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<document id="1003AE1F724DC7D0C135C4F81256BDEE" ID-DOI="10.5281/zenodo.5170203" ID-ISSN="1942-1354" ID-Zenodo-Dep="5170203" ID-ZooBank="AB1A57F0-7CB4-4830-920B-DF219740A596" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628281492031" checkinUser="felipe" docAuthor="Blahnik, Roger J. &amp; Holzenthal, Ralph W." docDate="2017" docId="03F687A7FF96F868FF01BC264409FBAF" docLanguage="en" docName="InsectaMundi.0602.1-251.pdf" docOrigin="Insecta Mundi 2017 (602)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Mortoniella (Nanotrichia) spangleri Blahnik &amp; Holzenthal 2017, new species" docType="treatment" docVersion="4" lastPageNumber="110" masterDocId="FFCFFFDFFFF8F807FFC1B865400BFFB4" masterDocTitle="Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *" masterLastPageNumber="251" masterPageNumber="1" pageNumber="109" updateTime="1699039969677" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="D56578D926E45EAC7604C6D960F45B77" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C5D47692037AA577168ECF7A086047CC">
<mods:title id="1002EC0D096C8A751098D71C951A55D6">Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *</mods:title>
</mods:titleInfo>
<mods:name id="F238CE369E8F1C2DFD96D65FC35A61F3" type="personal">
<mods:role id="B7995C6C2B91F85614C01AB11DF981FD">
<mods:roleTerm id="ABD2904DE383521C9F533DBBF90BCD48">Author</mods:roleTerm>
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<mods:namePart id="FB82BC879CD05A123FD6EF5F9C029A0B">Blahnik, Roger J.</mods:namePart>
</mods:name>
<mods:name id="36D5A26895510A6150A7544134BC89ED" type="personal">
<mods:role id="932EBC3CD4405A11392354E759CE83A3">
<mods:roleTerm id="17DC78AD9E37A2675C1CDBDB77E8C3D1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9931A9AE188E91C52D8AAB73833683A2">Holzenthal, Ralph W.</mods:namePart>
</mods:name>
<mods:typeOfResource id="C47277BDCAB45F88109E2620AB014FC9">text</mods:typeOfResource>
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<mods:title id="C306B7740AB649BFCE1D9C6BFC4180D6">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="663D583AB5D9F1D06987159F02C2680D">
<mods:date id="8010A013846D0A10DF117E5073F54196">2017</mods:date>
<mods:detail id="33D6D9950FC9ABB4AFB8174517312BAC" type="pubDate">
<mods:number id="D26C64F34505414525233CBA40AA235D">2017-12-29</mods:number>
</mods:detail>
<mods:detail id="FEC026E989BF3149B2AF68A4DD2EADD2" type="volume">
<mods:number id="40557C00F28C8545F391D6A60463017C">2017</mods:number>
</mods:detail>
<mods:detail id="16CCC771BE1BAB4A3FBFD3261C535C69" type="issue">
<mods:number id="38CA9B31D18060479C89C91E762DE91D">602</mods:number>
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<mods:start id="28FD817A5B23458F01F5B964E04D551A">1</mods:start>
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</mods:extent>
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<mods:classification id="250D24E33C1069EC38BF3278B7F545D6">journal article</mods:classification>
<mods:identifier id="FEA99F1B6B37156AB6B2EA056368DDB1" type="DOI">10.5281/zenodo.5170203</mods:identifier>
<mods:identifier id="58DED6C57835F8FBC38C1706DB9B3E15" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="893FA13F5AC930C53E038CDBC15CF28A" type="Zenodo-Dep">5170203</mods:identifier>
<mods:identifier id="B6362F2DB116602B29E0A8B744D76DDD" type="ZooBank">AB1A57F0-7CB4-4830-920B-DF219740A596</mods:identifier>
</mods:mods>
<treatment id="03F687A7FF96F868FF01BC264409FBAF" ID-DOI="http://doi.org/10.5281/zenodo.5186349" ID-Zenodo-Dep="5186349" LSID="urn:lsid:plazi:treatment:03F687A7FF96F868FF01BC264409FBAF" httpUri="http://treatment.plazi.org/id/03F687A7FF96F868FF01BC264409FBAF" lastPageId="111" lastPageNumber="110" pageId="110" pageNumber="109">
<subSubSection id="C345653AFF96F869FF01BC264365FBEF" box="[192,878,1091,1115]" pageId="110" pageNumber="109" type="nomenclature">
<paragraph id="8BE036B1FF96F869FF01BC264365FBEF" blockId="110.[192,878,1091,1147]" box="[192,878,1091,1115]" pageId="110" pageNumber="109">
<heading id="D0A881DDFF96F869FF01BC264365FBEF" bold="true" box="[192,878,1091,1115]" fontSize="10" level="2" pageId="110" pageNumber="109" reason="3">
<taxonomicName id="4C5F4D32FF96F869FF01BC2642BCFBEF" authority="Blahnik &amp; Holzenthal, 2017" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" box="[192,695,1091,1115]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="110" pageNumber="109" phylum="Tracheophyta" rank="species" species="spangleri" status="sp. nov." subGenus="Nanotrichia">
<emphasis id="B92BEAA3FF96F869FF01BC2642BCFBEF" bold="true" box="[192,695,1091,1115]" pageId="110" pageNumber="109">
<emphasis id="B92BEAA3FF96F869FF01BC264168FBEF" bold="true" box="[192,355,1091,1115]" italics="true" pageId="110" pageNumber="109">Mortoniella</emphasis>
(
<emphasis id="B92BEAA3FF96F869FEB7BC26422AFBEF" bold="true" box="[374,545,1091,1115]" italics="true" pageId="110" pageNumber="109">Nanotrichia</emphasis>
)
<emphasis id="B92BEAA3FF96F869FDF2BC2642BCFBEF" bold="true" box="[563,695,1091,1115]" italics="true" pageId="110" pageNumber="109">spangleri</emphasis>
</emphasis>
</taxonomicName>
,
<emphasis id="B92BEAA3FF96F869FD06BC264365FBEF" bold="true" box="[711,878,1091,1115]" pageId="110" pageNumber="109">
<taxonomicNameLabel id="A21857D8FF96F869FD06BC264365FBEF" box="[711,878,1091,1115]" pageId="110" pageNumber="109" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C345653AFF96F868FF01BC064305FE4F" lastPageId="111" lastPageNumber="110" pageId="110" pageNumber="109" type="description">
<paragraph id="8BE036B1FF96F869FF01BC064159FBCF" blockId="110.[192,878,1091,1147]" box="[192,338,1123,1147]" pageId="110" pageNumber="109">
<figureCitation id="13642A34FF96F869FF01BC064112FBCF" box="[192,281,1123,1147]" captionStart="Figure 94" captionStartId="234.[192,276,1836,1858]" captionTargetBox="[355,1284,244,1734]" captionTargetId="figure-59@234.[355,1284,244,1734]" captionTargetPageId="234" captionText="Figure 94. Mortoniella (Nanotrichia) spangleri, new species, male genitalia. A—lateral; B—segment IX and tergum X, dorsal; C—phallic ensemble, ventral; D—dorsal phallic spine and phallobase, dorsal; E—ventral process of segment VI, lateral." figureDoi="http://doi.org/10.5281/zenodo.5170403" httpUri="https://zenodo.org/record/5170403/files/figure.png" pageId="110" pageNumber="109">Fig. 94</figureCitation>
,
<figureCitation id="13642A34FF96F869FEE4BC064159FBCF" box="[293,338,1123,1147]" captionStart="Figures 121-125" captionStartId="248.[192,287,1807,1829]" captionTargetBox="[302,1326,269,1692]" captionTargetId="figure-75@248.[302,1327,269,1692]" captionTargetPageId="248" captionText="Figures 121-125. Mortoniella (Nanotrichia), velasquezi group species, female genitalia.121—Mortoniella eduardoi (Rueda Martín and Gibon): A—lateral; B—dorsal. 122—Mortoniella licina, n. sp., dorsal. 123—Mortoniella venezuelensis, n. sp., dorsal. 124—Mortoniella spangleri, n. sp. A—lateral; B—dorsal. 125—Mortoniella coheni, n. sp., dorsal." figureDoi="http://doi.org/10.5281/zenodo.5170423" httpUri="https://zenodo.org/record/5170423/files/figure.png" pageId="110" pageNumber="109">124</figureCitation>
</paragraph>
<paragraph id="8BE036B1FF96F869FF31BCC64559FAEF" blockId="110.[192,1441,1187,1372]" pageId="110" pageNumber="109">This is a distinctive species, unlikely to be confused with any other species in the velasquezi group. Especially diagnostic is the structure of the dorsal phallic spine, which has a patch of minute spines on its posteroventral surface. Other useful diagnostic characters include the strongly curved paramere appendages, relatively short inferior appendages, and the distinctively arched tergum X (as viewed laterally). Additionally, males have an extensive field of darkened scale-like setae on the hind wings. The female is also distinctive in that it has an arched and projecting tergum VIII.</paragraph>
<paragraph id="8BE036B1FF96F869FF31BDE6437EF9AF" blockId="110.[192,1440,1411,1915]" pageId="110" pageNumber="109">
<emphasis id="B92BEAA3FF96F869FF31BDE6413DFA2F" box="[240,310,1411,1435]" italics="true" pageId="110" pageNumber="109">Adult</emphasis>
—Length of forewing: male
<quantity id="4CA79B54FF96F869FD48BDE6431DFA2F" box="[649,790,1411,1435]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="110" pageNumber="109" unit="mm" value="3.0" valueMax="3.2" valueMin="2.8">2.8-3.2 mm</quantity>
; female 3.0-
<quantity id="4CA79B54FF96F869FC69BDE6440DFA2F" box="[936,1030,1411,1435]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="110" pageNumber="109" unit="mm" value="3.5">3.5 mm</quantity>
. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0:3:4. Overall color (in alcohol) light brown. Tibial spurs slightly darker than legs, contrasting in color. Forewing with distinct white wing bar at anastomosis. Males with scale-like setae paralleling veins in fore- and hind wing, hind wing with extensive field of darkened, semi-erect, scale-like setae.
</paragraph>
<paragraph id="8BE036B1FF96F868FF31BE464305FE4F" blockId="110.[192,1440,1411,1915]" lastBlockId="111.[192,1441,195,1052]" lastPageId="111" lastPageNumber="110" pageId="110" pageNumber="109">
<emphasis id="B92BEAA3FF96F869FF31BE4641A4F98F" box="[240,431,1571,1595]" italics="true" pageId="110" pageNumber="109">Male genitalia</emphasis>
—Ventral process of segment VI posteriorly projecting, narrow and slightly constricted basally, length about 4 times width at base. Segment IX with anterolateral margin rounded and distinctly produced in ventral ½, posterolateral margin nearly straight; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by about ½ width of segment. Tergum X, as viewed laterally, relatively short and narrow, distinctly arched in middle; as viewed dorsally, short and moderate in width, lateral margins subparallel, apicomesally with deep emargination, extending more than ½ length of tergum, basomesally with pair of setose sclerites; ventrolateral lobes obsolete. Inferior appendage with digitate, setose, dorsolateral projections, fused apicoventrally to spine-like projections of the mesal pockets. Mesal pockets of inferior appendage very large, with moderately elongate, thick, spine-like, apicoventral projections. Paramere appendage moderately elongate, narrow, distinctly curved, ventrally near base and dorsally at apex, apex acute. Phallobase very short, with elongate ventral rod-like projections, each strongly flared apically. Dorsal phallic spine, as viewed laterally, almost evenly dorsally curved from base, apex slightly inflated, ventral margin with distinct notch near middle, apparently to accommodate lightly sclerotized projection on endophallic membrane, upturned part with slight indentation bearing numerous small spines; as viewed dorsally, with distinct dorsolateral projections, widest in basal ½, narrowing apically, reflexed apex rounded. Phallicata short, continuous with endophallic membrane apically, laterally with broadly rounded sclerotized projections, surrounding mesal pockets of inferior appendage. Endophallic membrane continuous with phallicata, dorsomesally with lightly sclerotized projection, ventrally with pair of short sclerotized projections (modified phallotremal spines?), each bluntly rounded as viewed ventrally, apices with minute spines.
</paragraph>
</subSubSection>
<subSubSection id="C345653AFF97F868FF31BA664409FBAF" pageId="111" pageNumber="110" type="materials_examined">
<paragraph id="8BE036B1FF97F868FF31BA664222FD8F" blockId="111.[192,1441,195,1052]" pageId="111" pageNumber="110">
<materialsCitation id="3B373CECFF97F868FF31BA664222FD8F" collectingDate="1977-05-05" collectionCode="DR, NMNH" collectorName="PJ Spangler &amp; Givens" country="Ecuador" location="Puyo" pageId="111" pageNumber="110" specimenCode="UMSP000097077" specimenCount="1" stateProvince="Pastaza" typeStatus="holotype">
<emphasis id="B92BEAA3FF97F868FF31BA6641B1FDAF" bold="true" box="[240,442,515,539]" italics="true" pageId="111" pageNumber="110">
<typeStatus id="54E48813FF97F868FF31BA664161FDAF" box="[240,362,515,539]" pageId="111" pageNumber="110" type="holotype">Holotype</typeStatus>
male
</emphasis>
(alcohol)—
<collectingCountry id="F3487621FF97F868FD8ABA6642D5FDAF" box="[587,734,515,539]" name="Ecuador" pageId="111" pageNumber="110">ECUADOR</collectingCountry>
:
<collectingRegion id="499BF853FF97F868FD2DBA614358FDAF" box="[748,851,516,539]" country="Ecuador" name="Pastaza" pageId="111" pageNumber="110">Pastaza</collectingRegion>
:
<location id="8E80606AFF97F868FCA2BA6143AEFDA8" LSID="urn:lsid:plazi:treatment:03F687A7FF96F868FF01BC264409FBAF:8E80606AFF97F868FCA2BA6143AEFDA8" box="[867,933,516,540]" country="Ecuador" name="Puyo" pageId="111" pageNumber="110" stateProvince="Pastaza">Puyo</location>
,
<date id="FFE11071FF97F868FC74BA66442AFDAF" box="[949,1057,515,539]" pageId="111" pageNumber="110" value="1977-05-05">
<collectingDate id="EFA5E999FF97F868FC74BA66442AFDAF" box="[949,1057,515,539]" pageId="111" pageNumber="110" value="1977-05-05">5.v.1977</collectingDate>
</date>
,
<collectorName id="26AA5367FF97F868FBF3BA6144C4FDAF" box="[1074,1231,515,539]" pageId="111" pageNumber="110">PJ Spangler</collectorName>
and
<collectionCode id="ED4EAE74FF97F868FAD4BA614536FDAF" box="[1301,1341,516,539]" country="Germany" lsid="urn:lsid:biocol.org:col:15173" name="Technische Universität Dresden" pageId="111" pageNumber="110" type="Herbarium">DR</collectionCode>
<collectorName id="26AA5367FF97F868FA88BA6645ABFDAF" box="[1353,1440,515,539]" pageId="111" pageNumber="110">Givens</collectorName>
(
<specimenCode id="DBF99ECAFF97F868FF09BA4641A8FD8F" box="[200,419,547,571]" collectionCode="UMSP" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="111" pageNumber="110" type="University or college">UMSP000097077</specimenCode>
) (
<collectionCode id="ED4EAE74FF97F868FE7DBA464216FD8F" box="[444,541,547,571]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="111" pageNumber="110" type="Museum">NMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BE036B1FF97F868FF31BA26453FFC0F" blockId="111.[192,1441,195,1052]" pageId="111" pageNumber="110">
<materialsCitation id="3B373CECFF97F868FF31BA2641A9FDCF" collectingDate="1977-05-06" collectionCode="DR, NMNH" collectorName="Puyo &amp; PJ Spangler" country="Ecuador" location="Givens" pageId="111" pageNumber="110" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Napo" typeStatus="paratype">
<emphasis id="B92BEAA3FF97F868FF31BA26429AFDEF" bold="true" box="[240,657,579,603]" pageId="111" pageNumber="110">
<typeStatus id="54E48813FF97F868FF31BA264175FDEF" box="[240,382,579,603]" pageId="111" pageNumber="110" type="paratype">Paratypes</typeStatus>
—
<collectingCountry id="F3487621FF97F868FE58BA264239FDEF" box="[409,562,579,603]" name="Ecuador" pageId="111" pageNumber="110">ECUADOR</collectingCountry>
:
<collectingRegion id="499BF853FF97F868FD81BA264280FDEF" box="[576,651,579,603]" country="Ecuador" name="Napo" pageId="111" pageNumber="110">Napo</collectingRegion>
:
</emphasis>
<collectorName id="26AA5367FF97F868FD58BA2142D1FDE8" box="[665,730,580,604]" pageId="111" pageNumber="110">Puyo</collectorName>
,
<date id="FFE11071FF97F868FD29BA264359FDEF" box="[744,850,579,603]" pageId="111" pageNumber="110" value="1977-05-06">
<collectingDate id="EFA5E999FF97F868FD29BA264359FDEF" box="[744,850,579,603]" pageId="111" pageNumber="110" value="1977-05-06">6.v.1977</collectingDate>
</date>
,
<collectorName id="26AA5367FF97F868FCA1BA2143F1FDEF" box="[864,1018,579,603]" pageId="111" pageNumber="110">PJ Spangler</collectorName>
and
<collectionCode id="ED4EAE74FF97F868FBFBBA214469FDEF" box="[1082,1122,580,603]" country="Germany" lsid="urn:lsid:biocol.org:col:15173" name="Technische Universität Dresden" pageId="111" pageNumber="110" type="Herbarium">DR</collectionCode>
<location id="8E80606AFF97F868FBAABA2644CEFDEF" LSID="urn:lsid:plazi:treatment:03F687A7FF96F868FF01BC264409FBAF:8E80606AFF97F868FBAABA2644CEFDEF" box="[1131,1221,579,603]" country="Ecuador" name="Givens" pageId="111" pageNumber="110" stateProvince="Napo">Givens</location>
–
<specimenCount id="9D59FD38FF97F868FB17BA264523FDEF" box="[1238,1320,579,603]" pageId="111" pageNumber="110" type="male">1 male</specimenCount>
,
<specimenCount id="9D59FD38FF97F868FAF4BA2645ABFDEF" box="[1333,1440,579,603]" pageId="111" pageNumber="110" type="female">1 female</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FF97F868FEFBBA064191FDCF" box="[314,410,611,635]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="111" pageNumber="110" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CECFF97F868FE6EBA0141ABFD2F" collectingDate="1975-10-17" collectionCode="E, A, NMNH" collectorName="Lago Agrio &amp; Via &amp; Tarapoa" country="Ecuador" location="Langley" pageId="111" pageNumber="110" specimenCount="47" specimenCount-female="25" specimenCount-male="22" stateProvince="Napo" typeStatus="paratype">
<collectorName id="26AA5367FF97F868FE6EBA014233FDCF" box="[431,568,611,636]" pageId="111" pageNumber="110">Lago Agrio</collectorName>
(
<quantity id="4CA79B54FF97F868FD89BA06429CFDCF" box="[584,663,611,635]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="111" pageNumber="110" unit="km" value="30.0">30 km</quantity>
<collectionCode id="ED4EAE74FF97F868FD5EBA0642B8FDCF" box="[671,691,611,635]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="111" pageNumber="110" type="Herbarium">E</collectionCode>
)
<collectorName id="26AA5367FF97F868FD02BA0642E5FDCF" box="[707,750,611,635]" pageId="111" pageNumber="110">Via</collectorName>
a
<collectorName id="26AA5367FF97F868FCCFBA014372FDCF" box="[782,889,612,635]" pageId="111" pageNumber="110">Tarapoa</collectorName>
,
<date id="FFE11071FF97F868FC44BA0643F5FDCF" box="[901,1022,611,635]" pageId="111" pageNumber="110" value="1975-10-17">
<collectingDate id="EFA5E999FF97F868FC44BA0643F5FDCF" box="[901,1022,611,635]" pageId="111" pageNumber="110" value="1975-10-17">17.x.1975</collectingDate>
</date>
,
<collectionCode id="ED4EAE74FF97F868FBCDBA064414FDCF" box="[1036,1055,611,635]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="111" pageNumber="110" type="Herbarium">A</collectionCode>
<location id="8E80606AFF97F868FBE9BA064480FDCF" LSID="urn:lsid:plazi:treatment:03F687A7FF96F868FF01BC264409FBAF:8E80606AFF97F868FBE9BA064480FDCF" box="[1064,1163,611,635]" country="Ecuador" name="Langley" pageId="111" pageNumber="110" stateProvince="Napo">Langley</location>
–
<specimenCount id="9D59FD38FF97F868FB5ABA064506FDCF" box="[1179,1293,611,635]" pageId="111" pageNumber="110" type="male">22 males</specimenCount>
,
<specimenCount id="9D59FD38FF97F868FADBBA0645ABFDCF" box="[1306,1440,611,635]" pageId="111" pageNumber="110" type="female">25 females</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FF97F868FEF9BAE64193FD2F" box="[312,408,643,667]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="111" pageNumber="110" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CECFF97F868FE6ABAE6413BFD0F" collectingDate="1977-05-05" collectionCode="DR, NMNH" collectorName="Puyo &amp; PJ Spangler" country="Ecuador" location="Givens" pageId="111" pageNumber="110" specimenCount="12" specimenCount-female="7" specimenCount-male="5" stateProvince="Pastaza" typeStatus="paratype">
<emphasis id="B92BEAA3FF97F868FE6ABAE6422BFD2F" bold="true" box="[427,544,643,667]" pageId="111" pageNumber="110">
<collectingRegion id="499BF853FF97F868FE6ABAE64211FD2F" box="[427,538,643,667]" country="Ecuador" name="Pastaza" pageId="111" pageNumber="110">Pastaza</collectingRegion>
:
</emphasis>
<collectorName id="26AA5367FF97F868FDE7BAE1426CFD28" box="[550,615,644,668]" pageId="111" pageNumber="110">Puyo</collectorName>
,
<date id="FFE11071FF97F868FDB3BAE642D6FD2F" box="[626,733,643,667]" pageId="111" pageNumber="110" value="1977-05-05">
<collectingDate id="EFA5E999FF97F868FDB3BAE642D6FD2F" box="[626,733,643,667]" pageId="111" pageNumber="110" value="1977-05-05">5.v.1977</collectingDate>
</date>
,
<collectorName id="26AA5367FF97F868FD29BAE1438BFD2F" box="[744,896,643,667]" pageId="111" pageNumber="110">PJ Spangler</collectorName>
and
<collectionCode id="ED4EAE74FF97F868FC7ABAE143E8FD2F" box="[955,995,644,667]" country="Germany" lsid="urn:lsid:biocol.org:col:15173" name="Technische Universität Dresden" pageId="111" pageNumber="110" type="Herbarium">DR</collectionCode>
<location id="8E80606AFF97F868FC2BBAE6444AFD2F" LSID="urn:lsid:plazi:treatment:03F687A7FF96F868FF01BC264409FBAF:8E80606AFF97F868FC2BBAE6444AFD2F" box="[1002,1089,643,667]" country="Ecuador" name="Givens" pageId="111" pageNumber="110" stateProvince="Pastaza">Givens</location>
–
<specimenCount id="9D59FD38FF97F868FB90BAE644BBFD2F" box="[1105,1200,643,667]" pageId="111" pageNumber="110" type="male">5 males</specimenCount>
,
<specimenCount id="9D59FD38FF97F868FB7ABAE1453AFD2F" box="[1211,1329,643,667]" pageId="111" pageNumber="110" type="female">7 females</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FF97F868FF09BAC64123FD0F" box="[200,296,675,699]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="111" pageNumber="110" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CECFF97F868FEFFBAC144E6FD0F" box="[318,1261,675,699]" collectingDate="1977-05-06" collectionCode="UMSP" collectorName="Puyo &amp; PJ Spangler" country="Ecuador" location="Givens" pageId="111" pageNumber="110" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Pastaza" typeStatus="paratype">
same locality and collectors,
<date id="FFE11071FF97F868FD68BAC6431BFD0F" box="[681,784,675,699]" pageId="111" pageNumber="110" value="1977-05-06">
<collectingDate id="EFA5E999FF97F868FD68BAC6431BFD0F" box="[681,784,675,699]" pageId="111" pageNumber="110" value="1977-05-06">6.v.1977</collectingDate>
</date>
–
<specimenCount id="9D59FD38FF97F868FCE3BAC6438EFD0F" box="[802,901,675,699]" pageId="111" pageNumber="110" type="male">3 males</specimenCount>
,
<specimenCount id="9D59FD38FF97F868FC52BAC64400FD0F" box="[915,1035,675,699]" pageId="111" pageNumber="110" type="female">2 females</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FF97F868FB4EBAC644EEFD0F" box="[1167,1253,675,699]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="111" pageNumber="110" type="University or college">UMSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CECFF97F868FB3ABAC14334FD6F" collectingDate="1977-05-07" collectionCode="NMNH" collectorName="Puyo &amp; PJ Spangler" country="Ecuador" location="Givens" pageId="111" pageNumber="110" specimenCount="3" specimenCount-female="3" stateProvince="Pastaza" typeStatus="paratype">
same locality and collectors,
<date id="FFE11071FF97F868FEB5BAA641D1FD6F" box="[372,474,707,731]" pageId="111" pageNumber="110" value="1977-05-07">
<collectingDate id="EFA5E999FF97F868FEB5BAA641D1FD6F" box="[372,474,707,731]" pageId="111" pageNumber="110" value="1977-05-07">7.v.1977</collectingDate>
</date>
–
<specimenCount id="9D59FD38FF97F868FE2ABAA64256FD6F" box="[491,605,707,731]" pageId="111" pageNumber="110" type="female">3 females</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FF97F868FD19BAA6433CFD6F" box="[728,823,707,731]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="111" pageNumber="110" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CECFF97F868FC88BAA141A8FD4F" collectingDate="1976-01-30" collectionCode="NMNH" collectorName="Puyo &amp; Spangler" country="Ecuador" pageId="111" pageNumber="110" specimenCount="8" specimenCount-female="7" specimenCount-male="1" stateProvince="Pastaza" typeStatus="paratype">
<collectorName id="26AA5367FF97F868FC88BAA14381FD68" box="[841,906,708,732]" pageId="111" pageNumber="110">Puyo</collectorName>
,
<date id="FFE11071FF97F868FC55BAA6440DFD6F" box="[916,1030,707,731]" pageId="111" pageNumber="110" value="1976-01-30">
<collectingDate id="EFA5E999FF97F868FC55BAA6440DFD6F" box="[916,1030,707,731]" pageId="111" pageNumber="110" value="1976-01-30">30.i.1976</collectingDate>
</date>
,
<collectorName id="26AA5367FF97F868FBD1BAA64474FD6F" box="[1040,1151,707,731]" pageId="111" pageNumber="110">Spangler</collectorName>
et al.–
<specimenCount id="9D59FD38FF97F868FB12BAA64528FD6F" box="[1235,1315,707,731]" pageId="111" pageNumber="110" type="male">1 male</specimenCount>
,
<specimenCount id="9D59FD38FF97F868FAECBAA145ABFD6F" box="[1325,1440,707,731]" pageId="111" pageNumber="110" type="female">7 females</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FF97F868FEFABA864190FD4F" box="[315,411,739,763]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="111" pageNumber="110" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CECFF97F868FE71BA8144E5FD4F" box="[432,1262,739,763]" collectingDate="1976-02-01" collectingDateMax="1976-02-07" collectingDateMin="1976-02-01" collectionCode="NMNH" collectorName="Puyo &amp; Spangler" country="Ecuador" pageId="111" pageNumber="110" specimenCount="1" specimenCount-male="1" stateProvince="Pastaza" typeStatus="paratype">
same locality and collectors,
<date id="FFE11071FF97F868FCDABA864394FD4F" box="[795,927,739,763]" pageId="111" pageNumber="110" value="1976-02-01" valueMax="1976-02-07" valueMin="1976-02-01">
<collectingDate id="EFA5E999FF97F868FCDABA864394FD4F" box="[795,927,739,763]" pageId="111" pageNumber="110" value="1976-02-01" valueMax="1976-02-07" valueMin="1976-02-01">1-7.ii.1976</collectingDate>
</date>
–
<specimenCount id="9D59FD38FF97F868FC73BA864409FD4F" box="[946,1026,739,763]" pageId="111" pageNumber="110" type="male">1 male</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FF97F868FB47BA8644EDFD4F" box="[1158,1254,739,763]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="111" pageNumber="110" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CECFF97F868FB3ABA81436FFCAF" collectingDate="1976-02-08" collectingDateMax="1976-02-11" collectingDateMin="1976-02-08" collectionCode="NMNH" collectorName="Puyo &amp; Spangler" country="Ecuador" pageId="111" pageNumber="110" specimenCount="1" specimenCount-female="1" stateProvince="Pastaza" typeStatus="paratype">
same locality and collectors,
<date id="FFE11071FF97F868FEB7BB664202FCAF" box="[374,521,771,795]" pageId="111" pageNumber="110" value="1976-02-08" valueMax="1976-02-11" valueMin="1976-02-08">
<collectingDate id="EFA5E999FF97F868FEB7BB664202FCAF" box="[374,521,771,795]" pageId="111" pageNumber="110" value="1976-02-08" valueMax="1976-02-11" valueMin="1976-02-08">8-11.ii.1976</collectingDate>
</date>
–
<specimenCount id="9D59FD38FF97F868FDDDBB664274FCAF" box="[540,639,771,795]" pageId="111" pageNumber="110" type="female">1 female</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FF97F868FD3DBB664357FCAF" box="[764,860,771,795]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="111" pageNumber="110" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CECFF97F868FCAEBB6142EFFC8F" collectingDate="1976-02-04" collectionCode="N, NMNH" collectorName="Puyo &amp; Est. Fluvia Metrica" country="Ecuador" location="Spangler" pageId="111" pageNumber="110" specimenCount="3" specimenCount-female="3" stateProvince="Pastaza" typeStatus="paratype">
<collectorName id="26AA5367FF97F868FCAEBB6143A6FCA8" box="[879,941,772,796]" pageId="111" pageNumber="110">Puyo</collectorName>
(
<quantity id="4CA79B54FF97F868FC7ABB66440CFCAF" box="[955,1031,771,795]" metricMagnitude="4" metricUnit="m" metricValue="2.7" pageId="111" pageNumber="110" unit="km" value="27.0">27 km</quantity>
<collectionCode id="ED4EAE74FF97F868FBCCBB664428FCAF" box="[1037,1059,771,795]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="111" pageNumber="110" type="Herbarium">N</collectionCode>
)
<collectorName id="26AA5367FF97F868FBF0BB61452FFCAF" box="[1073,1316,771,795]" pageId="111" pageNumber="110">Est. Fluvia Metrica</collectorName>
,
<date id="FFE11071FF97F868FAEFBB664597FCAF" box="[1326,1436,771,795]" pageId="111" pageNumber="110" value="1976-02-04">
<collectingDate id="EFA5E999FF97F868FAEFBB664597FCAF" box="[1326,1436,771,795]" pageId="111" pageNumber="110" value="1976-02-04">4.ii.1976</collectingDate>
</date>
,
<location id="8E80606AFF97F868FF01BB46413BFC8F" LSID="urn:lsid:plazi:treatment:03F687A7FF96F868FF01BC264409FBAF:8E80606AFF97F868FF01BB46413BFC8F" box="[192,304,803,827]" country="Ecuador" name="Spangler" pageId="111" pageNumber="110" stateProvince="Pastaza">Spangler</location>
et al.–
<specimenCount id="9D59FD38FF97F868FE47BB4641F0FC8F" box="[390,507,803,827]" pageId="111" pageNumber="110" type="female">3 females</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FF97F868FDBDBB4642D7FC8F" box="[636,732,803,827]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="111" pageNumber="110" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CECFF97F868FD31BB414202FCEF" collectingDate="1976-02-03" collectionCode="W, NMNH" collectorName="Puyo &amp; Spangler" country="Ecuador" pageId="111" pageNumber="110" specimenCount="59" specimenCount-female="41" specimenCount-male="18" stateProvince="Pastaza" typeStatus="paratype">
<collectorName id="26AA5367FF97F868FD31BB414325FC88" box="[752,814,804,828]" pageId="111" pageNumber="110">Puyo</collectorName>
(
<quantity id="4CA79B54FF97F868FCFEBB464387FC8F" box="[831,908,803,827]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="111" pageNumber="110" unit="km" value="16.0">16 km</quantity>
<collectionCode id="ED4EAE74FF97F868FC52BB4643BAFC8F" box="[915,945,803,827]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="111" pageNumber="110" type="Herbarium">W</collectionCode>
),
<date id="FFE11071FF97F868FC07BB464438FC8F" box="[966,1075,803,827]" pageId="111" pageNumber="110" value="1976-02-03">
<collectingDate id="EFA5E999FF97F868FC07BB464438FC8F" box="[966,1075,803,827]" pageId="111" pageNumber="110" value="1976-02-03">3.ii.1976</collectingDate>
</date>
,
<collectorName id="26AA5367FF97F868FB81BB4644BBFC8F" box="[1088,1200,803,827]" pageId="111" pageNumber="110">Spangler</collectorName>
et al.–
<specimenCount id="9D59FD38FF97F868FAC5BB46457DFC8F" box="[1284,1398,803,827]" pageId="111" pageNumber="110" type="male">18 males</specimenCount>
,
<specimenCount id="9D59FD38FF97F868FA43BB46412BFCEF" pageId="111" pageNumber="110" type="female">41 females</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FF97F868FE60BB26420AFCEF" box="[417,513,835,859]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="111" pageNumber="110" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CECFF97F868FDD7BB264201FCCF" collectingDate="1975-05-28" collectionCode="NMNH" collectorName="Banos &amp; Cohen &amp; Langley" country="Ecuador" elevation="1798" location="Tungurahua" pageId="111" pageNumber="110" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Tungurahua" typeStatus="paratype">
<emphasis id="B92BEAA3FF97F868FDD7BB2642C7FCEF" bold="true" box="[534,716,835,859]" pageId="111" pageNumber="110">
<collectingRegion id="499BF853FF97F868FDD7BB2642CDFCEF" box="[534,710,835,859]" country="Ecuador" name="Tungurahua" pageId="111" pageNumber="110">Tungurahua</collectingRegion>
:
</emphasis>
<collectorName id="26AA5367FF97F868FD15BB214328FCEF" box="[724,803,836,859]" pageId="111" pageNumber="110">Baños</collectorName>
,
<quantity id="4CA79B54FF97F868FCF1BB264386FCEF" box="[816,909,835,859]" metricMagnitude="3" metricUnit="m" metricValue="1.798" pageId="111" pageNumber="110" unit="m" value="1798.0">
<elevation id="0072D182FF97F868FCF1BB264386FCEF" box="[816,909,835,859]" metricMagnitude="3" metricUnit="m" metricValue="1.798" pageId="111" pageNumber="110" unit="m" value="1798.0">1798 m</elevation>
</quantity>
,
<date id="FFE11071FF97F868FC5ABB26441FFCEF" box="[923,1044,835,859]" pageId="111" pageNumber="110" value="1975-05-28">
<collectingDate id="EFA5E999FF97F868FC5ABB26441FFCEF" box="[923,1044,835,859]" pageId="111" pageNumber="110" value="1975-05-28">28.v.1975</collectingDate>
</date>
,
<collectorName id="26AA5367FF97F868FBE0BB26447BFCEF" box="[1057,1136,835,859]" pageId="111" pageNumber="110">Cohen</collectorName>
and
<collectorName id="26AA5367FF97F868FB6EBB26451FFCEF" box="[1199,1300,835,859]" pageId="111" pageNumber="110">Langley</collectorName>
–
<specimenCount id="9D59FD38FF97F868FAE5BB26458EFCEF" box="[1316,1413,835,859]" pageId="111" pageNumber="110" type="male">3 males</specimenCount>
,
<specimenCount id="9D59FD38FF97F868FA50BB26412BFCCF" pageId="111" pageNumber="110" type="female">2 females</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FF97F868FE63BB064209FCCF" box="[418,514,867,891]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="111" pageNumber="110" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CECFF97F868FDD6BB0141ACFC2F" collectingDate="1976-01-25" collectionCode="E, NMNH" collectorName="Banos" country="Ecuador" location="Spangler" pageId="111" pageNumber="110" specimenCount="24" specimenCount-female="15" specimenCount-male="9" stateProvince="Tungurahua" typeStatus="paratype">
<collectorName id="26AA5367FF97F868FDD6BB01426FFCCF" box="[535,612,868,891]" pageId="111" pageNumber="110">Baños</collectorName>
(
<quantity id="4CA79B54FF97F868FDB5BB0642C8FCCF" box="[628,707,867,891]" metricMagnitude="4" metricUnit="m" metricValue="3.4" pageId="111" pageNumber="110" unit="km" value="34.0">34 km</quantity>
<collectionCode id="ED4EAE74FF97F868FD0ABB0642EBFCCF" box="[715,736,867,891]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="111" pageNumber="110" type="Herbarium">E</collectionCode>
),
<date id="FFE11071FF97F868FD36BB064360FCCF" box="[759,875,867,891]" pageId="111" pageNumber="110" value="1976-01-25">
<collectingDate id="EFA5E999FF97F868FD36BB064360FCCF" box="[759,875,867,891]" pageId="111" pageNumber="110" value="1976-01-25">25.i.1976</collectingDate>
</date>
, 1280 m,
<location id="8E80606AFF97F868FC25BB06445FFCCF" LSID="urn:lsid:plazi:treatment:03F687A7FF96F868FF01BC264409FBAF:8E80606AFF97F868FC25BB06445FFCCF" box="[996,1108,867,891]" country="Ecuador" name="Spangler" pageId="111" pageNumber="110" stateProvince="Tungurahua">Spangler</location>
et al.–
<specimenCount id="9D59FD38FF97F868FB6BBB064506FCCF" box="[1194,1293,867,891]" pageId="111" pageNumber="110" type="male">9 males</specimenCount>
,
<specimenCount id="9D59FD38FF97F868FADBBB0645ABFCCF" box="[1306,1440,867,891]" pageId="111" pageNumber="110" type="female">15 females</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FF97F868FEFFBBE64194FC2F" box="[318,415,899,923]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="111" pageNumber="110" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CECFF97F868FE76BBE641ABFC0F" collectingDate="1976-06-01" collectingDateMax="1976-06-05" collectingDateMin="1976-06-01" collectionCode="A, NMNH" collectorName="Zamora" country="Ecuador" location="Langley" pageId="111" pageNumber="110" specimenCount="1" specimenCount-male="1" stateProvince="Zamora-Chinchipe" typeStatus="paratype">
<emphasis id="B92BEAA3FF97F868FE76BBE642CEFC2F" bold="true" box="[439,709,899,923]" pageId="111" pageNumber="110">
<collectingRegion id="499BF853FF97F868FE76BBE642B4FC2F" box="[439,703,899,923]" country="Ecuador" name="Zamora-Chinchipe" pageId="111" pageNumber="110">Zamora-Chinchipe</collectingRegion>
:
</emphasis>
<collectorName id="26AA5367FF97F868FD11BBE14338FC2F" box="[720,819,900,923]" pageId="111" pageNumber="110">Zamora</collectorName>
, at lights,
<date id="FFE11071FF97F868FC02BBE64444FC2F" box="[963,1103,899,923]" pageId="111" pageNumber="110" value="1976-06-01" valueMax="1976-06-05" valueMin="1976-06-01">
<collectingDate id="EFA5E999FF97F868FC02BBE64444FC2F" box="[963,1103,899,923]" pageId="111" pageNumber="110" value="1976-06-01" valueMax="1976-06-05" valueMin="1976-06-01">1-5.vi.1976</collectingDate>
</date>
,
<collectionCode id="ED4EAE74FF97F868FBA1BBE64478FC2F" box="[1120,1139,899,923]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="111" pageNumber="110" type="Herbarium">A</collectionCode>
<location id="8E80606AFF97F868FBBEBBE644ECFC2F" LSID="urn:lsid:plazi:treatment:03F687A7FF96F868FF01BC264409FBAF:8E80606AFF97F868FBBEBBE644ECFC2F" box="[1151,1255,899,923]" country="Ecuador" name="Langley" pageId="111" pageNumber="110" stateProvince="Zamora-Chinchipe">Langley</location>
, et al.–
<specimenCount id="9D59FD38FF97F868FA8DBBE645ABFC2F" box="[1356,1440,899,923]" pageId="111" pageNumber="110" type="male">1 male</specimenCount>
(pinned) (
<collectionCode id="ED4EAE74FF97F868FEF9BBC64193FC0F" box="[312,408,931,955]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="111" pageNumber="110" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373CECFF97F868FE6CBBC1453BFC0F" box="[429,1328,931,955]" collectingDate="1978-12-04" collectionCode="NMNH" collectorName="Zamora &amp; JJ Anderson" country="Ecuador" pageId="111" pageNumber="110" specimenCount="13" specimenCount-female="10" specimenCount-male="3" stateProvince="Zamora-Chinchipe" typeStatus="paratype">
<collectorName id="26AA5367FF97F868FE6CBBC1421BFC0F" box="[429,528,932,955]" pageId="111" pageNumber="110">Zamora</collectorName>
,
<date id="FFE11071FF97F868FDDDBBC64293FC0F" box="[540,664,931,955]" pageId="111" pageNumber="110" value="1978-12-04">
<collectingDate id="EFA5E999FF97F868FDDDBBC64293FC0F" box="[540,664,931,955]" pageId="111" pageNumber="110" value="1978-12-04">4.xii.1978</collectingDate>
</date>
,
<collectorName id="26AA5367FF97F868FD64BBC14348FC0F" box="[677,835,931,955]" pageId="111" pageNumber="110">JJ Anderson</collectorName>
–
<specimenCount id="9D59FD38FF97F868FC92BBC643B8FC0F" box="[851,947,931,955]" pageId="111" pageNumber="110" type="male">3 males</specimenCount>
,
<specimenCount id="9D59FD38FF97F868FC01BBC6444DFC0F" box="[960,1094,931,955]" pageId="111" pageNumber="110" type="female">10 females</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FF97F868FB06BBC64523FC0F" box="[1223,1320,931,955]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="111" pageNumber="110" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BE036B1FF97F868FF31BBA64409FBAF" blockId="111.[192,1441,195,1052]" pageId="111" pageNumber="110">
<emphasis id="B92BEAA3FF97F868FF31BBA64179FC6F" box="[240,370,963,987]" italics="true" pageId="111" pageNumber="110">Etymology</emphasis>
—We take pleasure in naming this species for Dr. Paul Spangler, retired aquatic beetle specialist at the Smithsonian Institution, who also collected many specimens of
<taxonomicName id="4C5F4D32FF97F868FB70BB86454FFC4F" baseAuthorityName="Holzenthal and Calor" baseAuthorityYear="2017" box="[1201,1348,995,1019]" class="Insecta" kingdom="Animalia" order="Trichoptera" pageId="111" pageNumber="110" phylum="Arthropoda" rank="order">Trichoptera</taxonomicName>
during his career, including the majority of
<typeStatus id="54E48813FF97F868FD46BC6142B6FBA8" box="[647,701,1028,1052]" pageId="111" pageNumber="110">type</typeStatus>
specimens of this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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