413 lines
42 KiB
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413 lines
42 KiB
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<mods:title id="9CAD5CFDBC219E6C74CFF4895D0934B9">Systematic revision of Acanthodactylus busacki (Squamata: Lacertidae) with a description of a new species from Morocco</mods:title>
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<heading id="D0C36C7FFFC26B19FF19FC9AECC5FCBB" bold="true" box="[151,643,834,860]" fontSize="11" level="1" pageId="10" pageNumber="367" reason="1">
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<taxonomicName id="4C34A090FFC26B19FF19FC9AECC5FCBB" authority="Salvador, 1982" authorityName="Salvador" authorityYear="1982" box="[151,643,834,860]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="367" phylum="Chordata" rank="species" species="busacki">
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<emphasis id="B9400701FFC26B19FF19FC9AECC5FCBB" bold="true" box="[151,643,834,860]" pageId="10" pageNumber="367">
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<emphasis id="B9400701FFC26B19FF19FC9AEF83FCBB" bold="true" box="[151,453,834,860]" italics="true" pageId="10" pageNumber="367">Acanthodactylus busacki</emphasis>
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<bibRefCitation id="EFA5A6E2FFC26B19FE42FC9AECC5FCBB" author="Salvador" box="[460,643,834,860]" pageId="10" pageNumber="367" refString="Salvador, A. (1982) A revision of the lizards of the genus Acanthodactylus (Sauria: Lacertidae). Bonner Zoologisches Monographien, Nr. 16, Zoologisches Forschungsinstitut und Museum Alexander Koenig, 1 - 167." type="journal article" year="1982">Salvador, 1982</bibRefCitation>
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<paragraph id="8B8BDB13FFC26B19FF19FC51EC1FFC59" blockId="10.[151,1436,905,958]" pageId="10" pageNumber="367">
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<taxonomicName id="4C34A090FFC26B19FF19FC51EFD5FC78" authorityName="Tamar & Geniez & Brito & Crochet" authorityYear="2017" box="[151,403,905,927]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="367" phylum="Chordata" rank="species" species="busacki">
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<emphasis id="B9400701FFC26B19FF19FC51EFD5FC78" box="[151,403,905,927]" italics="true" pageId="10" pageNumber="367">Acanthodactylus busacki</emphasis>
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</taxonomicName>
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,
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<bibRefCitation id="EFA5A6E2FFC26B19FE2FFC51EC7AFC78" author="Salvador" box="[417,572,905,927]" pageId="10" pageNumber="367" refString="Salvador, A. (1982) A revision of the lizards of the genus Acanthodactylus (Sauria: Lacertidae). Bonner Zoologisches Monographien, Nr. 16, Zoologisches Forschungsinstitut und Museum Alexander Koenig, 1 - 167." type="journal article" year="1982">
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<collectingCountry id="F3239B83FFC26B19FE2FFC51EFBAFC78" box="[417,508,905,927]" name="El Salvador" pageId="10" pageNumber="367">Salvador</collectingCountry>
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, 1982
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</bibRefCitation>
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, A revision of the lizards of the genus
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<taxonomicName id="4C34A090FFC26B19FC44FC51EA35FC78" authorityName="Fitzinger" authorityYear="1834" box="[970,1139,905,927]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="367" phylum="Chordata" rank="genus">
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<emphasis id="B9400701FFC26B19FC44FC51EA35FC78" box="[970,1139,905,927]" italics="true" pageId="10" pageNumber="367">Acanthodactylus</emphasis>
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</taxonomicName>
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(Sauria:
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<taxonomicName id="4C34A090FFC26B19FB5FFC51EB06FC78" box="[1233,1344,905,927]" class="Reptilia" family="Lacertidae" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="367" phylum="Chordata" rank="family">Lacertidae</taxonomicName>
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), Bonner Zoologische Monographien, Nr. 16: 88.
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</paragraph>
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<subSubSection id="C32E8898FFC26B19FF19FC32ECDBFBF8" box="[151,669,1002,1055]" pageId="10" pageNumber="367" type="description">
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<paragraph id="8B8BDB13FFC26B19FF19FC32EF50FBE7" blockId="10.[151,1436,1002,1270]" box="[151,278,1002,1024]" pageId="10" pageNumber="367">Chresonyms</paragraph>
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<paragraph id="8B8BDB13FFC26B19FF19FBD1ECDBFBF8" blockId="10.[151,1436,1002,1270]" box="[151,669,1033,1055]" pageId="10" pageNumber="367">
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<treatmentCitationGroup id="AB24FC3DFFC26B19FF19FBD1ECDBFBF8" box="[151,669,1033,1055]" pageId="10" pageNumber="367">
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<taxonomicName id="4C34A090FFC26B19FF19FBD1EC1EFBF8" authority="Salvador, 1982: 88" authorityName="Salvador" authorityPageNumber="88" authorityYear="1982" box="[151,600,1033,1055]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="367" phylum="Chordata" rank="species" species="busacki">
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<emphasis id="B9400701FFC26B19FF19FBD1EFD5FBF8" box="[151,403,1033,1055]" italics="true" pageId="10" pageNumber="367">Acanthodactylus busacki</emphasis>
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<treatmentCitation id="0A95FD02FFC26B19FE14FBD1EC1EFBF8" author="Salvador" box="[410,600,1033,1055]" page="88" pageId="10" pageNumber="367" year="1982">
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<bibRefCitation id="EFA5A6E2FFC26B19FE14FBD1EC73FBF8" author="Salvador" box="[410,565,1033,1055]" pageId="10" pageNumber="367" refString="Salvador, A. (1982) A revision of the lizards of the genus Acanthodactylus (Sauria: Lacertidae). Bonner Zoologisches Monographien, Nr. 16, Zoologisches Forschungsinstitut und Museum Alexander Koenig, 1 - 167." type="journal article" year="1982">Salvador, 1982</bibRefCitation>
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: 88
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</treatmentCitation>
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</taxonomicName>
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(part.)
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</treatmentCitationGroup>
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</paragraph>
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<subSubSection id="C32E8898FFC26B19FF19FBF0EF59FAB8" pageId="10" pageNumber="367" type="reference_group">
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<paragraph id="8B8BDB13FFC26B19FF19FBF0ED41FBD9" blockId="10.[151,1436,1002,1270]" box="[151,775,1064,1086]" pageId="10" pageNumber="367">
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<treatmentCitationGroup id="AB24FC3DFFC26B19FF19FBF0ED41FBD9" box="[151,775,1064,1086]" pageId="10" pageNumber="367">
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<taxonomicName id="4C34A090FFC26B19FF19FBF0EC85FBD9" authority="Arnold, 1983: 319" authorityName="Arnold" authorityPageNumber="319" authorityYear="1983" box="[151,707,1064,1086]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="367" phylum="Chordata" rank="subSpecies" species="pardalis" subSpecies="bedriagai">
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<emphasis id="B9400701FFC26B19FF19FBF0EC44FBD9" box="[151,514,1064,1086]" italics="true" pageId="10" pageNumber="367">Acanthodactylus pardalis bedriagai</emphasis>
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<treatmentCitation id="0A95FD02FFC26B19FD86FBF0EC85FBD9" author="Arnold" box="[520,707,1064,1086]" page="319" pageId="10" pageNumber="367" year="1983">
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<bibRefCitation id="EFA5A6E2FFC26B19FD86FBF0ECD5FBD9" author="Arnold" box="[520,659,1064,1086]" pageId="10" pageNumber="367" refString="Arnold, E. N. (1983) Osteology, genitalia and the relationships of Acanthodactylus (Reptilia: Lacertidae). Bulletin of the British Museum (Natural History), Zoology, 44, 291 - 339." type="journal article" year="1983">Arnold, 1983</bibRefCitation>
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: 319
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</treatmentCitation>
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</taxonomicName>
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(part.)
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B8BDB13FFC26B19FF19FB9EED41FBBB" blockId="10.[151,1436,1002,1270]" box="[151,775,1094,1116]" pageId="10" pageNumber="367">
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<treatmentCitationGroup id="AB24FC3DFFC26B19FF19FB9EED41FBBB" box="[151,775,1094,1116]" pageId="10" pageNumber="367">
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<taxonomicName id="4C34A090FFC26B19FF19FB9EEC84FBBB" authority="Harris & Arnold 2000: 352" authorityName="Harris & Arnold" authorityPageNumber="352" authorityYear="2000" box="[151,706,1094,1116]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="367" phylum="Chordata" rank="species" species="bedriagai">
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<emphasis id="B9400701FFC26B19FF19FB9EEFEFFBBB" box="[151,425,1094,1116]" italics="true" pageId="10" pageNumber="367">Acanthodactylus bedriagai</emphasis>
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<treatmentCitation id="0A95FD02FFC26B19FE21FB9EEC84FBBB" author="Harris" box="[431,706,1094,1116]" page="352" pageId="10" pageNumber="367" year="2000">
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<bibRefCitation id="EFA5A6E2FFC26B19FE21FB9EECD5FBBB" author="Harris" box="[431,659,1094,1116]" pageId="10" pageNumber="367" refString="Harris, D. J. & Arnold, E. N. (2000) Elucidation of the relationships of spiny footed lizards, Acanthodactylus spp. (Reptilia: Lacertidae) using mitochondrial DNA sequence, with comments on their biogeography and evolution. Journal of Zoology, 252, 351 - 362." type="journal article" year="2000">Harris & Arnold 2000</bibRefCitation>
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: 352
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</treatmentCitation>
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</taxonomicName>
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(part.)
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B8BDB13FFC26B19FF19FBBDEC9CFB11" blockId="10.[151,1436,1002,1270]" pageId="10" pageNumber="367">
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<treatmentCitationGroup id="AB24FC3DFFC26B19FF19FBBDEC9CFB11" pageId="10" pageNumber="367">
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<taxonomicName id="4C34A090FFC26B19FF19FBBDECFFFB9C" authority="Mellado & Dakki 1988: 175" authorityName="Mellado & Dakki" authorityPageNumber="175" authorityYear="1988" box="[151,697,1125,1147]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="367" phylum="Chordata" rank="species" species="busacki">
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<emphasis id="B9400701FFC26B19FF19FBBDEFD5FB9C" box="[151,403,1125,1147]" italics="true" pageId="10" pageNumber="367">Acanthodactylus busacki</emphasis>
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<treatmentCitation id="0A95FD02FFC26B19FE14FBBDECFFFB9C" author="Mellado" box="[410,697,1125,1147]" page="175" pageId="10" pageNumber="367" year="1988">
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<bibRefCitation id="EFA5A6E2FFC26B19FE14FBBDECCFFB9C" author="Mellado" box="[410,649,1125,1147]" pageId="10" pageNumber="367" refString="Mellado, J. & Dakki, M. (1988) Inventaire commente des amphibiens et reptiles du Maroc. Bulletin de l'Institut Scientifique, Rabat, 12, 171 - 181." type="journal article" year="1988">Mellado & Dakki 1988</bibRefCitation>
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: 175
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</treatmentCitation>
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</taxonomicName>
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(part.);
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<treatmentCitation id="0A95FD02FFC26B19FC82FBBDEA06FB9C" author="Mellado" box="[780,1088,1125,1147]" page="133" pageId="10" pageNumber="367" year="1990">
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<bibRefCitation id="EFA5A6E2FFC26B19FC82FBBDEA56FB9C" author="Mellado" box="[780,1040,1125,1147]" pageId="10" pageNumber="367" refString="Mellado, J. & Olmedo, G. (1990) El genero Acanthodactylus en Marruecos: problemas de identificacion en los grupos de especies A. pardalis y A. scutellatus. Amphibia-Reptilia, 11, 131 - 146." type="journal article" year="1990">Mellado & Olmedo 1990</bibRefCitation>
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: 133
|
|||
|
|
</treatmentCitation>
|
|||
|
|
(part.);
|
|||
|
|
<treatmentCitation id="0A95FD02FFC26B19FB1CFBBEEBDDFB9C" author="Bons" box="[1170,1435,1125,1147]" page="162" pageId="10" pageNumber="367" year="1996">
|
|||
|
|
<bibRefCitation id="EFA5A6E2FFC26B19FB1CFBBEEB2DFB9C" author="Bons" box="[1170,1387,1125,1147]" pageId="10" pageNumber="367" refString="Bons, J. & Geniez, P. (1996) Amphibiens et reptiles du Maroc. Asociacion Herpetologica Espanola, Barcelona, 319 pp." type="book" year="1996">Bons & Geniez 1996</bibRefCitation>
|
|||
|
|
: 162
|
|||
|
|
</treatmentCitation>
|
|||
|
|
(part.);
|
|||
|
|
<treatmentCitation id="0A95FD02FFC26B19FE9BFB5CEC5AFB7D" author="Schleich" box="[277,540,1156,1178]" page="391" pageId="10" pageNumber="367" year="1996">
|
|||
|
|
<bibRefCitation id="EFA5A6E2FFC26B19FE9BFB5CEFAFFB7D" author="Schleich" box="[277,489,1156,1178]" pageId="10" pageNumber="367" refString="Schleich, H. H., Kastle, W. & Kabisch, K. (1996) Amphibians and reptiles of North Africa. Koeltz, Koenigstein, 627 pp." type="book" year="1996">
|
|||
|
|
Schleich
|
|||
|
|
<emphasis id="B9400701FFC26B19FEFBFB5DEFEDFB7D" box="[373,427,1156,1178]" italics="true" pageId="10" pageNumber="367">et al.</emphasis>
|
|||
|
|
1996
|
|||
|
|
</bibRefCitation>
|
|||
|
|
: 391
|
|||
|
|
</treatmentCitation>
|
|||
|
|
(part.);
|
|||
|
|
<treatmentCitation id="0A95FD02FFC26B19FDFDFB5CED2FFB7D" author="Donaire" box="[627,873,1156,1178]" page="10" pageId="10" pageNumber="367" year="2000">
|
|||
|
|
<bibRefCitation id="EFA5A6E2FFC26B19FDFDFB5CED04FB7D" author="Donaire" box="[627,834,1156,1178]" pageId="10" pageNumber="367" refString="Donaire, D., Mateo, J. A., Hasi, M. & Geniez, P. (2000) Nuevos datos sobre la fauna reptiliana de la Hamada de Tinduf (Argelia). Boletin de la Asociacion Herpetologica Espanola, 11, 8 - 12." type="journal article" year="2000">
|
|||
|
|
Donaire
|
|||
|
|
<emphasis id="B9400701FFC26B19FD43FB5DED45FB7D" box="[717,771,1156,1178]" italics="true" pageId="10" pageNumber="367">et al.</emphasis>
|
|||
|
|
2000
|
|||
|
|
</bibRefCitation>
|
|||
|
|
: 10
|
|||
|
|
</treatmentCitation>
|
|||
|
|
;
|
|||
|
|
<treatmentCitation id="0A95FD02FFC26B19FCF9FB5CEA28FB7D" author="Geniez" box="[887,1134,1156,1178]" page="102" pageId="10" pageNumber="367" year="2004">
|
|||
|
|
<bibRefCitation id="EFA5A6E2FFC26B19FCF9FB5CEA7DFB7D" author="Geniez" box="[887,1083,1156,1178]" pageId="10" pageNumber="367" refString="Geniez, P., Mateo, J. A., Geniez, M. & Pether, J. (2004) The amphibians and reptiles of the Western Sahara (former Spanish Sahara) and adjacent regions. Edition Chimaira, Frankfurt, 228 pp." type="book" year="2004">
|
|||
|
|
Geniez
|
|||
|
|
<emphasis id="B9400701FFC26B19FC49FB5DEDBBFB7D" box="[967,1021,1156,1178]" italics="true" pageId="10" pageNumber="367">et al.</emphasis>
|
|||
|
|
2004
|
|||
|
|
</bibRefCitation>
|
|||
|
|
: 102
|
|||
|
|
</treatmentCitation>
|
|||
|
|
(part.); Brito
|
|||
|
|
<emphasis id="B9400701FFC26B19FA8CFB5DEB7EFB7D" box="[1282,1336,1156,1178]" italics="true" pageId="10" pageNumber="367">et al.</emphasis>
|
|||
|
|
2008: 21 (part.);
|
|||
|
|
<treatmentCitation id="0A95FD02FFC26B19FE9DFB7AEC28FB5F" author="Sindaco" box="[275,622,1186,1208]" page="218" pageId="10" pageNumber="367" year="2008">
|
|||
|
|
<bibRefCitation id="EFA5A6E2FFC26B19FE9DFB7AEC7BFB5F" author="Sindaco" box="[275,573,1186,1208]" pageId="10" pageNumber="367" refString="Sindaco, R. & Jeremcenko, V. K. (2008) The reptiles of the Western Palearctic. Annotated checklist and distributional atlas of the turtles, crocodiles, amphisbaenians and lizards of Europe, North Africa, Middle East and Central Asia. Monografie della Societas Herpetologica Italica. Edizioni Belvedere, Latina, 579 pp." type="book" year="2008">Sindaco & Jeremčenko 2008</bibRefCitation>
|
|||
|
|
: 218
|
|||
|
|
</treatmentCitation>
|
|||
|
|
(part.);
|
|||
|
|
<treatmentCitation id="0A95FD02FFC26B19FD4CFB7BEDE2FB5F" author="Fonseca" box="[706,932,1186,1208]" page="9" pageId="10" pageNumber="367" year="2008">
|
|||
|
|
<bibRefCitation id="EFA5A6E2FFC26B19FD4CFB7BEDCBFB5F" author="Fonseca" box="[706,909,1186,1208]" pageId="10" pageNumber="367" refString="Fonseca, M. M., Brito, J. C., Rebelo, H., Kalboussi, M., Larbes, S., Carretero, M. A. & Harris, D. J. (2008) Genetic variation among spiny-footed lizards in the Acanthodactylus pardalis group from North Africa. African Zoology, 43, 8 - 15. https: // doi. org / 10.1080 / 15627020.2008.11407401" type="journal article" year="2008">
|
|||
|
|
Fonseca
|
|||
|
|
<emphasis id="B9400701FFC26B19FC92FB7BED17FB5F" box="[796,849,1186,1208]" italics="true" pageId="10" pageNumber="367">et al.</emphasis>
|
|||
|
|
2008
|
|||
|
|
</bibRefCitation>
|
|||
|
|
: 9
|
|||
|
|
</treatmentCitation>
|
|||
|
|
(part.);
|
|||
|
|
<bibRefCitation id="EFA5A6E2FFC26B19FC77FB7AEAFCFB5F" author="Geniez" box="[1017,1210,1186,1208]" pageId="10" pageNumber="367" refString="Geniez, P., Mateo Miras, J. A., Joger, U., Pleguezuelos, J., Slimani, T. & El Mouden, E. H. (2010) Acanthodactylus busacki. The IUCN Red List of Threatened Species 2010: e. T 178569 A 7572735. Available from: http: // www. iucnredlist. org / details / 178569 / 0 (accessed 24 April 2017)" type="journal article" year="2010">
|
|||
|
|
Geniez
|
|||
|
|
<emphasis id="B9400701FFC26B19FBC6FB7BEA3AFB5F" box="[1096,1148,1186,1208]" italics="true" pageId="10" pageNumber="367">et al.</emphasis>
|
|||
|
|
2010
|
|||
|
|
</bibRefCitation>
|
|||
|
|
: IUCN red list (part.); Harris
|
|||
|
|
<emphasis id="B9400701FFC26B19FE82FB19EF06FB30" box="[268,320,1217,1239]" italics="true" pageId="10" pageNumber="367">el al.</emphasis>
|
|||
|
|
2010: 22 (part.);
|
|||
|
|
<treatmentCitation id="0A95FD02FFC26B19FE7CFB19ECB0FB30" author="Carretero" box="[498,758,1217,1239]" page="139" pageId="10" pageNumber="367" year="2011">
|
|||
|
|
<bibRefCitation id="EFA5A6E2FFC26B19FE7CFB19EC80FB30" author="Carretero" box="[498,710,1217,1239]" pageId="10" pageNumber="367" refString="Carretero, M. A., Fonseca, M. M., Garcia-Munoz, E., Brito, J. C. & Harris, D. J. (2011) Adding Acanthodactylus beershebensis to the mtDNA phylogeny of the Acanthodactylus pardalis group. North-Western Journal of Zoology, 7, 138 - 142." type="journal article" year="2011">
|
|||
|
|
Carretero
|
|||
|
|
<emphasis id="B9400701FFC26B19FDD9FB1AECCDFB30" box="[599,651,1217,1239]" italics="true" pageId="10" pageNumber="367">et al.</emphasis>
|
|||
|
|
2011
|
|||
|
|
</bibRefCitation>
|
|||
|
|
: 139
|
|||
|
|
</treatmentCitation>
|
|||
|
|
(part.);
|
|||
|
|
<treatmentCitation id="0A95FD02FFC26B19FCC6FB1AEA6EFB30" author="Trape" box="[840,1064,1217,1239]" page="302" pageId="10" pageNumber="367" year="2012">
|
|||
|
|
<bibRefCitation id="EFA5A6E2FFC26B19FCC6FB1AEDBFFB30" author="Trape" box="[840,1017,1217,1239]" pageId="10" pageNumber="367" refString="Trape, J. - F., Trape, S. & Chirio, L. (2012) Lezards, crocodiles et tortues d'Afrique occidentale et du Sahara. IRD Orstom, Marseille, 503 pp." type="book" year="2012">
|
|||
|
|
Trape
|
|||
|
|
<emphasis id="B9400701FFC26B19FC07FB1AEDFAFB30" box="[905,956,1217,1239]" italics="true" pageId="10" pageNumber="367">et al.</emphasis>
|
|||
|
|
2012
|
|||
|
|
</bibRefCitation>
|
|||
|
|
: 302
|
|||
|
|
</treatmentCitation>
|
|||
|
|
(part.);
|
|||
|
|
<treatmentCitation id="0A95FD02FFC26B19FBF5FB1AEB16FB30" author="Pyron" box="[1147,1360,1217,1239]" page="17" pageId="10" pageNumber="367" year="2013">
|
|||
|
|
<bibRefCitation id="EFA5A6E2FFC26B19FBF5FB1AEB6AFB30" author="Pyron" box="[1147,1324,1217,1239]" pageId="10" pageNumber="367" refString="Pyron, R. A., Burbrink, F. T. & Wiens, J. J. (2013) A phylogeny and revised classification of Squamata, including 4161 species of lizards and snakes. BMC Evolutionary Biology, 13, 93." type="journal article" year="2013">
|
|||
|
|
Pyron
|
|||
|
|
<emphasis id="B9400701FFC26B19FB33FB1AEAB6FB30" box="[1213,1264,1217,1239]" italics="true" pageId="10" pageNumber="367">et al.</emphasis>
|
|||
|
|
2013
|
|||
|
|
</bibRefCitation>
|
|||
|
|
: 17
|
|||
|
|
</treatmentCitation>
|
|||
|
|
(part.);
|
|||
|
|
<treatmentCitation id="0A95FD02FFC26B19FF49FB38EFF9FB11" author="Crochet" box="[199,447,1248,1270]" page="584" pageId="10" pageNumber="367" year="2015">
|
|||
|
|
<bibRefCitation id="EFA5A6E2FFC26B19FF49FB38EFCBFB11" author="Crochet" box="[199,397,1248,1270]" pageId="10" pageNumber="367" refString="Crochet, P. - A., Leblois, R. & Renoult, J. P. (2015) New reptile records from Morocco and Western Sahara. Herpetology Notes, 8, 583 - 588." type="journal article" year="2015">
|
|||
|
|
Crochet
|
|||
|
|
<emphasis id="B9400701FFC26B19FE93FB39EF17FB11" box="[285,337,1248,1270]" italics="true" pageId="10" pageNumber="367">et al.</emphasis>
|
|||
|
|
2015
|
|||
|
|
</bibRefCitation>
|
|||
|
|
: 584
|
|||
|
|
</treatmentCitation>
|
|||
|
|
;
|
|||
|
|
<treatmentCitation id="0A95FD02FFC26B19FE47FB39ECD0FB11" author="Tamar" box="[457,662,1248,1270]" page="8" pageId="10" pageNumber="367" year="2016">
|
|||
|
|
<bibRefCitation id="EFA5A6E2FFC26B19FE47FB39ECC6FB11" author="Tamar" box="[457,640,1248,1270]" pageId="10" pageNumber="367" refString="Tamar, K., Carranza, S., Sindaco, R., Moravec, J., Trape, J. - F. & Meiri, S. (2016) Out of Africa: Phylogeny and biogeography of the widespread genus Acanthodactylus (Reptilia: Lacertidae). Molecular Phylogenetics and Evolution, 103, 6 - 18. https: // doi. org / 10.1016 / j. ympev. 2016.07.003" type="journal article" year="2016">
|
|||
|
|
Tamar
|
|||
|
|
<emphasis id="B9400701FFC26B19FD9EFB39EC02FB11" box="[528,580,1248,1270]" italics="true" pageId="10" pageNumber="367">et al.</emphasis>
|
|||
|
|
2016
|
|||
|
|
</bibRefCitation>
|
|||
|
|
: 8
|
|||
|
|
</treatmentCitation>
|
|||
|
|
(part.)
|
|||
|
|
</treatmentCitationGroup>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B8BDB13FFC26B19FF19FAFAEF59FAB8" blockId="10.[151,1437,1314,2023]" pageId="10" pageNumber="367">
|
|||
|
|
<emphasis id="B9400701FFC26B19FF19FAFAEFCCFADC" bold="true" box="[151,394,1314,1339]" pageId="10" pageNumber="367">Name-bearing type.</emphasis>
|
|||
|
|
Natural History Museum (London) BMNH1970.250, adult male, holotype by original designation.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C32E8898FFC26B19FF49FAB2EF9FF999" pageId="10" pageNumber="367" type="materials_examined">
|
|||
|
|
<paragraph id="8B8BDB13FFC26B19FF49FAB2EDA3FA65" blockId="10.[151,1437,1314,2023]" box="[199,997,1386,1411]" pageId="10" pageNumber="367">
|
|||
|
|
<materialsCitation id="3B5CD14EFFC26B19FF49FAB2EDA3FA65" ID-GBIF-Occurrence="1563872023" box="[199,997,1386,1411]" country="Morocco" location="Goulimine" municipality="Guelmim" pageId="10" pageNumber="367" specimenCount="1" typeStatus="holotype">
|
|||
|
|
<emphasis id="B9400701FFC26B19FF49FAB2EF23FA64" bold="true" box="[199,357,1386,1411]" pageId="10" pageNumber="367">
|
|||
|
|
<typeStatus id="548F65B1FFC26B19FF49FAB2EF44FA64" box="[199,258,1386,1411]" pageId="10" pageNumber="367">Type</typeStatus>
|
|||
|
|
locality.
|
|||
|
|
</emphasis>
|
|||
|
|
<quantity id="4CCC76F6FFC26B19FEE2FAB3EFF5FA64" box="[364,435,1386,1411]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="367" unit="km" value="30.0">30 km</quantity>
|
|||
|
|
south-west of
|
|||
|
|
<location id="8EEB8DC8FFC26B19FDD6FAB2EC96FA64" LSID="urn:lsid:plazi:treatment:039D6A05FFC26B18FF19FC9AEA9EFCBA:8EEB8DC8FFC26B19FDD6FAB2EC96FA64" box="[600,720,1386,1411]" country="Morocco" municipality="Guelmim" name="Goulimine" pageId="10" pageNumber="367">Goulimine</location>
|
|||
|
|
(=
|
|||
|
|
<collectingMunicipality id="6BEF4169FFC26B19FD79FAB2ED23FA64" box="[759,869,1386,1411]" pageId="10" pageNumber="367">Guelmim</collectingMunicipality>
|
|||
|
|
),
|
|||
|
|
<collectingCountry id="F3239B83FFC26B19FCF6FAB3EDA7FA65" box="[888,993,1387,1410]" name="Morocco" pageId="10" pageNumber="367">Morocco</collectingCountry>
|
|||
|
|
.
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B8BDB13FFC26B19FF49FA56EFF5F9F4" blockId="10.[151,1437,1314,2023]" pageId="10" pageNumber="367">
|
|||
|
|
<materialsCitation id="3B5CD14EFFC26B19FF49FA56EFF5F9F4" ID-GBIF-Occurrence="1563872022" collectionCode="BMNH, EBD" collectorName="Cape Bojador & Rio de Oro & Other & Appendix II" country="Morocco" pageId="10" pageNumber="367" specimenCode="BMNH1970.249, EBD2440" specimenCount="4" typeStatus="paratype">
|
|||
|
|
<emphasis id="B9400701FFC26B19FF49FA56EF00FA40" bold="true" box="[199,326,1422,1447]" pageId="10" pageNumber="367">
|
|||
|
|
<typeStatus id="548F65B1FFC26B19FF49FA56EF07FA40" box="[199,321,1422,1447]" pageId="10" pageNumber="367" type="paratype">Paratypes</typeStatus>
|
|||
|
|
.
|
|||
|
|
</emphasis>
|
|||
|
|
Among the
|
|||
|
|
<typeStatus id="548F65B1FFC26B19FE61FA57EC1CFA40" box="[495,602,1423,1447]" pageId="10" pageNumber="367" type="paratype">paratypes</typeStatus>
|
|||
|
|
originally listed by
|
|||
|
|
<bibRefCitation id="EFA5A6E2FFC26B19FCEBFA56EA61FA40" author="Salvador" box="[869,1063,1422,1447]" pageId="10" pageNumber="367" refString="Salvador, A. (1982) A revision of the lizards of the genus Acanthodactylus (Sauria: Lacertidae). Bonner Zoologisches Monographien, Nr. 16, Zoologisches Forschungsinstitut und Museum Alexander Koenig, 1 - 167." type="journal article" year="1982">Salvador (1982)</bibRefCitation>
|
|||
|
|
, MNHN-RA-1938.189 (Asrifa,
|
|||
|
|
<collectingCountry id="F3239B83FFC26B19FF19FA6BEF47FA2C" box="[151,257,1459,1483]" name="Morocco" pageId="10" pageNumber="367">Morocco</collectingCountry>
|
|||
|
|
),
|
|||
|
|
<specimenCode id="DB927368FFC26B19FE9AFA6BEF97FA2C" box="[276,465,1459,1483]" collectionCode="BMNH" pageId="10" pageNumber="367">
|
|||
|
|
<collectionCode id="ED2543D6FFC26B19FE9AFA6BEF2FFA2C" box="[276,361,1459,1483]" pageId="10" pageNumber="367">BMNH</collectionCode>
|
|||
|
|
1970.249
|
|||
|
|
</specimenCode>
|
|||
|
|
(same locality as the
|
|||
|
|
<typeStatus id="548F65B1FFC26B19FD45FA6AED74FA2C" box="[715,818,1458,1483]" pageId="10" pageNumber="367" type="holotype">holotype</typeStatus>
|
|||
|
|
), and
|
|||
|
|
<specimenCode id="DB927368FFC26B19FCF8FA6BEDA0FA2C" box="[886,998,1459,1483]" collectionCode="EBD" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="10" pageNumber="367">
|
|||
|
|
<collectionCode id="ED2543D6FFC26B19FCF8FA6BEDEBFA2C" box="[886,941,1459,1483]" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="10" pageNumber="367">EBD</collectionCode>
|
|||
|
|
2440
|
|||
|
|
</specimenCode>
|
|||
|
|
(
|
|||
|
|
<collectorName id="26C1BEC5FFC26B19FC78FA6BEAD3FA2C" box="[1014,1173,1458,1483]" pageId="10" pageNumber="367">Cape Bojador</collectorName>
|
|||
|
|
,
|
|||
|
|
<collectorName id="26C1BEC5FFC26B19FB2CFA6AEB64FA2C" box="[1186,1314,1458,1483]" pageId="10" pageNumber="367">Rio de Oro</collectorName>
|
|||
|
|
) belong to this species.
|
|||
|
|
<collectorName id="26C1BEC5FFC26B19FEA5FA0EEF2DFA08" box="[299,363,1494,1519]" pageId="10" pageNumber="367">Other</collectorName>
|
|||
|
|
<typeStatus id="548F65B1FFC26B19FEFBFA0FEFA6FA08" box="[373,480,1495,1519]" pageId="10" pageNumber="367" type="paratype">paratypes</typeStatus>
|
|||
|
|
listed in
|
|||
|
|
<bibRefCitation id="EFA5A6E2FFC26B19FDDFFA0EED4FFA08" author="Salvador" box="[593,777,1494,1519]" pageId="10" pageNumber="367" refString="Salvador, A. (1982) A revision of the lizards of the genus Acanthodactylus (Sauria: Lacertidae). Bonner Zoologisches Monographien, Nr. 16, Zoologisches Forschungsinstitut und Museum Alexander Koenig, 1 - 167." type="journal article" year="1982">
|
|||
|
|
<collectingCountry id="F3239B83FFC26B19FDDFFA0EECF3FA08" box="[593,693,1494,1519]" name="El Salvador" pageId="10" pageNumber="367">Salvador</collectingCountry>
|
|||
|
|
(1982)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
are allocated to the new species (see below) or to
|
|||
|
|
<taxonomicName id="4C34A090FFC26B19FAD0FA0FEF4CF9F4" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="367" phylum="Chordata" rank="species" species="maculatus">
|
|||
|
|
<emphasis id="B9400701FFC26B19FAD0FA0FEF4CF9F4" italics="true" pageId="10" pageNumber="367">A. cf. maculatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<collectorName id="26C1BEC5FFC26B19FE96FA22EFE0F9F4" box="[280,422,1530,1555]" pageId="10" pageNumber="367">Appendix II</collectorName>
|
|||
|
|
).
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B8BDB13FFC26B19FF49F9C6EF9FF999" blockId="10.[151,1437,1314,2023]" pageId="10" pageNumber="367">
|
|||
|
|
<emphasis id="B9400701FFC26B19FF49F9C6EFC5F9D0" bold="true" box="[199,387,1566,1591]" pageId="10" pageNumber="367">Other material.</emphasis>
|
|||
|
|
Ten voucher specimens listed in Appendix II under
|
|||
|
|
<taxonomicName id="4C34A090FFC26B19FC54F9C7EA09F9D1" authorityName="Mellado & Dakki" authorityYear="1988" box="[986,1103,1566,1590]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="367" phylum="Chordata" rank="species" species="busacki">
|
|||
|
|
<emphasis id="B9400701FFC26B19FC54F9C7EA09F9D1" box="[986,1103,1566,1590]" italics="true" pageId="10" pageNumber="367">A. busacki</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, apart from the holotype and paratypes. Photographic material of one voucher specimen from MB and of 25 individuals photographed in the wild is listed in Appendix II.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C32E8898FFC26B18FF49F952EC91FE6F" lastPageId="11" lastPageNumber="368" pageId="10" pageNumber="367" type="diagnosis">
|
|||
|
|
<paragraph id="8B8BDB13FFC26B18FF49F952EC91FE6F" blockId="10.[151,1437,1314,2023]" lastBlockId="11.[151,1436,151,861]" lastPageId="11" lastPageNumber="368" pageId="10" pageNumber="367">
|
|||
|
|
<emphasis id="B9400701FFC26B19FF49F952EF07F944" bold="true" box="[199,321,1674,1699]" pageId="10" pageNumber="367">Diagnosis.</emphasis>
|
|||
|
|
A species of the
|
|||
|
|
<taxonomicName id="4C34A090FFC26B19FD80F953EC2CF944" box="[526,618,1675,1699]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="367" phylum="Chordata" rank="species" species="pardalis">
|
|||
|
|
<emphasis id="B9400701FFC26B19FD80F953EC2CF944" box="[526,618,1675,1699]" italics="true" pageId="10" pageNumber="367">pardalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species-group (i.e., small flat or carinated dorsal scales; three series of scales on the fingers; three supraoculars; 12 and sometimes 14 straight longitudinal row of ventrals; slightly pectinate toes; body pattern combining longitudinal rows of light ocelli and black reticulation) characterized by the combination of the following characters: (1) maximum recorded SVL
|
|||
|
|
<quantity id="4CCC76F6FFC26B19FC24F92FEDBFF8E9" box="[938,1017,1783,1807]" metricMagnitude="-2" metricUnit="m" metricValue="7.3" pageId="10" pageNumber="367" unit="mm" value="73.0">73 mm</quantity>
|
|||
|
|
(
|
|||
|
|
<quantity id="4CCC76F6FFC26B19FB89F92EEAC7F8E9" box="[1031,1153,1782,1807]" metricMagnitude="-2" metricUnit="m" metricValue="6.2" metricValueMax="7.3" metricValueMin="5.1" pageId="10" pageNumber="367" unit="mm" value="62.0" valueMax="73.0" valueMin="51.0">51–73 mm</quantity>
|
|||
|
|
in adult males); (2) three supraoculars, the first supraocular is either entire or fragmented with usually one row of granules between the supraoculars and the superciliaries; (3) 13–27 granules around the supraoculars; (4) subocular with a distinct keel located between the fourth and fifth upper labials and not contacting the lip; (5) upper temporals small and pointed whereas the lower temporals are large and smooth; (6) 8–11 collar scales; (7) 12 longitudinal rows of ventral scales; (8) 29–34 transverse rows of ventral scales; (9) 20–26 femoral pores on each side; (10) three rows of scales on fingers with slight lateral pectination, with 16–23 lamellae underneath the fourth toe; (11) dorsal scales are pointed and smooth; (12) dorsal colour pattern of juveniles, females and sub-adult males consists of two lateral longitudinal lines of pale ocelli on each side with strong black reticulation among them (four longitudinal dorsal lines in total), adult males have intricate colouration of black reticulation that covers most of the dorsal area and the rows of pale ocelli become indistinct or may completely disappear; (13) the males exhibit a unique colouration: the neck, the posterior part of the throat, the anterior parts of the body and belly, and the forelimbs are more or less reddish; this tint becoming progressively yellowish, whitish or slightly bluish at the posterior parts and tail; (14) sub-adults have reddish or white ventral tail colour.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C32E8898FFC36B18FF49FE4CEA9EFCBA" pageId="11" pageNumber="368" type="distribution">
|
|||
|
|
<paragraph id="8B8BDB13FFC36B18FF49FE4CEF1EFD4F" blockId="11.[151,1436,151,861]" pageId="11" pageNumber="368">
|
|||
|
|
<emphasis id="B9400701FFC36B18FF49FE4CEF26FE4A" bold="true" box="[199,352,404,429]" pageId="11" pageNumber="368">Distribution.</emphasis>
|
|||
|
|
<taxonomicName id="4C34A090FFC36B18FEE7FE4CECC7FE4B" authorityName="Mellado & Dakki" authorityYear="1988" box="[361,641,404,428]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="368" phylum="Chordata" rank="species" species="busacki">
|
|||
|
|
<emphasis id="B9400701FFC36B18FEE7FE4CECC7FE4B" box="[361,641,404,428]" italics="true" pageId="11" pageNumber="368">Acanthodactylus busacki</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is found in the northern Saharan Atlantic coastal desert, from around Guelmim in the north to
|
|||
|
|
<quantity id="4CCC76F6FFC36B18FE38FE6FEFB8FE37" box="[438,510,439,464]" metricMagnitude="4" metricUnit="m" metricValue="6.7" pageId="11" pageNumber="368" unit="km" value="67.0">67 km</quantity>
|
|||
|
|
north-east of Dakhla (
|
|||
|
|
<bibRefCitation id="EFA5A6E2FFC36B18FC8FFE6FEDA7FE37" author="Crochet" box="[769,993,439,464]" pageId="11" pageNumber="368" refString="Crochet, P. - A., Leblois, R. & Renoult, J. P. (2015) New reptile records from Morocco and Western Sahara. Herpetology Notes, 8, 583 - 588." type="journal article" year="2015">
|
|||
|
|
Crochet
|
|||
|
|
<emphasis id="B9400701FFC36B18FCEAFE61EDD8FE37" box="[868,926,440,464]" italics="true" pageId="11" pageNumber="368">et al.</emphasis>
|
|||
|
|
2015
|
|||
|
|
</bibRefCitation>
|
|||
|
|
) in the south (
|
|||
|
|
<figureCitation id="130FC796FFC36B18FB06FE6FEA8BFE37" box="[1160,1229,439,464]" captionStart="FIGURE 1" captionStartId="2.[151,250,1221,1243]" captionTargetBox="[200,1393,351,1196]" captionTargetId="figure@2.[193,1394,339,1196]" captionTargetPageId="2" captionText="FIGURE 1. Localities of the material examined in this study for Acanthodactylus margaritae sp. nov. (green symbols) and A. busacki (orange symbols), with type localities (star), samples used for the genetic analyses (circle), voucher specimens (square), and photographic material (triangle). Numbers correspond to samples listed in Table 1. Taxon names correspond to changes proposed in this paper." httpUri="https://zenodo.org/record/806955/files/figure.png" pageId="11" pageNumber="368">Fig. 1</figureCitation>
|
|||
|
|
; Appendix II). Its range is bordered to the north by the southern extension of the Anti-Atlas Mountains. According to
|
|||
|
|
<bibRefCitation id="EFA5A6E2FFC36B18FA8FFE04EEA7FDFF" pageId="11" pageNumber="368" refString="Donaire, D., Mateo, J. A., Hasi, M. & Geniez, P. (2000) Nuevos datos sobre la fauna reptiliana de la Hamada de Tinduf (Argelia). Boletin de la Asociacion Herpetologica Espanola, 11, 8 - 12." type="journal article">
|
|||
|
|
Donaire
|
|||
|
|
<emphasis id="B9400701FFC36B18FAEDFE05EBDAFE13" box="[1379,1436,476,500]" italics="true" pageId="11" pageNumber="368">et al.</emphasis>
|
|||
|
|
(2000)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
and
|
|||
|
|
<bibRefCitation id="EFA5A6E2FFC36B18FE91FE27EC4FFDFF" box="[287,521,511,536]" pageId="11" pageNumber="368" refString="Geniez, P., Mateo, J. A., Geniez, M. & Pether, J. (2004) The amphibians and reptiles of the Western Sahara (former Spanish Sahara) and adjacent regions. Edition Chimaira, Frankfurt, 228 pp." type="book">
|
|||
|
|
Geniez
|
|||
|
|
<emphasis id="B9400701FFC36B18FEF7FDD9EFF2FDFF" box="[377,436,512,536]" italics="true" pageId="11" pageNumber="368">et al.</emphasis>
|
|||
|
|
(2004)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
the species reaches the Hammada of
|
|||
|
|
<collectingRegion id="49F015F1FFC36B18FC49FE27EA66FDFF" box="[967,1056,511,536]" country="Algeria" name="Tindouf" pageId="11" pageNumber="368">Tindouf</collectingRegion>
|
|||
|
|
in the extreme south-east of the Atlantic Sahara, entering
|
|||
|
|
<collectingCountry id="F3239B83FFC36B18FE4FFDFCEC53FDDA" box="[449,533,548,573]" name="Algeria" pageId="11" pageNumber="368">Algeria</collectingCountry>
|
|||
|
|
and approaching
|
|||
|
|
<collectingCountry id="F3239B83FFC36B18FD67FDFCED21FDDA" box="[745,871,548,573]" name="Mauritania" pageId="11" pageNumber="368">Mauritania</collectingCountry>
|
|||
|
|
, but these records need to be re-examined: they belong to a taxon of the
|
|||
|
|
<taxonomicName id="4C34A090FFC36B18FE39FD90EC52FD87" box="[439,532,584,608]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="368" phylum="Chordata" rank="species" species="pardalis">
|
|||
|
|
<emphasis id="B9400701FFC36B18FE39FD90EC52FD87" box="[439,532,584,608]" italics="true" pageId="11" pageNumber="368">pardalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species-group, but not necessarily to
|
|||
|
|
<taxonomicName id="4C34A090FFC36B18FC45FD90EA07FD87" authorityName="Mellado & Dakki" authorityYear="1988" box="[971,1089,584,608]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="368" phylum="Chordata" rank="species" species="busacki">
|
|||
|
|
<emphasis id="B9400701FFC36B18FC45FD90EA07FD87" box="[971,1089,584,608]" italics="true" pageId="11" pageNumber="368">A. busacki</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; they might instead involve a desert form of
|
|||
|
|
<taxonomicName id="4C34A090FFC36B18FEC8FDB4EC47FD63" box="[326,513,620,644]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="368" phylum="Chordata" rank="species" species="maculatus">
|
|||
|
|
<emphasis id="B9400701FFC36B18FEC8FDB4EC47FD63" box="[326,513,620,644]" italics="true" pageId="11" pageNumber="368">A. cf. maculatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
entering from the northern Sahara into
|
|||
|
|
<collectingCountry id="F3239B83FFC36B18FC5AFDB4EA6EFD62" box="[980,1064,620,645]" name="Algeria" pageId="11" pageNumber="368">Algeria</collectingCountry>
|
|||
|
|
and
|
|||
|
|
<collectingCountry id="F3239B83FFC36B18FBEBFDB5EA8CFD63" box="[1125,1226,621,644]" name="Morocco" pageId="11" pageNumber="368">Morocco</collectingCountry>
|
|||
|
|
(J.A. Mateo pers. com.; pers. obs.).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B8BDB13FFC36B18FF49FD6CEA9EFCBA" blockId="11.[151,1436,151,861]" pageId="11" pageNumber="368">
|
|||
|
|
<emphasis id="B9400701FFC36B18FF49FD6CEFCCFD2A" bold="true" box="[199,394,692,717]" pageId="11" pageNumber="368">Natural history.</emphasis>
|
|||
|
|
<taxonomicName id="4C34A090FFC36B18FE18FD6CECF4FD2B" authorityName="Mellado & Dakki" authorityYear="1988" box="[406,690,692,716]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="368" phylum="Chordata" rank="species" species="busacki">
|
|||
|
|
<emphasis id="B9400701FFC36B18FE18FD6CECF4FD2B" box="[406,690,692,716]" italics="true" pageId="11" pageNumber="368">Acanthodactylus busacki</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is a ground-dwelling, diurnal, oviparous, medium-sized lizard, relatively large and stout-bodied. It mostly inhabits habitats with hard substratum, most often clay deposits, more rarely rocky ground, always with scattered bushes. It is widely sympatric with
|
|||
|
|
<taxonomicName id="4C34A090FFC36B18FBAFFD24EACBFCF3" box="[1057,1165,764,788]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="368" phylum="Chordata" rank="species" species="aureus">
|
|||
|
|
<emphasis id="B9400701FFC36B18FBAFFD24EACBFCF3" box="[1057,1165,764,788]" italics="true" pageId="11" pageNumber="368">A. aureus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
which occurs on sandy substratum (pers. obs.;
|
|||
|
|
<bibRefCitation id="EFA5A6E2FFC36B18FE3EFCC7ECDEFCDF" author="Geniez" box="[432,664,799,824]" pageId="11" pageNumber="368" refString="Geniez, P., Mateo, J. A., Geniez, M. & Pether, J. (2004) The amphibians and reptiles of the Western Sahara (former Spanish Sahara) and adjacent regions. Edition Chimaira, Frankfurt, 228 pp." type="book" year="2004">
|
|||
|
|
Geniez
|
|||
|
|
<emphasis id="B9400701FFC36B18FD80FCF9EC0BFCDF" box="[526,589,800,824]" italics="true" pageId="11" pageNumber="368">et al.</emphasis>
|
|||
|
|
2004
|
|||
|
|
</bibRefCitation>
|
|||
|
|
). Reddish ventral tail colouration of sub-adults is present in all populations of this species, and not only in the southern populations (
|
|||
|
|
<emphasis id="B9400701FFC36B18FC2EFC9DEDAFFCBB" box="[928,1001,837,860]" italics="true" pageId="11" pageNumber="368">contra</emphasis>
|
|||
|
|
<bibRefCitation id="EFA5A6E2FFC36B18FC7FFC9CEA8AFCBA" author="Crochet" box="[1009,1228,836,861]" pageId="11" pageNumber="368" refString="Crochet, P. - A., Leblois, R. & Renoult, J. P. (2015) New reptile records from Morocco and Western Sahara. Herpetology Notes, 8, 583 - 588." type="journal article" year="2015">
|
|||
|
|
Crochet
|
|||
|
|
<emphasis id="B9400701FFC36B18FBDEFC9DEACEFCBB" box="[1104,1160,836,860]" italics="true" pageId="11" pageNumber="368">et al.</emphasis>
|
|||
|
|
2015
|
|||
|
|
</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|