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<document id="C192A782FEFD841733FFA1EE55FA077F" ID-CLB-Dataset="45913" ID-DOI="10.5281/zenodo.279281" ID-GBIF-Dataset="6a6204af-9f48-4992-8ec5-ba22c28b8413" ID-ISSN="1175-5326" ID-Zenodo-Dep="279281" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460413515392" checkinUser="plazi" docAuthor="Husana, Daniel Edison M., Tan, Swee Hee &amp; Kase, Tomoki" docDate="2011" docId="03C2082FFF80FF85FF75FBD0422CFC7C" docLanguage="en" docName="zt03109p059.pdf" docOrigin="Zootaxa 3109" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Samarplax Husana, Tan &amp; Kase, 2011, new genus" docType="treatment" docVersion="9" lastPageNumber="52" masterDocId="FFFB7057FF82FF86FFE2FFAC470EFF91" masterDocTitle="A new genus and species of anchialine Hymenosomatidae (Crustacea, Decapoda, Brachyura) from Samar, Philippines" masterLastPageNumber="59" masterPageNumber="49" pageNumber="51" updateTime="1698328222292" updateUser="ExternalLinkService">
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<mods:title id="D637E57031AA2A8A2AC2BD9E5AB81B6F">A new genus and species of anchialine Hymenosomatidae (Crustacea, Decapoda, Brachyura) from Samar, Philippines</mods:title>
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<mods:name id="122DF92F9124DB2FA72792EDC8C6642D" type="personal">
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<mods:namePart id="2B337CFBADE2B2A893FC20CCFED95C91">Husana, Daniel Edison M.</mods:namePart>
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<mods:namePart id="5E62A3D8479B6F4CF98AA4CFFA35E33E">Tan, Swee Hee</mods:namePart>
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<mods:roleTerm id="D884AB620F0E56E8F94915D29FFFADF2">Author</mods:roleTerm>
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<mods:namePart id="990B7001BE54459F1C076D60BAAE64A4">Kase, Tomoki</mods:namePart>
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<mods:title id="26DD69375A933533C7A6332EF287D804">Zootaxa</mods:title>
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<mods:part id="07A94BEC22017BCF23552CCA0C41D046">
<mods:date id="6E7154E85E2BB7B42D0EDE4D3D88F572">2011</mods:date>
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<treatment id="03C2082FFF80FF85FF75FBD0422CFC7C" ID-DOI="http://doi.org/10.5281/zenodo.6182668" ID-GBIF-Taxon="119424178" ID-Zenodo-Dep="6182668" LSID="urn:lsid:plazi:treatment:03C2082FFF80FF85FF75FBD0422CFC7C" httpUri="http://treatment.plazi.org/id/03C2082FFF80FF85FF75FBD0422CFC7C" lastPageId="3" lastPageNumber="52" pageId="2" pageNumber="51">
<subSubSection id="C371EAB2FF80FF84FF75FBD046AAFB04" box="[151,420,1147,1174]" pageId="2" pageNumber="51" type="nomenclature">
<paragraph id="8BD4B939FF80FF84FF75FBD046AAFB04" blockId="2.[151,420,1147,1174]" box="[151,420,1147,1174]" pageId="2" pageNumber="51">
<heading id="D09C0E55FF80FF84FF75FBD046AAFB04" bold="true" box="[151,420,1147,1174]" fontSize="11" level="1" pageId="2" pageNumber="51" reason="1">
<emphasis id="B91F652BFF80FF84FF75FBD046AAFB04" bold="true" box="[151,420,1147,1174]" pageId="2" pageNumber="51">
<taxonomicName id="4C6BC2BAFF80FF84FF75FBD04615FB07" ID-CoL="7PYL4" box="[151,283,1148,1174]" class="Malacostraca" family="Hymenosomatidae" genus="Samarplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91F652BFF80FF84FF75FBD04615FB07" bold="true" box="[151,283,1148,1174]" italics="true" pageId="2" pageNumber="51">Samarplax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22CD850FF80FF84FEC0FBD746AAFB04" box="[290,420,1147,1173]" pageId="2" pageNumber="51" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C371EAB2FF80FF84FF75FB6F44B6FB4B" box="[151,952,1218,1243]" pageId="2" pageNumber="51" type="materials_examined">
<paragraph id="8BD4B939FF80FF84FF75FB6F44B6FB4B" blockId="2.[151,1437,1218,2035]" box="[151,952,1218,1243]" pageId="2" pageNumber="51">
<emphasis id="B91F652BFF80FF84FF75FB6F4625FB4A" bold="true" box="[151,299,1218,1243]" pageId="2" pageNumber="51">
<typeStatus id="54D0079BFF80FF84FF75FB6F47DCFB4A" box="[151,210,1219,1243]" pageId="2" pageNumber="51">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="4C6BC2BAFF80FF84FEDBFB6F4518FB4A" box="[313,534,1219,1243]" class="Malacostraca" family="Hymenosomatidae" genus="Samarplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="principe" status="sp. nov.">
<emphasis id="B91F652BFF80FF84FEDBFB6F4518FB4A" box="[313,534,1219,1243]" italics="true" pageId="2" pageNumber="51">Samarplax principe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22CD850FF80FF84FDFFFB6F45A9FB4A" box="[541,679,1218,1243]" pageId="2" pageNumber="51" rank="species">
<emphasis id="B91F652BFF80FF84FDFFFB6F45A9FB4A" bold="true" box="[541,679,1218,1243]" pageId="2" pageNumber="51">new species</emphasis>
</taxonomicNameLabel>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="C371EAB2FF80FF84FF27FB4A46E7F9F7" pageId="2" pageNumber="51" type="diagnosis">
<paragraph id="8BD4B939FF80FF84FF27FB4A46E7F9F7" blockId="2.[151,1437,1218,2035]" pageId="2" pageNumber="51">
<emphasis id="B91F652BFF80FF84FF27FB4A4631FB6E" bold="true" box="[197,319,1254,1279]" pageId="2" pageNumber="51">Diagnosis.</emphasis>
Carapace sub-polygonal, as long as wide to slightly longer than wide; dorsal surface uneven due to regions, surrounded by continuous rim, regions slightly inflated, separated by deep grooves. Rostrum absent; frontal margin gently bilobed. Posterolateral margin entire. Antennular fossa very shallow. Epistome almost flat, not visible from dorsal view. Eyes strongly reduced, visible as vestigial structure only under scanning electron microscopy (SEM) (
<figureCitation id="1350A5BCFF80FF84FED4FADA4673FA1F" box="[310,381,1398,1422]" captionStart="FIGURE 7" captionStartId="7.[151,250,1322,1345]" captionTargetBox="[151,1436,337,1300]" captionTargetId="figure@7.[151,1436,337,1301]" captionTargetPageId="7" captionText="FIGURE 7. Samarplax principe new genus, new species. SEM micrograph of the frontal region showing the vestigial eyestalk. Paratype male, 4.09 x 3.98 mm (NSMT-Cr 2007 - 00040)." httpUri="https://zenodo.org/record/279288/files/figure.png" pageId="2" pageNumber="51">Fig. 7</figureCitation>
); cornea absent. Male sterno-abdominal cavity relatively narrow; abdominal locking button indistinct. Male abdomen-pleotelson with somites 3–5 fused; somites 1, 2 with rounded lobes on disto-lateral angle on both sides; fused somites narrowed distally; pleotelson distinctly narrower than half width of abdomen; tip of pleotelson reaching to mid-point of abdominal somite 4. Female abdomen dome-shaped, with undivided plate formed by fusion of somite 2 to pleotelson, somite 2 and pleotelson marked by shallow suture on outer side. G1 slender, slightly sinuous, twisted at base, tapering at tip, curving inwards, distal portion with long stiff setae. G2 short, about half length of G1.
</paragraph>
</subSubSection>
<subSubSection id="C371EAB2FF80FF84FF27F9DF47C1F943" pageId="2" pageNumber="51" type="etymology">
<paragraph id="8BD4B939FF80FF84FF27F9DF47C1F943" blockId="2.[151,1437,1218,2035]" pageId="2" pageNumber="51">
<emphasis id="B91F652BFF80FF84FF27F9DF4645F91A" bold="true" box="[197,331,1651,1675]" pageId="2" pageNumber="51">Etymology.</emphasis>
The genus name
<taxonomicName id="4C6BC2BAFF80FF84FDF1F9DF4583F91A" box="[531,653,1651,1675]" class="Malacostraca" family="Hymenosomatidae" genus="Samarplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91F652BFF80FF84FDF1F9DF4583F91A" box="[531,653,1651,1675]" italics="true" pageId="2" pageNumber="51">Samarplax</emphasis>
</taxonomicName>
is derived from the locality, Samar province and
<emphasis id="B91F652BFF80FF84FB5EF9DF43E2F91A" box="[1212,1260,1651,1675]" italics="true" pageId="2" pageNumber="51">plax</emphasis>
for “plate”, the suffix of the cavernicolous genera,
<taxonomicName id="4C6BC2BAFF80FF84FDC4F93A4586F93F" box="[550,648,1686,1710]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B91F652BFF80FF84FDC4F93A4586F93F" box="[550,648,1686,1710]" italics="true" pageId="2" pageNumber="51">Sulaplax</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6BC2BAFF80FF84FD22F93A442EF93F" box="[704,800,1686,1710]" class="Malacostraca" family="Hymenosomatidae" genus="Guaplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B91F652BFF80FF84FD22F93A442EF93F" box="[704,800,1686,1710]" italics="true" pageId="2" pageNumber="51">Guaplax</emphasis>
</taxonomicName>
, alluding to their superficial resemblance. Gender feminine.
</paragraph>
</subSubSection>
<subSubSection id="C371EAB2FF80FF85FF27F9734438FEF5" lastPageId="3" lastPageNumber="52" pageId="2" pageNumber="51" type="discussion">
<paragraph id="8BD4B939FF80FF85FF27F9734438FEF5" blockId="2.[151,1437,1218,2035]" lastBlockId="3.[151,1436,151,1005]" lastPageId="3" lastPageNumber="52" pageId="2" pageNumber="51">
<emphasis id="B91F652BFF80FF84FF27F9734637F966" bold="true" box="[197,313,1759,1783]" pageId="2" pageNumber="51">Remarks.</emphasis>
<taxonomicName id="4C6BC2BAFF80FF84FEA0F97246B5F967" box="[322,443,1758,1782]" class="Malacostraca" family="Hymenosomatidae" genus="Samarplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91F652BFF80FF84FEA0F97246B5F967" box="[322,443,1758,1782]" italics="true" pageId="2" pageNumber="51">Samarplax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22CD850FF80FF84FE26F973454CF966" box="[452,578,1759,1783]" pageId="2" pageNumber="51" rank="genus">
<emphasis id="B91F652BFF80FF84FE26F973454CF966" bold="true" box="[452,578,1759,1783]" pageId="2" pageNumber="51">new genus</emphasis>
</taxonomicNameLabel>
, differs from other epigean genera of
<taxonomicName id="4C6BC2BAFF80FF84FC1EF97243C0F967" box="[1020,1230,1758,1782]" class="Malacostraca" family="Hymenosomatidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="family">Hymenosomatidae</taxonomicName>
in having several troglobitic features such as the extreme reduction of eyes, the absence of pigment and the disproportionately long legs. Otherwise, it resembles the epigean genus
<taxonomicName id="4C6BC2BAFF80FF84FD4CF88A44FAF8AF" authority="Sakai, 1938" authorityName="Sakai" authorityYear="1938" box="[686,1012,1830,1854]" class="Malacostraca" family="Hymenosomatidae" genus="Neorhynchoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B91F652BFF80FF84FD4CF88A4469F8AF" box="[686,871,1830,1854]" italics="true" pageId="2" pageNumber="51">Neorhynchoplax</emphasis>
<bibRefCitation id="EFFAC4C8FF80FF84FC8FF88A44FAF8AF" author="Sakai" box="[877,1012,1830,1854]" pageId="2" pageNumber="51" refString="Sakai, T. (1938) Studies on the crabs of Japan. III. Brachygnatha, Oxyrhyncha. Yokendo, Tokyo. 193 - 364 pp., Pls. 20 - 41." type="book chapter" year="1938">Sakai, 1938</bibRefCitation>
</taxonomicName>
, in having well defined dorsal surface regions and the eggs being held in the brood cavity (see
<bibRefCitation id="EFFAC4C8FF80FF84FCEAF8E744EBF8F3" author="Ng" box="[776,997,1866,1891]" pageId="2" pageNumber="51" refString="Ng, P. K. L. &amp; Chuang, C. T. N. (1996) The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of Southeast Asia, with notes on other species. Raffles Bulletin of Zoology, Supplement 3, 1 - 82." type="book chapter" year="1996">Ng &amp; Chuang 1996</bibRefCitation>
;
<bibRefCitation id="EFFAC4C8FF80FF84FC13F8E743CFF8F3" author="Naruse" box="[1009,1217,1866,1891]" pageId="2" pageNumber="51" refString="Naruse, T., Shokita, S. &amp; Kawahara, T. (2005) Neorhynchoplax yaeyamaensis, a new false spider crab (Decapoda: Brachyura: Hymenosomatidae) from the Yaeyama Group, the Ryukyu Islands, Japan. Zootaxa, 877, 1 - 7." type="journal article" year="2005">
Naruse
<emphasis id="B91F652BFF80FF84FBA4F8E04370F8F2" box="[1094,1150,1867,1891]" italics="true" pageId="2" pageNumber="51">et al.</emphasis>
2005
</bibRefCitation>
). However, in addition to the above-mentioned troglobitic features,
<taxonomicName id="4C6BC2BAFF80FF84FD5CF8C24439F817" box="[702,823,1902,1926]" class="Malacostraca" family="Hymenosomatidae" genus="Samarplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91F652BFF80FF84FD5CF8C24439F817" box="[702,823,1902,1926]" italics="true" pageId="2" pageNumber="51">Samarplax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22CD850FF80FF84FCDDF8C344B5F816" box="[831,955,1903,1927]" pageId="2" pageNumber="51" rank="genus">
<emphasis id="B91F652BFF80FF84FCDDF8C344B5F816" bold="true" box="[831,955,1903,1927]" pageId="2" pageNumber="51">new genus</emphasis>
</taxonomicNameLabel>
, also differs from
<taxonomicName id="4C6BC2BAFF80FF84FB6EF8C2424BF817" box="[1164,1349,1902,1926]" class="Malacostraca" family="Hymenosomatidae" genus="Neorhynchoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B91F652BFF80FF84FB6EF8C2424BF817" box="[1164,1349,1902,1926]" italics="true" pageId="2" pageNumber="51">Neorhynchoplax</emphasis>
</taxonomicName>
in having no rostrum, the lateral walls of the carapace are not laterally expanded and the ambulatory dactyli are unarmed (see
<bibRefCitation id="EFFAC4C8FF80FF84FF28F81B4694F85F" author="Naruse" box="[202,410,1974,1999]" pageId="2" pageNumber="51" refString="Naruse, T., Ng, P. K. L. &amp; Guinot, D. (2008) Two new genera and new species of troglobitic false spider crabs (Crustacea: Decapoda: Brachyura: Hymenosomatidae) from Indonesia, with notes on the genus Cancrocaeca Ng, 1991. Zootaxa, 1739, 21 - 40." type="journal article" year="2008">
Naruse
<emphasis id="B91F652BFF80FF84FEC2F8144656F85F" box="[288,344,1974,1999]" italics="true" pageId="2" pageNumber="51">et al.</emphasis>
2008
</bibRefCitation>
: Table 1). Externally, the new genus most closely resembles
<taxonomicName id="4C6BC2BAFF80FF84FBDFF81A47DCF863" authority="Naruse, Ng &amp; Guinot, 2008" authorityName="Naruse, Ng &amp; Guinot" authorityYear="2008" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B91F652BFF80FF84FBDFF81A4391F85F" box="[1085,1183,1974,1998]" italics="true" pageId="2" pageNumber="51">Sulaplax</emphasis>
Naruse, Ng &amp;
<bibRefCitation id="EFFAC4C8FF80FF84FAA5F81A47DCF863" author="Guinot" pageId="2" pageNumber="51" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="book chapter" year="2008">Guinot, 2008</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C6BC2BAFF80FF84FEECF87745BAF863" authority="Naruse, Ng &amp; Guinot, 2008" authorityName="Naruse, Ng &amp; Guinot" authorityYear="2008" box="[270,692,2010,2035]" class="Malacostraca" family="Hymenosomatidae" genus="Guaplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B91F652BFF80FF84FEECF8774661F862" box="[270,367,2011,2035]" italics="true" pageId="2" pageNumber="51">Guaplax</emphasis>
Naruse, Ng &amp;
<bibRefCitation id="EFFAC4C8FF80FF84FDFFF87645BAF863" author="Guinot" box="[541,692,2010,2034]" pageId="2" pageNumber="51" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="book chapter" year="2008">Guinot, 2008</bibRefCitation>
</taxonomicName>
, from
<collectingCountry id="F37CF9A9FF80FF84FD1DF8764462F863" box="[767,876,2010,2034]" name="Indonesia" pageId="2" pageNumber="51">Indonesia</collectingCountry>
in possessing unpigmented eyes that are fused to the orbit, absence of a subterminal tooth on the ambulatory dactylus; having the dorsal carapace regions well defined, eggs contained in a brood cavity, a four-segmented pleotelson, and a G1 which is straight, tapering and having simple tip. The form of the third maxilliped is closer to that of
<taxonomicName id="4C6BC2BAFF81FF85FC48FF4C4304FF69" box="[938,1034,224,248]" class="Malacostraca" family="Hymenosomatidae" genus="Guaplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B91F652BFF81FF85FC48FF4C4304FF69" box="[938,1034,224,248]" italics="true" pageId="3" pageNumber="52">Guaplax</emphasis>
</taxonomicName>
but overall,
<taxonomicName id="4C6BC2BAFF81FF85FB7BFF4C421CFF69" box="[1177,1298,224,248]" class="Malacostraca" family="Hymenosomatidae" genus="Samarplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91F652BFF81FF85FB7BFF4C421CFF69" box="[1177,1298,224,248]" italics="true" pageId="3" pageNumber="52">Samarplax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22CD850FF81FF85FAFBFF4C429AFF69" box="[1305,1428,224,248]" pageId="3" pageNumber="52" rank="genus">
<emphasis id="B91F652BFF81FF85FAFBFF4C429AFF69" bold="true" box="[1305,1428,224,248]" pageId="3" pageNumber="52">new genus</emphasis>
</taxonomicNameLabel>
, appears to be more similar to
<taxonomicName id="4C6BC2BAFF81FF85FE13FEA8455DFE8D" box="[497,595,260,284]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B91F652BFF81FF85FE13FEA8455DFE8D" box="[497,595,260,284]" italics="true" pageId="3" pageNumber="52">Sulaplax</emphasis>
</taxonomicName>
, sharing characters such as the basal antennal article not been positioned directly below the base of the eye stalk but offset to the outer edge (versus directly below in
<taxonomicName id="4C6BC2BAFF81FF85FB4EFE844202FED1" box="[1196,1292,296,320]" class="Malacostraca" family="Hymenosomatidae" genus="Guaplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B91F652BFF81FF85FB4EFE844202FED1" box="[1196,1292,296,320]" italics="true" pageId="3" pageNumber="52">Guaplax</emphasis>
</taxonomicName>
) and in having very long chelipeds, with the merus especially elongate.
</paragraph>
</subSubSection>
<subSubSection id="C371EAB2FF81FF85FF27FEDC422CFC7C" pageId="3" pageNumber="52" type="reference_group">
<paragraph id="8BD4B939FF81FF85FF27FEDC4599FCCD" blockId="3.[151,1436,151,1005]" pageId="3" pageNumber="52">
<taxonomicName id="4C6BC2BAFF81FF85FF27FEDC4630FE19" ID-CoL="7PYL4" box="[197,318,368,392]" class="Malacostraca" family="Hymenosomatidae" genus="Samarplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91F652BFF81FF85FF27FEDC4630FE19" box="[197,318,368,392]" italics="true" pageId="3" pageNumber="52">Samarplax</emphasis>
</taxonomicName>
<emphasis id="B91F652BFF81FF85FEA7FEDC46CFFE19" bold="true" box="[325,449,368,392]" pageId="3" pageNumber="52">
<taxonomicNameLabel id="A22CD850FF81FF85FEA7FEDC46CFFE19" box="[325,449,368,392]" pageId="3" pageNumber="52" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
, nevertheless differs from
<taxonomicName id="4C6BC2BAFF81FF85FD0CFEDC445EFE19" ID-CoL="7Q3BP" box="[750,848,368,392]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B91F652BFF81FF85FD0CFEDC445EFE19" box="[750,848,368,392]" italics="true" pageId="3" pageNumber="52">Sulaplax</emphasis>
</taxonomicName>
in the following aspects: the inner distal edge of the ischium of the third maxilliped is slightly projected outwards and not as long as that observed on
<taxonomicName id="4C6BC2BAFF81FF85FB37FE384239FE3D" ID-CoL="7Q3BP" box="[1237,1335,404,428]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B91F652BFF81FF85FB37FE384239FE3D" box="[1237,1335,404,428]" italics="true" pageId="3" pageNumber="52">Sulaplax</emphasis>
</taxonomicName>
(
<figureCitation id="1350A5BCFF81FF85FAA4FE384284FE3D" box="[1350,1418,404,428]" captionStart="FIGURE 4" captionStartId="5.[151,250,1812,1835]" captionTargetBox="[159,1426,189,1788]" captionTargetId="figure@5.[151,1436,185,1796]" captionTargetPageId="5" captionText="FIGURE 4. Samarplax principe new genus, new species. a, paratype female (ZRC 2011.0879; ex. NSMT-Cr 2007 - 0002)," httpUri="https://zenodo.org/record/279285/files/figure.png" pageId="3" pageNumber="52">Fig. 4</figureCitation>
e, versus
<bibRefCitation id="EFFAC4C8FF81FF85FF07FE1446B6FE5E" author="Naruse" box="[229,440,439,464]" pageId="3" pageNumber="52" refString="Naruse, T., Ng, P. K. L. &amp; Guinot, D. (2008) Two new genera and new species of troglobitic false spider crabs (Crustacea: Decapoda: Brachyura: Hymenosomatidae) from Indonesia, with notes on the genus Cancrocaeca Ng, 1991. Zootaxa, 1739, 21 - 40." type="journal article" year="2008">
Naruse
<emphasis id="B91F652BFF81FF85FEDEFE15467BFE41" box="[316,373,440,464]" italics="true" pageId="3" pageNumber="52">et al.</emphasis>
2008
</bibRefCitation>
: fig. 4c); the epistome is almost flat (
<figureCitation id="1350A5BCFF81FF85FCB8FE1B4493FE5E" box="[858,925,439,463]" captionStart="FIGURE 7" captionStartId="7.[151,250,1322,1345]" captionTargetBox="[151,1436,337,1300]" captionTargetId="figure@7.[151,1436,337,1301]" captionTargetPageId="7" captionText="FIGURE 7. Samarplax principe new genus, new species. SEM micrograph of the frontal region showing the vestigial eyestalk. Paratype male, 4.09 x 3.98 mm (NSMT-Cr 2007 - 00040)." httpUri="https://zenodo.org/record/279288/files/figure.png" pageId="3" pageNumber="52">Fig. 7</figureCitation>
) (versus relatively long with produced posterior margin in
<taxonomicName id="4C6BC2BAFF81FF85FEDAFE704576FE65" ID-CoL="7Q3BP" authority="Naruse et al. 2008" authorityName="Naruse et al." authorityYear="2008" box="[312,632,476,500]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B91F652BFF81FF85FEDAFE704694FE65" box="[312,410,476,500]" italics="true" pageId="3" pageNumber="52">Sulaplax</emphasis>
,
<bibRefCitation id="EFFAC4C8FF81FF85FE45FE714576FE65" author="Naruse" box="[423,632,476,500]" pageId="3" pageNumber="52" refString="Naruse, T., Ng, P. K. L. &amp; Guinot, D. (2008) Two new genera and new species of troglobitic false spider crabs (Crustacea: Decapoda: Brachyura: Hymenosomatidae) from Indonesia, with notes on the genus Cancrocaeca Ng, 1991. Zootaxa, 1739, 21 - 40." type="journal article" year="2008">
Naruse
<emphasis id="B91F652BFF81FF85FE1FFE71453BFE65" box="[509,565,476,500]" italics="true" pageId="3" pageNumber="52">et al.</emphasis>
2008
</bibRefCitation>
</taxonomicName>
: fig. 4b); and the fused abdominal piece consisting of somites 3-pleotelson is prominently triangular in shape with the lateral margins of somite 4 converging towards the telson (
<figureCitation id="1350A5BCFF81FF85FAA1FE534288FD86" box="[1347,1414,511,535]" captionStart="FIGURE 5" captionStartId="6.[151,250,1096,1119]" captionTargetBox="[169,1418,211,1056]" captionTargetId="figure@6.[151,1436,193,1075]" captionTargetPageId="6" captionText="FIGURE 5. Samarplax principe new genus, new species. a, holotype male (NMCR- 39033), abdomen, ventral view. b-c, paratype female (ZRC 2011.0879; ex. NSMT-Cr 2007 - 0002), abdomen: b, pleotelson; c, abdomen. Scale bars = 1 mm .." httpUri="https://zenodo.org/record/279286/files/figure.png" pageId="3" pageNumber="52">Fig. 5</figureCitation>
a) (versus with fused piece not subtriangular, with the lateral margins of somite 4 subparallel in
<taxonomicName id="4C6BC2BAFF81FF85FB5DFD8847F9FDCE" ID-CoL="7Q3BP" authority="Naruse et al. 2008" authorityName="Naruse et al." authorityYear="2008" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B91F652BFF81FF85FB5DFD88422FFDAD" box="[1215,1313,548,572]" italics="true" pageId="3" pageNumber="52">Sulaplax</emphasis>
,
<bibRefCitation id="EFFAC4C8FF81FF85FAD2FD8947F9FDCE" author="Naruse" pageId="3" pageNumber="52" refString="Naruse, T., Ng, P. K. L. &amp; Guinot, D. (2008) Two new genera and new species of troglobitic false spider crabs (Crustacea: Decapoda: Brachyura: Hymenosomatidae) from Indonesia, with notes on the genus Cancrocaeca Ng, 1991. Zootaxa, 1739, 21 - 40." type="journal article" year="2008">
Naruse
<emphasis id="B91F652BFF81FF85FA6AFD8947BAFDF1" italics="true" pageId="3" pageNumber="52">et al.</emphasis>
2008
</bibRefCitation>
</taxonomicName>
: fig. 5d). In addition,
<taxonomicName id="4C6BC2BAFF81FF85FE0FFDE44568FDF1" ID-CoL="7PYL4" box="[493,614,584,608]" class="Malacostraca" family="Hymenosomatidae" genus="Samarplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91F652BFF81FF85FE0FFDE44568FDF1" box="[493,614,584,608]" italics="true" pageId="3" pageNumber="52">Samarplax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22CD850FF81FF85FD8FFDE445E7FDF1" box="[621,745,584,608]" pageId="3" pageNumber="52" rank="genus">
<emphasis id="B91F652BFF81FF85FD8FFDE445E7FDF1" bold="true" box="[621,745,584,608]" pageId="3" pageNumber="52">new genus</emphasis>
</taxonomicNameLabel>
, has a sub-polygonal carapace the length and width subequal (versus wider than long,
<bibRefCitation id="EFFAC4C8FF81FF85FE4FFDC14570FD15" author="Naruse" box="[429,638,620,644]" pageId="3" pageNumber="52" refString="Naruse, T., Ng, P. K. L. &amp; Guinot, D. (2008) Two new genera and new species of troglobitic false spider crabs (Crustacea: Decapoda: Brachyura: Hymenosomatidae) from Indonesia, with notes on the genus Cancrocaeca Ng, 1991. Zootaxa, 1739, 21 - 40." type="journal article" year="2008">
Naruse
<emphasis id="B91F652BFF81FF85FDE1FDC14535FD15" box="[515,571,620,644]" italics="true" pageId="3" pageNumber="52">et al.</emphasis>
2008
</bibRefCitation>
: figs. 3, 4a), a simple straight, tapering distal part of the G1 that is gently curved towards the midline (versus more prominently twisted,
<bibRefCitation id="EFFAC4C8FF81FF85FC6EFD3C436DFD36" author="Naruse" box="[908,1123,655,680]" pageId="3" pageNumber="52" refString="Naruse, T., Ng, P. K. L. &amp; Guinot, D. (2008) Two new genera and new species of troglobitic false spider crabs (Crustacea: Decapoda: Brachyura: Hymenosomatidae) from Indonesia, with notes on the genus Cancrocaeca Ng, 1991. Zootaxa, 1739, 21 - 40." type="journal article" year="2008">
Naruse
<emphasis id="B91F652BFF81FF85FC06FD3D4310FD39" box="[996,1054,656,680]" italics="true" pageId="3" pageNumber="52">et al.</emphasis>
2008
</bibRefCitation>
: fig. 5e, f), and a relatively long chela armed with brush-like setae along the inner margin of its fingers instead of cutting teeth (
<figureCitation id="1350A5BCFF81FF85FB1AFD184234FD5D" box="[1272,1338,692,716]" captionStart="FIGURE 7" captionStartId="7.[151,250,1322,1345]" captionTargetBox="[151,1436,337,1300]" captionTargetId="figure@7.[151,1436,337,1301]" captionTargetPageId="7" captionText="FIGURE 7. Samarplax principe new genus, new species. SEM micrograph of the frontal region showing the vestigial eyestalk. Paratype male, 4.09 x 3.98 mm (NSMT-Cr 2007 - 00040)." httpUri="https://zenodo.org/record/279288/files/figure.png" pageId="3" pageNumber="52">Fig. 7</figureCitation>
) (versus lined with low teeth and without brush-like setae,
<bibRefCitation id="EFFAC4C8FF81FF85FD29FD744490FD7E" author="Naruse" box="[715,926,727,752]" pageId="3" pageNumber="52" refString="Naruse, T., Ng, P. K. L. &amp; Guinot, D. (2008) Two new genera and new species of troglobitic false spider crabs (Crustacea: Decapoda: Brachyura: Hymenosomatidae) from Indonesia, with notes on the genus Cancrocaeca Ng, 1991. Zootaxa, 1739, 21 - 40." type="journal article" year="2008">
Naruse
<emphasis id="B91F652BFF81FF85FCC0FD754454FD61" box="[802,858,728,752]" italics="true" pageId="3" pageNumber="52">et al.</emphasis>
2008
</bibRefCitation>
: fig. 5b, c). This chela structure is not known in other hymenosomatids so far. The G2 of
<taxonomicName id="4C6BC2BAFF81FF85FD6EFD50440BFC85" ID-CoL="7PYL4" box="[652,773,764,788]" class="Malacostraca" family="Hymenosomatidae" genus="Samarplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91F652BFF81FF85FD6EFD50440BFC85" box="[652,773,764,788]" italics="true" pageId="3" pageNumber="52">Samarplax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22CD850FF81FF85FCECFD504482FC85" box="[782,908,764,788]" pageId="3" pageNumber="52" rank="genus">
<emphasis id="B91F652BFF81FF85FCECFD504482FC85" bold="true" box="[782,908,764,788]" pageId="3" pageNumber="52">new genus</emphasis>
</taxonomicNameLabel>
, also differs from that of
<taxonomicName id="4C6BC2BAFF81FF85FB57FD504219FC85" ID-CoL="7Q3BP" box="[1205,1303,764,788]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B91F652BFF81FF85FB57FD504219FC85" box="[1205,1303,764,788]" italics="true" pageId="3" pageNumber="52">Sulaplax</emphasis>
</taxonomicName>
in that it is only about half the G1 length with a stout distal tip (versus about three-quarters of the G1 length with a slender and tapering tip in
<taxonomicName id="4C6BC2BAFF81FF85FED9FCE84693FCCD" ID-CoL="7Q3BP" box="[315,413,836,860]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B91F652BFF81FF85FED9FCE84693FCCD" box="[315,413,836,860]" italics="true" pageId="3" pageNumber="52">Sulaplax</emphasis>
</taxonomicName>
, present observation).
</paragraph>
<paragraph id="8BD4B939FF81FF85FF27FCCB422CFC7C" blockId="3.[151,1436,151,1005]" pageId="3" pageNumber="52">Overall, the absence of rostrum and eyes, well-defined dorsal surface of the carapace with two small spines or sharp teeth on its lateral margin, proportionally shorter projected merus of the third maxilliped, an almost flat epistome, long chelipeds without cutting teeth but lined instead with brush-like setae, a four-segmented pleotelson, relatively simple G1 and eggs that are held in the brood cavity are diagnostic characters of this new genus.</paragraph>
</subSubSection>
</treatment>
</document>
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