treatments-xml/data/03/FE/C0/03FEC052FFF5DC78569F5DF0FEAD083D.xml

<document id="956D89C6EE03648FC24B4DF01A7A0E88" ID-DOI="10.5281/zenodo.189961" ID-GBIF-Dataset="f4918554-8847-4b2e-89af-ea92badaad40" ID-ISSN="1175-5326" ID-Zenodo-Dep="189961" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461106560087" checkinUser="plazi" docAuthor="Colloff, Matthew J. &amp; Perdomo, Giselle" docDate="2009" docId="03FEC052FFF5DC78569F5DF0FEAD083D" docLanguage="en" docName="zt02217p036.pdf" docOrigin="Zootaxa 2217" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Crotonia alpina Colloff &amp; Perdomo, 2009, sp. nov." docType="treatment" docVersion="9" lastPageNumber="5" masterDocId="FFC7B82AFFF7DC7C56085F5DFFFD0B69" masterDocTitle="New species of Crotonia (Acari: Oribatida: Camisiidae) from Nothofagus and Eucalyptus forests in Victoria, Australia, with a redescription of the fossil species Crotonia ramus (Womersley, 1957)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="3" updateTime="1698593351658" updateUser="plazi">
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<mods:title id="937B495E33E857337A8230DE91AD436A">New species of Crotonia (Acari: Oribatida: Camisiidae) from Nothofagus and Eucalyptus forests in Victoria, Australia, with a redescription of the fossil species Crotonia ramus (Womersley, 1957)</mods:title>
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<mods:namePart id="DC7E315DBACF8C0E4A42D995AE74B932">Colloff, Matthew J.</mods:namePart>
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<mods:namePart id="E6FAE8BAD889243175E4ACB117D78151">Perdomo, Giselle</mods:namePart>
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<mods:date id="B495DCE30B089B32BD1A6403BE34C1E8">2009</mods:date>
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<mods:number id="2AB49A8AEA9DD3F3156A5BE3CF382A76">2217</mods:number>
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<treatment id="03FEC052FFF5DC78569F5DF0FEAD083D" ID-DOI="http://doi.org/10.5281/zenodo.6225390" ID-GBIF-Taxon="119623399" ID-Zenodo-Dep="6225390" LSID="urn:lsid:plazi:treatment:03FEC052FFF5DC78569F5DF0FEAD083D" httpUri="http://treatment.plazi.org/id/03FEC052FFF5DC78569F5DF0FEAD083D" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
<subSubSection id="C34D22CFFFF5DC7E569F5DF0FE830983" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BE87144FFF5DC7E569F5DF0FE4609AE" blockId="2.[151,443,685,746]" box="[151,443,685,711]" pageId="2" pageNumber="3">
<heading id="D0A0C628FFF5DC7E569F5DF0FE4609AE" bold="true" box="[151,443,685,711]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="B923AD56FFF5DC7E569F5DF0FE4609AE" bold="true" box="[151,443,685,711]" pageId="2" pageNumber="3">
<taxonomicName id="4C570AC7FFF5DC7E569F5DF0FEAB09AE" ID-CoL="ZRG8" box="[151,342,685,711]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="alpina" status="sp. nov.">
<emphasis id="B923AD56FFF5DC7E569F5DF0FEAB09AE" bold="true" box="[151,342,685,711]" italics="true" pageId="2" pageNumber="3">Crotonia alpina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A210102DFFF5DC7E57555DF0FE4609AE" box="[349,443,685,711]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE87144FFF5DC7E569F5D8DFE830983" blockId="2.[151,443,685,746]" box="[151,382,720,746]" pageId="2" pageNumber="3">
(
<figureCitation id="136C6DC1FFF5DC7E56975D8DFF0C0983" box="[159,241,720,746]" captionStart="FIGURE 1" captionStartId="3.[151,255,1725,1749]" captionTargetBox="[212,1366,476,1675]" captionTargetId="figure@3.[199,1387,461,1700]" captionTargetPageId="3" captionText="FIGURE 1. Crotonia alpina sp. nov. holotype female a) dorsal; b) ventral." httpUri="https://zenodo.org/record/189962/files/figure.png" pageId="2" pageNumber="3">Figs. 1</figureCitation>
, 
<figureCitation id="136C6DC1FFF5DC7E56F75D8DFEE30983" box="[255,286,720,746]" captionStart="FIGURE 15" captionStartId="25.[151,257,1372,1396]" captionTargetBox="[216,1374,207,1333]" captionTargetId="figure@25.[193,1394,194,1348]" captionTargetPageId="25" captionText="FIGURE 15. Crotonia gadubanudi sp. nov. paratype female a) subcapitulum b) Oral setae (or). Oral setae of c) C. momitoi sp. nov. paratype female; d) C. blacki sp. nov. paratype female; e) C. alpina sp. nov.; f) C. cornuta sp. nov.; g) C. victoriae sp. nov. AS = adoral sclerite; G = gena; H = mentum; RU = rutellum." httpUri="https://zenodo.org/record/189972/files/figure.png" pageId="2" pageNumber="3">15</figureCitation>
, 
<figureCitation id="136C6DC1FFF5DC7E57235D8DFE880983" box="[299,373,720,746]" captionStart-0="FIGURE 17" captionStart-1="FIGURE 18" captionStart-2="FIGURE 19" captionStartId-0="28.[151,255,1525,1549]" captionStartId-1="29.[151,255,1842,1866]" captionStartId-2="30.[151,257,1379,1403]" captionTargetBox-0="[230,1360,297,1483]" captionTargetBox-1="[304,1277,294,1804]" captionTargetBox-2="[169,1413,494,1336]" captionTargetId-0="figure@28.[210,1377,274,1501]" captionTargetId-1="figure@29.[283,1304,277,1817]" captionTargetId-2="figure@30.[151,1436,476,1352]" captionTargetPageId-0="28" captionTargetPageId-1="29" captionTargetPageId-2="30" captionText-0="FIGURE 17. Posteriolateral views of a) Crotonia momitoi sp. nov., male paratype; b) Crotonia blacki sp. nov. female paratype; c) Crotonia gadubanudi sp. nov.; d) Crotonia alpina sp. nov.; e) Crotonia cornuta sp. nov.; f) Crotonia victoriae sp. nov." captionText-1="FIGURE 18. Details of bothridia of a) Crotonia momitoi sp. nov.; b) Crotonia blacki sp. nov.; c) Crotonia gadubanudi sp. nov.; d) Crotonia alpina sp. nov.; e) Crotonia cornuta sp. nov.; f) Crotonia victoriae sp. nov." captionText-2="FIGURE 19. Distribution map of Crotonia spp. in Victoria. Stippled area is the distribution of the broad vegetation group categorised as ‘ wet or damp forests’ (Department of Sustainability and Environment, 2005). NP = National Park; SF = State Forest. Black circles are towns and cities, provided for spatial reference. Inset: 30 - year means of annual rainfall (source: Australian Bureau of Meteorology, www. bom. gov. au)." httpUri-0="https://zenodo.org/record/189974/files/figure.png" httpUri-1="https://zenodo.org/record/189975/files/figure.png" httpUri-2="https://zenodo.org/record/189976/files/figure.png" pageId="2" pageNumber="3">17–19</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34D22CFFFF5DC7F569F5C40FE630C75" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="description">
<paragraph id="8BE87144FFF5DC7E569F5C40FC370837" blockId="2.[151,1437,797,2018]" pageId="2" pageNumber="3">
<emphasis id="B923AD56FFF5DC7E569F5C40FED5085E" bold="true" box="[151,296,797,823]" pageId="2" pageNumber="3">Dimensions</emphasis>
. 
<typeStatus id="54ECCFE6FFF5DC7E573F5C40FE59085E" box="[311,420,797,823]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
female length 1153, breadth 570; female 
<typeStatus id="54ECCFE6FFF5DC7E559A5C40FC0A085E" box="[914,1015,797,823]" pageId="2" pageNumber="3" type="paratype">paratype</typeStatus>
lengths 1067, 1082, 1122; breadths 537, 435, 474. Mean ratio of length of prodorsum to total length: 0.28
</paragraph>
<paragraph id="8BE87144FFF5DC7E56CD5C37FE250F89" blockId="2.[151,1437,797,2018]" pageId="2" pageNumber="3">
<emphasis id="B923AD56FFF5DC7E56CD5C37FEDC08ED" bold="true" box="[197,289,874,900]" pageId="2" pageNumber="3">Female</emphasis>
. 
<emphasis id="B923AD56FFF5DC7E57395C37FE4B08EA" box="[305,438,874,899]" italics="true" pageId="2" pageNumber="3">Prodorsum</emphasis>
: rostrum well-developed, with very prominent naso, lateral edges incurved; rostral setae (
<emphasis id="B923AD56FFF5DC7E56EC5CCCFF0108C3" box="[228,252,913,938]" italics="true" pageId="2" pageNumber="3">ro</emphasis>
) 41, straight, spiniform, smooth (
<figureCitation id="136C6DC1FFF5DC7E549B5CCCFD2A08C2" box="[659,727,913,939]" captionStart="FIGURE 1" captionStartId="3.[151,255,1725,1749]" captionTargetBox="[212,1366,476,1675]" captionTargetId="figure@3.[199,1387,461,1700]" captionTargetPageId="3" captionText="FIGURE 1. Crotonia alpina sp. nov. holotype female a) dorsal; b) ventral." httpUri="https://zenodo.org/record/189962/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
a). Lamellar setae (
<emphasis id="B923AD56FFF5DC7E55CB5CCCFC2508C3" box="[963,984,913,938]" italics="true" pageId="2" pageNumber="3">le</emphasis>
) 350, recurved, smooth, flagelliform. Lamellar apophyses 97, half as long as their mutual distance; extending anteriorly as far as apices of rostral setae (
<emphasis id="B923AD56FFF5DC7E56EC5C82FF010891" box="[228,252,991,1016]" italics="true" pageId="2" pageNumber="3">ro</emphasis>
). Interlamellar apophyses twice as long as broad; interlamellar setae (
<emphasis id="B923AD56FFF5DC7E52375C82FBA80891" box="[1087,1109,991,1016]" italics="true" pageId="2" pageNumber="3">in</emphasis>
) slender, 280, flagelliform, smooth; extending anteriorly just beyond apices of lamellar apophyses. Prodorsal ridges extending half the distance between interlamellar and lamellar apophyses. Diameter of each bothridium 35; anteriolateral auriculate ridge of bothridium a blunt, curved projection, longer than broad, bearing a V-shaped ridge of cuticle projecting anteriolaterally; bothridial membrane reticulate with sub-hexagonal cells medially (
<figureCitation id="136C6DC1FFF5DC7E53635B24FF4B0FD3" captionStart="FIGURE 17" captionStartId="28.[151,255,1525,1549]" captionTargetBox="[230,1360,297,1483]" captionTargetId="figure@28.[210,1377,274,1501]" captionTargetPageId="28" captionText="FIGURE 17. Posteriolateral views of a) Crotonia momitoi sp. nov., male paratype; b) Crotonia blacki sp. nov. female paratype; c) Crotonia gadubanudi sp. nov.; d) Crotonia alpina sp. nov.; e) Crotonia cornuta sp. nov.; f) Crotonia victoriae sp. nov." httpUri="https://zenodo.org/record/189974/files/figure.png" pageId="2" pageNumber="3">Fig. 17</figureCitation>
d). Inter-bothridial ridge almost transverse; very slightly bowed, crenellated. Median field of muscle sigilla present. Prodorsum porose.
</paragraph>
<paragraph id="8BE87144FFF5DC7E56CD5BB0FD4F0E58" blockId="2.[151,1437,797,2018]" pageId="2" pageNumber="3">
<emphasis id="B923AD56FFF5DC7E56CD5BB0FE9B0E6F" box="[197,358,1261,1286]" italics="true" pageId="2" pageNumber="3">Subcapitulum</emphasis>
: with three setae on gena: setae 
<emphasis id="B923AD56FFF5DC7E54EA5BB0FD0C0E6F" box="[738,753,1261,1286]" italics="true" pageId="2" pageNumber="3">a</emphasis>
10; 
<emphasis id="B923AD56FFF5DC7E552C5BB0FCC40E6F" box="[804,825,1261,1286]" italics="true" pageId="2" pageNumber="3">m</emphasis>
1 5; 
<emphasis id="B923AD56FFF5DC7E556F5BB0FC810E6F" box="[871,892,1261,1286]" italics="true" pageId="2" pageNumber="3">m</emphasis>
2 5. Oral setae 
<emphasis id="B923AD56FFF5DC7E522E5BB0FBBD0E6F" box="[1062,1088,1261,1286]" italics="true" pageId="2" pageNumber="3">or</emphasis>
1 bifurcate, barbed bilaterally; 
<emphasis id="B923AD56FFF5DC7E569F5A4AFF4C0E59" box="[151,177,1303,1328]" italics="true" pageId="2" pageNumber="3">or</emphasis>
2 and 
<emphasis id="B923AD56FFF5DC7E56FA5A4AFEF10E59" box="[242,268,1303,1328]" italics="true" pageId="2" pageNumber="3">or</emphasis>
3 spiniform, subequal, 22 (
<figureCitation id="136C6DC1FFF5DC7E54485A4AFD6B0E58" box="[576,662,1303,1329]" captionStart="FIGURE 15" captionStartId="25.[151,257,1372,1396]" captionTargetBox="[216,1374,207,1333]" captionTargetId="figure@25.[193,1394,194,1348]" captionTargetPageId="25" captionText="FIGURE 15. Crotonia gadubanudi sp. nov. paratype female a) subcapitulum b) Oral setae (or). Oral setae of c) C. momitoi sp. nov. paratype female; d) C. blacki sp. nov. paratype female; e) C. alpina sp. nov.; f) C. cornuta sp. nov.; g) C. victoriae sp. nov. AS = adoral sclerite; G = gena; H = mentum; RU = rutellum." httpUri="https://zenodo.org/record/189972/files/figure.png" pageId="2" pageNumber="3">Fig. 15</figureCitation>
e).
</paragraph>
<paragraph id="8BE87144FFF5DC7E56CD5A1CFB590C2D" blockId="2.[151,1437,797,2018]" pageId="2" pageNumber="3">
<emphasis id="B923AD56FFF5DC7E56CD5A1CFEB70E33" box="[197,330,1345,1370]" italics="true" pageId="2" pageNumber="3">Notogaster</emphasis>
: ratio of length to breadth 1.63; broadest at bases of setae 
<emphasis id="B923AD56FFF5DC7E521C5A1CFBDC0E33" box="[1044,1057,1345,1370]" italics="true" pageId="2" pageNumber="3">e</emphasis>
2 (
<figureCitation id="136C6DC1FFF5DC7E52345A1DFB7C0E33" box="[1084,1153,1344,1370]" captionStart="FIGURE 1" captionStartId="3.[151,255,1725,1749]" captionTargetBox="[212,1366,476,1675]" captionTargetId="figure@3.[199,1387,461,1700]" captionTargetPageId="3" captionText="FIGURE 1. Crotonia alpina sp. nov. holotype female a) dorsal; b) ventral." httpUri="https://zenodo.org/record/189962/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
a). Dorsosejugal suture discrete, simple. With 14 pairs of smooth notogastral setae. Pre-notogastral shield bearing thin setiform setae 
<emphasis id="B923AD56FFF5DC7E569F5ACCFF590EC3" box="[151,164,1425,1450]" italics="true" pageId="2" pageNumber="3">c</emphasis>
2, 90, and shorter 
<emphasis id="B923AD56FFF5DC7E57665ACCFE860EC3" box="[366,379,1425,1450]" italics="true" pageId="2" pageNumber="3">c</emphasis>
1, 55, and separated from notogastral shield by transverse hyaline strip. Apophyses of setae 
<emphasis id="B923AD56FFF5DC7E569F5AE7FF590EBA" box="[151,164,1466,1491]" italics="true" pageId="2" pageNumber="3">c</emphasis>
3 prominent, 25, at least twice as long as broad; setae 
<emphasis id="B923AD56FFF5DC7E554B5AE7FCAD0EBA" box="[835,848,1466,1491]" italics="true" pageId="2" pageNumber="3">c</emphasis>
3 flagelliform, 330, extending as far as bases of lamellar apophyses. Notogastral shield tuberculate except in central and lateral regions, bordered by two narrow strips of denser tubercles extending posteriorly almost as far as setae 
<emphasis id="B923AD56FFF5DC7E524C5956FBB10D4D" box="[1092,1100,1547,1572]" italics="true" pageId="2" pageNumber="3">f</emphasis>
1. Caudal region with dense tubercles. Lateral hyaline strip (suprapleural scissure) well developed, bearing tubercles of setae 
<emphasis id="B923AD56FFF5DC7E532F5969FAB90D24" box="[1319,1348,1588,1613]" italics="true" pageId="2" pageNumber="3">cp</emphasis>
and 
<emphasis id="B923AD56FFF5DC7E53775969FA710D24" box="[1407,1420,1588,1613]" italics="true" pageId="2" pageNumber="3">e</emphasis>
2. Tubercles of setae 
<emphasis id="B923AD56FFF5DC7E57795903FE840D1E" box="[369,377,1630,1655]" italics="true" pageId="2" pageNumber="3">f</emphasis>
2 on sub-triangular projections extending beyond lateral margin. Setae 
<emphasis id="B923AD56FFF5DC7E52B85903FB360D1E" box="[1200,1227,1630,1655]" italics="true" pageId="2" pageNumber="3">cp</emphasis>
, setiform, 190; 
<emphasis id="B923AD56FFF5DC7E538D5903FA690D1E" box="[1413,1428,1630,1655]" italics="true" pageId="2" pageNumber="3">d</emphasis>
2 flagelliform, 175; 
<emphasis id="B923AD56FFF5DC7E576659D5FE860DC8" box="[366,379,1672,1697]" italics="true" pageId="2" pageNumber="3">e</emphasis>
2 setiform, 135. Opisthosomal gland opening 
<emphasis id="B923AD56FFF5DC7E559B59D5FC450DC8" box="[915,952,1672,1697]" italics="true" pageId="2" pageNumber="3">gla</emphasis>
positioned at level slightly anterior of 
<emphasis id="B923AD56FFF5DC7E538C59D5FA710DC8" box="[1412,1420,1672,1697]" italics="true" pageId="2" pageNumber="3">f</emphasis>
2. Flagelliform setae 
<emphasis id="B923AD56FFF5DC7E577659ECFE7B0DA3" box="[382,390,1713,1738]" italics="true" pageId="2" pageNumber="3">f</emphasis>
1 90, their apophyses projecting posteriolaterally, adjacent to those of 
<emphasis id="B923AD56FFF5DC7E52EB59ECFB0F0DA3" box="[1251,1266,1713,1738]" italics="true" pageId="2" pageNumber="3">h</emphasis>
1. Setae 
<emphasis id="B923AD56FFF5DC7E535E59ECFA980DA3" box="[1366,1381,1713,1738]" italics="true" pageId="2" pageNumber="3">h</emphasis>
2 70, flagelliform, smooth; their apophyses 130, same length as their distance apart, parallel, diverging apically; caudal margin between them transverse, apophyses of setae 
<emphasis id="B923AD56FFF5DC7E5598585CFC620C73" box="[912,927,1793,1818]" italics="true" pageId="2" pageNumber="3">h</emphasis>
3 positioned posterior of 
<emphasis id="B923AD56FFF5DC7E52DD585CFB190C73" box="[1237,1252,1793,1818]" italics="true" pageId="2" pageNumber="3">h</emphasis>
1 when viewed dorsally. Apophyses of setae 
<emphasis id="B923AD56FFF5DC7E57E55876FE080C2D" box="[493,501,1835,1860]" italics="true" pageId="2" pageNumber="3">f</emphasis>
1 and 
<emphasis id="B923AD56FFF5DC7E543E5876FDB80C2D" box="[566,581,1835,1860]" italics="true" pageId="2" pageNumber="3">h</emphasis>
<date id="FFE95784FFF5DC7E544D5866FD5D0C2D" box="[581,672,1834,1865]" pageId="2" pageNumber="3" value="1925-01-30">1 25–30</date>
, cylindrical, subequal; those of 
<emphasis id="B923AD56FFF5DC7E521C5876FBDE0C2D" box="[1044,1059,1835,1860]" italics="true" pageId="2" pageNumber="3">h</emphasis>
3 15, squat.
</paragraph>
<paragraph id="8BE87144FFF5DC7F56CD5809FD170ACC" blockId="2.[151,1437,797,2018]" lastBlockId="3.[151,1436,152,421]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<emphasis id="B923AD56FFF5DC7E56CD5809FEF20C04" box="[197,271,1876,1901]" italics="true" pageId="2" pageNumber="3">Ve n t e r</emphasis>
: epimeres porose (
<figureCitation id="136C6DC1FFF5DC7E57F25809FDBD0C07" box="[506,576,1876,1902]" captionStart="FIGURE 1" captionStartId="3.[151,255,1725,1749]" captionTargetBox="[212,1366,476,1675]" captionTargetId="figure@3.[199,1387,461,1700]" captionTargetPageId="3" captionText="FIGURE 1. Crotonia alpina sp. nov. holotype female a) dorsal; b) ventral." httpUri="https://zenodo.org/record/189962/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
b); epimeral setae smooth, spiniform, formula 3-1-3-3; ca. 20–25 long except longer stout 
<emphasis id="B923AD56FFF5DC7E578F5826FE590CFD" box="[391,420,1915,1940]" italics="true" pageId="2" pageNumber="3">3c</emphasis>
on well-developed tubercle. Genital plates sub-circular. Perigenital region sparsely tuberculate. Each plate 207 long, 112 broad with eight spiniform setae; two pairs of aggenital setae, subequal in length to genital setae. Anal plate 60 broad, 257 long with three spiniform setae on posterior half of anal plate; three pairs of spiniform adanal setae. Ventral margin of notogaster surrounding anal plates U-shaped. Setae of 
<emphasis id="B923AD56FFF4DC7F56F45FE2FEF60BB1" box="[252,267,191,216]" italics="true" pageId="3" pageNumber="4">p</emphasis>
series smooth, flagelliform, 
<emphasis id="B923AD56FFF4DC7F54515FE2FD950BB1" box="[601,616,191,216]" italics="true" pageId="3" pageNumber="4">p</emphasis>
3 90, 
<emphasis id="B923AD56FFF4DC7F54AA5FE2FD4C0BB1" box="[674,689,191,216]" italics="true" pageId="3" pageNumber="4">p</emphasis>
2 71, 
<emphasis id="B923AD56FFF4DC7F54E35FE2FD070BB1" box="[747,762,191,216]" italics="true" pageId="3" pageNumber="4">p</emphasis>
1 118, on short tubercles located almost adjacent (Fig. 2a). 
<emphasis id="B923AD56FFF4DC7F569F5FB5FED00A68" box="[151,301,232,257]" italics="true" pageId="3" pageNumber="4">Lateral view</emphasis>
: Caudal margin almost perpendicular to notogastral shield (
<figureCitation id="136C6DC1FFF4DC7F55F65FB5FBAB0A6B" box="[1022,1110,232,258]" captionStart="FIGURE 17" captionStartId="28.[151,255,1525,1549]" captionTargetBox="[230,1360,297,1483]" captionTargetId="figure@28.[210,1377,274,1501]" captionTargetPageId="28" captionText="FIGURE 17. Posteriolateral views of a) Crotonia momitoi sp. nov., male paratype; b) Crotonia blacki sp. nov. female paratype; c) Crotonia gadubanudi sp. nov.; d) Crotonia alpina sp. nov.; e) Crotonia cornuta sp. nov.; f) Crotonia victoriae sp. nov." httpUri="https://zenodo.org/record/189974/files/figure.png" pageId="3" pageNumber="4">Fig. 17</figureCitation>
d); distance between dorsal and ventral surface ca. 435. Apophyses of setae 
<emphasis id="B923AD56FFF4DC7F54C45E52FD290A41" box="[716,724,271,296]" italics="true" pageId="3" pageNumber="4">f</emphasis>
1 and 
<emphasis id="B923AD56FFF4DC7F551C5E52FCDE0A41" box="[788,803,271,296]" italics="true" pageId="3" pageNumber="4">h</emphasis>
1 and 
<emphasis id="B923AD56FFF4DC7F556C5E52FC8E0A41" box="[868,883,271,296]" italics="true" pageId="3" pageNumber="4">h</emphasis>
2 pointing dorsolaterally, those of 
<emphasis id="B923AD56FFF4DC7F52F65E52FAF00A41" box="[1278,1293,271,296]" italics="true" pageId="3" pageNumber="4">h</emphasis>
2 prominent, horizontal, pointing posteriorly; those of 
<emphasis id="B923AD56FFF4DC7F54755E64FD710A3B" box="[637,652,313,338]" italics="true" pageId="3" pageNumber="4">h</emphasis>
3 positioned ventral of 
<emphasis id="B923AD56FFF4DC7F55915E64FC550A3B" box="[921,936,313,338]" italics="true" pageId="3" pageNumber="4">h</emphasis>
1, pointing posteriorly. Apophyses of setae 
<emphasis id="B923AD56FFF4DC7F569F5E3FFF5B0A12" box="[151,166,354,379]" italics="true" pageId="3" pageNumber="4">p</emphasis>
1 on caudal region, the most dorsal of the 
<emphasis id="B923AD56FFF4DC7F54995E3FFD5D0A12" box="[657,672,354,379]" italics="true" pageId="3" pageNumber="4">p</emphasis>
series, then 
<emphasis id="B923AD56FFF4DC7F553C5E3FFCBE0A12" box="[820,835,354,379]" italics="true" pageId="3" pageNumber="4">p</emphasis>
2, then 
<emphasis id="B923AD56FFF4DC7F559C5E3FFC5E0A12" box="[916,931,354,379]" italics="true" pageId="3" pageNumber="4">p</emphasis>
3. Distance between apophyses of 
<emphasis id="B923AD56FFF4DC7F533C5E3FFABE0A12" box="[1332,1347,354,379]" italics="true" pageId="3" pageNumber="4">p</emphasis>
1 and 
<emphasis id="B923AD56FFF4DC7F538D5E3FFA690A12" box="[1413,1428,354,379]" italics="true" pageId="3" pageNumber="4">h</emphasis>
3 230. Pleuraspis with dense tubercles in dorsal half.
</paragraph>
<caption id="DF2821CCFFF4DC7F569F59E0FC450DBC" box="[151,952,1725,1749]" httpUri="https://zenodo.org/record/189962/files/figure.png" pageId="3" pageNumber="4" targetBox="[212,1366,476,1675]" targetPageId="3">
<paragraph id="8BE87144FFF4DC7F569F59E0FC450DBC" blockId="3.[151,952,1725,1749]" box="[151,952,1725,1749]" pageId="3" pageNumber="4">
<emphasis id="B923AD56FFF4DC7F569F59E0FEE40DBC" bold="true" box="[151,281,1725,1749]" pageId="3" pageNumber="4">FIGURE 1.</emphasis>
<taxonomicName id="4C570AC7FFF4DC7F572859E3FE370DBC" box="[288,458,1726,1749]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="alpina" status="sp. nov.">
<emphasis id="B923AD56FFF4DC7F572859E3FE370DBC" box="[288,458,1726,1749]" italics="true" pageId="3" pageNumber="4">Crotonia alpina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A210102DFFF4DC7F57D859E0FDDB0DBC" box="[464,550,1725,1749]" pageId="3" pageNumber="4" rank="species">
<emphasis id="B923AD56FFF4DC7F57D859E0FDE20DBC" bold="true" box="[464,543,1725,1749]" pageId="3" pageNumber="4">sp. nov</emphasis>
.
</taxonomicNameLabel>
holotype female a) dorsal; b) ventral.
</paragraph>
</caption>
<paragraph id="8BE87144FFF4DC7F56CD585CFE630C75" blockId="3.[151,1438,1793,2013]" box="[197,414,1793,1820]" pageId="3" pageNumber="4">
<emphasis id="B923AD56FFF4DC7F56CD585CFEF90C72" bold="true" box="[197,260,1793,1819]" pageId="3" pageNumber="4">Male</emphasis>
. Not known.
</paragraph>
</subSubSection>
<subSubSection id="C34D22CFFFF4DC7856CD5875FDBE0A25" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="8BE87144FFF4DC7856CD5875FDFB0BB1" blockId="3.[151,1438,1793,2013]" lastBlockId="4.[151,1438,152,857]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<emphasis id="B923AD56FFF4DC7F56CD5875FD320C2B" bold="true" box="[197,719,1832,1858]" pageId="3" pageNumber="4">Material Examined and Locality Data.</emphasis>
<typeStatus id="54ECCFE6FFF4DC7F54DF5874FCB10C2A" box="[727,844,1833,1859]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
female and one 
<typeStatus id="54ECCFE6FFF4DC7F52295874FB730C2A" box="[1057,1166,1833,1859]" pageId="3" pageNumber="4" type="paratype">paratype</typeStatus>
female, sieved litter, 
<taxonomicName id="4C570AC7FFF4DC7F569F580DFEE40C00" box="[151,281,1872,1897]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="genus">
<emphasis id="B923AD56FFF4DC7F569F580DFEE40C00" box="[151,281,1872,1897]" italics="true" pageId="3" pageNumber="4">Eucalyptus</emphasis>
</taxonomicName>
forest, Eurobin Falls, Mount Buffalo National Park, Victoria, 
<geoCoordinate id="EE631783FFF4DC7F55E75812FB9B0C00" box="[1007,1126,1871,1897]" direction="south" orientation="latitude" pageId="3" pageNumber="4" precision="15" value="-36.7175">36°43'3&quot;S</geoCoordinate>
<geoCoordinate id="EE631783FFF4DC7F52655812FAF80C00" box="[1133,1285,1871,1897]" direction="east" orientation="longitude" pageId="3" pageNumber="4" precision="15" value="146.84138">146°50'29&quot;E</geoCoordinate>
, 
<quantity id="4CAFDCA1FFF4DC7F53195812FAA30C00" box="[1297,1374,1871,1897]" metricMagnitude="2" metricUnit="m" metricValue="4.6" pageId="3" pageNumber="4" unit="m" value="460.0">460 m</quantity>
, coll. D. Black. Two 
<typeStatus id="54ECCFE6FFF4DC7F5746582BFE480CF9" box="[334,437,1910,1936]" pageId="3" pageNumber="4" type="paratype">paratype</typeStatus>
females; soil, grass and litter on rocky slope, burnt area, 
<taxonomicName id="4C570AC7FFF4DC7F5264582BFB0C0CE6" box="[1132,1265,1910,1935]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="genus">
<emphasis id="B923AD56FFF4DC7F5264582BFB0C0CE6" box="[1132,1265,1910,1935]" italics="true" pageId="3" pageNumber="4">Eucalyptus</emphasis>
</taxonomicName>
forest, Mount Buffalo National Park, 
<geoCoordinate id="EE631783FFF4DC7F57A158C1FDCB0CDF" box="[425,566,1948,1974]" direction="south" orientation="latitude" pageId="3" pageNumber="4" precision="15" value="-36.710835">36°42’39”S</geoCoordinate>
, 146°50’06”, 680 meters above sea level (m, hereinafter), coll. J. Bloszyk &amp; S. Konwerski, 
<date id="FFE95784FFF4DC7F5775589EFDEE0CB4" box="[381,531,1987,2013]" pageId="3" pageNumber="4" value="2007-07-16">16.vii.2007</date>
(MTB-014). 
<typeStatus id="54ECCFE6FFF4DC7F54CE589EFCC20CB4" box="[710,831,1987,2013]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
and one 
<typeStatus id="54ECCFE6FFF4DC7F55B2589EFBD60CB4" box="[954,1067,1987,2013]" pageId="3" pageNumber="4" type="paratype">paratype</typeStatus>
deposited in Department of Entomology, Museum Victoria, Melbourne. Remaining 
<typeStatus id="54ECCFE6FFF3DC7855305FC5FC560BDB" box="[824,939,152,178]" pageId="4" pageNumber="5" type="paratype">paratypes</typeStatus>
in Australian National Insect Collection, 
<collectionCode id="ED46E981FFF3DC78569F5FE3FF130BB1" box="[151,238,190,216]" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="4" pageNumber="5">CSIRO</collectionCode>
Entomology, Canberra.
</paragraph>
<paragraph id="8BE87144FFF3DC7856CD5FB8FDBE0A25" blockId="4.[151,1438,152,857]" pageId="4" pageNumber="5">
<emphasis id="B923AD56FFF3DC7856CD5FB8FE8E0B97" box="[197,371,229,254]" italics="true" pageId="4" pageNumber="5">Other material</emphasis>
: one nymph, same locality data as 
<typeStatus id="54ECCFE6FFF3DC7855035FB8FC880B96" box="[779,885,229,255]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
; eight nymphs, MTB-014; one tritonymph, wet fern, 
<taxonomicName id="4C570AC7FFF3DC7856DD5E51FEAA0A4C" box="[213,343,268,293]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="genus">
<emphasis id="B923AD56FFF3DC7856DD5E51FEAA0A4C" box="[213,343,268,293]" italics="true" pageId="4" pageNumber="5">Eucalyptus</emphasis>
</taxonomicName>
forest, Mount Buffalo National Park, 
<geoCoordinate id="EE631783FFF3DC7855135E51FC550A4F" box="[795,936,268,294]" direction="south" orientation="latitude" pageId="4" pageNumber="5" precision="15" value="-36.710835">36°42’39”S</geoCoordinate>
, 146°50’06”, 
<quantity id="4CAFDCA1FFF3DC7852455E51FB660A4F" box="[1101,1179,268,294]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="4" pageNumber="5" unit="m" value="680.0">680 m</quantity>
, coll. J. Bloszyk &amp; S. Konwerski, 
<date id="FFE95784FFF3DC78572E5E6FFE560A25" box="[294,427,306,332]" pageId="4" pageNumber="5" value="2007-07-16">16.vii.2007</date>
(MTB-016).
</paragraph>
</subSubSection>
<subSubSection id="C34D22CFFFF3DC7856CD5E05FB880A1A" box="[197,1141,344,371]" pageId="4" pageNumber="5" type="etymology">
<paragraph id="8BE87144FFF3DC7856CD5E05FB880A1A" blockId="4.[151,1438,152,857]" box="[197,1141,344,371]" pageId="4" pageNumber="5">
<emphasis id="B923AD56FFF3DC7856CD5E05FEAF0A1B" bold="true" box="[197,338,344,370]" pageId="4" pageNumber="5">Etymology.</emphasis>
The species is named for its 
<typeStatus id="54ECCFE6FFF3DC7854AE5E04FD240A1A" box="[678,729,345,371]" pageId="4" pageNumber="5">type</typeStatus>
locality, within the Victorian Alps.
</paragraph>
</subSubSection>
<subSubSection id="C34D22CFFFF3DC7856CD5E22FEAD083D" pageId="4" pageNumber="5" type="discussion">
<paragraph id="8BE87144FFF3DC7856CD5E22FA9D0908" blockId="4.[151,1438,152,857]" pageId="4" pageNumber="5">
<emphasis id="B923AD56FFF3DC7856CD5E22FEC30AF0" bold="true" box="[197,318,383,409]" pageId="4" pageNumber="5">Remarks.</emphasis>
<taxonomicName id="4C570AC7FFF3DC78574E5EDDFDFE0AF0" box="[326,515,384,409]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="alpina" status="sp. nov.">
<emphasis id="B923AD56FFF3DC78574E5EDDFDFE0AF0" box="[326,515,384,409]" italics="true" pageId="4" pageNumber="5">Crotonia alpina</emphasis>
</taxonomicName>
<emphasis id="B923AD56FFF3DC7854035E22FD970AF0" bold="true" box="[523,618,383,409]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="A210102DFFF3DC7854035E22FD970AF0" box="[523,618,383,409]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from all other 
<taxonomicName id="4C570AC7FFF3DC78557A5EDDFC260AF0" box="[882,987,384,409]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B923AD56FFF3DC78557A5EDDFC260AF0" box="[882,987,384,409]" italics="true" pageId="4" pageNumber="5">Crotonia</emphasis>
</taxonomicName>
spp. by the following combination of characters: 1) with three pairs of 
<emphasis id="B923AD56FFF3DC7854205EFAFDC80AA9" box="[552,565,423,448]" italics="true" pageId="4" pageNumber="5">c</emphasis>
setae; 2) setae 
<emphasis id="B923AD56FFF3DC7854F85EFAFD000AA9" box="[752,765,423,448]" italics="true" pageId="4" pageNumber="5">c</emphasis>
3 extremely long, extending anteriorly as far as lamellar apophyses; 3) notogastral shield with narrow lateral strips of densely-packed small tubercles, and with extensive, diffuse tuberculate microsculpture on lateral, caudal and humeral regions; 4) bothridia without prominent anteriolateral auriculae; 5); setae 
<emphasis id="B923AD56FFF3DC7854955D43FD57095E" box="[669,682,542,567]" italics="true" pageId="4" pageNumber="5">c</emphasis>
1 about half the length of 
<emphasis id="B923AD56FFF3DC7855DB5D43FC1D095E" box="[979,992,542,567]" italics="true" pageId="4" pageNumber="5">c</emphasis>
2, which are a third of the length of 
<emphasis id="B923AD56FFF3DC7853775D43FA71095E" box="[1407,1420,542,567]" italics="true" pageId="4" pageNumber="5">c</emphasis>
3; 6) inter-bothridial ridge a distinctive crenellated shallow curve; 7) setae 
<emphasis id="B923AD56FFF3DC7855EC5D1AFC0F0909" box="[996,1010,583,608]" italics="true" pageId="4" pageNumber="5">d</emphasis>
2 very long, as long as setae 
<emphasis id="B923AD56FFF3DC7853355D1AFAA50909" box="[1341,1368,583,608]" italics="true" pageId="4" pageNumber="5">cp</emphasis>
.
</paragraph>
<paragraph id="8BE87144FFF3DC7856CD5D2CFEAD083D" blockId="4.[151,1438,152,857]" pageId="4" pageNumber="5">
<taxonomicName id="4C570AC7FFF3DC7856CD5D2CFE6509E3" box="[197,408,625,650]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="alpina" status="sp. nov.">
<emphasis id="B923AD56FFF3DC7856CD5D2CFE6509E3" box="[197,408,625,650]" italics="true" pageId="4" pageNumber="5">Crotonia alpina</emphasis>
</taxonomicName>
<emphasis id="B923AD56FFF3DC7857AA5D2DFDF009E3" bold="true" box="[418,525,624,650]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="A210102DFFF3DC7857AA5D2DFDF009E3" box="[418,525,624,650]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is a member of the 
<taxonomicName id="4C570AC7FFF3DC7855125D2CFC5C09E2" box="[794,929,625,651]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="34" phylum="Arthropoda" rank="species" species="capistrata">Capistrata</taxonomicName>
species-group (
<bibRefCitation id="EFC60CB5FFF3DC78527C5D2CFAC109E2" author="Colloff" box="[1140,1340,625,651]" pageId="4" pageNumber="35" refString="Colloff, M. J. (2009 b) Species-group concepts and biogeography of the genus Crotonia (Acari: Oribatida: Crotoniidae), with new species from South and Central America. Zootaxa, 2081, 1 - 30." type="journal article" year="2009" yearSuffix="b">Colloff, 2009b</bibRefCitation>
) and is morphologically most similar to 
<taxonomicName id="4C570AC7FFF3DC7854135DCAFC8F09D8" authority="Colloff, 2009" authorityName="Colloff" authorityYear="2009" box="[539,882,663,689]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="tasmaniana">
<emphasis id="B923AD56FFF3DC7854135DCAFD3A09D9" box="[539,711,663,688]" italics="true" pageId="4" pageNumber="5">C. tasmaniana</emphasis>
Colloff, 2009
</taxonomicName>
. It differs in having setae 
<emphasis id="B923AD56FFF3DC7852AE5DCAFB4E09D9" box="[1190,1203,663,688]" italics="true" pageId="4" pageNumber="5">c</emphasis>
3 and 
<emphasis id="B923AD56FFF3DC7852FD5DCAFAFF09D9" box="[1269,1282,663,688]" italics="true" pageId="4" pageNumber="5">c</emphasis>
1 about twice as long, and 
<emphasis id="B923AD56FFF3DC7857255D9CFEC109B3" box="[301,316,705,730]" italics="true" pageId="4" pageNumber="5">d</emphasis>
2 at least three times as long, as those of 
<taxonomicName id="4C570AC7FFF3DC7855155D9CFC3409B3" box="[797,969,705,730]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="tasmaniana">
<emphasis id="B923AD56FFF3DC7855155D9CFC3409B3" box="[797,969,705,730]" italics="true" pageId="4" pageNumber="5">C. tasmaniana</emphasis>
</taxonomicName>
, and 
<emphasis id="B923AD56FFF3DC7852035D9CFBE709B3" box="[1035,1050,705,730]" italics="true" pageId="4" pageNumber="5">h</emphasis>
2 curved, flagelliform rather than spiniform. The epimeres of 
<taxonomicName id="4C570AC7FFF3DC7857D45DB7FDB4086A" box="[476,585,746,771]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="alpina">
<emphasis id="B923AD56FFF3DC7857D45DB7FDB4086A" box="[476,585,746,771]" italics="true" pageId="4" pageNumber="5">C. alpina</emphasis>
</taxonomicName>
are porose whereas those of 
<taxonomicName id="4C570AC7FFF3DC7855945DB7FBBA086A" box="[924,1095,746,771]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="tasmaniana">
<emphasis id="B923AD56FFF3DC7855945DB7FBBA086A" box="[924,1095,746,771]" italics="true" pageId="4" pageNumber="5">C. tasmaniana</emphasis>
</taxonomicName>
are tuberculate laterally. The apophyses of setae 
<emphasis id="B923AD56FFF3DC7857735C4CFE770843" box="[379,394,785,810]" italics="true" pageId="4" pageNumber="5">h</emphasis>
3 of 
<taxonomicName id="4C570AC7FFF3DC7857B25C4CFDD50843" box="[442,552,785,810]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="alpina">
<emphasis id="B923AD56FFF3DC7857B25C4CFDD50843" box="[442,552,785,810]" italics="true" pageId="4" pageNumber="5">C. alpina</emphasis>
</taxonomicName>
are markedly posterior of those of setae 
<emphasis id="B923AD56FFF3DC7852025C4CFBE40843" box="[1034,1049,785,810]" italics="true" pageId="4" pageNumber="5">p</emphasis>
1 whereas in 
<taxonomicName id="4C570AC7FFF3DC7852A75C4CFAA60843" box="[1199,1371,785,810]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="tasmaniana">
<emphasis id="B923AD56FFF3DC7852A75C4CFAA60843" box="[1199,1371,785,810]" italics="true" pageId="4" pageNumber="5">C. tasmaniana</emphasis>
</taxonomicName>
, they are lateral of 
<emphasis id="B923AD56FFF3DC78573A5C66FEBD083D" box="[306,320,827,852]" italics="true" pageId="4" pageNumber="5">p</emphasis>
1.
</paragraph>
</subSubSection>
</treatment>
</document>