<documentID-CLB-Dataset="281715"ID-DOI="http://dx.doi.org/10.3897/zitteliana.97.115682"ID-GBIF-Dataset="57209cd6-ddaf-4d5f-b8c8-004d0468a59e"ID-Pensoft-Pub="2747-8106-97-53"ID-Pensoft-UUID="4A3318772B185D35B1E7EC9578EF4A47"ID-ZooBank="933EC356F21C45AF9CFA563E64D27953"ModsDocID="2747-8106-97-53"checkinTime="1702462880200"checkinUser="pensoft"docAuthor="Neubauer, Thomas A. & Wesselingh, Frank P."docDate="2023"docId="DB4FB853CD0056BBBBFFBFD893962178"docLanguage="en"docName="Zitteliana 97: 53-88"docOrigin="Zitteliana 97"docPubDate="2023-12-12"docSource="http://dx.doi.org/10.3897/zitteliana.97.115682"docTitle="Esperiana esperi"docType="treatment"docVersion="4"id="4A3318772B185D35B1E7EC9578EF4A47"lastPageNumber="53"masterDocId="4A3318772B185D35B1E7EC9578EF4A47"masterDocTitle="The Early Pleistocene freshwater mollusks of the Denizli Basin (Turkey): a new long-lived lake fauna at the crossroads of Pontocaspian and Aegean-Anatolian realms"masterLastPageNumber="88"masterPageNumber="53"pageNumber="53"updateTime="1732588348792"updateUser="ExternalLinkService">
<mods:titleid="255A7DDA97D2827BF9C344C735FBA5A1">The Early Pleistocene freshwater mollusks of the Denizli Basin (Turkey): a new long-lived lake fauna at the crossroads of Pontocaspian and Aegean-Anatolian realms</mods:title>
<mods:affiliationid="EFA96F04E6C3B397730DF7F51B5A7820">SNSB - Bavarian State Collection for Palaeontology and Geology, Richard-Wagner-Strasse 10, 80333 Munich, Germany & Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, Netherlands</mods:affiliation>
<mods:affiliationid="FEEC25512A606C8BD7FE4B51971A2F1E">Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, Netherlands & Department of Earth Sciences, Utrecht University, Princetonlaan 8 a, 3584 CB Utrecht, Netherlands</mods:affiliation>
<taxonomicNameid="6D9F0242F5AF5E9E6BD58F0CF1D86045"baseAuthorityName="Adams & Adams"baseAuthorityYear="1854"class="Bivalvia"family="Melanopsidae"higherTaxonomySource="GBIF"kingdom="Animalia"lsidName=""order="Sorbeoconcha"pageId="0"pageNumber="53"phylum="Mollusca"rank="family">Melanopsidae</taxonomicName>
concerning the elongate, smooth shell with adpressed whorls, lanceolate aperture with thickened inner lip and the slight concavity between base of the last whorl and base of the aperture (
<bibRefCitationid="30D9A44C74A1245215A30F37518F7CC8"author="Welter-Schultes, FW"journalOrPublisher="Planet Poster Editions, Goettingen"pageId="0"pageNumber="53"refId="B170"refString="Welter-Schultes, FW, 2012. European non-marine molluscs, a guide for species identification. Planet Poster Editions, Goettingen"title="European non-marine molluscs, a guide for species identification."year="2012">Welter-Schultes 2012</bibRefCitation>
;
<bibRefCitationid="94335DDB26D815A17AEB97CA5292BDE3"author="Gloeer, P"journalOrPublisher="Identification key, Anatomy, Ecology, Distribution. Privately published"pageId="0"pageNumber="53"refId="B61"refString="Gloeer, P, 2019. The Freshwater Gastropods of the West-Palaearctis. Volume I. Fresh- and brackish waters except spring and subterranean snails. Identification key, Anatomy, Ecology, Distribution. Privately published"title="The Freshwater Gastropods of the West-Palaearctis. Volume I. Fresh- and brackish waters except spring and subterranean snails."year="2019">
). Even the color pattern, consisting of dissected zigzag lines to a chess-like arrangement of orange spots, and the corroded apex are typical of the modern species.
Stefanescu, 1896 from the Dacian Basin (Romania) (
<bibRefCitationid="411AF0DECE863ACAD64561CACDBF1CED"author="Stefanescu, S"journalOrPublisher="Paleontologique"pageId="0"pageNumber="53"pagination="1 - 147"refId="B147"refString="Stefanescu, S, 1896. Etudes sur les Terrains tertiaires de Roumanie. Contribution a l'etude des faunes sarmatique, pontique et levantine. Memoirs de la Societe Geologique de France, Memoir 15. Paleontologique 6: 1 - 147"title="Etudes sur les Terrains tertiaires de Roumanie. Contribution a l'etude des faunes sarmatique, pontique et levantine. Memoirs de la Societe Geologique de France, Memoir 15."volume="6"year="1896">Stefanescu 1896</bibRefCitation>
;
<bibRefCitationid="619DAF5FD8D0A0B3B647D50FF69ED3D1"author="Pană, I"journalOrPublisher="Institutul de cercetǎri, inginerie tehnologicǎ și proiectǎri miniere pentru lignit, Craiova"pageId="0"pageNumber="53"refId="B105"refString="Pană, I, Enache, C, Andreescu, I, 1981. Fauna de moluste a depozitelor cu ligniti din Oltenia. Institutul de cercetǎri, inginerie tehnologicǎ și proiectǎri miniere pentru lignit, Craiova"title="Fauna de moluste a depozitelor cu ligniti din Oltenia."year="1981">
<bibRefCitationid="D00D26EDA52E3AC35A96180663D1A14F"author="Pană, I"editor="Papaianopol, I"journalOrPublisher="Neogen der Zentrale Paratethys, Bd. X, Pl 2. Romanien. Editura Academiei Romane, Bucuresti"pageId="0"pageNumber="53"pagination="296 - 349"refId="B104"refString="Pană, I, 2003. Les nannogastropodes. In: Papaianopol, I, Marinescu, F, Krstic, N, Macalet, R, Eds., Chronostratigraphie und Neostratotypen. Neogen der Zentrale Paratethys, Bd. X, Pl2. Romanien. Editura Academiei Romane, Bucuresti: 296 - 349"title="Les nannogastropodes."volumeTitle="Chronostratigraphie und Neostratotypen."year="2003">
<bibRefCitationid="587155A5166C0A40B70093017CBFCEEF"author="Stefanescu, S"journalOrPublisher="Paleontologique"pageId="0"pageNumber="53"pagination="1 - 147"refId="B147"refString="Stefanescu, S, 1896. Etudes sur les Terrains tertiaires de Roumanie. Contribution a l'etude des faunes sarmatique, pontique et levantine. Memoirs de la Societe Geologique de France, Memoir 15. Paleontologique 6: 1 - 147"title="Etudes sur les Terrains tertiaires de Roumanie. Contribution a l'etude des faunes sarmatique, pontique et levantine. Memoirs de la Societe Geologique de France, Memoir 15."volume="6"year="1896">Stefanescu 1896</bibRefCitation>
). It might well be that both species range within the morphological variability of
<bibRefCitationid="13ABC3325D3D221B2315425F85D97A7A"author="Taner, G"journalOrPublisher="Bulletin of Mineral Research and Exploration Institute of Turkey (MTA)"pageId="0"pageNumber="53"pagination="89 - 126"refId="B148"refString="Taner, G, 1974a. Denizli boelgesi Neojeninin paleontolojik ve stratigrafik etuedue. Bulletin of Mineral Research and Exploration Institute of Turkey (MTA) 82: 89 - 126"title="Denizli boelgesi Neojeninin paleontolojik ve stratigrafik etuedue."volume="82"year="1974 a">Taner (1974a)</bibRefCitation>
Today in central to southeastern Europe (western Black Sea region to Hungary, Slovakia, Austria, and Slovenia) and northwestern Anatolia (
<bibRefCitationid="48642CC19E2394D6A96FC1A243EF78E5"author="Welter-Schultes, FW"journalOrPublisher="Planet Poster Editions, Goettingen"pageId="0"pageNumber="53"refId="B170"refString="Welter-Schultes, FW, 2012. European non-marine molluscs, a guide for species identification. Planet Poster Editions, Goettingen"title="European non-marine molluscs, a guide for species identification."year="2012">Welter-Schultes 2012</bibRefCitation>
;
<bibRefCitationid="2B1B9814F65E80228C3ABC623CBB3A74"author="Gloeer, P"journalOrPublisher="Identification key, Anatomy, Ecology, Distribution. Privately published"pageId="0"pageNumber="53"refId="B61"refString="Gloeer, P, 2019. The Freshwater Gastropods of the West-Palaearctis. Volume I. Fresh- and brackish waters except spring and subterranean snails. Identification key, Anatomy, Ecology, Distribution. Privately published"title="The Freshwater Gastropods of the West-Palaearctis. Volume I. Fresh- and brackish waters except spring and subterranean snails."year="2019">
). Also found in Pleistocene strata of the North Caspian Basin (
<bibRefCitationid="A99AE00AB9DA548090DB794727A8AD81"DOI="https://doi.org/10.3897/zookeys.770.25365"author="Neubauer, TA"journalOrPublisher="ZooKeys"pageId="0"pageNumber="53"pagination="43 - 103"refId="B91"refString="Neubauer, TA, van de Velde, S, Yanina, T, Wesselingh, FP, 2018. A late Pleistocene gastropod fauna from the northern Caspian Sea with implications for Pontocaspian gastropod taxonomy. ZooKeys 770: 43 - 103, DOI: https://doi.org/10.3897/zookeys.770.25365"title="A late Pleistocene gastropod fauna from the northern Caspian Sea with implications for Pontocaspian gastropod taxonomy."url="https://doi.org/10.3897/zookeys.770.25365"volume="770"year="2018">Neubauer et al. 2018</bibRefCitation>