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<document id="5DFF4FC7C5B74022906EB5811F79D1FC" ID-CLB-Dataset="2603" ID-DOI="10.11646/zootaxa.5091.2.7" ID-GBIF-Dataset="1f11a62b-6c25-497a-8506-9dc282daadb6" ID-ISSN="1175-5326" ID-Zenodo-Dep="5843784" ID-ZooBank="68EB0115-26FA-4DEF-9564-699C5CCEFC66" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1642061483800" checkinUser="plazi" docAuthor="Zhang, Meng, Irfan, Muhammad, Wang, Lu-Yu &amp; Zhang, Zhi-Sheng" docDate="2022" docId="03B6F12C861FF95DFF70FB5ED6CE8037" docLanguage="en" docName="zootaxa.5091.2.7.pdf" docOrigin="Zootaxa 5091 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tonsilla subtruculenta Zhang &amp; Irfan &amp; Wang &amp; Zhang 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="367" masterDocId="FF8F89548617F957FFE7FFD8D45A822E" masterDocTitle="Six new species of Tonsilla Wang &amp; Yin, 1992 (Araneae: Agelenidae) from southern China, with the first description of the male of T. yanlingensis (Zhang, Yin &amp; Kim, 2000)" masterLastPageNumber="372" masterPageNumber="357" pageNumber="365" updateTime="1699302077523" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="410BE397C00041654EE1982D9F232DAF">Six new species of Tonsilla Wang &amp; Yin, 1992 (Araneae: Agelenidae) from southern China, with the first description of the male of T. yanlingensis (Zhang, Yin &amp; Kim, 2000)</mods:title>
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<mods:namePart id="F5862A0CCEC2153814DE167E02653D5A">Zhang, Meng</mods:namePart>
<mods:nameIdentifier id="023C8CFF7BEC3F1A9264E826B6DE9B5D" type="ORCID">0000-0003-2561-225X</mods:nameIdentifier>
<mods:affiliation id="294A81A013B76FBE404E699592855CDA">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Sciences, Southwest University, Chongqing 400715, China. &amp; 675891014 @ qq. com; https: // orcid. org / 0000 - 0003 - 2561 - 225 X</mods:affiliation>
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<mods:namePart id="375C1EE4283694342EBC7DC22991C345">Irfan, Muhammad</mods:namePart>
<mods:nameIdentifier id="F6BAB11F64BE66D19ADAB8BD548F0882" type="ORCID">0000-0003-0445-9612</mods:nameIdentifier>
<mods:affiliation id="622E34AB0C09D175C8944572292212F0">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Sciences, Southwest University, Chongqing 400715, China. &amp; irfanuos 94 @ yahoo. com; https: // orcid. org / 0000 - 0003 - 0445 - 9612</mods:affiliation>
<mods:nameIdentifier id="331E3B507D8E0A511E81B72F0DCC61D1" type="email">irfanuos94@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="AE66A27A7264A6AD0D0E524C4C89F610">Wang, Lu-Yu</mods:namePart>
<mods:nameIdentifier id="48657E10D8A3882F2967546AA91ADFAE" type="ORCID">0000-0002-5250-3473</mods:nameIdentifier>
<mods:affiliation id="1B13D73EF14105B0208FB49594B080D5">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Sciences, Southwest University, Chongqing 400715, China. &amp; wangluyu 1989 @ 163. com; https: // orcid. org / 0000 - 0002 - 5250 - 3473</mods:affiliation>
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<mods:namePart id="58522524CCAF809CBE40C258A39F1AB5">Zhang, Zhi-Sheng</mods:namePart>
<mods:affiliation id="9AD73E909A85E7F477F8C1148872520A">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Sciences, Southwest University, Chongqing 400715, China.</mods:affiliation>
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<mods:date id="D8F6FB1E66063C9FC9220FD491009AA9">2022</mods:date>
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<treatment id="03B6F12C861FF95DFF70FB5ED6CE8037" ID-DOI="http://doi.org/10.5281/zenodo.5844049" ID-GBIF-Taxon="192084953" ID-Zenodo-Dep="5844049" LSID="urn:lsid:plazi:treatment:03B6F12C861FF95DFF70FB5ED6CE8037" httpUri="http://treatment.plazi.org/id/03B6F12C861FF95DFF70FB5ED6CE8037" lastPageId="10" lastPageNumber="367" pageId="8" pageNumber="365">
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<taxonomicName id="4C1F3BB9861FF95FFF70FB5ED5FD868F" ID-CoL="98YK3" authority="Zhang &amp; Irfan &amp; Wang &amp; Zhang, 2022" authorityName="Zhang &amp; Irfan &amp; Wang &amp; Zhang" authorityYear="2022" box="[151,423,1158,1185]" class="Arachnida" family="Agelenidae" genus="Tonsilla" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="365" phylum="Arthropoda" rank="species" species="subtruculenta" status="sp. nov.">
<emphasis id="B96B9C28861FF95FFF70FB5ED5FD868F" bold="true" box="[151,423,1158,1185]" italics="true" pageId="8" pageNumber="365">Tonsilla subtruculenta</emphasis>
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</paragraph>
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<subSubSection id="C30513B1861FF95FFF70FB73D5C686EB" box="[151,412,1195,1221]" pageId="8" pageNumber="365" type="description">
<paragraph id="8BA0403A861FF95FFF70FB73D5C686EB" blockId="8.[151,524,1158,1221]" box="[151,412,1195,1221]" pageId="8" pageNumber="365">
<figureCitation id="13245CBF861FF95FFF70FB73D56C86EB" box="[151,310,1195,1221]" captionStart="FIGURE 6" captionStartId="9.[151,250,1823,1847]" captionTargetBox="[151,1432,453,1795]" captionTargetId="figure-151@9.[151,1435,453,1800]" captionTargetPageId="9" captionText="FIGURE 6. Tonsilla subtruculenta sp. nov. A. Male holotype, left palp, prolateral view; B. Same, ventral view; C. Same, retrolateral view; D. Epigyne, ventral view; E. Same, dorsal view. Abbreviations: CD, copulatory duct; CDA, Conductor’s dorsal apophysis; CF, cymbial furrow; CL, conductor lamella; Co, conductor; CO, copulatory opening; EH, epigynal hood; Em, embolus; ET, epigynal teeth; FD, fertilization ducts; LTA, lateral tibial apophysis; MA, median apophysis; PA, patellar apophysis; RTA, retrolateral tibial apophysis; SH, spermathecal heads; Sp, spermathecae." figureDoi="http://doi.org/10.5281/zenodo.5843804" httpUri="https://zenodo.org/record/5843804/files/figure.png" pageId="8" pageNumber="365">Figures 6A–E</figureCitation>
,
<figureCitation id="13245CBF861FF95FFEA5FB73D52F86EB" box="[322,373,1195,1221]" captionStart="FIGURE 9" captionStartId="13.[151,250,1902,1926]" captionTargetBox="[197,1388,181,1879]" captionTargetId="figure-19@13.[197,1390,181,1879]" captionTargetPageId="13" captionText="FIGURE 9. Habitus of Tonsilla spp. A–B, male holotype and female paratype of T. jinyunensis sp. nov.; C–D, male holotype and female paratype of T. jiugongensis sp. nov.; E–F, male holotype and female paratype of T. rutunda sp. nov.; G–H, male holotype and female paratype of T. subrostrum sp. nov.; I–J, male holotype and female paratype of T. subtruculenta sp. nov.; K–L, male and female of T. yanlingensis (Zhang, Yin &amp; Kim, 2000); M, male holotype of T. yueliangensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5843814" httpUri="https://zenodo.org/record/5843814/files/figure.png" pageId="8" pageNumber="365">9I–J</figureCitation>
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<figureCitation id="13245CBF861FF95FFE67FB73D5C686EB" box="[384,412,1195,1221]" captionStart="FIGURE 10" captionStartId="14.[151,250,1414,1438]" captionTargetBox="[167,1422,187,1386]" captionTargetId="figure-25@14.[161,1426,181,1390]" captionTargetPageId="14" captionText="FIGURE 10. Distribution of Tonsilla species treated here." figureDoi="http://doi.org/10.5281/zenodo.5843816" httpUri="https://zenodo.org/record/5843816/files/figure.png" pageId="8" pageNumber="365">10</figureCitation>
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<emphasis id="B96B9C28861FF95FFF70FB2BD5138722" bold="true" box="[151,329,1266,1293]" pageId="8" pageNumber="365">Type material.</emphasis>
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<typeStatus id="54A4FE98861FF95FFEB3FB2BD59B8723" box="[340,449,1267,1293]" pageId="8" pageNumber="365" type="holotype">Holotype</typeStatus>
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♂,
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♀
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;
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,
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,
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,
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,
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,
<date id="FFA166FA861FF95FFF1FFA5BD52A87B3" box="[248,368,1411,1437]" pageId="8" pageNumber="365" value="2012-10-18">
<collectingDate id="EFE59F12861FF95FFF1FFA5BD52A87B3" box="[248,368,1411,1437]" pageId="8" pageNumber="365" value="2012-10-18">18.X.2012</collectingDate>
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<collectorName id="26EA25EC861FF95FFE65FA5BD65887B3" box="[386,514,1411,1437]" pageId="8" pageNumber="365">L.Y. Wang</collectorName>
,
<collectorName id="26EA25EC861FF95FFDF4FA5BD6CB87B3" box="[531,657,1411,1437]" pageId="8" pageNumber="365">X.K. Jiang</collectorName>
&amp;
<collectorName id="26EA25EC861FF95FFD26FA5BD71387B3" box="[705,841,1411,1437]" pageId="8" pageNumber="365">X.W. Meng</collectorName>
leg. (SWUC-T-AG-53-04~05)
</materialsCitation>
;
<specimenCount id="9D198BB3861FF95FFB23FA5BD0AE87B3" box="[1220,1268,1411,1437]" pageId="8" pageNumber="365" type="female">2 ♀</specimenCount>
,
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,
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,
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,
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</quantity>
,
<date id="FFA166FA861FF95FFDB4FA7ED69187EF" box="[595,715,1446,1473]" pageId="8" pageNumber="365" value="2005-10-14">
<collectingDate id="EFE59F12861FF95FFDB4FA7ED69187EF" box="[595,715,1446,1473]" pageId="8" pageNumber="365" value="2005-10-14">14.X.2005</collectingDate>
</date>
,
<collectorName id="26EA25EC861FF95FFD3EFA7FD70287EF" box="[729,856,1447,1473]" pageId="8" pageNumber="365">Z.S. Zhang</collectorName>
&amp;
<collectorName id="26EA25EC861FF95FFC99FA7FD7A787EF" box="[894,1021,1447,1473]" pageId="8" pageNumber="365">H.M. Chen</collectorName>
leg. (SWUC-T-AG-53-06~07)
</materialsCitation>
;
<specimenCount id="9D198BB3861FF95FFA8DFA7FD1CF87EF" box="[1386,1429,1447,1473]" pageId="8" pageNumber="365" type="female">1 ♀</specimenCount>
,
<materialsCitation id="3B774A67861FF95FFF70FA13D6CB87CB" ID-GBIF-Occurrence="3436475301" box="[151,657,1482,1509]" collectingDate="2012-10-21" collectorName="L. Y. Wang &amp; X. K. Jiang &amp; X. W. Meng" country="China" elevation="1494" latitude="26.367933" location="Xian'nvtang" longLatPrecision="1" longitude="108.20137" pageId="8" pageNumber="365" specimenCount="1" stateProvince="Leigong Mountain" typeStatus="paratype">same data as holotype (SWUC-T-AG-53-08)</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30513B1861FF95FFF20FA37D55E8403" pageId="8" pageNumber="365" type="etymology">
<paragraph id="8BA0403A861FF95FFF20FA37D55E8403" blockId="8.[151,1437,1266,2013]" pageId="8" pageNumber="365">
<emphasis id="B96B9C28861FF95FFF20FA37D5148427" bold="true" box="[199,334,1519,1545]" pageId="8" pageNumber="365">Etymology.</emphasis>
The specific epithet refers to similarity of the male to
<taxonomicName id="4C1F3BB9861FF95FFC30FA37D1CD8427" authority="Wang &amp; Yin, 1992" authorityName="Wang &amp; Yin" authorityYear="1992" box="[983,1431,1519,1545]" class="Arachnida" family="Agelenidae" genus="Tonsilla" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="365" phylum="Arthropoda" rank="species" species="truculenta">
<emphasis id="B96B9C28861FF95FFC30FA37D0F48427" box="[983,1198,1519,1545]" italics="true" pageId="8" pageNumber="365">Tonsilla truculenta</emphasis>
<bibRefCitation id="EF8E3DCB861FF95FFB5FFA37D1CD8427" author="Wang, J. F. &amp; Yin, C. M." box="[1208,1431,1519,1545]" pageId="8" pageNumber="365" pagination="263 - 272" refId="ref8817" refString="Wang, J. F. &amp; Yin, C. M. (1992) A new genus and three new species of funnel-web spiders from south China (Araneae: Agelenidae). Journal of Natural Science of Hunan Normal University, 15, 263 - 272." type="journal article" year="1992">Wang &amp; Yin, 1992</bibRefCitation>
</taxonomicName>
; adjective.
</paragraph>
</subSubSection>
<subSubSection id="C30513B1861FF95FFF20F9EED730855F" pageId="8" pageNumber="365" type="diagnosis">
<paragraph id="8BA0403A861FF95FFF20F9EED730855F" blockId="8.[151,1437,1266,2013]" pageId="8" pageNumber="365">
<emphasis id="B96B9C28861FF95FFF20F9EED518847E" bold="true" box="[199,322,1590,1616]" pageId="8" pageNumber="365">Diagnosis.</emphasis>
The male of this new species resembles
<taxonomicName id="4C1F3BB9861FF95FFCF5F9EFD031847F" authority="(Zhu et al. 2017)" baseAuthorityName="Zhu" baseAuthorityYear="2017" box="[786,1131,1591,1617]" class="Arachnida" family="Agelenidae" genus="Tonsilla" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="365" phylum="Arthropoda" rank="species" species="truculenta">
<emphasis id="B96B9C28861FF95FFCF5F9EFD7F8847F" box="[786,930,1591,1617]" italics="true" pageId="8" pageNumber="365">T. truculenta</emphasis>
(
<bibRefCitation id="EF8E3DCB861FF95FFC56F9EFD039847F" author="Zhu, M. S. &amp; Wang, X. P. &amp; Zhang, Z. S." box="[945,1123,1591,1617]" pageId="8" pageNumber="365" refId="ref9297" refString="Zhu, M. S., Wang, X. P. &amp; Zhang, Z. S. (2017) Fauna Sinica: Invertebrata. Vol. 59. Arachnida: Araneae: Agelenidae and Amaurobiidae. Science Press, Beijing, 727 pp." type="book" year="2017">
Zhu
<emphasis id="B96B9C28861FF95FFC0FF9EFD040847F" box="[1000,1050,1591,1617]" italics="true" pageId="8" pageNumber="365">et al</emphasis>
. 2017
</bibRefCitation>
)
</taxonomicName>
in having a similar tegular sclerite, conductor’s dorsal apophysis, tibial apophysis of male palp. The females of both species have epigynes with long epigynal teeth. Both species can be distinguished by the following characters: patellar apophysis orientated parallel to tibial base in retrolateral view in
<taxonomicName id="4C1F3BB9861FF95FFD62F97BD7618493" authority="Zhang &amp; Irfan &amp; Wang &amp; Zhang, 2022" authorityName="Zhang &amp; Irfan &amp; Wang &amp; Zhang" authorityYear="2022" box="[645,827,1699,1725]" class="Arachnida" family="Agelenidae" genus="Tonsilla" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="365" phylum="Arthropoda" rank="species" species="subtruculenta" status="sp. nov.">
<emphasis id="B96B9C28861FF95FFD62F97BD7618493" box="[645,827,1699,1725]" italics="true" pageId="8" pageNumber="365">T. subtruculenta</emphasis>
</taxonomicName>
<emphasis id="B96B9C28861FF95FFCA4F97BD7C78493" bold="true" box="[835,925,1699,1725]" pageId="8" pageNumber="365">
<taxonomicNameLabel id="A2582153861FF95FFCA4F97BD7C78493" box="[835,925,1699,1725]" pageId="8" pageNumber="365" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13245CBF861FF95FFC4AF97BD07C8493" box="[941,1062,1699,1725]" captionStart="FIGURE 6" captionStartId="9.[151,250,1823,1847]" captionTargetBox="[151,1432,453,1795]" captionTargetId="figure-151@9.[151,1435,453,1800]" captionTargetPageId="9" captionText="FIGURE 6. Tonsilla subtruculenta sp. nov. A. Male holotype, left palp, prolateral view; B. Same, ventral view; C. Same, retrolateral view; D. Epigyne, ventral view; E. Same, dorsal view. Abbreviations: CD, copulatory duct; CDA, Conductor’s dorsal apophysis; CF, cymbial furrow; CL, conductor lamella; Co, conductor; CO, copulatory opening; EH, epigynal hood; Em, embolus; ET, epigynal teeth; FD, fertilization ducts; LTA, lateral tibial apophysis; MA, median apophysis; PA, patellar apophysis; RTA, retrolateral tibial apophysis; SH, spermathecal heads; Sp, spermathecae." figureDoi="http://doi.org/10.5281/zenodo.5843804" httpUri="https://zenodo.org/record/5843804/files/figure.png" pageId="8" pageNumber="365">Fig. 6B, C</figureCitation>
), vs. extending above the tibia in
<taxonomicName id="4C1F3BB9861FF95FFF70F91FD57C84CF" box="[151,294,1735,1761]" class="Arachnida" family="Agelenidae" genus="Tonsilla" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="365" phylum="Arthropoda" rank="species" species="truculenta">
<emphasis id="B96B9C28861FF95FFF70F91FD57C84CF" box="[151,294,1735,1761]" italics="true" pageId="8" pageNumber="365">T. truculenta</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8E3DCB861FF95FFED3F91FD5BE84CF" author="Zhu, M. S. &amp; Wang, X. P. &amp; Zhang, Z. S." box="[308,484,1735,1761]" pageId="8" pageNumber="365" refId="ref9297" refString="Zhu, M. S., Wang, X. P. &amp; Zhang, Z. S. (2017) Fauna Sinica: Invertebrata. Vol. 59. Arachnida: Araneae: Agelenidae and Amaurobiidae. Science Press, Beijing, 727 pp." type="book" year="2017">
Zhu
<emphasis id="B96B9C28861FF95FFE8DF91FD5C184CF" box="[362,411,1735,1761]" italics="true" pageId="8" pageNumber="365">et al</emphasis>
. 2017
</bibRefCitation>
, fig. 361E). Upper part of conductor apex somewhat rectangular in ventral view in
<taxonomicName id="4C1F3BB9861FF95FFA6FF91FD56B852B" authority="Zhang &amp; Irfan &amp; Wang &amp; Zhang, 2022" authorityName="Zhang &amp; Irfan &amp; Wang &amp; Zhang" authorityYear="2022" class="Arachnida" family="Agelenidae" genus="Tonsilla" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="365" phylum="Arthropoda" rank="species" species="subtruculenta" status="sp. nov.">
<emphasis id="B96B9C28861FF95FFA6FF91FD56B852B" italics="true" pageId="8" pageNumber="365">T. subtruculenta</emphasis>
</taxonomicName>
<emphasis id="B96B9C28861FF95FFEDEF933D5CE852B" bold="true" box="[313,404,1771,1797]" pageId="8" pageNumber="365">
<taxonomicNameLabel id="A2582153861FF95FFEDEF933D5CE852B" box="[313,404,1771,1797]" pageId="8" pageNumber="365" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13245CBF861FF95FFE43F933D5A78528" box="[420,509,1771,1798]" captionStart="FIGURE 6" captionStartId="9.[151,250,1823,1847]" captionTargetBox="[151,1432,453,1795]" captionTargetId="figure-151@9.[151,1435,453,1800]" captionTargetPageId="9" captionText="FIGURE 6. Tonsilla subtruculenta sp. nov. A. Male holotype, left palp, prolateral view; B. Same, ventral view; C. Same, retrolateral view; D. Epigyne, ventral view; E. Same, dorsal view. Abbreviations: CD, copulatory duct; CDA, Conductor’s dorsal apophysis; CF, cymbial furrow; CL, conductor lamella; Co, conductor; CO, copulatory opening; EH, epigynal hood; Em, embolus; ET, epigynal teeth; FD, fertilization ducts; LTA, lateral tibial apophysis; MA, median apophysis; PA, patellar apophysis; RTA, retrolateral tibial apophysis; SH, spermathecal heads; Sp, spermathecae." figureDoi="http://doi.org/10.5281/zenodo.5843804" httpUri="https://zenodo.org/record/5843804/files/figure.png" pageId="8" pageNumber="365">Fig. 6B</figureCitation>
), vs. thumb-shaped in
<taxonomicName id="4C1F3BB9861FF95FFCE7F933D7CA852B" box="[768,912,1771,1797]" class="Arachnida" family="Agelenidae" genus="Tonsilla" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="365" phylum="Arthropoda" rank="species" species="truculenta">
<emphasis id="B96B9C28861FF95FFCE7F933D7CA852B" box="[768,912,1771,1797]" italics="true" pageId="8" pageNumber="365">T. truculenta</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8E3DCB861FF95FFC78F933D009852B" author="Zhu, M. S. &amp; Wang, X. P. &amp; Zhang, Z. S." box="[927,1107,1771,1797]" pageId="8" pageNumber="365" refId="ref9297" refString="Zhu, M. S., Wang, X. P. &amp; Zhang, Z. S. (2017) Fauna Sinica: Invertebrata. Vol. 59. Arachnida: Araneae: Agelenidae and Amaurobiidae. Science Press, Beijing, 727 pp." type="book" year="2017">
Zhu
<emphasis id="B96B9C28861FF95FFC31F933D053852B" box="[982,1033,1771,1797]" italics="true" pageId="8" pageNumber="365">et al</emphasis>
. 2017
</bibRefCitation>
, fig. 361E). The female of
<taxonomicName id="4C1F3BB9861FF95FFA6FF933D56B8507" authority="Zhang &amp; Irfan &amp; Wang &amp; Zhang, 2022" authorityName="Zhang &amp; Irfan &amp; Wang &amp; Zhang" authorityYear="2022" class="Arachnida" family="Agelenidae" genus="Tonsilla" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="365" phylum="Arthropoda" rank="species" species="subtruculenta" status="sp. nov.">
<emphasis id="B96B9C28861FF95FFA6FF933D56B8507" italics="true" pageId="8" pageNumber="365">T. subtruculenta</emphasis>
</taxonomicName>
<emphasis id="B96B9C28861FF95FFEDFF8D7D5CB8507" bold="true" box="[312,401,1807,1833]" pageId="8" pageNumber="365">
<taxonomicNameLabel id="A2582153861FF95FFEDFF8D7D5CB8507" box="[312,401,1807,1833]" pageId="8" pageNumber="365" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from all congeners by the following characters: atrial ridges round with copulatory opening at the base of septum posteriorly; copulatory ducts oval, anteriorly covered by spermathecal head; spermathecae robust, situated ventro-mesally (
<figureCitation id="13245CBF861FF95FFD03F88FD707855F" box="[740,861,1879,1905]" captionStart="FIGURE 6" captionStartId="9.[151,250,1823,1847]" captionTargetBox="[151,1432,453,1795]" captionTargetId="figure-151@9.[151,1435,453,1800]" captionTargetPageId="9" captionText="FIGURE 6. Tonsilla subtruculenta sp. nov. A. Male holotype, left palp, prolateral view; B. Same, ventral view; C. Same, retrolateral view; D. Epigyne, ventral view; E. Same, dorsal view. Abbreviations: CD, copulatory duct; CDA, Conductor’s dorsal apophysis; CF, cymbial furrow; CL, conductor lamella; Co, conductor; CO, copulatory opening; EH, epigynal hood; Em, embolus; ET, epigynal teeth; FD, fertilization ducts; LTA, lateral tibial apophysis; MA, median apophysis; PA, patellar apophysis; RTA, retrolateral tibial apophysis; SH, spermathecal heads; Sp, spermathecae." figureDoi="http://doi.org/10.5281/zenodo.5843804" httpUri="https://zenodo.org/record/5843804/files/figure.png" pageId="8" pageNumber="365">Fig. 6D, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30513B1861FF95DFF20F8A2D15683DB" lastPageId="10" lastPageNumber="367" pageId="8" pageNumber="365" type="description">
<paragraph id="8BA0403A861FF95EFF20F8A2D01B82D7" blockId="8.[151,1437,1266,2013]" lastBlockId="9.[151,1437,150,429]" lastPageId="9" lastPageNumber="366" pageId="8" pageNumber="365">
<emphasis id="B96B9C28861FF95FFF20F8A2D50385BA" bold="true" box="[199,345,1914,1940]" pageId="8" pageNumber="365">Description.</emphasis>
Male (
<typeStatus id="54A4FE98861FF95FFE4BF8A3D64885BB" box="[428,530,1915,1941]" pageId="8" pageNumber="365" type="holotype">holotype</typeStatus>
,
<figureCitation id="13245CBF861FF95FFDC7F8A3D62A85BB" box="[544,624,1915,1941]" captionStart="FIGURE 9" captionStartId="13.[151,250,1902,1926]" captionTargetBox="[197,1388,181,1879]" captionTargetId="figure-19@13.[197,1390,181,1879]" captionTargetPageId="13" captionText="FIGURE 9. Habitus of Tonsilla spp. A–B, male holotype and female paratype of T. jinyunensis sp. nov.; C–D, male holotype and female paratype of T. jiugongensis sp. nov.; E–F, male holotype and female paratype of T. rutunda sp. nov.; G–H, male holotype and female paratype of T. subrostrum sp. nov.; I–J, male holotype and female paratype of T. subtruculenta sp. nov.; K–L, male and female of T. yanlingensis (Zhang, Yin &amp; Kim, 2000); M, male holotype of T. yueliangensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5843814" httpUri="https://zenodo.org/record/5843814/files/figure.png" pageId="8" pageNumber="365">Fig. 9I</figureCitation>
). Total length 14.87. Carapace 7.66 long, 4.71 wide; opisthosoma 7.49 long, 4.83 wide. Eye sizes and interdistances: AME 0.33, ALE 0.32, PME 0.28, PLE 0.34; AME–AME 0.13, AME– ALE 0.13, PME–PME 0.19, PME–PLE 0.35, ALE–PLE 0.08. MOA 0.88 long, front width 0.70, back width 0.84. Clypeus height 0.28. Leg measurements: I 22.20 (5.88, 7.46, 5.42, 3.44); II 20.14 (5.32, 6.50, 5.01, 3.31); III 17.52 (4.61, 5.45, 4.83, 2.63); IV 22.88 (5.97, 7.27, 6.35, 3.29). Spination of legs: femur I 220, II 400, III 600, IV 220; patella I-IV 000; tibia I-II 026, III-IV 046; metatarsus I 046, II 056, III 066, IV 067.
</paragraph>
<paragraph id="8BA0403A861EF95EFF20FEDBD17E8383" blockId="9.[151,1437,150,429]" pageId="9" pageNumber="366">
Palp (
<figureCitation id="13245CBF861EF95EFEEFFEDBD5DE8330" box="[264,388,259,286]" captionStart="FIGURE 6" captionStartId="9.[151,250,1823,1847]" captionTargetBox="[151,1432,453,1795]" captionTargetId="figure-151@9.[151,1435,453,1800]" captionTargetPageId="9" captionText="FIGURE 6. Tonsilla subtruculenta sp. nov. A. Male holotype, left palp, prolateral view; B. Same, ventral view; C. Same, retrolateral view; D. Epigyne, ventral view; E. Same, dorsal view. Abbreviations: CD, copulatory duct; CDA, Conductor’s dorsal apophysis; CF, cymbial furrow; CL, conductor lamella; Co, conductor; CO, copulatory opening; EH, epigynal hood; Em, embolus; ET, epigynal teeth; FD, fertilization ducts; LTA, lateral tibial apophysis; MA, median apophysis; PA, patellar apophysis; RTA, retrolateral tibial apophysis; SH, spermathecal heads; Sp, spermathecae." figureDoi="http://doi.org/10.5281/zenodo.5843804" httpUri="https://zenodo.org/record/5843804/files/figure.png" pageId="9" pageNumber="366">Fig. 6A–C</figureCitation>
). Patella longer than patellar apophysis; patellar apophysis almost parallel to the tibial base in retrolateral view. Retrolateral and lateral tibial apophysis distinct with blunt end. Cymbium approximately 3 times longer than wide; cymbial furrow extensively sclerotized, 1/3 of the cymbial length. Median apophysis spoon-like. Conductor apex bifurcated, upper part somewhat rectangular and lower part round. Conductor lamella distinct. Conductor’s dorsal apophysis curved clockwise. Embolus filiform, originating at the 7 o’clock position.
</paragraph>
<caption id="DF6010B2861EF95EFF70F8C7D07B85E6" ID-DOI="http://doi.org/10.5281/zenodo.5843804" ID-Zenodo-Dep="5843804" httpUri="https://zenodo.org/record/5843804/files/figure.png" pageId="9" pageNumber="366" startId="9.[151,250,1823,1847]" targetBox="[151,1432,453,1795]" targetPageId="9">
<paragraph id="8BA0403A861EF95EFF70F8C7D07B85E6" blockId="9.[151,1436,1823,1992]" pageId="9" pageNumber="366">
<emphasis id="B96B9C28861EF95EFF70F8C7D54C8519" bold="true" box="[151,278,1823,1847]" pageId="9" pageNumber="366">FIGURE 6.</emphasis>
<taxonomicName id="4C1F3BB9861EF95EFEF8F8F8D65E8516" authority="Zhang &amp; Irfan &amp; Wang &amp; Zhang, 2022" authorityName="Zhang &amp; Irfan &amp; Wang &amp; Zhang" authorityYear="2022" box="[287,516,1824,1848]" class="Arachnida" family="Agelenidae" genus="Tonsilla" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="366" phylum="Arthropoda" rank="species" species="subtruculenta" status="sp. nov.">
<emphasis id="B96B9C28861EF95EFEF8F8F8D65E8516" box="[287,516,1824,1848]" italics="true" pageId="9" pageNumber="366">Tonsilla subtruculenta</emphasis>
</taxonomicName>
<emphasis id="B96B9C28861EF95EFDE9F8F8D63B8516" bold="true" box="[526,609,1824,1848]" pageId="9" pageNumber="366">
<taxonomicNameLabel id="A2582153861EF95EFDE9F8F8D63B8516" box="[526,609,1824,1848]" pageId="9" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. Male holotype, left palp, prolateral view; B. Same, ventral view; C. Same, retrolateral view; D. Epigyne, ventral view; E. Same, dorsal view. Abbreviations: CD, copulatory duct; CDA, Conductor’s dorsal apophysis; CF, cymbial furrow; CL, conductor lamella; Co, conductor; CO, copulatory opening; EH, epigynal hood; Em, embolus; ET, epigynal teeth; FD, fertilization ducts; LTA, lateral tibial apophysis; MA, median apophysis; PA, patellar apophysis; RTA, retrolateral tibial apophysis; SH, spermathecal heads; Sp, spermathecae.
</paragraph>
</caption>
<paragraph id="8BA0403A861DF95DFF20FF4FD010834B" blockId="10.[151,1437,151,538]" pageId="10" pageNumber="367">
Female (one of the
<typeStatus id="54A4FE98861DF95DFE4FFF4FD64C829F" box="[424,534,151,177]" pageId="10" pageNumber="367" type="paratype">paratypes</typeStatus>
,
<figureCitation id="13245CBF861DF95DFDC4FF4FD62E829F" box="[547,628,151,177]" captionStart="FIGURE 9" captionStartId="13.[151,250,1902,1926]" captionTargetBox="[197,1388,181,1879]" captionTargetId="figure-19@13.[197,1390,181,1879]" captionTargetPageId="13" captionText="FIGURE 9. Habitus of Tonsilla spp. A–B, male holotype and female paratype of T. jinyunensis sp. nov.; C–D, male holotype and female paratype of T. jiugongensis sp. nov.; E–F, male holotype and female paratype of T. rutunda sp. nov.; G–H, male holotype and female paratype of T. subrostrum sp. nov.; I–J, male holotype and female paratype of T. subtruculenta sp. nov.; K–L, male and female of T. yanlingensis (Zhang, Yin &amp; Kim, 2000); M, male holotype of T. yueliangensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5843814" httpUri="https://zenodo.org/record/5843814/files/figure.png" pageId="10" pageNumber="367">Fig. 9J</figureCitation>
). Total length 13.06. Carapace 6.63 long, 4.17 wide; opisthosoma 6.93 long, 4.16 wide. Eye sizes and interdistances: AME 0.18, ALE 0.27, PME 0.23, PLE 0.24; AME–AME 0.10, AME– ALE 0.12, PME–PME 0.17, PME–PLE 0.26, ALE–PLE 0.11. MOA 0.73 long, front width 0.46, back width 0.68. Clypeus height 0.22. Leg measurements: I 16.22 (4.53, 5.59, 3.64, 2.46); II 14.32 (4.19, 4.53, 3.24, 2.36); III 13.04 (3.75, 4.22, 3.14, 1.93); IV 17.53 (4.80, 5.75, 4.68, 2.30). Spination of legs: femur I-II 210, III 600, IV 400; patella I-III 010, IV 000; tibia I-II 026, III-IV 046; metatarsus I 026, II 046, III 026, IV 426.
</paragraph>
<paragraph id="8BA0403A861DF95DFF20FEB7D15683DB" blockId="10.[151,1437,151,538]" pageId="10" pageNumber="367">
Epigyne (
<figureCitation id="13245CBF861DF95DFED5FEB7D5F283A7" box="[306,424,367,393]" captionStart="FIGURE 6" captionStartId="9.[151,250,1823,1847]" captionTargetBox="[151,1432,453,1795]" captionTargetId="figure-151@9.[151,1435,453,1800]" captionTargetPageId="9" captionText="FIGURE 6. Tonsilla subtruculenta sp. nov. A. Male holotype, left palp, prolateral view; B. Same, ventral view; C. Same, retrolateral view; D. Epigyne, ventral view; E. Same, dorsal view. Abbreviations: CD, copulatory duct; CDA, Conductor’s dorsal apophysis; CF, cymbial furrow; CL, conductor lamella; Co, conductor; CO, copulatory opening; EH, epigynal hood; Em, embolus; ET, epigynal teeth; FD, fertilization ducts; LTA, lateral tibial apophysis; MA, median apophysis; PA, patellar apophysis; RTA, retrolateral tibial apophysis; SH, spermathecal heads; Sp, spermathecae." figureDoi="http://doi.org/10.5281/zenodo.5843804" httpUri="https://zenodo.org/record/5843804/files/figure.png" pageId="10" pageNumber="367">Fig. 6D, E</figureCitation>
). Epigynal plate almost round, teeth long, reaching ventro-mesally. Epigynal hoods shallow, distinct, located ventro-mesally.Atrial ridges distinct and round.Atrium posteriorly divided with septum. Copulatory opening situated around the septum. Copulatory ducts oval. Spermathecal head extending anteriorly, curved around the copulatory ducts. Spermathecae robust, longer than wide, slightly curved, situated ventro-mesally.
</paragraph>
</subSubSection>
<subSubSection id="C30513B1861DF95DFF20FE26D6CE8037" box="[199,660,510,538]" pageId="10" pageNumber="367" type="distribution">
<paragraph id="8BA0403A861DF95DFF20FE26D6CE8037" blockId="10.[151,1437,151,538]" box="[199,660,510,538]" pageId="10" pageNumber="367">
<emphasis id="B96B9C28861DF95DFF20FE26D53B8036" bold="true" box="[199,353,510,536]" pageId="10" pageNumber="367">Distribution.</emphasis>
<collectingCountry id="F30800AA861DF95DFE80FE27D5F08037" box="[359,426,511,537]" name="China" pageId="10" pageNumber="367">China</collectingCountry>
(
<collectingRegion id="49DB8ED8861DF95DFE5EFE27D6478034" box="[441,541,511,538]" country="China" name="Guizhou" pageId="10" pageNumber="367">Guizhou</collectingRegion>
) (
<figureCitation id="13245CBF861DF95DFDD3FE27D6DD8037" box="[564,647,511,537]" captionStart="FIGURE 10" captionStartId="14.[151,250,1414,1438]" captionTargetBox="[167,1422,187,1386]" captionTargetId="figure-25@14.[161,1426,181,1390]" captionTargetPageId="14" captionText="FIGURE 10. Distribution of Tonsilla species treated here." figureDoi="http://doi.org/10.5281/zenodo.5843816" httpUri="https://zenodo.org/record/5843816/files/figure.png" pageId="10" pageNumber="367">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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