<documentid="37D6563317A2EFD47E07E5012D0C3D49"ID-DOI="10.1206/4015.1"ID-ISSN="0003-0082"ID-Zenodo-Dep="12171426"IM.bibliography_approvedBy="felipe"IM.illustrations_approvedBy="felipe"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.tables_requiresApprovalFor="GgImagineBatch"IM.taxonomicNames_approvedBy="felipe"IM.treatmentCitations_approvedBy="felipe"IM.treatments_approvedBy="felipe"checkinTime="1718827081284"checkinUser="felipe"docAuthor="Grimaldi, David A."docDate="2024"docId="0387351BFFDFFF9AF6585ABDFE91FE06"docLanguage="en"docName="AmMusNovit.2024.4015.1-43.pdf"docOrigin="American Museum Novitates 2024 (4015)"docSource="https://bioone.org/journals/american-museum-novitates/volume-2024/issue-4015/4015.1/The-Drosophila-Sophophora-obscura-species-group-in-the-Americas-Diptera/10.1206/4015.1.full"docStyle="DocumentStyle:26C3580A17D4BBE58BD8D8AC6AE09AC4.6:AmMusNovit.2010-.journal_article"docStyleId="26C3580A17D4BBE58BD8D8AC6AE09AC4"docStyleName="AmMusNovit.2010-.journal_article"docStyleVersion="6"docTitle="Drosophila (Sophophora) maya Heed and O'Grady 2000"docType="treatment"docVersion="3"lastPageNumber="35"masterDocId="FFBE4D63FFFDFFB9F7345809FFBBFF97"masterDocTitle="The Drosophila (Sophophora) obscura species group in the Americas (Diptera: Drosophilidae): review, revisions, and three new species"masterLastPageNumber="44"masterPageNumber="1"pageNumber="35"updateTime="1718834106313"updateUser="ExternalLinkService"zenodo-license-document="CC-BY-4.0">
<mods:titleid="F06B24771ACF2ED1060E5DB606C2F9E2">The Drosophila (Sophophora) obscura species group in the Americas (Diptera: Drosophilidae): review, revisions, and three new species</mods:title>
<taxonomicNameid="4C2EFF8EFFDFFF9BF6585ABDFC07FD59"ID-CoL="37S37"authority="Heed and O'Grady"authorityName="Heed and O'Grady"authorityYear="2000"box="[364,956,692,718]"class="Insecta"family="Drosophilidae"genus="Drosophila"kingdom="Animalia"order="Diptera"pageId="34"pageNumber="35"phylum="Arthropoda"rank="species"species="maya"subGenus="Sophophora">
<figureCitationid="13159888FFDFFF9BF6DD5AE5FDD2FC91"box="[489,617,748,774]"captionStart="FIG"captionStartId="5.[109,149,1138,1159]"captionTargetBox="[108,1205,224,1111]"captionTargetId="figure-185@5.[108,1212,224,1111]"captionTargetPageId="5"captionText="FIG. 2. Heads and thoraces (lateral view) of representative New World species of the obscura group (A–D), and a male midleg of D. cuauhtemoci (E). A. D. chibcha holotype, n. sp. (ASG 05); B. D. maya holotype. C. D. narragansett paralectotype male. D. D. seminole holotype. E. D. cuauhtemoci, male midleg (lateral view), paratype."figureDoi="http://doi.org/10.5281/zenodo.12171434"httpUri="https://zenodo.org/record/12171434/files/figure.png"pageId="34"pageNumber="35">Figures 2B</figureCitation>
,
<figureCitationid="13159888FFDFFF9BF5415AE5FD2DFC91"box="[629,662,748,774]"captionStart="FIG"captionStartId="7.[108,150,1306,1327]"captionTargetBox="[108,1212,224,1280]"captionTargetId="figure-174@7.[108,1212,224,1280]"captionTargetPageId="7"captionText="FIG. 3. Frontal view of heads of representative species in the D. pseudoobscura subgroup. A. D. lowei, male paratype (A19.2); B. D. maya, male holotype (44.12A). C. D. pseudoobscura, male. D. cuauhtemoci, male paratype (face and antennae only) (ASG 39)."figureDoi="http://doi.org/10.5281/zenodo.12171439"httpUri="https://zenodo.org/record/12171439/files/figure.png"pageId="34"pageNumber="35">3B</figureCitation>
,
<figureCitationid="13159888FFDFFF9BF5975AE5FD7DFC91"box="[675,710,748,774]"captionStart="FIG"captionStartId="16.[109,151,1145,1166]"captionTargetBox="[178,1177,243,1108]"captionTargetId="figure-185@16.[108,1212,224,1118]"captionTargetPageId="16"captionText="FIG. 8. Oviscapts and spermathecae (lateral views) of species in the pseudoobscura subgroup, with a detail for D. persimilis. A. D. cuauhtemoci (ASG 39); B. D. lowei (ASG 26); C. D. maya (ASG 19); D. D. persimilis (ASG 37); E. D. pseudoobscura (ASG 34); F. D. zapoteca (ASG 21)."figureDoi="http://doi.org/10.5281/zenodo.12171463"httpUri="https://zenodo.org/record/12171463/files/figure.png"pageId="34"pageNumber="35">8C</figureCitation>
,
<figureCitationid="13159888FFDFFF9BF5E75AE5FCBFFC91"box="[723,772,748,774]"captionStart="FIG"captionStartId="18.[108,150,856,877]"captionTargetBox="[202,1152,244,823]"captionTargetId="figure-329@18.[108,1212,224,830]"captionTargetPageId="18"captionText="FIG. 10. Protarsal male sex combs of species in the D. pseudoobscura subgroup. A. D. cuauhtemoci; B. D. lowei (ASG 40); C. D. maya (ASG 41); D. D. persimilis (ASG 36); E. D. pseudoobscura (ASG 35); F. D. zapoteca (ASG 01). All to the same scale."figureDoi="http://doi.org/10.5281/zenodo.12171467"httpUri="https://zenodo.org/record/12171467/files/figure.png"pageId="34"pageNumber="35">10C</figureCitation>
,
<figureCitationid="13159888FFDFFF9BF4255AE5FC85FC91"box="[785,830,748,774]"captionStart="FIG"captionStartId="33.[108,150,1668,1689]"captionTargetBox="[238,1026,241,1641]"captionTargetId="figure-6@33.[108,1212,224,1642]"captionTargetPageId="33"captionText="FIG. 16. Hypandrium, aedeagus and other appendages of representative species of the pseudoobscura subgroup. A. D. cuauhtemoci (ASG 38); B. D. maya (ASG 02: Guatemala), with details of pregonite lobes; C. D. persimilis (ASG 36); D. D. zapoteca, n. sp. (ASG 20: holotype)"figureDoi="http://doi.org/10.5281/zenodo.12171479"httpUri="https://zenodo.org/record/12171479/files/figure.png"pageId="34"pageNumber="35">16B</figureCitation>
<treatmentCitationid="0A8FA21CFFDFFF9BF7585B2FFD55FCA9"author="Heed, W. B. & P. M. O'Grady"box="[108,750,806,830]"page="98"pageId="34"pageNumber="35"year="2000">
<taxonomicNameid="4C2EFF8EFFDFFF9BF7585B2FFD55FCA9"ID-CoL="37S37"authority="Heed and O'Grady, 2000: 98"authorityName="Heed and O'Grady"authorityPageNumber="98"authorityYear="2000"box="[108,750,806,830]"class="Insecta"family="Drosophilidae"genus="Drosophila"kingdom="Animalia"order="Diptera"pageId="34"pageNumber="35"phylum="Arthropoda"rank="species"species="maya"subGenus="Sophophora">
<bibRefCitationid="EFBFF9FCFFDFFF9BF6895B2FFD55FCA9"author="Heed, W. B. & P. M. O'Grady"box="[445,750,806,830]"pageId="34"pageNumber="35"pagination="98 - 105"refId="ref19215"refString="Heed, W. B., and P. M. O'Grady. 2000. Drosophila maya, a new Neotropical member of the Drosophila obscura species group (Diptera: Drosophilidae). Journal of the New York Entomological Society 108: 98 - 105."type="journal article"year="2000">Heed and O’Grady, 2000: 98</bibRefCitation>
<paragraphid="8B91840DFFDFFF9BF7A85B55FC8EFBA6"blockId="34.[108,1214,860,1745]"pageId="34"pageNumber="35">DIAGNOSIS: Acrostichals in 6 rows; carina narrow above, broadened at base; sex comb with ta1 having 3–5 teeth and ta2 having 2–4; ta1 1.5× length of ta2; surstylus with row of 7–9 prensisetae, inner “hook” short and blunt; posterior margin of hypandrium with small pair of raised, sclerotized lobes (height equal to width); anterior margin of hypandrium narrowed, flat. Oviscapt with small ovisensilla pegs; spermatheca unique in group: dome shaped, no apical indentation, basal sleeve very short (~1/4 length of capsule).</paragraph>
<materialsCitationid="3B468E50FFDFFF9BF7A85C34FD0BFA86"collectingDate="1954-02-05"collectionCode="CUIC"collectorName="WB Heed & Genitalia & The & Heed & Slide & Monte Cristo & El Salvador & W. B. Heed"country="El Salvador"location="Cerro Monte Cristo"pageId="34"pageNumber="35"specimenCode="CUIC 88673"specimenCount="2"specimenCount-male="2"typeStatus="holotype">
<locationid="8EF1D2D6FFDFFF9BF5975C35FC36FBC1"LSID="urn:lsid:plazi:treatment:0387351BFFDFFF9AF6585ABDFE91FE06:8EF1D2D6FFDFFF9BF5975C35FC36FBC1"box="[675,909,1084,1111]"country="El Salvador"name="Cerro Monte Cristo"pageId="34"pageNumber="35">Cerro Monte Cristo</location>
<taxonomicNameid="4C2EFF8EFFDFFF9BF30C5C68FE44FB36"authority="Heed & O'Grady. In AMNH."class="Insecta"family="Drosophilidae"genus="Drosophila"kingdom="Animalia"order="Diptera"pageId="34"pageNumber="35"phylum="Arthropoda"rank="species"species="maya">Drosophila maya Heed & O’Grady. In AMNH.</taxonomicName>
<specimenCodeid="DB882C76FFDFFF9BF52C5CFEFD0BFA86"box="[536,688,1271,1297]"collectionCode="CUIC"country="USA"httpUri="http://grbio.org/cool/sxrc-6x8s"name="Cornell University Insect Collection"pageId="34"pageNumber="35"type="University or college">CUIC 88673</specimenCode>
<materialsCitationid="3B468E50FFDFFF9BF7A85D15FEF8F9AC"collectingDate="1954-02-06"collectingDateMax="1954-03"collectingDateMin="1954-02-06"collectionCode="AMNH, CUIC"collectorName="W. B. Heed. In & WB Heed & M. Sharkey & In & David A. Grimaldi"country="El Salvador"county="Rep. de Honduras"elevation="2200"location="Cerro Monte Cristo"municipality="Monte Vyuca"pageId="34"pageNumber="35"specimenCount="8"specimenCount-female="2"specimenCount-male="3"stateProvince="Zacapa"typeStatus="paratype">
<collectionCodeid="ED3F1CC8FFDFFF9BF7E35D6EFE8FFA16"box="[215,308,1383,1409]"country="USA"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925"lsid="urn:lsid:biocol.org:col:34925"name="American Museum of Natural History"pageId="34"pageNumber="35">AMNH</collectionCode>
<locationid="8EF1D2D6FFDFFF9BF4BB5D6EFBCFFA16"LSID="urn:lsid:plazi:treatment:0387351BFFDFFF9AF6585ABDFE91FE06:8EF1D2D6FFDFFF9BF4BB5D6EFBCFFA16"box="[911,1140,1383,1409]"country="El Salvador"county="Rep. de Honduras"municipality="Monte Vyuca"name="Cerro Monte Cristo"pageId="34"pageNumber="35"stateProvince="Zacapa">Cerro Monte Cristo</location>
<locationid="8EF1D2D6FFDFFF9BF6995DB8FE50FA5C"LSID="urn:lsid:plazi:treatment:0387351BFFDFFF9AF6585ABDFE91FE06:8EF1D2D6FFDFFF9BF6995DB8FE50FA5C"box="[429,491,1457,1483]"country="El Salvador"county="Rep. de Honduras"municipality="Monte Vyuca"name="Heed"pageId="34"pageNumber="35"stateProvince="Zacapa">Heed</location>
“genitalia 44.12B, sex combs 4/3, 3/3” / ASG41.
<locationid="8EF1D2D6FFDFFF9BF30B5DBBFBE2FA5B"LSID="urn:lsid:plazi:treatment:0387351BFFDFFF9AF6585ABDFE91FE06:8EF1D2D6FFDFFF9BF30B5DBBFBE2FA5B"box="[1087,1113,1458,1484]"country="El Salvador"county="Rep. de Honduras"municipality="Monte Vyuca"name="In"pageId="34"pageNumber="35"stateProvince="Zacapa">In</location>
<collectionCodeid="ED3F1CC8FFDFFF9BF3565DBBFB06FA5B"box="[1122,1213,1458,1484]"country="USA"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925"lsid="urn:lsid:biocol.org:col:34925"name="American Museum of Natural History"pageId="34"pageNumber="35">AMNH</collectionCode>
(also slide mounted dissections: in
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<collectionCodeid="ED3F1CC8FFDFFF9BF4F25DF5FB98F981"box="[966,1059,1532,1558]"country="USA"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925"lsid="urn:lsid:biocol.org:col:34925"name="American Museum of Natural History"pageId="34"pageNumber="35">AMNH</collectionCode>
; dissected by
<collectorNameid="26DBE1DBFFDFFF9BF7595E28FF1FF9AC"box="[109,164,1569,1595]"name="David A. Grimaldi"pageId="34"pageNumber="35">D.G.</collectorName>
<bibRefCitationid="EFBFF9FCFFDFFF9BF6C25E4EFC94F9F6"author="Heed, W. B. & P. M. O'Grady"box="[502,815,1607,1633]"pageId="34"pageNumber="35"pagination="98 - 105"refId="ref19215"refString="Heed, W. B., and P. M. O'Grady. 2000. Drosophila maya, a new Neotropical member of the Drosophila obscura species group (Diptera: Drosophilidae). Journal of the New York Entomological Society 108: 98 - 105."type="journal article"year="2000">Heed and O’Grady (2000)</bibRefCitation>
<collectionCodeid="ED3F1CC8FFDFFF9BF5C35EBEFCFBF946"box="[759,832,1719,1745]"country="USA"httpUri="http://grbio.org/cool/sxrc-6x8s"name="Cornell University Insect Collection"pageId="34"pageNumber="35"type="University or college">CUIC</collectionCode>
, have the hypandrium distorted by the weight of the coverslip.