treatments-xml/data/7E/13/87/7E13878EFF9A1B56FF321E64FD1F8DE1.xml

<document ID-DOI="10.3897/zookeys.36.306" ID-GBIF-Dataset="4db6b327-7482-432e-a5f6-36f91c79fef3" ID-ISSN="1313–2970" ID-Zenodo-Dep="576620" ID-ZooBank="ADCACC88-6C78-4386-8E33-3F98234ECE92" approvalRequired="105" approvalRequired_for_textStreams="105" checkinTime="1587967053581" checkinUser="plazi" docAuthor="Rix, Michael &amp; Harvey, Mark" docDate="2010" docId="7E13878EFF9A1B56FF321E64FD1F8DE1" docLanguage="en" docName="ZK_article_2154.pdf" docOrigin="ZooKeys 36 (36)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@6ed9075d" docStyleName="zookeys.2008.journal_article" docTitle="Epigastrina typhlops Rix &amp; Harvey 2010, sp. n." docType="treatment" docUuid="39623469-D1A5-4EDD-974A-1F3AD909EBC8" docUuidSource="ZooBank" docVersion="5" lastPageNumber="46" masterDocId="822AFFF6FFB61B7BFFBE1F57FFE98D05" masterDocTitle="The spider family Micropholcommatidae (Arachnida: Araneae: Araneoidea): a relimitation and revision at the generic level" masterLastPageNumber="321" masterPageNumber="1" pageNumber="45" updateTime="1669157185164" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:titleInfo>
<mods:title>The spider family Micropholcommatidae (Arachnida: Araneae: Araneoidea): a relimitation and revision at the generic level</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rix, Michael</mods:namePart>
<mods:affiliation>Western Australian Museum, Welshpool DC, Perth, Australia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Harvey, Mark</mods:namePart>
<mods:affiliation>Western Australian Museum, Perth, Welshpool, Australia</mods:affiliation>
</mods:name>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2010</mods:date>
<mods:detail type="pubDate">
<mods:number>2010-02-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>36</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>36</mods:number>
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<mods:start>1</mods:start>
<mods:end>321</mods:end>
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<mods:identifier type="DOI">10.3897/zookeys.36.306</mods:identifier>
<mods:identifier type="GBIF-Dataset">4db6b327-7482-432e-a5f6-36f91c79fef3</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576620</mods:identifier>
<mods:identifier type="ZooBank">ADCACC88-6C78-4386-8E33-3F98234ECE92</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3789397" ID-GBIF-Taxon="163749035" ID-Zenodo-Dep="3789397" LSID="urn:lsid:zoobank.org:act:39623469-D1A5-4EDD-974A-1F3AD909EBC8" httpUri="http://treatment.plazi.org/id/7E13878EFF9A1B56FF321E64FD1F8DE1" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
<subSubSection pageId="44" pageNumber="45" type="nomenclature">
<paragraph blockId="44.[140,986,307,405]" box="[140,633,307,334]" pageId="44" pageNumber="45">
<heading box="[140,633,307,334]" fontSize="11" level="2" pageId="44" pageNumber="45" reason="3">
<emphasis bold="true" box="[140,633,307,334]" pageId="44" pageNumber="45">
<taxonomicName authority="Rix &amp; Harvey" authorityName="Rix &amp; Harvey" authorityYear="2010" box="[140,556,307,334]" class="Arachnida" family="Micropholcommatidae" genus="Epigastrina" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="typhlops" status="sp. n.">
<emphasis bold="true" box="[140,383,307,334]" italics="true" pageId="44" pageNumber="45">
<emphasis box="[140,278,307,333]" inLineHeading="true" pageId="44" pageNumber="45" reason="1">Epigastrina</emphasis>
typhlops
</emphasis>
Rix &amp; Harvey
</taxonomicName>
, 
<taxonomicNameLabel box="[568,633,307,333]" pageId="44" pageNumber="45" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="44.[140,986,307,405]" box="[140,986,342,369]" pageId="44" pageNumber="45">
<uri box="[140,986,342,369]" pageId="44" pageNumber="45">
urn:lsid:zoobank.org:act: 
<uuid box="[422,986,342,369]" pageId="44" pageNumber="45">39623469-D1A5-4EDD-974A-1F3AD909EBC8</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection box="[140,385,378,405]" pageId="44" pageNumber="45" type="description">
<paragraph blockId="44.[140,986,307,405]" box="[140,385,378,405]" pageId="44" pageNumber="45">
<figureCitation box="[140,241,378,404]" captionStart="Figure 63" captionStartId="185.[140,212,1104,1126]" captionTargetBox="[140,1095,293,1081]" captionTargetId="figure@185.[140,1108,293,1084]" captionTargetPageId="185" captionText="Figure 63. Epigastrina species, female epigynes and receptacula. A–B E. fulva (Hickman) from Mount Wellington, Tasmania: A external epigyne, ventral view, showing the very large epigastric plate B cleared receptacula, dorsal view. C–E external epigynes, ventral view, at scale-identical size for comparison: C E. fulva from Mount Wellington, Tasmania D holotype E. loongana sp. n. from the Loongana karst, Tasmania; E, allotype E. typhlops sp. n. from the Mole Creek karst, Tasmania. Arrows indicate the trajectory of insemination and proximal fertilisation ducts. Scale bars = 0.065 mm (65 µm)." figureDoi="http://doi.org/10.5281/zenodo.3767783" httpUri="https://zenodo.org/record/3767783/files/figure.png" pageId="44" pageNumber="45">Figs 63E</figureCitation>
, 
<figureCitation box="[253,329,378,405]" captionStart-0="Figure 65" captionStart-1="Figure 66" captionStartId-0="187.[140,212,856,878]" captionStartId-1="187.[140,212,1622,1644]" captionTargetBox-0="[140,1108,165,837]" captionTargetBox-1="[190,1037,999,1588]" captionTargetId-0="figure@187.[140,1109,165,839]" captionTargetId-1="figure@187.[187,1060,978,1602]" captionTargetPageId-0="187" captionTargetPageId-1="187" captionText-0="Figure 65. Epigastrina typhlops sp. n. from the Mole Creek karst, Tasmania. A, C holotype male (AMS KS29793): A habitus, dorsal view C cephalothorax and right pedipalp (left removed), antero-dorsal view. B, D allotype female (QVM 13:12765): B habitus, dorsal view D abdomen, ventral view. Note the pale body colouration and complete absence of eyes." captionText-1="Figure 66. Epigastrina typhlops sp. n., holotype male from the Mole Creek karst, Tasmania (AMS KS29793): A left pedipalp, ventral view B left pedipalp, retrolateral view. Scale bar = 0.065 mm (65 µm)." figureDoi-0="http://doi.org/10.5281/zenodo.3767787" figureDoi-1="http://doi.org/10.5281/zenodo.3767789" httpUri-0="https://zenodo.org/record/3767787/files/figure.png" httpUri-1="https://zenodo.org/record/3767789/files/figure.png" pageId="44" pageNumber="45">65–66</figureCitation>
, 213
</paragraph>
</subSubSection>
<subSubSection pageId="44" pageNumber="45" type="materials_examined">
<paragraph blockId="44.[140,1108,448,1671]" pageId="44" pageNumber="45">
<emphasis bold="true" box="[140,315,448,474]" pageId="44" pageNumber="45">
<emphasis box="[140,198,448,474]" inLineHeading="true" pageId="44" pageNumber="45" reason="1">Type</emphasis>
material.
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2608825978" collectingDate="1990-10-24" collectionCode="AMS" collectorName="S. Eberhard" country="Australia" latitude="-41.553333" location="Kubla Khan" longLatPrecision="19" longitude="146.28111" municipality="Mole Creek" pageId="44" pageNumber="45" specimenCode="KS29793" specimenCount="1" stateProvince="Tasmania" typeStatus="holotype">
<typeStatus box="[326,433,448,474]" pageId="44" pageNumber="45">Holotype</typeStatus>
male: 
<emphasis box="[516,654,448,474]" italics="true" pageId="44" pageNumber="45">
<location LSID="urn:lsid:plazi:treatment:7E13878EFF9A1B56FF321E64FD1F8DE1:F3656043FF9A1B57FDBA1E97FD678CDF" box="[516,654,448,474]" country="Australia" latitude="-41.553333" longLatPrecision="19" longitude="146.28111" municipality="Mole Creek" name="Kubla Khan" pageId="44" pageNumber="45" stateProvince="Tasmania">Kubla Khan</location>
</emphasis>
(MC-1), 
<collectingMunicipality box="[773,910,448,474]" pageId="44" pageNumber="45">Mole Creek</collectingMunicipality>
karst, 
<collectingRegion box="[993,1103,448,474]" country="Australia" name="Tasmania" pageId="44" pageNumber="45">Tasmania</collectingRegion>
, 
<collectingCountry box="[140,243,483,509]" name="Australia" pageId="44" pageNumber="45">Australia</collectingCountry>
, from sheet web with vertical stay lines in cracks of wall above pool, near lower entrance (dark zone), 
<geoCoordinate box="[456,588,519,545]" degrees="41" direction="south" minutes="33" orientation="latitude" pageId="44" pageNumber="45" precision="15" seconds="12" value="-41.553333">41°33'12&quot;S</geoCoordinate>
, 
<geoCoordinate box="[600,750,518,545]" degrees="146" direction="east" minutes="16" orientation="longitude" pageId="44" pageNumber="45" precision="15" seconds="52" value="146.28111">146°16'52&quot;E</geoCoordinate>
, 
<date box="[762,886,519,545]" pageId="44" pageNumber="45" value="1990-10-24">
<collectingDate box="[762,886,519,545]" pageId="44" pageNumber="45" value="1990-10-24">24.X.1990</collectingDate>
</date>
, 
<collectorName box="[898,1031,519,545]" pageId="44" pageNumber="45">S. Eberhard</collectorName>
(
<collectionCode box="[1046,1107,519,545]" pageId="44" pageNumber="45">AMS</collectionCode>
<specimenCode box="[140,253,554,580]" collectionCode="AMS" pageId="44" pageNumber="45">KS29793</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph blockId="44.[140,1108,448,1671]" box="[185,971,589,616]" pageId="44" pageNumber="45">
<typeStatus box="[185,295,590,616]" pageId="44" pageNumber="45">Paratypes</typeStatus>
: 
<materialsCitation ID-GBIF-Occurrence="2608825883" box="[306,968,589,616]" collectingDate="1990-10-24" collectionCode="QVM" collectorName="S. Eberhard" country="Australia" latitude="-41.553333" location="Kubla Khan" longLatPrecision="19" longitude="146.28111" municipality="Mole Creek" pageId="44" pageNumber="45" specimenCode="QVM 13" specimenCount="1" stateProvince="Tasmania" typeStatus="allotype">
<typeStatus box="[306,401,589,615]" pageId="44" pageNumber="45">Allotype</typeStatus>
female, same data as holotype (
<specimenCode box="[759,869,589,615]" collectionCode="QVM" country="Australia" httpUri="http://grbio.org/cool/2s8k-2m7w" name="Queen Victoria Museum" pageId="44" pageNumber="45">QVM 13</specimenCode>
: 12765)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="44" pageNumber="45" type="etymology">
<paragraph blockId="44.[140,1108,448,1671]" pageId="44" pageNumber="45">
<emphasis bold="true" box="[185,320,624,650]" pageId="44" pageNumber="45">
<emphasis box="[185,315,624,650]" inLineHeading="true" pageId="44" pageNumber="45" reason="1">Etymology</emphasis>
.
</emphasis>
The specific epithet is derived from the Greek ‘typhlos’, meaning ‘blind’ (
<bibRefCitation author="Brown RW" box="[228,375,659,686]" pageId="44" pageNumber="45" refId="ref69600" refString="Brown RW (1956) Composition of Scientific Words: A Manual of Methods and a Lexicon of Materials for the Practice of Logotechnics. Smithsonian Books, Washington, DC, 882 pp." type="book" year="1956">Brown 1956</bibRefCitation>
), and refers to the absence of eyes in this species.
</paragraph>
</subSubSection>
<subSubSection pageId="44" pageNumber="45" type="diagnosis">
<paragraph blockId="44.[140,1108,448,1671]" pageId="44" pageNumber="45">
<emphasis bold="true" box="[185,312,694,720]" pageId="44" pageNumber="45">
<emphasis box="[185,307,694,720]" inLineHeading="true" pageId="44" pageNumber="45" reason="1">Diagnosis</emphasis>
.
</emphasis>
Males and females of 
<taxonomicName authorityName="Rix &amp; Harvey" authorityYear="2010" box="[560,776,695,721]" class="Arachnida" family="Micropholcommatidae" genus="Epigastrina" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="typhlops">
<emphasis box="[560,776,695,721]" italics="true" pageId="44" pageNumber="45">Epigastrina typhlops</emphasis>
</taxonomicName>
can be distinguished from all other described congeners by the complete absence of eyes (
<figureCitation box="[813,918,730,757]" captionStart="Figure 65" captionStartId="187.[140,212,856,878]" captionTargetBox="[140,1108,165,837]" captionTargetId="figure@187.[140,1109,165,839]" captionTargetPageId="187" captionText="Figure 65. Epigastrina typhlops sp. n. from the Mole Creek karst, Tasmania. A, C holotype male (AMS KS29793): A habitus, dorsal view C cephalothorax and right pedipalp (left removed), antero-dorsal view. B, D allotype female (QVM 13:12765): B habitus, dorsal view D abdomen, ventral view. Note the pale body colouration and complete absence of eyes." figureDoi="http://doi.org/10.5281/zenodo.3767787" httpUri="https://zenodo.org/record/3767787/files/figure.png" pageId="44" pageNumber="45">Fig. 65C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="44" pageNumber="45" type="description">
<paragraph blockId="44.[140,1108,448,1671]" pageId="44" pageNumber="45">
<emphasis bold="true" box="[185,339,765,791]" pageId="44" pageNumber="45">
<emphasis box="[185,334,765,791]" inLineHeading="true" pageId="44" pageNumber="45" reason="1">Description</emphasis>
.
</emphasis>
<emphasis box="[348,507,765,791]" italics="true" pageId="44" pageNumber="45">
<typeStatus box="[348,446,765,791]" pageId="44" pageNumber="45">Holotype</typeStatus>
male
</emphasis>
: Total length 1.18. Carapace 0.53 long, 0.39 wide. Abdomen 0.70 long, 0.47 wide. Leg I femur 0.48. Cephalothorax, legs very pale tan-yellow; abdomen pale cream. Carapace raised anteriorly, not fused to sternum except around petiole; dorsal surface of pars cephalica slightly convex in lateral view. Eyes absent. Chelicerae each with large, bulging anterior projection; promargin with three peg teeth. Legs relatively short (leg I femur-carapace ratio 0.91); distal, prolateral macroseta present on tibia I. Abdomen oval, covered with hair-like setae; dorsal scute large, indistinct, covering most of dorsal surface of abdomen; lateral sclerotic strips absent. Pedipalpal patella with retrolaterally-directed, hooked lRPA and ornate, ridged cuticular microstructure; tegulum large, bulging posteriorly, with excavate ETR and very large, divergent, fleshy medial conductor bearing rod-like prolateral process and massive, fleshy retrolateral process; embolus short (length &lt;5× width), spur-like, distally hooked, situated between divergent processes of conductor (
<figureCitation box="[232,320,1223,1250]" captionStart="Figure 66" captionStartId="187.[140,212,1622,1644]" captionTargetBox="[190,1037,999,1588]" captionTargetId="figure@187.[187,1060,978,1602]" captionTargetPageId="187" captionText="Figure 66. Epigastrina typhlops sp. n., holotype male from the Mole Creek karst, Tasmania (AMS KS29793): A left pedipalp, ventral view B left pedipalp, retrolateral view. Scale bar = 0.065 mm (65 µm)." figureDoi="http://doi.org/10.5281/zenodo.3767789" httpUri="https://zenodo.org/record/3767789/files/figure.png" pageId="44" pageNumber="45">Fig. 66</figureCitation>
).
</paragraph>
<paragraph blockId="44.[140,1108,448,1671]" pageId="44" pageNumber="45">
<emphasis box="[185,349,1258,1284]" italics="true" pageId="44" pageNumber="45">
<typeStatus box="[185,271,1258,1284]" pageId="44" pageNumber="45">Allotype</typeStatus>
female
</emphasis>
: Total length 1.12. Carapace 0.49 long, 0.40 wide. Abdomen 0.76 long, 0.55 wide. Leg I femur 0.43. Cephalothorax, legs very pale tan-yellow; abdomen pale cream, with darker tan-yellow epigyne. Carapace raised anteriorly, not fused to sternum except around petiole; dorsal surface of pars cephalica slightly convex in lateral view. Eyes absent. Chelicerae without bulging anterior projections; promargin without peg teeth. Legs relatively short (leg I femur-carapace ratio 0.88); macrosetae absent. Abdomen oval, covered with hair-like setae; dorsal scute and lateral sclerotic strips absent. Pedipalp entire, five-segmented. Epigyne heavily sclerotised externally, with distinctive, wedge-shaped epigastric plate (
<figureCitation box="[706,817,1539,1566]" captionStart="Figure 65" captionStartId="187.[140,212,856,878]" captionTargetBox="[140,1108,165,837]" captionTargetId="figure@187.[140,1109,165,839]" captionTargetPageId="187" captionText="Figure 65. Epigastrina typhlops sp. n. from the Mole Creek karst, Tasmania. A, C holotype male (AMS KS29793): A habitus, dorsal view C cephalothorax and right pedipalp (left removed), antero-dorsal view. B, D allotype female (QVM 13:12765): B habitus, dorsal view D abdomen, ventral view. Note the pale body colouration and complete absence of eyes." figureDoi="http://doi.org/10.5281/zenodo.3767787" httpUri="https://zenodo.org/record/3767787/files/figure.png" pageId="44" pageNumber="45">Fig. 65D</figureCitation>
); spermathecae globular, connecting to parallel, filiform insemination and fertilisation ducts (
<figureCitation box="[909,1011,1575,1602]" captionStart="Figure 63" captionStartId="185.[140,212,1104,1126]" captionTargetBox="[140,1095,293,1081]" captionTargetId="figure@185.[140,1108,293,1084]" captionTargetPageId="185" captionText="Figure 63. Epigastrina species, female epigynes and receptacula. A–B E. fulva (Hickman) from Mount Wellington, Tasmania: A external epigyne, ventral view, showing the very large epigastric plate B cleared receptacula, dorsal view. C–E external epigynes, ventral view, at scale-identical size for comparison: C E. fulva from Mount Wellington, Tasmania D holotype E. loongana sp. n. from the Loongana karst, Tasmania; E, allotype E. typhlops sp. n. from the Mole Creek karst, Tasmania. Arrows indicate the trajectory of insemination and proximal fertilisation ducts. Scale bars = 0.065 mm (65 µm)." figureDoi="http://doi.org/10.5281/zenodo.3767783" httpUri="https://zenodo.org/record/3767783/files/figure.png" pageId="44" pageNumber="45">Fig. 63E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="44" pageNumber="45" type="distribution">
<paragraph blockId="44.[140,1108,448,1671]" pageId="44" pageNumber="45">
<emphasis bold="true" box="[185,344,1609,1635]" pageId="44" pageNumber="45">
<emphasis box="[185,339,1609,1635]" inLineHeading="true" pageId="44" pageNumber="45" reason="1">Distribution</emphasis>
.
</emphasis>
Known only from the cave 
<emphasis box="[664,798,1610,1636]" italics="true" pageId="44" pageNumber="45">Kubla Khan</emphasis>
in the Mole Creek karst of north-central 
<collectingRegion box="[296,403,1645,1671]" country="Australia" name="Tasmania" pageId="44" pageNumber="45">Tasmania</collectingRegion>
(Fig. 213).
</paragraph>
</subSubSection>
<subSubSection pageId="45" pageNumber="46" type="discussion">
<paragraph blockId="45.[140,1107,167,228]" pageId="45" pageNumber="46">
<emphasis bold="true" box="[185,298,167,193]" pageId="45" pageNumber="46">Remarks.</emphasis>
<taxonomicName authorityName="Rix &amp; Harvey" authorityYear="2010" box="[304,520,167,193]" class="Arachnida" family="Micropholcommatidae" genus="Epigastrina" kingdom="Animalia" order="Araneae" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="typhlops">
<emphasis box="[304,520,167,193]" italics="true" pageId="45" pageNumber="46">Epigastrina typhlops</emphasis>
</taxonomicName>
is a highly troglomorphic and completely blind species, otherwise very similar to 
<taxonomicName authorityName="Rix &amp; Harvey" authorityYear="2010" baseAuthorityName="Rix &amp; Harvey" baseAuthorityYear="2010" box="[479,566,202,228]" class="Arachnida" family="Micropholcommatidae" genus="Epigastrina" kingdom="Animalia" order="Araneae" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="fulva">
<emphasis box="[479,566,202,228]" italics="true" pageId="45" pageNumber="46">E. fulva</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Rix &amp; Harvey" authorityYear="2010" box="[622,752,202,228]" class="Arachnida" family="Micropholcommatidae" genus="Epigastrina" kingdom="Animalia" order="Araneae" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="loongana">
<emphasis box="[622,752,202,228]" italics="true" pageId="45" pageNumber="46">E. loongana</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>