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<document id="B88BA2F6350A100EEC7B918C16499308" ID-DOI="10.11646/zootaxa.4994.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5043261" ID-ZooBank="107FCA64-345F-40A4-99D3-5C1441EEAD93" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624993528766" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2021" docId="03A187DEFFDFFFDB6194FCB207CEF646" docLanguage="en" docName="zootaxa.4994.1.1.pdf" docOrigin="Zootaxa 4994 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus triangularis Perkins 2021, new species" docType="treatment" docVersion="4" lastPageNumber="37" masterDocId="FF98FFA6FFFCFFFF6103FFF0045FF707" masterDocTitle="Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)" masterLastPageNumber="93" masterPageNumber="1" pageNumber="36" updateTime="1698978858223" updateUser="plazi">
<mods:mods id="5BAA286DE0B0F3D0B492EEE41EFD14E0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4CC5C9B4DACF444C9B22D3839FF4B307">
<mods:title id="D8AA1F1A3FBAA1EBE431A1FF0DBE1532">Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)</mods:title>
</mods:titleInfo>
<mods:name id="F74D32A897D874B49D3B7A38FBC4DF2A" type="personal">
<mods:role id="756B4BF6301F013796C67F4E68B69A0E">
<mods:roleTerm id="55630A10511127DEDDF7323420F769A6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A796A6ED966EEAFC70FFB25C04F0DD38">Perkins, Philip D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="19D06E3CDB5650289C9288850687E77F">text</mods:typeOfResource>
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<mods:title id="E2B9D5EE47DF2E3F0B7706090238B3B4">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5914A0395721642770022CE4D07950B6">
<mods:date id="6F3D6E1D9ABFE50B59B76AB38D9AB390">2021</mods:date>
<mods:detail id="0AFBB34B4E68AF056D1CA50E41750F00" type="pubDate">
<mods:number id="0C9B662AA19A3E1508E27E5149763D69">2021-06-28</mods:number>
</mods:detail>
<mods:detail id="BC256E88AE6D5A80FF955A8B496CA55D" type="volume">
<mods:number id="4FBF68C1456C86DF2411CEBE34CF2DD6">4994</mods:number>
</mods:detail>
<mods:detail id="DB3CCD81C69B4CEEAF5D0CAED0332055" type="issue">
<mods:number id="302A94B968ECB87AF1EB080887D7AB58">1</mods:number>
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<mods:start id="C05412CD5FFE3A4D6EDE6FE2FDBC08BC">1</mods:start>
<mods:end id="B7EEAFA090900AE3C6FD463FBF50377F">93</mods:end>
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<mods:classification id="0344D284697D1A9603C9566E72103573">journal article</mods:classification>
<mods:identifier id="29B31DCCDC5E3FD75988CE3C61138D4E" type="DOI">10.11646/zootaxa.4994.1.1</mods:identifier>
<mods:identifier id="94587F2B6855B5BB7E1C9D79DEC044FE" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="BE904407E5DDEB6EE043BAAB03B2FBF5" type="Zenodo-Dep">5043261</mods:identifier>
<mods:identifier id="1644FE0658EE05518E6BB70D079CD64F" type="ZooBank">107FCA64-345F-40A4-99D3-5C1441EEAD93</mods:identifier>
</mods:mods>
<treatment id="03A187DEFFDFFFDB6194FCB207CEF646" ID-DOI="http://doi.org/10.5281/zenodo.5043982" ID-Zenodo-Dep="5043982" LSID="urn:lsid:plazi:treatment:03A187DEFFDFFFDB6194FCB207CEF646" httpUri="http://treatment.plazi.org/id/03A187DEFFDFFFDB6194FCB207CEF646" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
<subSubSection id="C3126543FFDFFFDC6194FCB2060DF45B" box="[151,594,834,861]" pageId="35" pageNumber="36" type="nomenclature">
<paragraph id="8BB736C8FFDFFFDC6194FCB2060DF45B" blockId="35.[151,594,834,897]" box="[151,594,834,861]" pageId="35" pageNumber="36">
<heading id="D0FF81A4FFDFFFDC6194FCB2060DF45B" bold="true" box="[151,594,834,861]" fontSize="11" level="2" pageId="35" pageNumber="36" reason="4">
<emphasis id="B97CEADAFFDFFFDC6194FCB2060DF45B" bold="true" box="[151,594,834,861]" pageId="35" pageNumber="36">
<taxonomicName id="4C084D4BFFDFFFDC6194FCB205ECF45A" ID-CoL="8Y4RN" authority="Perkins, 2021" authorityName="Perkins" authorityYear="2021" box="[151,435,834,861]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="triangularis" status="sp. nov.">
<emphasis id="B97CEADAFFDFFFDC6194FCB205ECF45A" bold="true" box="[151,435,834,861]" italics="true" pageId="35" pageNumber="36">Hydrochus triangularis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24F57A1FFDFFFDC60C1FCB2060DF45B" box="[450,594,834,860]" pageId="35" pageNumber="36" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3126543FFDFFFDC6194FC960576F486" box="[151,297,870,897]" pageId="35" pageNumber="36" type="description">
<paragraph id="8BB736C8FFDFFFDC6194FC960576F486" blockId="35.[151,594,834,897]" box="[151,297,870,897]" pageId="35" pageNumber="36">
<figureCitation id="13332A4DFFDFFFDC6194FC9604BBF487" box="[151,228,870,896]" captionStart="FIGURE 3" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[169,1397,189,1912]" captionTargetId="figure-28@8.[154,1433,181,1919]" captionTargetPageId="8" captionText="FIGURE 3. Dorsal habitus of Hydrochus species in study area." figureDoi="http://doi.org/10.5281/zenodo.5043269" httpUri="https://zenodo.org/record/5043269/files/figure.png" pageId="35" pageNumber="36">Figs. 3</figureCitation>
,
<figureCitation id="13332A4DFFDFFFDC61F2FC97055EF486" box="[241,257,871,897]" captionStart="FIGURES 6–7" captionStartId="55.[151,264,1944,1968]" captionTargetBox="[163,1427,205,1900]" captionTargetId="figure-15@55.[151,1435,181,1920]" captionTargetPageId="55" captionText="FIGURES 6–7. Habitus and male genitalia of holotypes. 6. Hydrochus distinctus n. sp. 7. Hydrochus triangularis n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043278" httpUri="https://zenodo.org/record/5043278/files/figure.png" pageId="35" pageNumber="36">7</figureCitation>
,
<figureCitation id="13332A4DFFDFFFDC600EFC960576F486" box="[269,297,870,897]" captionStart="FIGURES 57–58" captionStartId="81.[151,264,1928,1952]" captionTargetBox="[151,1435,181,1905]" captionTargetId="figure-15@81.[151,1435,181,1905]" captionTargetPageId="81" captionText="FIGURES 57–58. Geographical distributions. 57. Endemics and near endemics. 58. H. bituberculatus." figureDoi="http://doi.org/10.5281/zenodo.5043356" httpUri="https://zenodo.org/record/5043356/files/figure.png" pageId="35" pageNumber="36">57</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3126543FFDFFFDC6194FC5E006AF35E" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="8BB736C8FFDFFFDC6194FC5E006AF35E" blockId="35.[151,1437,942,2012]" pageId="35" pageNumber="36">
<emphasis id="B97CEADAFFDFFFDC6194FC5E0512F4CF" bold="true" box="[151,333,942,968]" pageId="35" pageNumber="36">Type Material.</emphasis>
<materialsCitation id="3B603C95FFDFFFDC605AFC5E006AF35E" collectingDate="1969-06-23" collectingDateMax="1998-07-23" collectingDateMin="1969-06-23" collectionCode="NMNH, SEMC, NMW" collectorName="Rio Caraguata, M &amp; Fritz Plaumann &amp; K. B. Miller &amp; Corumba &amp; F. Plaumann &amp; P. &amp; P. Spangler &amp; San Bernadino &amp; U. Drechsel" country="Brazil" county="Rio Caraguata" elevation="163" latitude="-21.8" location="Central Dept." longLatPrecision="1262" longitude="-52.45" municipality="Brazil" pageId="35" pageNumber="36" specimenCount="2" stateProvince="Cordillera" typeStatus="holotype">
<emphasis id="B97CEADAFFDFFFDC605AFC5E0623F4CF" bold="true" box="[345,636,942,968]" pageId="35" pageNumber="36">
<typeStatus id="54B3886AFFDFFFDC605AFC5E0599F4CF" box="[345,454,942,968]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
(male):
<collectingCountry id="F31F7658FFDFFFDC6330FC5E0623F4CF" box="[563,636,942,968]" name="Brazil" pageId="35" pageNumber="36">Brazil</collectingCountry>
</emphasis>
: Mato Grosso, “
<collectorName id="26FD531EFFDFFFDC6240FC5E0045F4CE" box="[835,1050,942,969]" pageId="35" pageNumber="36">
<collectingCounty id="62D64E44FFDFFFDC6240FC5E07B1F4CF" box="[835,1006,942,968]" pageId="35" pageNumber="36">Rio Caraguata</collectingCounty>
, M
</collectorName>
. Grosso,
<collectingMunicipality id="6BD3ACB2FFDFFFDC658FFC5E008CF4CF" box="[1164,1235,942,968]" pageId="35" pageNumber="36">Brazil</collectingMunicipality>
, IX-53,
<geoCoordinate id="EE3C500FFFDFFFDC643AFC5E01C8F4CF" box="[1337,1431,942,968]" degrees="21" direction="south" minutes="48" orientation="latitude" pageId="35" pageNumber="36" precision="925" value="-21.8">21°48’S</geoCoordinate>
,
<geoCoordinate id="EE3C500FFFDFFFDC6194FC2204A3F4EA" box="[151,252,978,1005]" degrees="52" direction="west" minutes="27" orientation="longitude" pageId="35" pageNumber="36" precision="925" value="-52.45">52°27’W</geoCoordinate>
, 400M alt.,
<collectorName id="26FD531EFFDFFFDC6080FC220670F4EB" box="[387,559,978,1004]" pageId="35" pageNumber="36">Fritz Plaumann</collectorName>
leg.” (
<collectionCode id="ED19AE0DFFDFFFDC6379FC220689F4EB" box="[634,726,978,1004]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="35" pageNumber="36" type="Museum">NMNH</collectionCode>
).
<emphasis id="B97CEADAFFDFFFDC63EBFC2207B4F4EB" bold="true" box="[744,1003,978,1004]" pageId="35" pageNumber="36">
<typeStatus id="54B3886AFFDFFFDC63EBFC22073FF4EB" box="[744,864,978,1004]" pageId="35" pageNumber="36" type="paratype">Paratypes</typeStatus>
(5):
<collectingCountry id="F31F7658FFDFFFDC6294FC2207B4F4EB" box="[919,1003,978,1004]" name="Bolivia" pageId="35" pageNumber="36">Bolivia</collectingCountry>
</emphasis>
: Beni Department,
<locationDeviation id="9A052FEAFFDFFFDC65C4FC2301C7F4EB" box="[1223,1432,978,1005]" location="Trinidad" pageId="35" pageNumber="36">
<quantity id="4CF09B2DFFDFFFDC65C4FC23015FF4EB" box="[1223,1280,978,1005]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="35" pageNumber="36" unit="km" value="7.0">7 km</quantity>
SW Trinidad
</locationDeviation>
,
<geoCoordinate id="EE3C500FFFDFFFDC6194FC06057DF316" box="[151,290,1014,1041]" degrees="14" direction="south" minutes="52" orientation="latitude" pageId="35" pageNumber="36" precision="15" seconds="12" value="-14.87">14°52’12’’S</geoCoordinate>
,
<geoCoordinate id="EE3C500FFFDFFFDC602EFC0605E0F316" box="[301,447,1014,1041]" degrees="64" direction="west" minutes="57" orientation="longitude" pageId="35" pageNumber="36" precision="15" seconds="32" value="-64.958885">64°57’32’’W</geoCoordinate>
,
<quantity id="4CF09B2DFFDFFFDC60CFFC060651F317" box="[460,526,1014,1040]" metricMagnitude="2" metricUnit="m" metricValue="1.63" pageId="35" pageNumber="36" unit="m" value="163.0">
<elevation id="0025D1FBFFDFFFDC60CFFC060651F317" box="[460,526,1014,1040]" metricMagnitude="2" metricUnit="m" metricValue="1.63" pageId="35" pageNumber="36" unit="m" value="163.0">163m</elevation>
</quantity>
,
<date id="FFB61008FFDFFFDC6319FC0606C4F317" box="[538,667,1014,1040]" pageId="35" pageNumber="36" value="1998-07-18">
<collectingDate id="EFF2E9E0FFDFFFDC6319FC0606C4F317" box="[538,667,1014,1040]" pageId="35" pageNumber="36" value="1998-07-18">18.vii.1998</collectingDate>
</date>
, leg.
<collectorName id="26FD531EFFDFFFDC63D4FC07073DF317" box="[727,866,1014,1041]" pageId="35" pageNumber="36">K. B. Miller</collectorName>
” (1
<collectionCode id="ED19AE0DFFDFFFDC6293FC0607BEF317" box="[912,993,1014,1040]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="35" pageNumber="36" type="Museum">SEMC</collectionCode>
).
<emphasis id="B97CEADAFFDFFFDC62F0FC060063F317" bold="true" box="[1011,1084,1014,1040]" pageId="35" pageNumber="36">
<collectingCountry id="F31F7658FFDFFFDC62F0FC060063F317" box="[1011,1084,1014,1040]" name="Brazil" pageId="35" pageNumber="36">Brazil</collectingCountry>
</emphasis>
: Mato Grosso,
<collectorName id="26FD531EFFDFFFDC65EAFC06010EF317" box="[1257,1361,1014,1040]" pageId="35" pageNumber="36">Corumba</collectorName>
leg.
<collectorName id="26FD531EFFDFFFDC648BFC070558F333" pageId="35" pageNumber="36">F. Plaumann</collectorName>
[no date] (1
<collectionCode id="ED19AE0DFFDFFFDC6098FBEB05A9F332" box="[411,502,1051,1077]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="35" pageNumber="36" type="Museum">NMNH</collectionCode>
).
<emphasis id="B97CEADAFFDFFFDC630BFBEA0624F333" bold="true" box="[520,635,1050,1076]" pageId="35" pageNumber="36">
<collectingCountry id="F31F7658FFDFFFDC630BFBEA0624F333" box="[520,635,1050,1076]" name="Paraguay" pageId="35" pageNumber="36">Paraguay</collectingCountry>
</emphasis>
:
<location id="8ED76013FFDFFFDC6388FBEA077AF332" LSID="urn:lsid:plazi:treatment:03A187DEFFDFFFDB6194FCB207CEF646:8ED76013FFDFFFDC6388FBEA077AF332" box="[651,805,1050,1077]" country="Brazil" county="Rio Caraguata" latitude="-21.8" longLatPrecision="1262" longitude="-52.45" municipality="Brazil" name="Central Dept." pageId="35" pageNumber="36" stateProvince="Cordillera">Central Dept.</location>
,
<locationDeviation id="9A052FEAFFDFFFDC6232FBEA0787F332" box="[817,984,1050,1077]" location="Luque" pageId="35" pageNumber="36">
<quantity id="4CF09B2DFFDFFFDC6232FBEA0734F333" box="[817,875,1050,1076]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="36" unit="km" value="2.0">2 km</quantity>
N Luque
</locationDeviation>
,
<date id="FFB61008FFDFFFDC62E7FBEA00D6F333" box="[996,1161,1050,1076]" pageId="35" pageNumber="36" value="1969-06-23" valueMax="1969-06-24" valueMin="1969-06-23">
<collectingDate id="EFF2E9E0FFDFFFDC62E7FBEA00D6F333" box="[996,1161,1050,1076]" pageId="35" pageNumber="36" value="1969-06-23" valueMax="1969-06-24" valueMin="1969-06-23">23–24.vi.1969</collectingDate>
</date>
, leg.
<collectorName id="26FD531EFFDFFFDC65C5FBEB0121F333" box="[1222,1406,1050,1077]" pageId="35" pageNumber="36">P. &amp; P. Spangler</collectorName>
(1
<collectionCode id="ED19AE0DFFDFFFDC6194FBCF04ADF35E" box="[151,242,1087,1113]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="35" pageNumber="36" type="Museum">NMNH</collectionCode>
);
<collectingRegion id="49CCF82AFFDFFFDC6006FBCF05EAF35F" box="[261,437,1086,1113]" country="Paraguay" name="Cordillera" pageId="35" pageNumber="36">Dep. Cordillera</collectingRegion>
,
<collectorName id="26FD531EFFDFFFDC60C2FBCE0635F35F" box="[449,618,1086,1112]" pageId="35" pageNumber="36">San Bernadino</collectorName>
,
<date id="FFB61008FFDFFFDC6376FBCE06A9F35F" box="[629,758,1086,1112]" pageId="35" pageNumber="36" value="1998-07-23">
<collectingDate id="EFF2E9E0FFDFFFDC6376FBCE06A9F35F" box="[629,758,1086,1112]" pageId="35" pageNumber="36" value="1998-07-23">23.vii.1998</collectingDate>
</date>
, leg.
<collectorName id="26FD531EFFDFFFDC6231FBCF07E6F35F" box="[818,953,1086,1113]" pageId="35" pageNumber="36">U. Drechsel</collectorName>
(2
<collectionCode id="ED19AE0DFFDFFFDC62DDFBCF0079F35E" box="[990,1062,1087,1113]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="35" pageNumber="36">NMW</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3126543FFDFFFDC61C4FB9207A5F237" pageId="35" pageNumber="36" type="diagnosis">
<paragraph id="8BB736C8FFDFFFDC61C4FB9207A5F237" blockId="35.[151,1437,942,2012]" pageId="35" pageNumber="36">
<emphasis id="B97CEADAFFDFFFDC61C4FB92058CF37B" bold="true" box="[199,467,1122,1148]" pageId="35" pageNumber="36">Differential Diagnosis.</emphasis>
Among species in the study area, recognized by the combination of moderate size (ca.
<quantity id="4CF09B2DFFDFFFDC6194FB76055DF3A6" box="[151,258,1158,1185]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="35" pageNumber="36" unit="mm" value="3.2">3.20 mm</quantity>
), the brown dorsum without black spots, the nearly parallel-sided body form, the pronotal depressions moderately deep, much more densely punctate than the low reliefs between depressions, and the male genitalia (
<figureCitation id="13332A4DFFDFFFDC646CFB5A04F8F3EE" captionStart="FIGURES 6–7" captionStartId="55.[151,264,1944,1968]" captionTargetBox="[163,1427,205,1900]" captionTargetId="figure-15@55.[151,1435,181,1920]" captionTargetPageId="55" captionText="FIGURES 6–7. Habitus and male genitalia of holotypes. 6. Hydrochus distinctus n. sp. 7. Hydrochus triangularis n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043278" httpUri="https://zenodo.org/record/5043278/files/figure.png" pageId="35" pageNumber="36">Fig. 7</figureCitation>
; described below). This species has a generalized
<taxonomicName id="4C084D4BFFDFFFDC63DFFB3E0709F3EF" authorityName="Leach" authorityYear="1817" box="[732,854,1230,1256]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEADAFFDFFFDC63DFFB3E0709F3EF" box="[732,854,1230,1256]" italics="true" pageId="35" pageNumber="36">Hydrochus</emphasis>
</taxonomicName>
habitus; reliable determinations will require dissection of males. The paramere tips are shaped similarly to those of
<taxonomicName id="4C084D4BFFDFFFDC6266FB0307AEF20B" authorityName="Perkins" authorityYear="2021" box="[869,1009,1266,1292]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis id="B97CEADAFFDFFFDC6266FB0307AEF20B" box="[869,1009,1266,1292]" italics="true" pageId="35" pageNumber="36">H. distinctus</emphasis>
</taxonomicName>
(
<figureCitation id="13332A4DFFDFFFDC6503FB02001BF20A" box="[1024,1092,1266,1293]" captionStart="FIGURES 6–7" captionStartId="55.[151,264,1944,1968]" captionTargetBox="[163,1427,205,1900]" captionTargetId="figure-15@55.[151,1435,181,1920]" captionTargetPageId="55" captionText="FIGURES 6–7. Habitus and male genitalia of holotypes. 6. Hydrochus distinctus n. sp. 7. Hydrochus triangularis n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043278" httpUri="https://zenodo.org/record/5043278/files/figure.png" pageId="35" pageNumber="36">Fig. 6</figureCitation>
); however, the other aspects of the male genitalia, and especially the dorsal habitus, differ in the two species.
</paragraph>
</subSubSection>
<subSubSection id="C3126543FFDFFFDB61C4FACA055AF61A" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="description">
<paragraph id="8BB736C8FFDFFFDC61C4FACA0095F10F" blockId="35.[151,1437,942,2012]" pageId="35" pageNumber="36">
<emphasis id="B97CEADAFFDFFFDC61C4FACA050EF253" bold="true" box="[199,337,1338,1364]" pageId="35" pageNumber="36">Description</emphasis>
. Size:
<typeStatus id="54B3886AFFDFFFDC60A7FACA0659F253" box="[420,518,1338,1364]" pageId="35" pageNumber="36" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 3.20/1.02; head width 0.75; pronotum l/w 0.69/0.70; PA 0.70; PB 0.59; elytra 2.06/1.02. Habitus as illustrated (
<figureCitation id="13332A4DFFDFFFDC6536FAAE0026F27E" box="[1077,1145,1374,1401]" captionStart="FIGURES 6–7" captionStartId="55.[151,264,1944,1968]" captionTargetBox="[163,1427,205,1900]" captionTargetId="figure-15@55.[151,1435,181,1920]" captionTargetPageId="55" captionText="FIGURES 6–7. Habitus and male genitalia of holotypes. 6. Hydrochus distinctus n. sp. 7. Hydrochus triangularis n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043278" httpUri="https://zenodo.org/record/5043278/files/figure.png" pageId="35" pageNumber="36">Fig. 7</figureCitation>
). Body size moderate (ca.
<quantity id="4CF09B2DFFDFFFDC6194FA7204A1F29A" box="[151,254,1410,1437]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="35" pageNumber="36" unit="mm" value="3.2">3.20 mm</quantity>
). Dorsum brown, with very slight to no iridescence, elytra without black spots. Legs testaceous, with tibiofemoral joints darker. Body form narrow (l/w ratio ca. 86/27). Punctation on elytra ca. 1–2x that of pronotum. Elytra interstriae ca. 1–1.5x strial puncture diameter. Interstria 9
<superScript id="7C7D9B80FFDFFFDC622AFA3A0769F2DF" attach="left" box="[809,822,1482,1496]" fontSize="6" pageId="35" pageNumber="36">th</superScript>
more convex than others, overhanging 10
<superScript id="7C7D9B80FFDFFFDC641FFA3A0176F2DF" attach="left" box="[1308,1321,1482,1496]" fontSize="6" pageId="35" pageNumber="36">th</superScript>
interstria. Usual area of callus on 5
<superScript id="7C7D9B80FFDFFFDC60AEFA1E05E5F2FB" attach="left" box="[429,442,1518,1532]" fontSize="6" pageId="35" pageNumber="36">th</superScript>
moderately raised, moderately elongate, ca. 5 punctures on each side.
</paragraph>
<paragraph id="8BB736C8FFDFFFDC61C4F9E30046F173" blockId="35.[151,1437,942,2012]" pageId="35" pageNumber="36">Pronotum very slightly wider than long (ratio ca. 19/18), widest at anterior margin, narrowed at base, sides slightly arcuate or very weakly sinuate, minutely denticulate; anterior margin slightly bisinuate; depressions moderately deep, much more densely punctate than low reliefs between depressions.</paragraph>
<paragraph id="8BB736C8FFDFFFDC61C4F98E007AF1BB" blockId="35.[151,1437,942,2012]" pageId="35" pageNumber="36">Elytra with spaces between strial punctures narrow walls to ca. 0.5x their diameter; apices rather sharply separately rounded in dorsal view, in lateral view outer margin with slight angulation.</paragraph>
<paragraph id="8BB736C8FFDFFFDC61C4F9360095F1E7" blockId="35.[151,1437,942,2012]" box="[199,1226,1734,1761]" pageId="35" pageNumber="36">Ventral characters: Mentum punctate, with central depression; submentum with two foveae.</paragraph>
<paragraph id="8BB736C8FFDFFFDC61C4F91A0531F0DB" blockId="35.[151,1437,942,2012]" pageId="35" pageNumber="36">
Male genitalia (
<figureCitation id="13332A4DFFDFFFDC607FF91A059EF002" box="[380,449,1770,1797]" captionStart="FIGURES 6–7" captionStartId="55.[151,264,1944,1968]" captionTargetBox="[163,1427,205,1900]" captionTargetId="figure-15@55.[151,1435,181,1920]" captionTargetPageId="55" captionText="FIGURES 6–7. Habitus and male genitalia of holotypes. 6. Hydrochus distinctus n. sp. 7. Hydrochus triangularis n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043278" httpUri="https://zenodo.org/record/5043278/files/figure.png" pageId="35" pageNumber="36">Fig. 7</figureCitation>
) general characters: genitalia large, elongate, length ca. 5x greatest width, basal piece very slightly shorter than parameres (ratio ca. 39/41); paramere tips extending slightly beyond aedeagus tip, medial margins beyond aedeagus tip parallel, plsa large, rounded, lateral margins between plsa and apex slightly arcuate; in lateral view parameres slightly arcuate, wide basally, tapering to sharply pointed apically; aedeagus wide, agw at ca. proximal 1/3, narrowing from there to apex and to base; basal piece in dorsal/ventral views parallel-sided, width in dorsal/ventral views and lateral view ca. equal, in lateral view very slightly arcuate, orifice slightly asymmetrical, margin sclerotized.
</paragraph>
<paragraph id="8BB736C8FFD8FFDB61C4FF67064EF7D2" blockId="36.[151,1437,150,321]" pageId="36" pageNumber="37">Dorsal surface: adtl at ca. distal 1/5; pdmm well separated in distal 1/5, remainder sinuate and overlapping alm; adbl very large, rounded laterally.</paragraph>
<paragraph id="8BB736C8FFD8FFDB61C4FF2F055AF61A" blockId="36.[151,1437,150,321]" pageId="36" pageNumber="37">Ventral surface: avtl not apparent; pvmm more widely separated than pdmm, overlapping alm except for ca. distal 1/5.</paragraph>
</subSubSection>
<subSubSection id="C3126543FFD8FFDB61C4FED707CEF646" box="[199,913,295,321]" pageId="36" pageNumber="37" type="etymology">
<paragraph id="8BB736C8FFD8FFDB61C4FED707CEF646" blockId="36.[151,1437,150,321]" box="[199,913,295,321]" pageId="36" pageNumber="37">
<emphasis id="B97CEADAFFD8FFDB61C4FED70511F646" bold="true" box="[199,334,295,321]" pageId="36" pageNumber="37">Etymology.</emphasis>
Named in reference to the triangular paramere tips.
</paragraph>
</subSubSection>
</treatment>
</document>
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