasis>
Lengths (in μm) and proportions of legs of
<taxonomicName id="4C094D21263B9F44FD4FE61753B21537" authority="Dantas &amp; Huamantinco Araujo &amp; Hamada, 2020" authorityName="Dantas &amp; Huamantinco Araujo &amp; Hamada" authorityYear="2020" box="[735,977,1810,1834]" class="Insecta" family="Chironomidae" genus="Rheotanytarsus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="species" species="amaru" status="sp. nov.">
<emphasis id="B97DEAB0263B9F44FD4FE61753B21537" box="[735,977,1810,1834]" italics="true" pageId="1" pageNumber="196">Rheotanytarsus amaru</emphasis>
</taxonomicName>
<emphasis id="B97DEAB0263B9F44FC47E61754701537" bold="true" box="[983,1043,1810,1834]" pageId="1" pageNumber="196">
<taxonomicNameLabel id="A24E57CB263B9F44FC47E61754701537" box="[983,1043,1810,1834]" pageId="1" pageNumber="196" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, male (n = 10).
</paragraph>
</caption>
<paragraph id="8BB636A2263B9F44FF49E63255EE15DD" pageId="1" pageNumber="196">
<table id="F909C402263B60BAFF36E63355EE15D8" box="[166,1421,1846,1989]" gridcols="11" gridrows="4" pageId="1" pageNumber="196">
<tr id="353934E0263B60BAFF36E63355EE154E" box="[166,1421,1846,1875]" gridrow="0" pageId="1" pageNumber="196" rowspan-0="1">
<th id="76E85D9C263B60BAFF49E633515D154E" box="[217,318,1846,1875]" gridcol="1" gridrow="0" pageId="1" pageNumber="196">Fe</th>
<th id="76E85D9C263B60BAFECBE63351D7154E" box="[347,436,1846,1875]" gridcol="2" gridrow="0" pageId="1" pageNumber="196">Ti</th>
<th id="76E85D9C263B60BAFE42E6335248154E" box="[466,555,1846,1875]" gridcol="3" gridrow="0" pageId="1" pageNumber="196">ta1</th>
<th id="76E85D9C263B60BAFDD8E63352C2154E" box="[584,673,1846,1875]" gridcol="4" gridrow="0" pageId="1" pageNumber="196">ta2</th>
<th id="76E85D9C263B60BAFD2FE633537B154E" box="[703,792,1846,1875]" gridcol="5" gridrow="0" pageId="1" pageNumber="196">ta3</th>
<th id="76E85D9C263B60BAFCA6E63353EC154E" box="[822,911,1846,1875]" gridcol="6" gridrow="0" pageId="1" pageNumber="196">ta4</th>
<th id="76E85D9C263B60BAFC3CE6335466154E" box="[940,1029,1846,1875]" gridcol="7" gridrow="0" pageId="1" pageNumber="196">ta5</th>
<th id="76E85D9C263B60BAFBB4E63354E9154E" box="[1060,1162,1846,1875]" gridcol="8" gridrow="0" pageId="1" pageNumber="196">LR</th>
<th id="76E85D9C263B60BAFB36E633556F154E" box="[1190,1292,1846,1875]" gridcol="9" gridrow="0" pageId="1" pageNumber="196">BV</th>
<th id="76E85D9C263B60BAFAB8E63355EE154E" box="[1320,1421,1846,1875]" gridcol="10" gridrow="0" pageId="1" pageNumber="196">SV</th>
</tr>
<tr id="353934E0263B60BAFF36E65955EE1564" box="[166,1421,1884,1913]" gridrow="1" pageId="1" pageNumber="196">
<th id="76E85D9C263B60BAFF36E65950DA1564" box="[166,185,1884,1913]" gridcol="0" gridrow="1" pageId="1" pageNumber="196">p1</th>
<td id="76E85D9C263B60BAFF49E659515D1564" box="[217,318,1884,1913]" gridcol="1" gridrow="1" pageId="1" pageNumber="196">843–886</td>
<td id="76E85D9C263B60BAFECBE65951D71564" box="[347,436,1884,1913]" gridcol="2" gridrow="1" pageId="1" pageNumber="196">512–550</td>
<td id="76E85D9C263B60BAFE42E65952481564" box="[466,555,1884,1913]" gridcol="3" gridrow="1" pageId="1" pageNumber="196">890–928</td>
<td id="76E85D9C263B60BAFDD8E65952C21564" box="[584,673,1884,1913]" gridcol="4" gridrow="1" pageId="1" pageNumber="196">463–492</td>
<td id="76E85D9C263B60BAFD2FE659537B1564" box="[703,792,1884,1913]" gridcol="5" gridrow="1" pageId="1" pageNumber="196">342–365</td>
<td id="76E85D9C263B60BAFCA6E65953EC1564" box="[822,911,1884,1913]" gridcol="6" gridrow="1" pageId="1" pageNumber="196">275–296</td>
<td id="76E85D9C263B60BAFC3CE65954661564" box="[940,1029,1884,1913]" gridcol="7" gridrow="1" pageId="1" pageNumber="196">125–127</td>
<td id="76E85D9C263B60BAFBB4E65954E91564" box="[1060,1162,1884,1913]" gridcol="8" gridrow="1" pageId="1" pageNumber="196">1.69–1.74</td>
<td id="76E85D9C263B60BAFB36E659556F1564" box="[1190,1292,1884,1913]" gridcol="9" gridrow="1" pageId="1" pageNumber="196">1.85–1.86</td>
<td id="76E85D9C263B60BAFAB8E65955EE1564" box="[1320,1421,1884,1913]" gridcol="10" gridrow="1" pageId="1" pageNumber="196">1.52–1.55</td>
</tr>
<tr id="353934E0263B60BAFF36E68455EE1582" box="[166,1421,1921,1951]" gridrow="2" pageId="1" pageNumber="196">
<th id="76E85D9C263B60BAFF36E68450DA1582" box="[166,185,1921,1951]" gridcol="0" gridrow="2" pageId="1" pageNumber="196">p2</th>
<td id="76E85D9C263B60BAFF49E684515D1582" box="[217,318,1921,1951]" gridcol="1" gridrow="2" pageId="1" pageNumber="196">970–1020</td>
<td id="76E85D9C263B60BAFECBE68451D71582" box="[347,436,1921,1951]" gridcol="2" gridrow="2" pageId="1" pageNumber="196">846–910</td>
<td id="76E85D9C263B60BAFE42E68452481582" box="[466,555,1921,1951]" gridcol="3" gridrow="2" pageId="1" pageNumber="196">513–542</td>
<td id="76E85D9C263B60BAFDD8E68452C21582" box="[584,673,1921,1951]" gridcol="4" gridrow="2" pageId="1" pageNumber="196">305–326</td>
<td id="76E85D9C263B60BAFD2FE684537B1582" box="[703,792,1921,1951]" gridcol="5" gridrow="2" pageId="1" pageNumber="196">258–275</td>
<td id="76E85D9C263B60BAFCA6E68453EC1582" box="[822,911,1921,1951]" gridcol="6" gridrow="2" pageId="1" pageNumber="196">167–173</td>
<td id="76E85D9C263B60BAFC3CE68454661582" box="[940,1029,1921,1951]" gridcol="7" gridrow="2" pageId="1" pageNumber="196">102–110</td>
<td id="76E85D9C263B60BAFBB4E68454E91582" box="[1060,1162,1921,1951]" gridcol="8" gridrow="2" pageId="1" pageNumber="196">0.60–0.61</td>
<td id="76E85D9C263B60BAFB36E684556F1582" box="[1190,1292,1921,1951]" gridcol="9" gridrow="2" pageId="1" pageNumber="196">2.76–2.88</td>
<td id="76E85D9C263B60BAFAB8E68455EE1582" box="[1320,1421,1921,1951]" gridcol="10" gridrow="2" pageId="1" pageNumber="196">3.54–3.56</td>
</tr>
<tr id="353934E0263B60BAFF36E6A255EE15D8" box="[166,1421,1959,1989]" gridrow="3" pageId="1" pageNumber="196">
<th id="76E85D9C263B60BAFF36E6A250DA15D8" box="[166,185,1959,1989]" gridcol="0" gridrow="3" pageId="1" pageNumber="196">p3</th>
<td id="76E85D9C263B60BAFF49E6A2515D15D8" box="[217,318,1959,1989]" gridcol="1" gridrow="3" pageId="1" pageNumber="196">845–895</td>
<td id="76E85D9C263B60BAFECBE6A251D715D8" box="[347,436,1959,1989]" gridcol="2" gridrow="3" pageId="1" pageNumber="196">665–712</td>
<td id="76E85D9C263B60BAFE42E6A2524815D8" box="[466,555,1959,1989]" gridcol="3" gridrow="3" pageId="1" pageNumber="196">347–384</td>
<td id="76E85D9C263B60BAFDD8E6A252C215D8" box="[584,673,1959,1989]" gridcol="4" gridrow="3" pageId="1" pageNumber="196">200–210</td>
<td id="76E85D9C263B60BAFD2FE6A2537B15D8" box="[703,792,1959,1989]" gridcol="5" gridrow="3" pageId="1" pageNumber="196">165–186</td>
<td id="76E85D9C263B60BAFCA6E6A253EC15D8" box="[822,911,1959,1989]" gridcol="6" gridrow="3" pageId="1" pageNumber="196">105–120</td>
<td id="76E85D9C263B60BAFC3CE6A2546615D8" box="[940,1029,1959,1989]" gridcol="7" gridrow="3" pageId="1" pageNumber="196">84–100</td>
<td id="76E85D9C263B60BAFBB4E6A254E915D8" box="[1060,1162,1959,1989]" gridcol="8" gridrow="3" pageId="1" pageNumber="196">0.52–0.54</td>
<td id="76E85D9C263B60BAFB36E6A2556F15D8" box="[1190,1292,1959,1989]" gridcol="9" gridrow="3" pageId="1" pageNumber="196">3.22–3.52</td>
<td id="76E85D9C263B60BAFAB8E6A255EE15D8" box="[1320,1421,1959,1989]" gridcol="10" gridrow="3" pageId="1" pageNumber="196">4.18–4.35</td>
</tr>
</table>
</paragraph>
<caption id="DF76662A26389F47FF07E6BC53D315EB" ID-DOI="http://doi.org/10.5281/zenodo.3605867" ID-Zenodo-Dep="3605867" httpUri="https://zenodo.org/record/3605867/files/figure.png" pageId="2" pageNumber="197" startId="2.[151,250,1977,2002]" targetBox="[186,1401,166,1958]" targetPageId="2">
<paragraph id="8BB636A226389F47FF07E6BC53D315EB" blockId="2.[151,1436,1977,2038]" pageId="2" pageNumber="197">
<emphasis id="B97DEAB026389F47FF07E6BC517115CF" bold="true" box="[151,274,1977,2002]" pageId="2" pageNumber="197">FIGURE 1.</emphasis>
<taxonomicName id="4C094D2126389F47FE87E6BF519F15CF" authority="Dantas &amp; Huamantinco Araujo &amp; Hamada, 2020" authorityName="Dantas &amp; Huamantinco Araujo &amp; Hamada" authorityYear="2020" box="[279,508,1978,2002]" class="Insecta" family="Chironomidae" genus="Rheotanytarsus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="amaru" status="sp. nov.">
<emphasis id="B97DEAB026389F47FE87E6BF519F15CF" box="[279,508,1978,2002]" italics="true" pageId="2" pageNumber="197">Rheotanytarsus amaru</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57CB26389F47FD92E6BF525A15CC" box="[514,569,1978,2002]" pageId="2" pageNumber="197" rank="species">
<emphasis id="B97DEAB026389F47FD92E6BF525015CF" bold="true" box="[514,563,1978,2002]" pageId="2" pageNumber="197">sp. n</emphasis>
.
</taxonomicNameLabel>
, Adult male: A. Head, in frontal view. B. Thorax, in lateral view. C. Wing. D. Hypopygium, in dorsal view. E. Hypopygium in dorsal view, with tergite IX removed.
</paragraph>
</caption>
<paragraph id="8BB636A226399F46FF57E19C553113CE" blockId="3.[151,1437,153,611]" pageId="3" pageNumber="198">
Hypopygium (
<figureCitation id="13322A2726399F46FEFCE19C51C812AE" box="[364,427,153,179]" captionStart="FIGURE 1" captionStartId="2.[151,250,1977,2002]" captionTargetBox="[186,1401,166,1958]" captionTargetId="figure@2.[185,1403,166,1958]" captionTargetPageId="2" captionText="FIGURE 1. Rheotanytarsus amaru sp. n., Adult male:A. Head, in frontal view. B. Thorax, in lateral view. C. Wing. D. Hypopygium, in dorsal view. E. Hypopygium in dorsal view, with tergite IX removed." figureDoi="http://doi.org/10.5281/zenodo.3605867" httpUri="https://zenodo.org/record/3605867/files/figure.png" pageId="3" pageNumber="198">Fig. 1</figureCitation>
D–E) Anal tergite with open, V-shaped bands. Posterior margin of tergite IX with about 10–13 setae. Anal point 48–52 μm long, 12–14 μm wide, parallel-sided; crest well developed and V-shaped with 2–5 median setae (
<figureCitation id="13322A2726399F46FEA8E1E451F212E6" box="[312,401,225,251]" captionStart="FIGURE 1" captionStartId="2.[151,250,1977,2002]" captionTargetBox="[186,1401,166,1958]" captionTargetId="figure@2.[185,1403,166,1958]" captionTargetPageId="2" captionText="FIGURE 1. Rheotanytarsus amaru sp. n., Adult male:A. Head, in frontal view. B. Thorax, in lateral view. C. Wing. D. Hypopygium, in dorsal view. E. Hypopygium in dorsal view, with tergite IX removed." figureDoi="http://doi.org/10.5281/zenodo.3605867" httpUri="https://zenodo.org/record/3605867/files/figure.png" pageId="3" pageNumber="198">Fig. 1D</figureCitation>
). Superior volsella 45–50 μm long, oval, apically tapered, with 6–7 dorsal and 2–3 marginal setae arranged as shown in
<figureCitation id="13322A2726399F46FE5BE00052711302" box="[459,530,261,287]" captionStart="FIGURE 1" captionStartId="2.[151,250,1977,2002]" captionTargetBox="[186,1401,166,1958]" captionTargetId="figure@2.[185,1403,166,1958]" captionTargetPageId="2" captionText="FIGURE 1. Rheotanytarsus amaru sp. n., Adult male:A. Head, in frontal view. B. Thorax, in lateral view. C. Wing. D. Hypopygium, in dorsal view. E. Hypopygium in dorsal view, with tergite IX removed." figureDoi="http://doi.org/10.5281/zenodo.3605867" httpUri="https://zenodo.org/record/3605867/files/figure.png" pageId="3" pageNumber="198">Figs. 1</figureCitation>
D–E. Stem of median volsella curved, 28–30 μm long,
<date id="FFB7106226399F46FB13E00154CC1302" box="[1155,1199,260,287]" pageId="3" pageNumber="198" value="2014-10">5–6</date>
,
<date id="FFB7106226399F46FB2CE00055131302" box="[1212,1392,261,287]" pageId="3" pageNumber="198" value="2014-10-08" valueMax="2014-12-08" valueMin="2014-10-08">8–10 and 12–14</date>
μm wide at base, middle and apex, respectively, swollen and rounded at apex, bearing a brush of setiform lamellae not fused into a plate (
<figureCitation id="13322A2726399F46FEF9E04851DC137A" box="[361,447,333,359]" captionStart="FIGURE 1" captionStartId="2.[151,250,1977,2002]" captionTargetBox="[186,1401,166,1958]" captionTargetId="figure@2.[185,1403,166,1958]" captionTargetPageId="2" captionText="FIGURE 1. Rheotanytarsus amaru sp. n., Adult male:A. Head, in frontal view. B. Thorax, in lateral view. C. Wing. D. Hypopygium, in dorsal view. E. Hypopygium in dorsal view, with tergite IX removed." figureDoi="http://doi.org/10.5281/zenodo.3605867" httpUri="https://zenodo.org/record/3605867/files/figure.png" pageId="3" pageNumber="198">Fig. 1E</figureCitation>
). Inferior volsella 80–85 μm long, subcylindrical, bearing about 12–15 apical/subapical bristles; internal margin covered with microtrichia (
<figureCitation id="13322A2726399F46FD75E074535F1396" box="[741,828,369,395]" captionStart="FIGURE 1" captionStartId="2.[151,250,1977,2002]" captionTargetBox="[186,1401,166,1958]" captionTargetId="figure@2.[185,1403,166,1958]" captionTargetPageId="2" captionText="FIGURE 1. Rheotanytarsus amaru sp. n., Adult male:A. Head, in frontal view. B. Thorax, in lateral view. C. Wing. D. Hypopygium, in dorsal view. E. Hypopygium in dorsal view, with tergite IX removed." figureDoi="http://doi.org/10.5281/zenodo.3605867" httpUri="https://zenodo.org/record/3605867/files/figure.png" pageId="3" pageNumber="198">Fig. 1E</figureCitation>
). Phallapodeme 80–90 μm long; transverse sternapo- deme 47–54 μm long, without distinct oral projections. Gonocoxite 132–150 μm long. Gonostylus 138–153 μm long, slightly arched, broadest subbasally, evenly tapering toward blunt apex. HR 0.95–0.98, HV 2.25–2.45.
</paragraph>
<paragraph id="8BB636A226399F46FF57E0D8523E13EA" blockId="3.[151,1437,153,611]" box="[199,605,476,503]" pageId="3" pageNumber="198">
<emphasis id="B97DEAB026399F46FF57E0D8518213EA" bold="true" box="[199,481,476,503]" pageId="3" pageNumber="198">Female and immatures.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C313652926399F46FF57E3055340107E" pageId="3" pageNumber="198" type="distribution">
<paragraph id="8BB636A226399F46FF57E3055340107E" blockId="3.[151,1437,153,611]" pageId="3" pageNumber="198">
<emphasis id="B97DEAB026399F46FF57E30552351006" bold="true" box="[199,598,512,539]" pageId="3" pageNumber="198">Distribution and ecological notes.</emphasis>
The new species is known only for the
<typeStatus id="54B2880026399F46FB83E30454201006" box="[1043,1091,513,539]" pageId="3" pageNumber="198">type</typeStatus>
locality in the Peruvian Andes (
<figureCitation id="13322A2726399F46FF30E32050851022" box="[160,230,549,575]" captionStart="FIGURE 2" captionStartId="4.[151,250,1662,1687]" captionTargetBox="[273,1314,246,1640]" captionTargetId="figure@4.[273,1314,246,1640]" captionTargetPageId="4" captionText="FIGURE 2. Distribution of Rheotanytarsus Thienemann &amp; Bause species in South America." figureDoi="http://doi.org/10.5281/zenodo.3605869" httpUri="https://zenodo.org/record/3605869/files/figure.png" pageId="3" pageNumber="198">Fig. 2</figureCitation>
). The adults of
<taxonomicName id="4C094D2126399F46FE08E32052F41022" authority="Dantas &amp; Huamantinco Araujo &amp; Hamada, 2020" authorityName="Dantas &amp; Huamantinco Araujo &amp; Hamada" authorityYear="2020" box="[408,663,549,575]" class="Insecta" family="Chironomidae" genus="Rheotanytarsus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="amaru" status="sp. nov.">
<emphasis id="B97DEAB026399F46FE08E32052F41022" box="[408,663,549,575]" italics="true" pageId="3" pageNumber="198">Rheotanytarsus amaru</emphasis>
</taxonomicName>
<emphasis id="B97DEAB026399F46FD0FE32052BC1022" bold="true" box="[671,735,549,575]" pageId="3" pageNumber="198">
<taxonomicNameLabel id="A24E57CB26399F46FD0FE32052BC1022" box="[671,735,549,575]" pageId="3" pageNumber="198" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
were collected next to a sand bottom river
<quantity id="4CF19B4726399F46FB5BE32054951022" box="[1227,1270,549,575]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="198" unit="m" value="6.0">6 m</quantity>
in width, with low temperature (about 15 °C) at about
<quantity id="4CF19B4726399F46FDC5E34C52D2107E" box="[597,689,585,611]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="198" unit="m" value="3.0">3.000 m</quantity>
above s.l.
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue Copy permalink