treatments-xml/data/03/BC/87/03BC87C5FFF08221FF5DF90D5F8697DE.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.6329707" ID-GBIF-Dataset="2de37bc1-3782-435b-b655-0f9d77f7afe9" ID-GBIF-Taxon="193873956" ID-Zenodo-Dep="6329707" approvedBy="tatiana" checkinTime="1646409764810" checkinUser="tatiana" docAuthor="Tepe, EJ &amp; Bohs, L" docDate="2011" docId="03BC87C5FFF08221FF5DF90D5F8697DE" docLanguage="en" docName="SystBot.36.4.1068-1087.pdf.imf" docOrigin="Systematic Botany 36 (4)" docSource="https://doi.org/10.1600/036364411X605074" docTitle="Solanum phaseoloides Pol., Linnaea" docType="treatment" docVersion="9" lastPageId="17" lastPageNumber="1084" masterDocId="FF85FFBDFFE08230FFFDFFB45C5B9424" masterDocTitle="A revision of Solanum section Herpystichum" masterLastPageNumber="1087" masterPageNumber="1068" pageId="16" pageNumber="1083" updateTime="1646746551081" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>A revision of Solanum section Herpystichum</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tepe, EJ</mods:namePart>
<mods:affiliation>Department of Biology, 257 South 1400 East, University of Utah, Salt Lake City, Utah 84112, U. S. A.; Department of Biological Sciences, 614 Rieveschl Hall, University of Cincinnati, Cincinnati, Ohio 45221, U. S. A.</mods:affiliation>
<mods:nameIdentifier type="email">eric.tepe@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bohs, L</mods:namePart>
<mods:affiliation>Department of Biology, 257 South 1400 East, University of Utah, Salt Lake City, Utah 84112, U. S. A.</mods:affiliation>
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<mods:title>Systematic Botany</mods:title>
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<mods:date>2011</mods:date>
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<mods:number>2011-12-31</mods:number>
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<mods:number>36</mods:number>
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<paragraph blockId="16.[160,809,1721,1831]" box="[160,520,1721,1743]" pageId="16" pageNumber="1083">
9. 
<taxonomicName authority="Pol.," authorityName="Pol., Linnaea" box="[191,520,1721,1743]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="1083" phylum="Tracheophyta" rank="species" species="phaseoloides">SOLANUM PHASEOLOIDES Pol.,</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection box="[533,783,1721,1743]" pageId="16" pageNumber="1083" type="reference_group">
<paragraph blockId="16.[160,809,1721,1831]" box="[533,783,1721,1743]" pageId="16" pageNumber="1083">
<treatmentCitation author="Pol." box="[533,779,1721,1743]" journal="Linnaea" page="585" pageId="16" pageNumber="1083" score="4.864797368871433" volume="41" year="1877">
<bibCitation author="Pol." box="[533,779,1721,1743]" journal="Linnaea" page="585" pageId="16" pageNumber="1083" volume="41" year="1877">Linnaea 41: 585. 1877</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="1083" type="materials_examined">
<paragraph blockId="16.[160,809,1721,1831]" pageId="16" pageNumber="1083">
— TYPE: 
<materialsCitation ID-GBIF-Occurrence="3698005303" collectingDate="1875-06" collectionCode="BM" collectorName="H. Polakowsky" country="Costa Rica" location="Desengano" pageId="16" pageNumber="1083" specimenCode="BM-BM000579755" specimenCount="1" stateProvince="Alajuela" typeStatus="lectotype">
<collectingCountry box="[288,445,1751,1772]" name="Costa Rica" pageId="16" pageNumber="1083">COSTA RICA</collectingCountry>
. 
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: 
<location LSID="urn:lsid:plazi:treatment:03BC87C5FFF08221FF5DF90D5F8697DE:8ECA6008FFF08220FDC7F9635EE992C8" box="[570,690,1751,1772]" country="Costa Rica" name="Desengano" pageId="16" pageNumber="1083" stateProvince="Alajuela">Desengaño</location>
, 
<date box="[709,808,1750,1772]" pageId="16" pageNumber="1083" value="1875-06">
<collectingDate box="[709,808,1750,1772]" pageId="16" pageNumber="1083" value="1875-06">Jun 1875</collectingDate>
</date>
(fl), 
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<collectorName box="[263,419,1780,1802]" pageId="16" pageNumber="1083">H. Polakowsky</collectorName>
147
</emphasis>
(
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, here designated: 
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!;
</materialsCitation>
isolectotype: B [destroyed]).
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="1083" type="reference_group">
<paragraph blockId="16.[160,809,1861,2001]" pageId="16" pageNumber="1083">
<treatmentCitation author="Donn.Sm." page="28" pageId="16" pageNumber="1083" year="1889">
<emphasis box="[160,358,1861,1882]" italics="true" pageId="16" pageNumber="1083">Solanum olivaeforme</emphasis>
Donn.Sm., 
<bibCitation journal="Bot. Gaz. (Crawfordsville)" page="28" pageId="16" pageNumber="1083" score="0.2895991392480618" volume="14" year="1889">Bot.Gaz.(Crawfordsville) 14:28. 1889</bibCitation>
.
</treatmentCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="1083" type="materials_examined">
<paragraph blockId="16.[160,809,1861,2001]" pageId="16" pageNumber="1083">
—TYPE: 
<materialsCitation ID-GBIF-Occurrence="3698005307" collectingDate="1887-05" collectionCode="US" collectorName="H. von Tuerckheim" country="Guatemala" elevation="1158" location="Pansamala" originalDetermination="Solanum olivaeforme Donn. Sm." pageId="16" pageNumber="1083" specimenCode="US-US00027712" specimenCount="1" stateProvince="Alta Verapaz">
<collectingCountry box="[361,521,1891,1912]" name="Guatemala" pageId="16" pageNumber="1083">GUATEMALA</collectingCountry>
. 
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: 
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, 
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<elevation box="[208,292,1920,1942]" metricMagnitude="3" metricUnit="m" metricValue="1.15824" pageId="16" pageNumber="1083" unit="ft" value="3800.0">3,800 ft</elevation>
</quantity>
, 
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<collectingDate box="[308,416,1920,1942]" pageId="16" pageNumber="1083" value="1887-05">May 1887</collectingDate>
</date>
(fl), 
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<collectorName box="[476,671,1920,1942]" pageId="16" pageNumber="1083">H. von Tuerckheim</collectorName>
1226
</emphasis>
(
<typeStatus pageId="16" pageNumber="1083">lectotype</typeStatus>
, here designated: 
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!;
</materialsCitation>
isolectotypes: GH–GH00077524!, K–K000449425!, NY–NY00169750!).
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="1083" type="description">
<paragraph blockId="16.[856,1510,160,2001]" pageId="16" pageNumber="1083">
Herbaceous vine, terrestrial. Stems slender, glabrous or rarely pubescent with slender trichomes 
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long. Sympodial units plurifoliate. Leaves 3-pinnate, the blades 1.5–7.5 × 
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, about as long as wide, membranous to chartaceous, glabrous adaxially or with widely scattered to dense, wide-diameter trichomes, 0.3–1.2 × 
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on the leaf blade, with trichomes like those of the stem more abundant along veins, glabrous abaxially, the margins entire to undulate, minutely revolute on some leaves, the rachis glabrous; lateral leaflets 0.5–5 × 
<quantity box="[1011,1102,424,446]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="5.5" metricValueMin="1.0" pageId="16" pageNumber="1083" unit="cm" value="3.25" valueMax="5.5" valueMin="1.0">1–5.5 cm</quantity>
, elliptical to rounded, the base oblique, cuneate to cordate on the proximal side, the apex obtuse to acuminate, the petiolules 
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, glabrous; apical leaflet 0.8–4 × 
<quantity box="[936,1012,512,534]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="6.0" metricValueMin="1.0" pageId="16" pageNumber="1083" unit="cm" value="3.5" valueMax="6.0" valueMin="1.0">1–6 cm</quantity>
, broadly elliptical to rhomboid, the base cuneate, the apex obtuse to acute, apiculate, the petiolule 
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, glabrous; petioles 
<quantity box="[1063,1176,571,593]" metricMagnitude="-2" metricUnit="m" metricValue="7.25" metricValueMax="13.0" metricValueMin="1.5" pageId="16" pageNumber="1083" unit="cm" value="7.25" valueMax="13.0" valueMin="1.5">1.5–13 cm</quantity>
, sparsely sand-punctate, glabrous or rarely pubescent. Internodes 
<quantity box="[1257,1346,600,622]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="12.0" metricValueMin="3.0" pageId="16" pageNumber="1083" unit="cm" value="7.5" valueMax="12.0" valueMin="3.0">3–12 cm</quantity>
. Inflorescences 
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, unbranched, extra-axillary, with (2–)3(–7) flowers, the axes glabrous; peduncle 
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, slender; rachis 
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; pedicels 
<quantity box="[1053,1161,688,710]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="16" pageNumber="1083" unit="mm" value="17.5" valueMax="25.0" valueMin="10.0">10–25 mm</quantity>
in flower and fruit, slender, glabrous, spaced 
<quantity box="[1011,1096,718,739]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="16" pageNumber="1083" unit="mm" value="6.0" valueMax="8.0" valueMin="4.0">4–8 mm</quantity>
apart. Calyx 
<quantity box="[1243,1347,717,739]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="16" pageNumber="1083" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
long, the tube 
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long, the lobes 0.5–1 × ca. 
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, deltoid to truncate, acute to acuminate at tips, glabrous; fruiting calyx slightly accrescent, the lobes 1–1.2 × 
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, truncate-acuminate. Corolla 
<quantity box="[942,1169,835,857]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="16" pageNumber="1083" unit="cm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 cmindiameter</quantity>
, ca. 
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long, stellate, white, the tube 
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, the lobes 6–7 × 
<quantity box="[1212,1317,864,886]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="3.0" metricValueMin="1.2" pageId="16" pageNumber="1083" unit="mm" value="2.1" valueMax="3.0" valueMin="1.2">1.2–3 mm</quantity>
, lanceolate, acute at the tips, glabrous adaxially and abaxially, the margins ciliate. Stamens with filaments 
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, glabrous; anthers 2–2.5 × ca. 
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. Ovary glabrous; style 4–5 × ca. 
<quantity box="[1317,1400,952,974]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="1083" unit="mm" value="0.2">0.2 mm</quantity>
, cylindrical, glabrous; stigma capitate. Fruits 2–4 × 
<quantity box="[1325,1422,981,1003]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="16" pageNumber="1083" unit="cm" value="1.75" valueMax="2.5" valueMin="1.0">1–2.5 cm</quantity>
, ovoidrhomboid, flattened, pointed at apex, greenish to yellowish, often mottled, glabrous. Seeds ca. 2 × 
<quantity box="[1250,1327,1040,1062]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="1083" unit="mm" value="1.5">1.5 mm</quantity>
in diameter, lenticular, light brown, the surface smooth. 
<figureCitation box="[1284,1388,1069,1091]" captionStart="FIG" captionStartId="2.[184,220,1816,1835]" captionTargetBox="[160,1504,158,1791]" captionTargetId="figure-10@2.[160,1504,158,1791]" captionTargetPageId="2" captionText="FIG.1. Flowers fruits, and habit of Solanum sect. Herpystichum: A. Solanum limoncochaense, habit, flower, and bud. B. S. limoncochaense fruit in situ and C. in cross section (E. J. Tepe &amp; S. Stern 2627, QCNE). D. S. trifolium flower and leaf (E. J. Tepe &amp; S. Stern 2682, QCNE). E. S. phaseoloides habit, flower, and leaf (photo by W. H. Haber). F. S. crassinervium habit (E. J. Tepe &amp; S. Stern 2729, QCNE), and G. flowers (S. Stern &amp; E. J. Tepe 400, QCNE). H. S. evolvulifolium habit and flower, and I. fruit (immature) (E. J. Tepe &amp; S. Stern 2671, QCNE). J. S. pacificum inflorescence with flowers, buds, and fruit (immature), and K. habit (E. J. Tepe et al. 2696, QCNE). L. S. loxophyllum habit, note adventitious roots at nodes, and M. older stem with inflorescences, note figure eightshaped stem (E. J. Tepe &amp; S. Stern 2726, QCNE). Inflorescences emerge from the cleft between the two lobes of the stem." figureDoi="http://doi.org/10.5281/zenodo.6327848" httpUri="https://zenodo.org/record/6327848/files/figure.png" pageId="16" pageNumber="1083">Figure 1E</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="1083" type="distribution">
<paragraph blockId="16.[856,1510,160,2001]" pageId="16" pageNumber="1083">
<emphasis box="[880,1396,1098,1120]" italics="true" pageId="16" pageNumber="1083">
<emphasis bold="true" box="[880,1181,1098,1120]" italics="true" pageId="16" pageNumber="1083">Habitat and Distribution—</emphasis>
<taxonomicName box="[1182,1396,1099,1120]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="1083" phylum="Tracheophyta" rank="species" species="phaseoloides">Solanum phaseoloides</taxonomicName>
</emphasis>
occurs in the understory of wet forests and in clearings from southern 
<collectingCountry box="[856,933,1157,1179]" name="Mexico" pageId="16" pageNumber="1083">Mexico</collectingCountry>
(
<collectingRegion box="[948,1038,1157,1179]" country="Mexico" name="Chiapas" pageId="16" pageNumber="1083">Chiapas</collectingRegion>
), 
<collectingCountry box="[1057,1175,1157,1179]" name="Guatemala" pageId="16" pageNumber="1083">Guatemala</collectingCountry>
, 
<collectingCountry box="[1186,1252,1157,1179]" name="Belize" pageId="16" pageNumber="1083">Belize</collectingCountry>
, 
<collectingCountry box="[1264,1374,1157,1179]" name="Honduras" pageId="16" pageNumber="1083">Honduras</collectingCountry>
, 
<collectingCountry box="[1385,1500,1157,1179]" name="Costa Rica" pageId="16" pageNumber="1083">Costa Rica</collectingCountry>
, to 
<collectingCountry box="[883,972,1187,1208]" name="Panama" pageId="16" pageNumber="1083">Panama</collectingCountry>
. It is usually terrestrial, but occasionally grows on fallen logs or as an epiphyte; 
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in elevation. Asingle collection is known from 
<collectingCountry box="[1162,1212,1246,1267]" name="Peru" pageId="16" pageNumber="1083">Peru</collectingCountry>
(
<figureCitation box="[1226,1289,1245,1268]" captionStart="FIG" captionStartId="17.[152,188,904,923]" captionTargetBox="[128,776,158,879]" captionTargetId="figure-1216@17.[128,776,158,879]" captionTargetPageId="17" captionText="FIG.9. Distribution of Solanum phaseoloides and S. trifolium." figureDoi="http://doi.org/10.5281/zenodo.6327864" httpUri="https://zenodo.org/record/6327864/files/figure.png" pageId="16" pageNumber="1083">Fig. 9</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="1083" type="biology_ecology">
<paragraph blockId="16.[856,1510,160,2001]" pageId="16" pageNumber="1083">
<emphasis bold="true" box="[880,1018,1274,1296]" italics="true" pageId="16" pageNumber="1083">Phenology—</emphasis>
Flowering and fruiting occur throughout the year.
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="1083" type="etymology">
<paragraph blockId="16.[856,1510,160,2001]" pageId="16" pageNumber="1083">
<emphasis bold="true" box="[880,1021,1333,1355]" italics="true" pageId="16" pageNumber="1083">Etymology—</emphasis>
The name, 
<taxonomicName box="[1148,1267,1333,1354]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="1083" phylum="Tracheophyta" rank="species" species="phaseoloides">
<emphasis box="[1148,1267,1333,1354]" italics="true" pageId="16" pageNumber="1083">phaseoloides</emphasis>
</taxonomicName>
, refers to the similar leaves and viny habit of the genus 
<taxonomicName authorityName="Monterosato" authorityYear="1875" box="[1224,1320,1363,1384]" class="Magnoliopsida" family="Fabaceae" genus="Phaseolus" kingdom="Plantae" order="Fabales" pageId="16" pageNumber="1083" phylum="Tracheophyta" rank="genus">
<emphasis box="[1224,1320,1363,1384]" italics="true" pageId="16" pageNumber="1083">Phaseolus</emphasis>
</taxonomicName>
L. (
<taxonomicName authorityName="Lindley &amp; John" authorityYear="1830" box="[1360,1461,1363,1385]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="16" pageNumber="1083" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="17" lastPageNumber="1084" pageId="16" pageNumber="1083" type="discussion">
<paragraph blockId="16.[856,1510,160,2001]" lastBlockId="17.[128,782,976,2002]" lastPageId="17" lastPageNumber="1084" pageId="16" pageNumber="1083">
<emphasis box="[880,1185,1392,1413]" italics="true" pageId="16" pageNumber="1083">
<emphasis bold="true" box="[880,967,1392,1413]" italics="true" pageId="16" pageNumber="1083">Notes—</emphasis>
<taxonomicName box="[967,1185,1392,1413]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="1083" phylum="Tracheophyta" rank="species" species="phaseoloides">Solanum phaseoloides</taxonomicName>
</emphasis>
is a ground-trailing species characterized by 3-foliate leaves and stellate flowers. It is most similar to 
<taxonomicName baseAuthorityName="Bitter" box="[963,1123,1451,1472]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="1083" phylum="Tracheophyta" rank="species" species="pentaphyllum">
<emphasis box="[963,1123,1451,1472]" italics="true" pageId="16" pageNumber="1083">S. pentaphyllum</emphasis>
</taxonomicName>
from which it differs by its 3-foliate vs. 5-foliate leaves. It also resembles 
<emphasis box="[1260,1381,1480,1501]" italics="true" pageId="16" pageNumber="1083">
<taxonomicName baseAuthorityName="Dunal" box="[1260,1377,1480,1501]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="1083" phylum="Tracheophyta" rank="species" species="trifolium">S. trifolium</taxonomicName>
,
</emphasis>
but can be differentiated by its white, stellate flowers and pointed leaf apices vs. blue-violet, rotate flowers and rounded leaf apices. Vegetative parts of most collections are glabrous; however, plants with pubescent stems and petioles appear to be localized in the Cordillera de Talamanca in 
<collectingCountry box="[1286,1401,1627,1649]" name="Costa Rica" pageId="16" pageNumber="1083">Costa Rica</collectingCountry>
at 
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.45" metricValueMax="3.2" metricValueMin="1.7" pageId="16" pageNumber="1083" unit="m" value="2450.0" valueMax="3200.0" valueMin="1700.0">
<geoCoordinate box="[1438,1491,1627,1648]" degrees="1,700" direction="north" orientation="latitude" pageId="16" pageNumber="1083" precision="55" value="1.7">1,700</geoCoordinate>
<geoCoordinate degrees="3,200" direction="west" orientation="longitude" pageId="16" pageNumber="1083" precision="55" value="-3.2">– 3,200</geoCoordinate>
m
</quantity>
. Asingle collection of 
<taxonomicName box="[1190,1336,1656,1677]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="1083" phylum="Tracheophyta" rank="species" species="phaseoloides">
<emphasis box="[1190,1336,1656,1677]" italics="true" pageId="16" pageNumber="1083">S. phaseoloides</emphasis>
</taxonomicName>
is known from north-central 
<collectingCountry box="[1000,1053,1686,1707]" name="Peru" pageId="16" pageNumber="1083">Peru</collectingCountry>
. The leaves of this collection are somewhat fleshier than typical plants from Central America, but it corresponds to them in other characters. If this specimen truly represents a population of 
<taxonomicName box="[1143,1288,1773,1795]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="1083" phylum="Tracheophyta" rank="species" species="phaseoloides">
<emphasis box="[1143,1288,1773,1795]" italics="true" pageId="16" pageNumber="1083">S. phaseoloides</emphasis>
</taxonomicName>
, then it is a remarkable disjunction from the species’ native range in Central America. Alternatively, it is possible that this collection represents an additional species of sect. 
<taxonomicName baseAuthorityName="Bitter" box="[1254,1393,1861,1882]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="1086" phylum="Tracheophyta" rank="section" section="Herpystichum">
<emphasis box="[1254,1393,1861,1882]" italics="true" pageId="16" pageNumber="1083">Herpystichum</emphasis>
</taxonomicName>
; however, we must reserve judgment until additional collections are available. We have been unable to successfully obtain a DNA sequence from this Peruvian collection. Finally, we have not seen collections of 
<taxonomicName box="[1051,1194,1979,2000]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="1083" phylum="Tracheophyta" rank="species" species="phaseoloides">
<emphasis box="[1051,1194,1979,2000]" italics="true" pageId="16" pageNumber="1083">S. phaseoloides</emphasis>
</taxonomicName>
from 
<collectingCountry box="[1255,1368,1979,2001]" name="Nicaragua" pageId="16" pageNumber="1083">Nicaragua</collectingCountry>
; however, its presence there is expected based on the overall distribution of this species.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.6327864" ID-Zenodo-Dep="6327864" box="[152,690,904,924]" httpUri="https://zenodo.org/record/6327864/files/figure.png" pageId="17" pageNumber="1084" startId="17.[152,188,904,923]" targetBox="[128,776,158,879]" targetPageId="17">
<paragraph blockId="17.[152,690,904,924]" box="[152,690,904,924]" pageId="17" pageNumber="1084">
FIG.9. Distribution of 
<taxonomicName box="[367,543,904,923]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="17" pageNumber="1084" phylum="Tracheophyta" rank="species" species="phaseoloides">
<emphasis box="[367,543,904,923]" italics="true" pageId="17" pageNumber="1084">Solanum phaseoloides</emphasis>
</taxonomicName>
and 
<taxonomicName baseAuthorityName="Dunal" box="[586,680,904,923]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="17" pageNumber="1084" phylum="Tracheophyta" rank="species" species="trifolium">
<emphasis box="[586,680,904,923]" italics="true" pageId="17" pageNumber="1084">S. trifolium</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph blockId="17.[128,782,976,2002]" pageId="17" pageNumber="1084">
The location Polakowsky’s 
<typeStatus box="[442,536,1034,1056]" pageId="17" pageNumber="1084">holotype</typeStatus>
is not explicitly stated in the protologue; the existence of a specimen at Bis implied, but it has almost certainly been destroyed and no photos of the specimen are known to exist. A duplicate of this collection exists at BM and this specimen (BM000579755) has been chosen as the 
<typeStatus box="[237,337,1181,1203]" pageId="17" pageNumber="1084">lectotype</typeStatus>
.
</paragraph>
<paragraph blockId="17.[128,782,976,2002]" pageId="17" pageNumber="1084">
Donnell Smith did not designate a single sheet of 
<emphasis italics="true" pageId="17" pageNumber="1084">Tuerckheim 1226</emphasis>
as the 
<typeStatus box="[254,348,1240,1262]" pageId="17" pageNumber="1084">holotype</typeStatus>
of 
<emphasis box="[385,523,1240,1261]" italics="true" pageId="17" pageNumber="1084">S. olivaeforme</emphasis>
in his 1889 protologue. The specimen from 
<collectingCountry box="[336,367,1269,1290]" name="United States of America" pageId="17" pageNumber="1084">US</collectingCountry>
(
<collectingCountry box="[379,409,1269,1291]" name="United States of America" pageId="17" pageNumber="1084">US</collectingCountry>
00027712) is here chosen as the 
<typeStatus pageId="17" pageNumber="1084">lectotype</typeStatus>
. Donnell Smith’s herbarium is now held at 
<collectingCountry box="[654,685,1299,1320]" name="United States of America" pageId="17" pageNumber="1084">US</collectingCountry>
and this specimen has the original label written in Donnell Smith’s hand.
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="1084" type="materials_examined">
<paragraph blockId="17.[128,782,976,2002]" pageId="17" pageNumber="1084">
<emphasis bold="true" box="[160,529,1383,1403]" italics="true" pageId="17" pageNumber="1084">Representative Specimens Examined—</emphasis>
<collectingCountry box="[529,613,1383,1402]" name="Mexico" pageId="17" pageNumber="1084">MEXICO</collectingCountry>
. 
<collectingRegion box="[633,707,1383,1402]" country="Mexico" name="Chiapas" pageId="17" pageNumber="1084">Chiapas</collectingRegion>
: Finca Irelanda, 
<date box="[215,295,1407,1426]" pageId="17" pageNumber="1084" value="1914-06">Jun 1914</date>
(fl), 
<emphasis box="[343,507,1407,1426]" italics="true" pageId="17" pageNumber="1084">C. A. Purpus 7310</emphasis>
(BM, F, GH, NY, 
<collectionCode box="[678,710,1407,1426]" country="USA" name="Upjohn Culture Collection" pageId="17" pageNumber="1084">UC</collectionCode>
); Finca Mexiquito, 
<date box="[229,305,1431,1450]" pageId="17" pageNumber="1084" value="1913-06">Jun 1913</date>
(fl), 
<emphasis box="[345,498,1431,1450]" italics="true" pageId="17" pageNumber="1084">C. A. Purpus 7460</emphasis>
(
<collectionCode box="[510,546,1431,1450]" country="USA" name="Upjohn Culture Collection" pageId="17" pageNumber="1084">UC</collectionCode>
).
</paragraph>
<paragraph blockId="17.[128,782,976,2002]" pageId="17" pageNumber="1084">
<collectingCountry box="[160,294,1455,1474]" name="Guatemala" pageId="17" pageNumber="1084">GUATEMALA</collectingCountry>
. 
<collectingRegion box="[299,414,1455,1474]" country="Guatemala" name="Alta Verapaz" pageId="17" pageNumber="1084">Alta Verapaz</collectingRegion>
:Barrancade Rubelcruz, 
<geoCoordinate box="[626,696,1455,1474]" degrees="15" direction="north" minutes="29" orientation="latitude" pageId="17" pageNumber="1084" precision="925" value="15.483334">15°29’N</geoCoordinate>
<geoCoordinate box="[699,773,1455,1474]" degrees="90" direction="west" minutes="08" orientation="longitude" pageId="17" pageNumber="1084" precision="925" value="-90.13333">90°08’W</geoCoordinate>
, 
<quantity box="[128,195,1479,1498]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="17" pageNumber="1084" unit="ft" value="2500.0">2,500 ft</quantity>
, 
<date box="[205,287,1479,1498]" pageId="17" pageNumber="1084" value="1889-04">Apr 1889</date>
(fr), 
<emphasis box="[334,522,1479,1498]" italics="true" pageId="17" pageNumber="1084">J. Donnell Smith 1785</emphasis>
(G, GH, M, 
<collectingCountry box="[638,666,1479,1498]" name="United States of America" pageId="17" pageNumber="1084">US</collectingCountry>
); Vicinityof Laguna Sapalá (Chajvuvuch), 
<quantity box="[405,463,1503,1522]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="17" pageNumber="1084" unit="m" value="280.0">280 m</quantity>
, 
<date box="[475,588,1503,1522]" pageId="17" pageNumber="1084" value="1942-03-11">11 Mar 1942</date>
(fl), 
<emphasis italics="true" pageId="17" pageNumber="1084">J. A. Steyermark 44895</emphasis>
(F, 
<collectionCode box="[217,250,1527,1546]" country="USA" name="Upjohn Culture Collection" pageId="17" pageNumber="1084">UC</collectionCode>
). Izabal: Montañas del Mico, 
<quantity box="[527,595,1527,1546]" metricMagnitude="3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="17" pageNumber="1084" unit="km" value="7.5" valueMax="8.0" valueMin="7.0">7–8 km</quantity>
<collectionCode box="[604,651,1527,1546]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="17" pageNumber="1084" type="Herbarium">Wof</collectionCode>
Santo Tomás de Castilla on road to microwave tower, 
<geoCoordinate box="[511,581,1551,1570]" degrees="15" direction="north" minutes="40" orientation="latitude" pageId="17" pageNumber="1084" precision="925" value="15.666667">15°40’N</geoCoordinate>
<geoCoordinate box="[589,663,1551,1570]" degrees="88" direction="west" minutes="40" orientation="longitude" pageId="17" pageNumber="1084" precision="925" value="-88.666664">88°40’W</geoCoordinate>
, 
<quantity box="[675,772,1551,1570]" metricMagnitude="2" metricUnit="m" metricValue="6.25" metricValueMax="6.5" metricValueMin="6.0" pageId="17" pageNumber="1084" unit="m" value="625.0" valueMax="650.0" valueMin="600.0">600–650 m</quantity>
, 
<date box="[128,234,1575,1594]" pageId="17" pageNumber="1084" value="1988-08-19">19 Aug 1988</date>
(fl, fr), 
<emphasis box="[296,503,1575,1594]" italics="true" pageId="17" pageNumber="1084">
<collectionCode box="[296,316,1576,1594]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="17" pageNumber="1084" type="Herbarium">W</collectionCode>
. D. Stevens et al. 25577
</emphasis>
(
<collectionCode box="[515,547,1575,1594]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="17" pageNumber="1084" type="Herbarium">NY</collectionCode>
). Peten: La Cumbre, Pusila road, in high forest, 
<date box="[315,428,1599,1618]" pageId="17" pageNumber="1084" value="1976-08-17">17 Aug 1976</date>
(fl), 
<emphasis box="[473,776,1599,1618]" italics="true" pageId="17" pageNumber="1084">C. L. Lundell &amp; E. Contreras 20189</emphasis>
(F, 
<collectionCode box="[160,188,1623,1642]" country="USA" lsid="urn:lsid:biocol.org:col:14627" name="University of Texas at Austin" pageId="17" pageNumber="1084" type="Herbarium">LL</collectionCode>
). Quetzaltenango: Finca Pirineos, lower south-facing slopes of Volcán Santa María, between Santa María de Jesús and Calahuaché, 
<quantity box="[128,254,1671,1690]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="17" pageNumber="1084" unit="m" value="1400.0" valueMax="1500.0" valueMin="1300.0">
<geoCoordinate box="[128,173,1671,1690]" degrees="1,300" direction="north" orientation="latitude" pageId="17" pageNumber="1084" precision="55" value="1.3">1,300</geoCoordinate>
<geoCoordinate box="[173,228,1671,1690]" degrees="1,500" direction="west" orientation="longitude" pageId="17" pageNumber="1084" precision="55" value="-1.5">–1,500</geoCoordinate>
m
</quantity>
, 
<date box="[265,373,1671,1690]" pageId="17" pageNumber="1084" value="1939-12-31">31 Dec 1939</date>
(infl), 
<emphasis box="[436,631,1671,1690]" italics="true" pageId="17" pageNumber="1084">J. A. Steyermark 33254</emphasis>
(F). Retalhuleu: Barrancade Salamá, 
<quantity box="[328,399,1695,1714]" metricMagnitude="2" metricUnit="m" metricValue="5.1816" pageId="17" pageNumber="1084" unit="ft" value="1700.0">1,700 ft</quantity>
, 
<date box="[412,497,1695,1714]" pageId="17" pageNumber="1084" value="1892-04">Apr 1892</date>
(fl, fr), 
<emphasis box="[579,776,1695,1714]" italics="true" pageId="17" pageNumber="1084">J. Donnell Smith 2673</emphasis>
(F, GH, K, M, MO, NY, 
<collectingCountry box="[324,352,1719,1738]" name="United States of America" pageId="17" pageNumber="1084">US</collectingCountry>
, 
<collectionCode box="[359,397,1719,1738]" country="USA" name="Wayland University" pageId="17" pageNumber="1084">WU</collectionCode>
); Near Chivolandia (Dept. Quezaltenango), along road to San Felipe, 
<quantity box="[351,405,1743,1762]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="17" pageNumber="1084" unit="m" value="650.0">650 m</quantity>
, 
<date box="[413,515,1743,1762]" pageId="17" pageNumber="1084" value="1941-02-15">15 Feb 1941</date>
(infl), 
<emphasis box="[573,742,1743,1762]" italics="true" pageId="17" pageNumber="1084">P. C. Standley 87182</emphasis>
(F). San Marcos: Finca El Porvenir, along Río Chopal, south-facing slopes of Volcán Tajumulco, 
<quantity box="[297,423,1791,1810]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="17" pageNumber="1084" unit="m" value="1400.0" valueMax="1500.0" valueMin="1300.0">
<geoCoordinate box="[297,342,1791,1810]" degrees="1,300" direction="north" orientation="latitude" pageId="17" pageNumber="1084" precision="55" value="1.3">1,300</geoCoordinate>
<geoCoordinate box="[342,398,1791,1810]" degrees="1,500" direction="west" orientation="longitude" pageId="17" pageNumber="1084" precision="55" value="-1.5">–1,500</geoCoordinate>
m
</quantity>
, 
<date box="[433,541,1791,1810]" pageId="17" pageNumber="1084" value="1940-03-11">11 Mar 1940</date>
(fl), 
<emphasis box="[584,776,1791,1810]" italics="true" pageId="17" pageNumber="1084">J. A. Steyermark 37528</emphasis>
(F). Suchitepéquez: Southern lower slopes of Volcán Zunil, vicinity of Finca Las Nubes, along Quebrada Chita, Eof Pueblo Nuevo, 
<quantity box="[677,772,1839,1858]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="17" pageNumber="1084" unit="m" value="650.0" valueMax="800.0" valueMin="500.0">500–800 m</quantity>
, 
<date box="[128,220,1863,1882]" pageId="17" pageNumber="1084" value="1940-02-02">2 Feb 1940</date>
(fl), 
<emphasis box="[261,449,1863,1882]" italics="true" pageId="17" pageNumber="1084">J. A. Steyermark 35395</emphasis>
(F).
</paragraph>
<paragraph blockId="17.[128,782,976,2002]" pageId="17" pageNumber="1084">
<collectingCountry box="[160,232,1887,1906]" name="Belize" pageId="17" pageNumber="1084">BELIZE</collectingCountry>
. Esperanza Rd, 
<quantity box="[379,442,1887,1906]" metricMagnitude="2" metricUnit="m" metricValue="7.9248" pageId="17" pageNumber="1084" unit="ft" value="2600.0">2600 ft</quantity>
, 
<date box="[453,543,1887,1906]" pageId="17" pageNumber="1084" value="1934-07-02">2 Jul 1934</date>
(fl), 
<emphasis box="[589,739,1887,1906]" italics="true" pageId="17" pageNumber="1084">
<collectionCode box="[589,609,1888,1906]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="17" pageNumber="1084" type="Herbarium">W</collectionCode>
. A. Schipp 727
</emphasis>
(F). 
<collectingRegion box="[128,177,1911,1930]" country="Belize" name="Cayo" pageId="17" pageNumber="1084">Cayo</collectingRegion>
: Chiquibul, Ceibo Grande to Main Divide track, 
<geoCoordinate box="[678,776,1910,1930]" degrees="16" direction="north" minutes="31" orientation="latitude" pageId="17" pageNumber="1084" precision="15" seconds="49" value="16.530277">16°31′49″N</geoCoordinate>
<geoCoordinate box="[128,229,1934,1954]" degrees="89" direction="west" minutes="05" orientation="longitude" pageId="17" pageNumber="1084" precision="15" seconds="06" value="-89.085">89°05′06″W</geoCoordinate>
, 
<quantity box="[239,294,1935,1954]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="1084" unit="m" value="500.0">500 m</quantity>
, 
<date box="[304,415,1935,1954]" pageId="17" pageNumber="1084" value="1998-08-26">26 Aug 1998</date>
(fl), 
<emphasis box="[458,727,1935,1954]" italics="true" pageId="17" pageNumber="1084">A. K. Monro &amp; S. Cafferty 2689</emphasis>
(BM, 
<collectionCode box="[128,165,1959,1978]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="17" pageNumber="1084" type="Herbarium">MO</collectionCode>
). 
<collectingRegion box="[178,240,1959,1978]" country="Belize" name="Toledo" pageId="17" pageNumber="1084">Toledo</collectingRegion>
:in high ridge, on hill slope, Edwards road beyond Columbia, 
<geoCoordinate box="[128,198,1983,2002]" degrees="16" direction="north" minutes="14" orientation="latitude" pageId="17" pageNumber="1084" precision="925" value="16.233334">16°14’N</geoCoordinate>
<geoCoordinate box="[203,277,1983,2002]" degrees="88" direction="west" minutes="14" orientation="longitude" pageId="17" pageNumber="1084" precision="925" value="-88.23333">88°14’W</geoCoordinate>
, 
<date box="[285,387,1983,2002]" pageId="17" pageNumber="1084" value="1948-02-18">18 Feb 1948</date>
(fl), 
<emphasis box="[427,571,1983,2002]" italics="true" pageId="17" pageNumber="1084">P. H. Gentle 6417</emphasis>
(
<collectionCode box="[583,614,1983,2002]" country="USA" lsid="urn:lsid:biocol.org:col:14627" name="University of Texas at Austin" pageId="17" pageNumber="1084" type="Herbarium">LL</collectionCode>
).
</paragraph>
<paragraph blockId="17.[824,1478,159,2003]" pageId="17" pageNumber="1084">
<collectingCountry box="[856,978,159,178]" name="Honduras" pageId="17" pageNumber="1084">HONDURAS</collectingCountry>
. La Muralla visitors center and environs, 
<quantity box="[1346,1391,159,178]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="17" pageNumber="1084" unit="km" value="8.0">8 km</quantity>
NNW of La Union, 
<geoCoordinate box="[920,990,183,202]" degrees="15" direction="north" minutes="05" orientation="latitude" pageId="17" pageNumber="1084" precision="925" value="15.083333">15°05’N</geoCoordinate>
<geoCoordinate box="[997,1071,183,202]" degrees="86" direction="west" minutes="44" orientation="longitude" pageId="17" pageNumber="1084" precision="925" value="-86.73333">86°44’W</geoCoordinate>
, 
<quantity box="[1081,1207,183,202]" metricMagnitude="3" metricUnit="m" metricValue="1.4975" metricValueMax="1.58" metricValueMin="1.415" pageId="17" pageNumber="1084" unit="m" value="1497.5" valueMax="1580.0" valueMin="1415.0">
<geoCoordinate box="[1081,1125,183,202]" degrees="1,415" direction="north" orientation="latitude" pageId="17" pageNumber="1084" precision="55" value="1.415">1,415</geoCoordinate>
<geoCoordinate box="[1125,1181,183,202]" degrees="1,580" direction="west" orientation="longitude" pageId="17" pageNumber="1084" precision="55" value="-1.58">–1,580</geoCoordinate>
m
</quantity>
, 
<date box="[1217,1311,183,202]" pageId="17" pageNumber="1084" value="1992-06-04">4 Jun 1992</date>
(fl), 
<emphasis italics="true" pageId="17" pageNumber="1084">
<collectionCode box="[1355,1375,184,202]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="17" pageNumber="1084" type="Herbarium">W</collectionCode>
. G. D’Arcy 18123
</emphasis>
(MO, 
<collectionCode box="[934,967,207,226]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="17" pageNumber="1084" type="Herbarium">NY</collectionCode>
). 
<collectingRegion box="[982,1025,207,226]" country="Honduras" name="Yoro" pageId="17" pageNumber="1084">Yoro</collectingRegion>
: Ca. 
<quantity box="[1070,1126,207,226]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="17" pageNumber="1084" unit="km" value="16.0">16 km</quantity>
from Yarucha on Quebrada de Oro to Cerro Bufalo, 
<quantity box="[947,1041,231,250]" metricMagnitude="2" metricUnit="m" metricValue="9.25" metricValueMax="9.5" metricValueMin="9.0" pageId="17" pageNumber="1084" unit="m" value="925.0" valueMax="950.0" valueMin="900.0">900–950 m</quantity>
, 
<date box="[1049,1157,231,250]" pageId="17" pageNumber="1084" value="1982-08-16">16 Aug 1982</date>
(fl, fr), 
<emphasis box="[1223,1380,231,250]" italics="true" pageId="17" pageNumber="1084">
<collectionCode box="[1223,1243,232,250]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="17" pageNumber="1084" type="Herbarium">W</collectionCode>
. C. Holmes 4407
</emphasis>
(
<collectionCode box="[1392,1429,231,250]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="17" pageNumber="1084" type="Herbarium">NY</collectionCode>
).
</paragraph>
<paragraph blockId="17.[824,1478,159,2003]" pageId="17" pageNumber="1084">
<collectingCountry box="[856,982,255,274]" name="Costa Rica" pageId="17" pageNumber="1084">COSTA RICA</collectingCountry>
. Rampante - Forets de la Palma, 
<date box="[1291,1373,255,274]" pageId="17" pageNumber="1084" value="1898-09">Sep 1898</date>
, 
<emphasis italics="true" pageId="17" pageNumber="1084">A. Tonduz 12611</emphasis>
(
<collectingCountry box="[887,916,279,298]" name="United States of America" pageId="17" pageNumber="1084">US</collectingCountry>
). Alajuela: Primary forest on ridge top between Río Gorrión and Río Toro, 
<geoCoordinate box="[960,1030,303,322]" degrees="10" direction="north" minutes="12" orientation="latitude" pageId="17" pageNumber="1084" precision="925" value="10.2">10°12’N</geoCoordinate>
<geoCoordinate box="[1039,1113,303,322]" degrees="84" direction="west" minutes="19" orientation="longitude" pageId="17" pageNumber="1084" precision="925" value="-84.316666">84°19’W</geoCoordinate>
, 
<quantity box="[1125,1198,303,322]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="17" pageNumber="1084" unit="m" value="1700.0">1,700 m</quantity>
, 
<date box="[1211,1310,303,322]" pageId="17" pageNumber="1084" value="1972-10-07">7 Oct 1972</date>
(fl), 
<emphasis box="[1359,1472,303,322]" italics="true" pageId="17" pageNumber="1084">R. Lent 2998</emphasis>
(F). Cartago: Paraíso, P. N. Tapantí-Macizo de La Muerte, Cuenca del Reventazón, 
<geoCoordinate box="[935,1022,350,370]" degrees="9" direction="north" minutes="45" orientation="latitude" pageId="17" pageNumber="1084" precision="15" seconds="20" value="9.755555">9°45′20”N</geoCoordinate>
<geoCoordinate box="[1027,1124,350,370]" degrees="83" direction="west" minutes="47" orientation="longitude" pageId="17" pageNumber="1084" precision="15" seconds="00" value="-83.78333">83°47′00″W</geoCoordinate>
, 
<quantity box="[1132,1200,351,370]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="17" pageNumber="1084" unit="m" value="1250.0">1,250 m</quantity>
, 
<date box="[1208,1314,351,370]" pageId="17" pageNumber="1084" value="2000-03-15">15 Mar 2000</date>
(fl, fr), 
<emphasis italics="true" pageId="17" pageNumber="1084">L. Acosta &amp; V. Ramirez 646</emphasis>
(
<collectionCode box="[966,999,375,394]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="17" pageNumber="1084" type="Herbarium">NY</collectionCode>
); El Muñeco, Sof Cartago and Navarro Valley, near boundary of San Joséand Cartago provinces, 
<quantity box="[1222,1286,399,418]" metricMagnitude="3" metricUnit="m" metricValue="1.3716" pageId="17" pageNumber="1084" unit="ft" value="4500.0">4,500 ft</quantity>
, 
<date box="[1294,1393,399,418]" pageId="17" pageNumber="1084" value="1928-06-19">19 Jun 1928</date>
(fr), 
<emphasis italics="true" pageId="17" pageNumber="1084">H.E. Stork 2659</emphasis>
(F, 
<collectionCode box="[941,974,423,442]" country="USA" name="Upjohn Culture Collection" pageId="17" pageNumber="1084">UC</collectionCode>
). Heredia: Braulio Carrillo Park, 
<quantity box="[1260,1383,423,442]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="17" pageNumber="1084" unit="m" value="1850.0" valueMax="2000.0" valueMin="1700.0">
<geoCoordinate box="[1260,1304,423,442]" degrees="1,700" direction="north" orientation="latitude" pageId="17" pageNumber="1084" precision="55" value="1.7">1,700</geoCoordinate>
<geoCoordinate box="[1304,1360,423,442]" degrees="2,000" direction="west" orientation="longitude" pageId="17" pageNumber="1084" precision="55" value="-2.0">–2,000</geoCoordinate>
m
</quantity>
, 
<date box="[1391,1472,423,442]" pageId="17" pageNumber="1084" value="1983-03">Mar 1983</date>
(st), 
<emphasis box="[863,1024,447,466]" italics="true" pageId="17" pageNumber="1084">L. D. Gómez 20200</emphasis>
(MO, 
<collectionCode box="[1082,1114,447,466]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="17" pageNumber="1084" type="Herbarium">NY</collectionCode>
); Vara Blanca de Sarapiquí, North slope of Central Cordillera, 
<quantity box="[1020,1145,471,490]" metricMagnitude="3" metricUnit="m" metricValue="1.625" metricValueMax="1.75" metricValueMin="1.5" pageId="17" pageNumber="1084" unit="m" value="1625.0" valueMax="1750.0" valueMin="1500.0">
<geoCoordinate box="[1020,1065,471,490]" degrees="1,500" direction="north" orientation="latitude" pageId="17" pageNumber="1084" precision="55" value="1.5">1,500</geoCoordinate>
<geoCoordinate box="[1064,1120,471,490]" degrees="1,750" direction="west" orientation="longitude" pageId="17" pageNumber="1084" precision="55" value="-1.75">–1,750</geoCoordinate>
m
</quantity>
, 
<date box="[1154,1225,471,490]" pageId="17" pageNumber="1084" value="1937-07">Jul 1937</date>
(fl, fr), 
<emphasis box="[1293,1440,471,490]" italics="true" pageId="17" pageNumber="1084">A. F. Skutch 3177</emphasis>
(K, MO, NY, 
<collectingCountry box="[908,937,495,514]" name="United States of America" pageId="17" pageNumber="1084">US</collectingCountry>
). Puntarenas: Ca. 
<quantity box="[1098,1142,495,514]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="1084" unit="km" value="2.0">2 km</quantity>
SE of Monteverde, 
<geoCoordinate box="[1319,1389,495,514]" degrees="10" direction="north" minutes="18" orientation="latitude" pageId="17" pageNumber="1084" precision="925" value="10.3">10°18’N</geoCoordinate>
<geoCoordinate box="[1395,1469,495,514]" degrees="84" direction="west" minutes="48" orientation="longitude" pageId="17" pageNumber="1084" precision="925" value="-84.8">84°48’W</geoCoordinate>
, 
<quantity box="[824,947,519,538]" metricMagnitude="3" metricUnit="m" metricValue="1.525" metricValueMax="1.55" metricValueMin="1.5" pageId="17" pageNumber="1084" unit="m" value="1525.0" valueMax="1550.0" valueMin="1500.0">
<geoCoordinate box="[824,869,519,538]" degrees="1,500" direction="north" orientation="latitude" pageId="17" pageNumber="1084" precision="55" value="1.5">1,500</geoCoordinate>
<geoCoordinate box="[869,924,519,538]" degrees="1,550" direction="west" orientation="longitude" pageId="17" pageNumber="1084" precision="55" value="-1.55">–1,550</geoCoordinate>
m
</quantity>
, 
<date box="[955,1060,519,538]" pageId="17" pageNumber="1084" value="1973-03-18">18 Mar 1973</date>
(fl, fr), 
<emphasis box="[1123,1391,519,538]" italics="true" pageId="17" pageNumber="1084">
J.L. Gentry &amp; 
<collectionCode box="[1241,1261,520,538]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="17" pageNumber="1084" type="Herbarium">W</collectionCode>
. C.Burger 2696
</emphasis>
(AAU, F, 
<collectionCode box="[824,861,543,562]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="17" pageNumber="1084" type="Herbarium">MO</collectionCode>
); Cantón de Osa, Fila Costeña, Fila Cruces, cabeceras del Río Piedras Blancas, Cerro Anguciana, faldas al Oeste, 
<geoCoordinate box="[1211,1300,566,586]" degrees="8" direction="north" minutes="48" orientation="latitude" pageId="17" pageNumber="1084" precision="15" seconds="56" value="8.815556">8°48′56″N</geoCoordinate>
<geoCoordinate box="[1306,1407,566,586]" degrees="83" direction="west" minutes="10" orientation="longitude" pageId="17" pageNumber="1084" precision="15" seconds="37" value="-83.17694">83°10′37″W</geoCoordinate>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="17" pageNumber="1084" unit="m" value="1500.0" valueMax="1600.0" valueMin="1400.0">
<geoCoordinate box="[1417,1461,567,586]" degrees="1,400" direction="north" orientation="latitude" pageId="17" pageNumber="1084" precision="55" value="1.4">1,400</geoCoordinate>
<geoCoordinate degrees="1,600" direction="west" orientation="longitude" pageId="17" pageNumber="1084" precision="55" value="-1.6">– 1,600</geoCoordinate>
m
</quantity>
, 
<date box="[908,1019,591,610]" pageId="17" pageNumber="1084" value="1993-12-10">10 Dec 1993</date>
, 
<emphasis box="[1030,1182,591,610]" italics="true" pageId="17" pageNumber="1084">B. Hammel 19293</emphasis>
(
<collectionCode box="[1196,1233,591,610]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="17" pageNumber="1084" type="Herbarium">MO</collectionCode>
); Fila las Cruces near San Vito de Java, 
<quantity box="[942,1011,615,634]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="17" pageNumber="1084" unit="m" value="1400.0">1,400 m</quantity>
, 
<date box="[1020,1129,615,634]" pageId="17" pageNumber="1084" value="1974-08-22">22 Aug 1974</date>
(fl, fr), 
<emphasis box="[1196,1446,615,634]" italics="true" pageId="17" pageNumber="1084">P. J. Maas &amp; B. McAlpin 1400</emphasis>
(F, GH, U). San José: Cordillera Talamanca, mountain of Cerro de la Muerte, Panamerican highway between San Isidro El General and Division, 
<quantity box="[1402,1469,663,682]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="17" pageNumber="1084" unit="m" value="1900.0">1,900 m</quantity>
, 
<date box="[824,924,687,706]" pageId="17" pageNumber="1084" value="1966-03-04">4 Mar 1966</date>
(fl), 
<emphasis box="[969,1223,687,706]" italics="true" pageId="17" pageNumber="1084">
A. 
<taxonomicName authority="Rositto et al." authorityName="Rositto" baseAuthorityName="Rositto et al." box="[995,1168,687,706]" class="Malacostraca" family="Paramelitidae" genus="Molina" kingdom="Animalia" order="Amphipoda" pageId="17" pageNumber="1084" phylum="Arthropoda" rank="genus">Molina Rositto et al.</taxonomicName>
18354
</emphasis>
(BM, F, GH, 
<collectionCode box="[1344,1376,687,706]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="17" pageNumber="1084" type="Herbarium">NY</collectionCode>
); San José, slopes of Cordillera de Talamanca Nof San Isidro El General, 
<quantity box="[1349,1469,711,730]" metricMagnitude="3" metricUnit="m" metricValue="1.875" metricValueMax="2.0" metricValueMin="1.75" pageId="17" pageNumber="1084" unit="m" value="1875.0" valueMax="2000.0" valueMin="1750.0">
<geoCoordinate box="[1349,1391,711,730]" degrees="1,750" direction="north" orientation="latitude" pageId="17" pageNumber="1084" precision="55" value="1.75">1,750</geoCoordinate>
<geoCoordinate box="[1391,1445,711,730]" degrees="2,000" direction="west" orientation="longitude" pageId="17" pageNumber="1084" precision="55" value="-2.0">–2,000</geoCoordinate>
m
</quantity>
, 
<date box="[824,916,735,754]" pageId="17" pageNumber="1084" value="1963-02-05">5 Feb 1963</date>
(fl), 
<emphasis box="[957,1175,735,754]" italics="true" pageId="17" pageNumber="1084">L. O. Williams et al. 24342</emphasis>
(F, G, GH, 
<collectionCode box="[1273,1306,735,754]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="17" pageNumber="1084" type="Herbarium">NY</collectionCode>
).
</paragraph>
<paragraph blockId="17.[824,1478,159,2003]" pageId="17" pageNumber="1084">
<collectingCountry box="[856,951,759,778]" name="Panama" pageId="17" pageNumber="1084">PANAMA</collectingCountry>
. 
<collectingRegion box="[960,1037,759,778]" country="Panama" name="Chiriqui" pageId="17" pageNumber="1084">Chiriqui</collectingRegion>
: Vicinity of Fortuna Dam, Sof lake on Eside of river valley across river, 
<quantity box="[1039,1108,783,802]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="17" pageNumber="1084" unit="m" value="1400.0">1,400 m</quantity>
, 
<date box="[1116,1207,783,802]" pageId="17" pageNumber="1084" value="1987-02-07">7 Feb 1987</date>
(fl, fr), 
<emphasis italics="true" pageId="17" pageNumber="1084">L. Bohs &amp; G. McPherson 2311</emphasis>
(
<collectionCode box="[880,914,807,826]" country="USA" lsid="urn:lsid:biocol.org:col:15631" name="Harvard University - Gray Herbarium" pageId="17" pageNumber="1084" type="Herbarium">GH</collectionCode>
); 
<quantity box="[934,992,807,826]" metricMagnitude="3" metricUnit="m" metricValue="5.632704" pageId="17" pageNumber="1084" unit="mi" value="3.5">3.5 mi</quantity>
NE of Boquete, end of road along Río Palo Alto, 
<date box="[824,930,831,850]" pageId="17" pageNumber="1084" value="1978-11-17">17 Nov 1978</date>
(fr), 
<emphasis box="[971,1104,831,850]" italics="true" pageId="17" pageNumber="1084">B. Hammel 5679</emphasis>
(
<collectionCode box="[1114,1151,831,850]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="17" pageNumber="1084" type="Herbarium">MO</collectionCode>
); Vicinityof Fortuna Dam, 
<geoCoordinate box="[1384,1472,830,850]" degrees="8" direction="north" minutes="45" orientation="latitude" pageId="17" pageNumber="1084" precision="15" seconds="04" value="8.751111">8°45′04″N</geoCoordinate>
<geoCoordinate box="[824,925,854,874]" degrees="82" direction="west" minutes="15" orientation="longitude" pageId="17" pageNumber="1084" precision="15" seconds="04" value="-82.251114">82°15′04″W</geoCoordinate>
, 
<quantity box="[937,1066,855,874]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="17" pageNumber="1084" unit="m" value="1350.0" valueMax="1400.0" valueMin="1300.0">
<geoCoordinate box="[937,982,855,874]" degrees="1,300" direction="north" orientation="latitude" pageId="17" pageNumber="1084" precision="55" value="1.3">1,300</geoCoordinate>
<geoCoordinate box="[982,1038,855,874]" degrees="1,400" direction="west" orientation="longitude" pageId="17" pageNumber="1084" precision="55" value="-1.4">–1,400</geoCoordinate>
m
</quantity>
, 
<date box="[1078,1178,855,874]" pageId="17" pageNumber="1084" value="1987-02-07">7 Feb 1987</date>
(fr), 
<emphasis box="[1230,1410,855,874]" italics="true" pageId="17" pageNumber="1084">G. McPherson 10387</emphasis>
(
<collectionCode box="[1426,1467,855,874]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="17" pageNumber="1084" type="Herbarium">MO</collectionCode>
). 
<collectingRegion box="[824,908,879,898]" country="Panama" name="Veraguas" pageId="17" pageNumber="1084">Veraguas</collectingRegion>
: 
<quantity box="[917,956,879,898]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="17" pageNumber="1084" unit="mi" value="5.0">5 mi</quantity>
<collectionCode box="[963,1007,879,898]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="17" pageNumber="1084" type="Herbarium">Wof</collectionCode>
Santa Fe on road past Escuela Agrícola Alto Piedra on Pacific side of divide, 
<quantity box="[1054,1164,903,922]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="17" pageNumber="1084" unit="m" value="1000.0" valueMax="1200.0" valueMin="800.0">800–1,200 m</quantity>
, 
<date box="[1175,1285,903,922]" pageId="17" pageNumber="1084" value="1973-03-18">18 Mar 1973</date>
(fl), 
<emphasis box="[1329,1472,903,922]" italics="true" pageId="17" pageNumber="1084">R. L. Liesner 948</emphasis>
(C, F, L, LL, 
<collectionCode box="[931,964,927,946]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="17" pageNumber="1084" type="Herbarium">NY</collectionCode>
).
</paragraph>
<paragraph blockId="17.[824,1478,159,2003]" pageId="17" pageNumber="1084">
<collectingCountry box="[856,912,951,970]" name="Peru" pageId="17" pageNumber="1084">PERU</collectingCountry>
. 
<collectingRegion box="[927,1034,951,970]" country="Peru" name="San Martin" pageId="17" pageNumber="1084">San Martín</collectingRegion>
: Cataratas de Ahuashiyacu, km 15 Tarapoto- Yurimaguas Road, 
<geoCoordinate box="[995,1050,975,994]" degrees="6" direction="south" minutes="29" orientation="latitude" pageId="17" pageNumber="1084" precision="925" value="-6.483333">6°29’S</geoCoordinate>
<geoCoordinate box="[1056,1130,975,994]" degrees="76" direction="west" minutes="21" orientation="longitude" pageId="17" pageNumber="1084" precision="925" value="-76.35">76°21’W</geoCoordinate>
, 
<quantity box="[1139,1195,975,994]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="17" pageNumber="1084" unit="m" value="700.0">700 m</quantity>
, 
<date box="[1205,1309,975,994]" pageId="17" pageNumber="1084" value="1986-06-10">10 Jun 1986</date>
(fl), 
<emphasis italics="true" pageId="17" pageNumber="1084">S. Knapp &amp; P. Alcorn 7791</emphasis>
(
<collectionCode box="[937,978,999,1018]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="17" pageNumber="1084" type="Herbarium">MO</collectionCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>