152 lines
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152 lines
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.753.22366" ID-GBIF-Dataset="fe320c78-20cd-45de-babe-5c2fd18c7a1d" ID-PMC="PMC5934354" ID-Pensoft-Pub="1313-2970-753-23" ID-PubMed="29731681" ID-ZBK="BBC59C6140DD44F3B9F34C011E0D0B75" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-753-23" ModsDocOrigin="ZooKeys 753" ModsDocTitle="Taxonomic revision of the Graphipterusserrator (Forskål) group (Coleoptera, Carabidae): an increase from five to 15 valid species" checkinTime="1524810864104" checkinUser="pensoft" docAuthor="Renan, Ittai, Assmann, Thorsten & Freidberg, Amnon" docDate="2018" docId="FDC2AE3C9B69567D0B58B4197B399078" docLanguage="en" docName="ZooKeys 753: 23-82" docOrigin="ZooKeys 753" docSource="http://dx.doi.org/10.3897/zookeys.753.22366" docTitle="Graphipterus stagonopsis Renan & Assmann, sp. n." docType="treatment" docUuid="435E6626-47B0-4701-A57C-45E23959B7AD" docUuidSource="ZooBank" docVersion="6" lastPageNumber="62" masterDocId="123E1514FFF23830FFA1FFB0FFE0DA05" masterDocTitle="Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species" masterLastPageNumber="82" masterPageNumber="23" pageNumber="58" updateTime="1668165743174" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Renan, Ittai</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Assmann, Thorsten</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Freidberg, Amnon</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="volume">
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<mods:number>753</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>23</mods:start>
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<mods:end>82</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.753.22366</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.753.22366</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-753-23</mods:identifier>
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<mods:identifier type="ZBK">BBC59C6140DD44F3B9F34C011E0D0B75</mods:identifier>
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<mods:identifier type="ZooBank">BBC59C6140DD44F3B9F34C011E0D0B75</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="143706385" LSID="urn:lsid:zoobank.org:act:435E6626-47B0-4701-A57C-45E23959B7AD" httpUri="http://treatment.plazi.org/id/FDC2AE3C9B69567D0B58B4197B399078" lastPageId="39" lastPageNumber="62" pageId="35" pageNumber="58">
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<subSubSection pageId="35" pageNumber="58" type="nomenclature">
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<paragraph pageId="35" pageNumber="58">
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<taxonomicName LSID="http://zoobank.org/435E6626-47B0-4701-A57C-45E23959B7AD" authority="Renan & Assmann" class="Insecta" family="Carabidae" genus="Graphipterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Graphipterus stagonopsis" order="Coleoptera" pageId="35" pageNumber="58" phylum="Arthropoda" rank="species" species="stagonopsis">Graphipterus stagonopsis Renan & Assmann</taxonomicName>
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<taxonomicNameLabel pageId="35" pageNumber="58">sp. n.</taxonomicNameLabel>
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Figs 18, 27b
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="36" lastPageNumber="59" pageId="35" pageNumber="58" type="types">
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<paragraph pageId="35" pageNumber="58">Types.</paragraph>
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<paragraph pageId="35" pageNumber="58">Holotype.♂ (White label, black handwritten): <Beni Abbes/23.III.48 F. Pierre>. (red label): <Holotype> (ae). Deposited in NHMB [examined].</paragraph>
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<paragraph lastPageId="36" lastPageNumber="59" pageId="35" pageNumber="58">
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Paratypes.(11♂, 3♀), NHMB (Colas collection): Gardhaia (Ghardaia), Sahara, G. Mahoux, 19.5.60 (2♂); Beni Abbes, 23.3.48, F. Paiu (2♂, 1- ae). (Negre collection): Beni Abbes, Sahara argelino, J. Mateu (3♀). (Antoine collection): Beni Abes, south Algerien (
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<taxonomicName genus="Graphipterina" lsidName="Graphipterina reymondi" pageId="35" pageNumber="58" rank="species" species="reymondi">reymondi</taxonomicName>
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) (♂ae). ZMUC Algerie, Beni Abbes, 11.3.1984, Tilg. 4-12.1948, Tentens-Nielsen [
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<taxonomicName lsidName="G. serrator" pageId="35" pageNumber="58" rank="species" species="serrator">G. serrator</taxonomicName>
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<taxonomicName genus="Graphipterina" lsidName="Graphipterina valdani" pageId="35" pageNumber="58" rank="species" species="valdani">valdani</taxonomicName>
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Guer. P. Basilewsky 1985] ♂. NMP: Algeria, Igli,
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<pageBreakToken pageId="36" pageNumber="59" start="start">12</pageBreakToken>
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.IV. 1988, Igt. Kepler, 11/1988. Ex call. M. Dvorak, National Museum, Prague, Czech Republlic. MRAC: Aoulel el Arab Tidicelt Sahara Cen., J. Mateu (♂); Pozo zug (R.O.)
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<normalizedToken originalValue="Sa’hara">Sa'hara</normalizedToken>
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espanol, J. Mateu (2♂); Oasis de la-Salah Tidikelt Sahara Cen, J. Mateu (3♂).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="37" lastPageNumber="60" pageId="36" pageNumber="59" type="diagnosis">
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<paragraph pageId="36" pageNumber="59">Diagnosis.</paragraph>
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<paragraph lastPageId="37" lastPageNumber="60" pageId="36" pageNumber="59">
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Large species with 16 white rounded to elongated spots on elytra, anterior and posterior pair of spots larger than others; six marginal extensions, extension
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<pageBreakToken pageId="37" pageNumber="60" start="start">I</pageBreakToken>
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triangular, extension I and II elongated. Elytra widest at interior rear third, drop-like shape. Median lobe of aedeagus with slightly bent tip.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="37" pageNumber="60" type="comparisons">
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<paragraph pageId="37" pageNumber="60">Comparisons.</paragraph>
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<paragraph pageId="37" pageNumber="60">
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<taxonomicName class="Insecta" family="Carabidae" genus="Graphipterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Graphipterus stagonopsis" order="Coleoptera" pageId="37" pageNumber="60" phylum="Arthropoda" rank="species" species="stagonopsis">Graphipterus stagonopsis</taxonomicName>
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sp. n. resembles
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<taxonomicName lsidName="G. reymondi" pageId="37" pageNumber="60" rank="species" species="reymondi">G. reymondi</taxonomicName>
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from which it differs mainly by the following characters: G.
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<taxonomicName genus="Graphipterina" lsidName="Graphipterina stagonopsis" pageId="37" pageNumber="60" rank="species" species="stagonopsis">stagonopsis</taxonomicName>
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sp. n.: mentum with two teeth; eight spots on elytra; scales of elytral disc brown; claws of hind legs and metatibial secondary spur dark.
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<taxonomicName lsidName="G. reymondi" pageId="37" pageNumber="60" rank="species" species="reymondi">G. reymondi</taxonomicName>
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: mentum with three teeth; 10-12 spots on elytra; scales of elytral disc black; claws of hind legs and metatibial secondary spur brown.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="38" lastPageNumber="61" pageId="37" pageNumber="60" type="description">
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<paragraph pageId="37" pageNumber="60">Description.</paragraph>
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<paragraph pageId="37" pageNumber="60">
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BL male: 17.2-20.1 mm, average 18.8
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<normalizedToken originalValue="±">+/-</normalizedToken>
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1 mm; BL female: 18.4-19.8 mm, average 18.9
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.6 mm.
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</paragraph>
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<paragraph pageId="37" pageNumber="60">Head slender; HW/PW: 0.7; EYL 1.1-1.7 mm; EYL/EL: 0.16. Mentum with two teeth (Fig. 3b). Frontal ridge slightly developed. Male, apical white frons stripes wider than exposed frons (Fig. 4b).</paragraph>
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<paragraph pageId="37" pageNumber="60">Pronotum strongly cordiform; PL/PW: 0.66. BPW/PW: 0.64; posteromedially concave and without white margin; white lateral margin as wide as antennomere I long.</paragraph>
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<paragraph lastPageId="38" lastPageNumber="61" pageId="37" pageNumber="60">
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Elytra droplet-like, humeri strongly narrowed; EL: 9.1-11.1 mm, average 8.4 mm; EW: 7.5-9.0 mm, average 8.4 mm; EL/EW: 1.1-1.5. Lateral cross section convex.
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<pageBreakToken pageId="38" pageNumber="61" start="start">Dense</pageBreakToken>
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black scales, disc not visible between them (Fig. 6a). White lateral margin nearly as half wide as antennomere I long and with six extensions; extension I triangular with rounded angels, as wide as lateral margin, posteriorly oriented; extension II small, often constricted at base, as wide as lateral margin; extension III large, elongated, posteriorly oriented. White posterior margin becomes narrower towards suture; gap at suture wider than lateral margin. Disc usually with 16 spots; anterior pair of spots elongate, as wide as extension I; posterior pair of spots rounded and larger than other ones; six spots forming arch pattern anterior and lateral to posterior rounded larger spots. Apical sinuation slightly developed to straight, apex not protuberant, broadly rounded, especially on the median side (Fig. 7c). Suture inconspicuous.
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</paragraph>
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<paragraph pageId="38" pageNumber="61">Legs long; MTIL: 6.0-7.0 mm, average 6.5 mm; El/MTIL: 1.6. Metatibial secondary spur black. MTAL: 4.7-5.3 mm, average 4.9 mm; MTAL/MTIL: 0.8. Claws of hind legs black at base.</paragraph>
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<paragraph pageId="38" pageNumber="61">Median lobe of aedeagus with bent tip (Fig. 9o).</paragraph>
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</subSubSection>
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<subSubSection pageId="38" pageNumber="61" type="etymology">
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<paragraph pageId="38" pageNumber="61">Etymology.</paragraph>
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<paragraph pageId="38" pageNumber="61">
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The name is derived from ancient Greek (σταγών,
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<normalizedToken originalValue="óψις">oψις</normalizedToken>
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) and means
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<normalizedToken originalValue="”drop-like”">"drop-like"</normalizedToken>
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which refers to the shape of the elytra.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="39" pageNumber="62" type="habitat">
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<paragraph pageId="39" pageNumber="62">
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<pageBreakToken pageId="39" pageNumber="62" start="start">Habitat</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="39" pageNumber="62">Unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="39" pageNumber="62" type="co-occurring species">
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<paragraph pageId="39" pageNumber="62">Co-occurring species.</paragraph>
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<paragraph pageId="39" pageNumber="62">
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<taxonomicName class="Insecta" family="Carabidae" genus="Graphipterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Graphipterus stagonopsis" order="Coleoptera" pageId="39" pageNumber="62" phylum="Arthropoda" rank="species" species="stagonopsis">Graphipterus stagonopsis</taxonomicName>
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lives in sympatry with
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<taxonomicName lsidName="G. luctuosus" pageId="39" pageNumber="62" rank="species" species="luctuosus">G. luctuosus</taxonomicName>
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,
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<taxonomicName lsidName="G. peletieri" pageId="39" pageNumber="62" rank="species" species="peletieri">G. peletieri</taxonomicName>
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, and
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<taxonomicName lsidName="G. valdanii" pageId="39" pageNumber="62" rank="species" species="valdanii">G. valdanii</taxonomicName>
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in Ghardaia, Algeria.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="39" pageNumber="62" type="distribution">
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<paragraph pageId="39" pageNumber="62">Distribution.</paragraph>
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<paragraph pageId="39" pageNumber="62">Central and west Algeria (Fig. 18).</paragraph>
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</subSubSection>
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<subSubSection pageId="39" pageNumber="62" type="conservation">
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<paragraph pageId="39" pageNumber="62">Conservation.</paragraph>
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<paragraph pageId="39" pageNumber="62">Unknown.</paragraph>
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</subSubSection>
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</treatment>
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</document>
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